Here are five common forest flowers that the average visitor to these environs may easily overlook during an early April visit.
The flowers of the Skunk Cabbage, a native member of the arum family, are contained within a spadix which is partially hidden inside the mottled maroon spathe at the base of the plant. In late winter, the closed spathe generates its own heat to melt through frozen water and soils to make its appearance in streamside and spring-fed wooded wetlands, just ahead of the emergence of the large green leaves.Common Spicebush is a native flowering shrub of damp woodland understories. Later in the season, its foliage provides food for Spicebush Swallowtail caterpillars. Pollinated blossoms yield bright-red oblong berries relished by a variety of birds in fall and winter.The flowers of the indigenous Red Maple will soon generate the familiar helicopter-like winged seeds which readily distribute this native tree into new ground ranging from lowlands to the crests of our highest ridges.Coltsfoot (Tussilago farfara) is non-native wildflower of Eurasian origin. Reminding one at first glance of a dandelion, it is commonly seen blooming in disturbed areas of woodlands. Coltsfoot often grows where it has little competition from other plants, such as among the debris left behind due to snow plowing and grading along forest roads.Though not particularly abundant, the non-native Sweet Cherry, an escape from cultivation, is widespread in forests and woodlots throughout the lower Susquehanna valley. Sweet Cherry is believed to be an ancestor of the Sour Cherry (Prunus cerasus) and is frequently used as rootstock for orchard and garden varieties of this popular fruit. Sweet Cherry is often called “Wild Cherry”, a name also applied to the Black Cherry, a native tree which blooms later in the spring.
Be certain to get out and enjoy this year’s blooming seasons of our hundreds of varieties of flowering plants. But, particularly when it comes to native species,…
Our wildlife has been having a tough winter. The local species not only contend with cold and stormy weather, but they also need to find food and shelter in a landscape that we’ve rendered sterile of these essentials throughout much of the lower Susquehanna valley’s farmlands, suburbs, and cities.
Planting trees, shrubs, flowers, and grasses that benefit our animals can go a long way, often turning a ho-hum parcel of property into a privately owned oasis. Providing places for wildlife to feed, rest, and raise their young can help assure the survival of many of our indigenous species. With a little dedication, you can be liberated from the chore of manicuring a lawn and instead spend your time enjoying the birds, mammals, insects, and other creatures that will visit your custom-made habitat.
What makes some neighborhoods so appealing? It’s the foresight property owners had a half a century or more ago when they planted their lawns and gardens with a variety of sturdy, long-lived trees and shrubs. They’ve not only minimized the need for mowing grass, they’ve provided the present-day residents of their home with added thermal stability during both the blazing heat of summer and the chilling cold of winter.
Fortunately for us, our local county conservation districts are again conducting springtime tree sales offering a variety of native and beneficial cultivated plants at discount prices. Listed here are links to information on how to pre-order your plants for pickup in April. Click away to check out the species each county is offering in 2025!
Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025
During its 2025 tree seedling sale, the Cumberland County Conservation District is offering Northeast Native Wildflower seed mix for four dollars per ounce. One ounce plants approximately 200 square feet of bare soil. This is a Zebra Swallowtail visiting nectar-rich flowers during July of the first year after sowing this mix at a site along the Susquehanna.
Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025
Able to thrive in wet soils, Red-osier and Silky Dogwood shrubs are ideal plants for intercepting and polishing stormwater in swales, detention/retention basins, and rain gardens. With their crimson twigs in winter, they look great along borders among clusters of cedars, pines, spruces, and other evergreens. They make an excellent choice for soil stabilization along the shorelines of streams, ponds, and other bodies of water too. Buy a dozen or more to create a showy mass planting in your soggy spot.
The Franklin County Conservation District is offering American Elm seedlings in bundles of 25 for 36 dollars. Start them in pots for a couple of years to really get ’em going, then find places with damp soil and plenty of room to give ’em a try. During autumn, they look great in the company of spruces, white pines, and other large evergreens.
We purchased these Eastern White Pine, Norway Spruce, and Common Winterberry plants from the Lancaster County Conservation District Tree Sale about four years ago. They’re filling in as understory growth in the margins beneath some thirty-year-old Eastern Hemlocks to create dense cover for resident and visiting fauna at susquehannawildlife.net headquarters.
For 2025, the Lebanon County Conservation District is offering Dawn Redwood (Metasequoia glyptostroboides) trees in packs of ten for twelve dollars. Though native to Asia, these adaptable trees present little threat of naturalizing and have many positive attributes in a conservation planting. Given ample space, the Dawn Redwood, a relative of the sequoias, will rapidly grow to a towering giant. They create a particularly dramatic landscape feature when planted in clumps of three to five trees or more. With age, the trunks become stout and very sturdy. Don’t like raking? The finely divided deciduous foliage can be left where it falls in autumn. It usually disintegrates by spring to enrich the soil and promote more growth.The genus Metasequoia was first described in 1941 based upon fossils collected in Jurassic and earlier strata from widespread locations in the northern hemisphere. Metasequoia were believed extinct until just a few years later when a small number of living Dawn Redwoods were first discovered in southern China. Now distributed around the world for cultivation, direct descendants of this wild population of Metasequoia glyptostroboides are available for nearly anyone in a temperate climate to plant and grow to exceptional size. (National Park Service image)The Lebanon County Conservation District is also selling Bald Cypress trees. They’re offered in bundles of ten for ten dollars. These long-lived trees resemble the Dawn Redwood. Both are tolerant of damp ground, but the native Bald Cypress is the species to choose for placement along streams, in wetlands, and on other sites with standing water or saturated soil.Wildlife rich Bald Cypress swamps currently occur on the Atlantic Coastal Plain as far north as Sussex County, Delaware. Just to the south, they’re also found along Chesapeake Bay in areas that, during the last glacial maximum when sea level was 300 to 400 feet below today’s tide lines, were the Lower Susquehanna River Watershed in portions of present-day Maryland and Virginia. The northward post-glacial range extension of Bald Cypresses is now blocked by centuries of human intervention that has eliminated, isolated, or fragmented the wetland habitats where they could potentially become established. Why not lend them a hand? Plant a cypress swamp in your flood-prone bottomland. (National Park Service image by Andrew Bennett)
A privacy planting of sturdy, native Eastern White Pines and Northern Red Oaks thriving around the border of a parking area where they also provide shade from the sun and help infiltrate a share of the stormwater that would otherwise become runoff.This year, the York County Conservation District is offering a Showy Northeast Native Wildflower and Grass seed mix for $19.99 per quarter pound. Sure beats mowing!
If you live in Adams County, Pennsylvania, you may be eligible to receive free trees and shrubs for your property from the Adams County Planting Partnership (Adams County Conservation District and the Watershed Alliance of Adams County). These trees are provided by the Chesapeake Bay Foundation’s Keystone 10-Million Trees Partnership which aims to close a seven-year project in 2025 by realizing the goal of planting 10 million trees to protect streams by stabilizing soils, taking up nutrients, reducing stormwater runoff, and providing shade. If you own property located outside of Adams County, but still within the Chesapeake Bay Watershed (which includes all of the Susquehanna, Juniata, and Potomac River drainages), you still may have an opportunity to get involved. Contact your local county conservation district office or watershed organization for information.
As they mature, tree and shrub plantings along streams return pollution-controlling functions to floodplains and provide critical habitat for wildlife. These riparian buffers not only improve water quality for fisheries, they also create travel corridors that prevent terrestrial animal populations from becoming isolated.Do you own a parcel of streamside or wetland acreage that you’d like to set aside and plant for the benefit of wildlife and water quality? Contact your local county conservation district office and ask them to tell you about CREP (Conservation Reserve Enhancement Program) and other programs that may offer incentives including payment of all or a portion of the costs of plantings and other habitat improvements.
We hope you’re already shopping. Need help making your selections? Click on the “Trees, Shrubs, and Woody Vines” tab at the top of this page to check out Uncle Tyler Dyer’s leaf collection. He has most of the species labelled with their National Wetland Plant List Indicator Rating. You can consult these ratings to help find species suited to the soil moisture on your planting site(s). For example: if your site has sloped upland ground and/or the soils sometimes dry out in summer, select plants with a rating such as UPL or FACU. If your planting in soils that remain moist or wet, select plants with the OBL or FACW rating. Plants rated FAC are generally adaptable and can usually go either way, but may not thrive or survive under stressful conditions in extremely wet or dry soils.
NATIONAL WETLAND PLANT LIST INDICATOR RATING DEFINITIONS
OBL (Obligate Wetland Plants)—Almost always occur in wetlands.
FACW (Facultative Wetland Plants)—Usually occur in wetlands, but may occur in non-wetlands.
FAC (Facultative Wetland Plants)—Occur in wetlands and non-wetlands.
FACU (Facultative Upland Plants)—Usually occur in non-wetlands, but may occur in wetlands.
UPL (Upland Plants)—Almost never occur in wetlands.
Using these ratings, you might choose to plant Pin Oaks (FACW) and Swamp White Oaks (FACW) in your riparian buffer along a stream; Northern Red Oaks (FACU) and White Oaks (FACU) in the lawn or along the street, driveway, or parking area; and Chestnut Oaks (UPL) on your really dry hillside with shallow soil. Give it a try.
Each spring and fall, Purple Finches are regular migrants through the Lower Susquehanna River Watershed. Northbound movements usually peak in April and early May. During the summer, these birds nest primarily in cool coniferous forests to our north. Then, in October and November each year, they make another local appearance on their way to wintering grounds in the southeastern United States. A significant population of Purple Finches remains to our north through the colder months, inhabiting spruce-pine and mixed forests from the Great Lakes east through New England and southeastern Canada. This population can be irruptive, moving south in conspicuous numbers to escape inclement weather, food shortages, and other environmental conditions. Every few years, these irruptive birds can be found visiting suburbs, parks, and feeding stations—sometimes lingering in areas not often visited by Purple Finches. Right now, Purple Finches in flocks larger than those that moved through earlier in the fall are being seen throughout the lower Susquehanna valley. Snow and blustery weather to our north may be prompting these birds to shift south for a visit. Here are some looks at members of a gathering of more than four dozen Purple Finches we’ve been watching in Lebanon County this month…
Purple Finches are easily distinguished from House Finches by their notched tails, darker streaking, and well-defined facial markings. Though most frequently found in proximity to conifers, they also visit native deciduous trees and shrubs to snack on berries, seeds, and buds. Purple Finches can be particularly fond of maple, birch, poplar, sumac, and viburnum.Adult male Purple Finches are unmistakable in their wine-colored plumage.Purple Finches join a Dark-eyed Junco, a White-throated Sparrow, and a Black-capped Chickadee at a feeding station.Purple Finches eating black oil sunflower seed.A male Purple Finch in successional forest edge habitat consumes seeds from a dried stand of goldenrod.Purple Finches with juncos and Northern Cardinals.Purple Finches filling up on supplemental foods.
During the past week, Uncle Tyler Dyer has been out searching for autumn leaves to add to his collection. One of the species he had not encountered in previous outings was the American Elm (Ulmus americana), so he made a special trip to see a rare mammoth specimen in a small neighborhood park (Park Place) along Chestnut Street between 5th and Quince Streets in Lebanon, Pennsylvania.
Possibly the largest and oldest remaining American Elm in the lower Susquehanna watershed, the Park Place tree in Lebanon exceeds 60 feet in height and may be more than 300 years of age. In the nearly one hundred years since Dutch elm disease (DED) first began killing elms in eastern North America, this tree has been spared the fatal effects of the infection. DED is caused by several species of microfungi (Ophiostoma ulmi, O. himal-ulmi, and O. novo-ulmi) spread by numerous bark beetle (Curculionidae) species. As early as 1928, infected beetles arrived in the United States from the Netherlands among shipments of logs.Leaves of the American Elm.
There’s still time to get out and see autumn foliage. With warmer weather upon us—at least temporarily—it’s a good time to go for a stroll. Who knows, you might find some spectacular leaves like these collected by Uncle Ty earlier this week. All were found adorning native plants!
Winged Sumac (Rhus copallina), a small native tree. In case you’re wondering, it’s not poisonous.Black Chokeberry (Photinia melanocarpa), a native shrub also known as Aronia. It produces black-colored fruits in summer.Red-osier Dogwood (Cornus sericea), a shrub that prefers wet or damp soils. It yields white fruits.Maple-leafed Viburnum (Viburnum acerifolium), a native shrub of upland forest understories.Highbush Blueberry (Vaccinium corymbosum), a native shrub and the source of many cultivated forms of the popular fruit.To identify your finds, be certain to click the “Trees, Shrubs, and Woody Vines” tab at the top of this page to check out Uncle Ty’s extensive collection.
Bathed in glowing sunshine, a very large fallout of migrating Neotropical songbirds enlivened the forest edge atop Second Mountain in northern Lebanon County, Pennsylvania, this morning. While last night’s flight was widespread beneath a dome of atmospheric high pressure covering the Mid-Atlantic States, this is a look at some of the 500 -1,000 migrants observed feeding on insects and other natural foods at just this single location.
As the composition of the nightly flights begins to transition to species that start their migration later in the season, numbers of birds like the Blackpoll Warbler (top) are beginning to rival those of the heretofore very common Black-throated Green Warbler (bottom).A Blackpoll Warbler.The Bay-breasted Warbler, another common species found among the fallout flock.The Tennessee Warbler was today’s most abundant migrant with at least 200 birds seen moving among the other species.A newly arrived Tennessee Warbler at sunrise.A Tennessee Warbler snatches breakfast.The Magnolia Warbler was another common find among today’s migrants.A Magnolia Warbler.A Nashville Warbler among tangles of Mile-a-minute Weed.A Nashville Warbler.Black-throated Blue Warbler.An adult male Black-throated Blue Warbler.An acrobatic Northern Parula.Blackburnian Warblers continue their southerly trek.An American Redstart.A Chestnut-sided Warbler feeding on Mile-a-minute Weed berries.A Cape May Warbler.A handsome adult male Cape May Warbler.A Swainson’s Thrush.Scarlet Tanagers continue to work their way south through the Lower Susquehanna River Watershed.A composite image of an uncommon find, a large-headed Olive-sided Flycatcher seen perched atop a dead tree snag.A brief encounter with a Philadelphia Vireo, another uncommon species.A Blue-headed Vireo, typically the last of its genus to move south in the fall, put in a welcome appearance among the hundreds of migrants.A favorite diurnal migrant, the Ruby-throated Hummingbird, was seen feeding on flowers that grew from the remains of garden waste dumped illegally along the mountain road last autumn.
Many of the observers at the Second Mountain Hawk Watch commented that this morning’s fallout was by far the best they had seen anywhere in the region during recent years. Others believed it to be the best they had ever seen. It was indeed less like a “wave” of migrants and more like a “tsunami”. Choosing a good viewing location and being there at the right time can improve your chances of seeing a spectacle like this. The good news is, it looks like another big flight is currently underway, so finding a forest edge on a ridgetop or along a utility right-of-way just might pay off for you early tomorrow morning.
Nocturnal migrating birds, a southwest-bound flight down the ridges of central Pennsylvania as indicated by Doppler radar between 10:30 and 11:30 PM EDT on September 11, 2024. (NOAA/National Weather Service image)
Back on March 24th, we took a detailed look at the process involved in administering prescribed fire as a tool for managing grassland and early successional habitat. Today we’re going turn back the hands of time to give you a glimpse of how the treated site fared during the five months since the controlled burn. Let’s go back to Middle Creek Wildlife Management Area for a photo tour to see how things have come along…
Pennsylvania Game Commission crews administering prescribed fire on a grassland demonstration site back on March 16.By late May, native herbaceous perennial wildflowers including Joe-pye Weed had re-greened the site. One of the goals of the burn was to kill fire-sensitive woody plants, thus preventing the process of succession from reforesting the site.The scorched, lifeless remains of small trees and shrubs indicate that that goal was met.Because prescribed fire is administered in a mosaic pattern that permits some early successional growth to remain until the next burn, birds including this thicket-nesting Yellow-breasted Chat are able to take advantage of the mixed habitat during their breeding season in May and June.By August, the site is a haven for native plants and animals.The burn has promoted the growth and late-summer bloom of fire-tolerant native wildflowers and warm-season grasses……including Indiangrass,……Big Bluestem……Thin-leaved Coneflower (Rudbeckia triloba), a plant also known as Brown-eyed Susan,……and Joe-pye Weed, a plant butterflies find irresistible.Eastern Tiger Swallowtail collecting nectar from Joe-pye Weed.A black-morph Eastern Tiger Swallowtail collecting nectar from Spotted Knapweed (Centaurea steobe micranthos), a non-native invasive plant found growing in an area of the burn site missed by this year’s fire. While many non-native plants are unable to survive the flames and heat produced by prescribed fire, it isn’t an absolute cure-all. It doesn’t eliminate all invasive plants, it just keeps them from dominating a landscape by out-competing native species. Left unmanaged, Spotted Knapweed is a tough perennial invasive that can easily become one the species able to overtake a vulnerable grassland. It can be a stubborn survivor of some prescribed burns. On the plus side, butterflies really like it.By August, native grassland plants in the prescribed fire area were already providing an abundance of seeds for birds including this American Goldfinch.For larger birds like turkeys and pheasants, an abundance of Carolina Grasshoppers are providing a protein-dense food source in managed grasslands.And tiny flying insects, a nuisance to us as we take a stroll alongside the grasslands, are a meal taken on the wing by dragonflies including this Black Saddlebags.
Elsewhere around the refuge at Middle Creek, prescribed fire and other management techniques are providing high-quality grassland habitat for numerous species of nesting birds…
Bobolinks nested both in areas subjected to controlled burns……and in hay fields where mowing was delayed until the nesting season, including the fledging process, was completed earlier this month.As advertised, Grasshopper Sparrows nested in these fields as well.
We hope you enjoyed this short photo tour of grassland management practices. Now, we’d like to leave you with one last set of pictures—a set you may find as interesting as we found them. Each is of a different Eastern Cottontail, a species we found to be particularly common on prescribed fire sites when we took these images in late May. The first two are of the individuals we happened to be able to photograph in areas subjected to fire two months earlier in March. The latter two are of cottontails we happened to photograph elsewhere on the refuge in areas not in proximity to ground treated with a prescribed burn or exposed to accidental fire in recent years.
Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.
These first two rabbits are living the good life in a warm-season grass wonderland.
Eastern Cottontail at a site not subjected to any recent fire activity.Eastern Cottontail at a site not subjected to any recent fire activity.
Oh Deer! Oh Deer! These last two rabbits have no clock to track the time; they have only ticks. Better not go for a stroll with them Alice—that’s no wonderland! I know, I know, it’s time to go. See ya later.
The Aphrodite Fritillary (Speyeria aphrodite), also known simply as the Aphrodite, is a brush-footed butterfly of deciduous, coniferous, and mixed forests. We found this female in a grassland margin between woodlots where prescribed fire was administered during the autumn of 2022 to reduce accumulations of natural fuels and an overabundance of invasive vegetation. A goal of the burn was to promote the growth of native species including the violets (Viola species) favored as larval host plants by this and other fritillaries.
A female Aphrodite Fritillary collecting nectar from a thistle flower.
By the time these adult butterflies make their reproductive flights in late summer, the violets that serve as larval host plants have gone dormant. To find patches of ground where the violets will come to life in spring, the female Aphrodite Fritillary has an ability to sense the presence of dormant roots, probably by smell. Upon finding an area where suitable violets will begin greening up next year, she’ll deposit her eggs. The eggs overwinter, then hatch to feed on the tender new violet leaves of spring.
A female Aphrodite Fritillary. A prescribed burn, when administered during spring to manage fritillary habitat, is applied only to a portion of the land parcel each year to avoid decimating an entire population of the larvae during the first instar of their life cycle, a time when they are vulnerable to fire.Our female Aphrodite Fritillary busily gathers nutritious nectar to provide sufficient energy for the critical process of mating and egg production. What’s the thistle that this goddess of love and procreation is pollinating?…It’s a non-native invasive, the Bull Thistle (Cirsium vulgare), a species that readily colonizes new areas by producing an abundance of airborne seeds. Continued management of this site with periodic applications of prescribed fire will prevent Bull Thistle and other invasives from overtaking the habitat during coming years.
Here’s a native plant you don’t see very often, but Ruby-throated Hummingbirds have a knack for finding and adoring it within the limited habitat where it still exists.
Allegheny Vine (Adlumia fungosa), also known as Climbing Fumitory or Mountain Fringe, is a seldom-found biennial herbaceous plant in the poppy family (Papaveraceae). Both its divided leaves and drooping clusters of flowers might remind the observer of a more familiar member of the poppy family, the spring-blooming Dutchman’s Breeches. But unlike the low-growing Dutchman’s Breeches, Allegheny Vine ascends nearby shrubs and small trees with trailing stems that add a garland-like pizzazz to the summertime foliage.Seen here creeping atop a small Sassafras (Sassafras albidum) tree, Allegheny Vine seldom escapes notice by the Ruby-throated Hummingbird, a primary pollinator of the plant.Allegheny Vine grows on moist, rocky slopes and blooms in late summer, just in time to refuel migrating Ruby-throated Hummingbirds.
Not likely to thrive in deep forest cover, Allegheny Vine is a denizen of edge habitat. The species is dependent on some type of disturbance to maintain suitable growing conditions. The plants seen here were found flowering in an area that had experienced a fire prior to last year’s growing season. It is therefore quite possible that the fire completed the stratification process and triggered long-dormant seeds to germinate last spring or summer to develop a basal plant which matured and flowered during this second summer of the biennial’s growth. On a forested slope opposite this site, a first-ever prescribed burn was conducted in March of this year to eliminate accumulated fuel and replicate a natural fire cycle. With a little luck, this forest management approach may prompt additional long-dormant Allegheny Vine seeds to germinate and form basal plants this year for maturation in 2025. Without man-made disturbances, Allegheny Vine may remain viable as a species by enduring seed dormancy periods of 40-100 years or more between fires, lightning strikes, wind storms, ice accumulations, and other events which clear the canopy and encourage growth.
Many years in the making, Allegheny Vine nectar must be delectable because the hummers can’t leave it alone. And another plus, with a little help from prescribed fire, this native creeping plant seems to be holding its own while in direct competition with invasive Mile-a-minute Weed (leaves in lower right).
In the forests and woodlots of the lower Susquehanna watershed, the Black Gum, also known as the Black Tupelo, is always the first of our native trees to pop with autumn color. Its crimson leaves glow in an otherwise green landscape to help lure hungry birds including waxwings, American Robins, and Pileated Woodpeckers to its soon-to-be-ripe berries. The seeds contained in the fruit are thus well-distributed as a by-product of this avian feast. While not among the most massive of our trees, the Black Gum can live for over five hundred years, an age not exceeded by any other non-clonal indigenous flowering plant.
Do you recall our “Photo of the Day” from seven months ago…
Here’s something to look forward to in the new year. The good citizens of East Donegal Township in Lancaster County have partnered with Alliance for the Chesapeake Bay to establish an extensive wildflower meadow on what had been a mowed field of turf grass at Riverside Park in the Susquehanna floodplain near Marietta. As the photo shows, the lawn plants have been eliminated in preparation for seeding with a diverse assortment of native grasses and wildflowers to provide habitat for birds and pollinators including butterflies, bees, and other insects. Once established, the meadow’s extensive vegetative growth will help reduce stormwater runoff by better infiltrating rainfall to recharge the aquifer. During flood events, the plantings will provide soil stabilization and increase the ability of the acreage to uptake nutrients, thus reducing the negative impact of major storms on the quality of water in the river and in Chesapeake Bay. Check the project’s progress by stopping by from time to time in 2024!
Well, here’s what that site looks like today…
The wildflowers, thousands of them, are now in bloom!Black-eyed Susan and Blue Vervain (Verbena hastata) dominate the assortment currently in flower.
And there are pollinating insects galore, most notably butterflies…
A Cabbage White collecting nectar on Blue Vervain.A Clouded or Orange Sulphur among the grasses in the meadow.A Silver-spotted Skipper.The Least Skipper is our tiniest butterfly.The Little Glassywing (Vernia verna), this one feeding on vervain nectar, deposits its eggs on Purpletop grass, which then functions as the host plant for this butterfly’s larvae.A Summer Azure (Celastrina neglecta) feeding on the nectar of a Black-eyed Susan.A Common Buckeye on a “Gloriosa Daisy”, a showy, large-flowered cultivar of Black-eyed Susan.A Red-winged Blackbird with a caterpillar found among the meadow’s lush growth.An Eastern Tiger Swallowtail on Blue Vervain. Nearby Yellow (Tulip) Poplars and other trees serve as host plants for this butterfly’s larvae.The black morph of the Eastern Tiger Swallowtail shows subdued shading in the wings that closely resembles the brilliant color patterns of the more familiar yellow form.The Sachem, this one visiting a Black-eyed Susan, is a variable species with a range that normally lies south of the 40th parallel, the line of latitude that intersects the Susquehanna in the area of the Conejohela Flats at Washington Boro.A Sachem visiting the blooms of Oxeye sunflower. During recent weeks, scorching winds from the south and southwest have transported an abundance of these vagrant skippers into the lower Susquehanna valley and beyond.A male Sachem approaching the bloom of a “Gloriosa Daisy”. Miles north of the 40th parallel, wandering Sachems are currently the most numerous of the butterflies at the Riverfront Park wildflower meadow.Here and there among the meadow’s plantings we noticed one of our favorites starting to flower, the Partridge Pea.Partridge Pea happens to be a host plant for another vagrant from the south, the big, lime-yellow Cloudless Sulphur. We saw at least half a dozen patrolling the meadow.The stars of the show are the Zebra Swallowtails, gorgeous butterflies that rely on stands of native Common Paw-paw trees in the river floodplain to host their eggs and larvae.The red-white-and-blue underside of a Zebra Swallowtail.WOW!
Why on earth would anyone waste their time, energy, and money mowing grass when they could have this? Won’t you please consider committing graminicide this fall? That’s right, kill that lawn—at least the majority of it. Then visit the Ernst Seed website, buy some “Native Northeast Wildlflower Mix” and/or other blends, and get your meadow planted in time for the 2025 growing season. Just think of all the new kinds of native plants and animals you’ll be seeing. It could change your life as well as theirs.
A Snowberry Clearwing (Hemaris diffinis), more commonly known as a hummingbird moth, visits the flowers of Blue Vervain in the Riverfront Park wildflower meadow.
Resilient to the pressures of flooding, ice scour, drought, and oft times really poor water quality, Water Willow (Dianthera americana, formerly Justicia americana) is the most common herbaceous plant on the Susquehanna’s non-forested alluvial islands. Yet, few know this native wildflower by name or reputation.
Pure stands of emergent Water Willow endure at times brutal conditions on non-forested islands in the Susquenanna.Alluvial deposits of sand, clay, gravel, and silt create ideal substrate for mats of Water Willow along shorelines of the Susquehanna and its larger tributaries. Provided the loose substrate remains moist, this emergent thrives even when water levels retreat during periods of dry weather. The woody plant in the background, the native Black Willow (Salix nigra), shares similar soil preferences but is found growing on slightly higher ground as a non-emergent tree or shrub. It is a member of the willow family (Salicaceae).In bloom now, the orchid-like flower of the Water Willow is a quick giveaway that it is not a close relative of the willow trees but is instead a member of the acanthus family (Acanthaceae) and is allied with the genus Ruellia, the wild petunias.
The spring of 2024 has been very kind to our beds of Water Willow. Rainfall in the Susquehanna watershed has been frequent enough to maintain river levels just high enough to keep the roots of the plants wet. During the interludes in storm activity, dry spells have rolled back any threat of flooding on the river’s main stem, thus eliminating chances of submerging the plants in muddy water and preventing the sun from keeping them warm, happy, and flowering early. Thundershowers throughout the basin earlier this week have now raised the river a few inches to inundate the base of the plants and make mats of Water Willow favorable places for newly hatched fry and other young fish to take refuge while they grow. Here’s a look…
The Spottail Shiner (Notropis hudsonius) is a common native minnow of the Susquehanna. This juvenile was found among several dozen small fish taking refuge in the cover of Water Willow below Conewago Falls.The Mimic Shiner (Notropis volucellus) is generally regarded to be a native transplant from the Mississippi drainage that has become established in the Susquehanna and many of its tributaries, possibly after introduction by way of bait buckets. However, the fish tends to be very fragile and dies quickly upon handling, so its use and transport as a bait species may be impractical. The Mimic Shiner is very common in around Conewago Falls.A juvenile Mimic Shiner less than one inch in length found among flooded Water Willow below Conewago Falls earlier this week.One of about a dozen juvenile Quillbacks (Carpiodes cyprinus) found in Water Willow just below Conewago Falls. For spawning, local populations of this compact native species of carpsucker favor the gravel-bottomed pools among the Jurassic-Triassic boulders of the falls’ pothole rocks. Probably hatched within the last eight weeks, this specimen was just one inch long.As summer progresses, stands of emergent Water Willow begin to expand their size by sending out rhizomes. Increasing numbers of small fish like this Spotfin/Satinfin Shiner (Cyprinella species) concentrate in the cover of the thickening vegetation.The importance of these patches of emergent wildflowers (sounds weird, doesn’t it?) is demonstrated by the numbers of fish gathered within their underwater forest of stems and leaves by summertime.To protect them from burial by silt and to prevent them from being swept away by current, spawning Spotfin/Satinfin Shiners deposit their eggs in crevices of submerged rocks and wood, often in or near mats of Water Willow. Males guard the eggs until hatching. The fry must then take shelter among boulders, cobble, and plant cover. Note the breeding-condition male in the upper right.Panfish like this non-native Green Sunfish (Lepomis cyanellus) will often choose nesting sites in deeper water adjacent to beds of Water Willow, particularly if submerged growth like this Water Stargrass adds to the availability of cover for their young after hatching.Smallmouth Bass gather in a pool adjacent to a Water Willow-covered island. These non-native predators rely on beds of these indigenous plants to provide habitat for their young, then, after spawning, lurk in the waters surrounding them to ambush less-than-vigilant minnows and other victims.
By now you’ve come to appreciate the importance of Water Willow to the sustainability of our populations of fish and other aquatic life. Like similar habitat features that reduce sediment runoff and nutrient pollution, undisturbed stands of terrestrial, emergent, and submerged native plant species are essential to the viability of our freshwater food webs.
The minuscule Long-legged Fly (Condylostylus species) is a predatory consumer of soft-bodied invertebrates. In the headquarters garden, we found this individual and others scurrying around on the leaves of Common Milkweed where they may be seeking to gobble up infesting aphids.
While in flower, this extensive stand of Princess Trees (Paulownia tomentosa) straddling Second Mountain north of Harrisburg might be mistaken for a series of boulder outcrops. Native to eastern Asia, the fast-growing Princess Tree has escaped cultivation to become naturalized in many parts of eastern North America. You’ll currently notice the showy purple blooms on many forested ridges and hilltops throughout the Lower Susquehanna River Watershed. Paulownia tomentosa is also known as the Phoenix Tree, a name derived from its ability, due to its extensive root system, to regenerate following fire. This fast-growing invasive therefore calls for measures in addition to prescribed burns for control within infected forests. Mechanical and/or chemical methods of removal are frequently required.
Homo sapiens owes much of its success as a species to an acquired knowledge of how to make, control, and utilize fire. Using fire to convert the energy stored in combustible materials into light and heat has enabled humankind to expand its range throughout the globe. Indeed, humans in their furless incomplete mammalian state may have never been able to expand their populations outside of tropical latitudes without mastery of fire. It is fire that has enabled man to exploit more of the earth’s resources than any other species. From cooking otherwise unpalatable foods to powering the modern industrial society, fire has set man apart from the rest of the natural world.
In our modern civilizations, we generally look at the unplanned outbreak of fire as a catastrophe requiring our immediate intercession. A building fire, for example, is extinguished as quickly as possible to save lives and property. And fires detected in fields, brush, and woodlands are promptly controlled to prevent their exponential growth. But has fire gone to our heads? Do we have an anthropocentric view of fire? Aren’t there naturally occurring fires that are essential to the health of some of the world’s ecosystems? And to our own safety? Indeed there are. And many species and the ecosystems they inhabit rely on the periodic occurrence of fire to maintain their health and vigor.
For the war effort- The campaign to reduce the frequency of forest fires got its start during World War II with distribution of this poster in 1942. The goal was to protect the nation’s timber resources from accidental or malicious loss due to fire caused by man-made ignition sources. The release of the Walt Disney film “Bambi” during the same year and the adoption of the Smokey the Bear mascot in 1944 softened the message’s delivery, but the public relations outreach continued to be a key element of a no-fire policy to save trees for lumber. Protection and management of healthy forest ecosystems in their entirety has only recently become a priority. (National Archives image)
Man has been availed of the direct benefits of fire for possibly 40,000 years or more. Here in the Lower Susquehanna River Watershed, the earliest humans arrived as early as 12,000 years ago—already possessing skills for using fire. Native plants and animals on the other hand, have been part of the ever-changing mix of ecosystems found here for a much longer period of time—millions to tens of millions of years. Many terrestrial native species are adapted to the periodic occurrence of fire. Some, in fact, require it. Most upland ecosystems need an occasional dose of fire, usually ignited by lightning (though volcanism and incoming cosmic projectiles are rare possibilities), to regenerate vegetation, release nutrients, and maintain certain non-climax habitat types.
But much of our region has been deprived of natural-type fires since the time of the clearcutting of the virgin forests during the eighteenth and nineteenth centuries. This absence of a natural fire cycle has contributed to degradation and/or elimination of many forest and non-forest habitats. Without fire, a dangerous stockpile of combustible debris has been collecting, season after season, in some areas for a hundred years or more. Lacking periodic fires or sufficient moisture to sustain prompt decomposition of dead material, wildlands can accumulate enough leaf litter, thatch, dry brush, tinder, and fallen wood to fuel monumentally large forest fires—fires similar to those recently engulfing some areas of the American west. So elimination of natural fire isn’t just a problem for native plants and animals, its a potential problem for humans as well.
Indiangrass (seen here), Switchgrass, Big Bluestem, and Little Bluestem are native species requiring periodic forms of disturbance to eliminate competition by woody plants. These warm-season grasses develop roots that penetrate deep into the soil, sometimes to depths of six feet or more, allowing them to survive severe drought and flash fire events. In the tall grass prairies, these extensive root systems allow these grasses to return following heavy grazing by roaming herds of American Bison (Bison bison). Without these habitat disturbances, warm season grasslands succumb to succession in about seven years. With their periodic occurrence, the plants thrive and provide excellent wildlife habitat, erosion control, and grazing forage.
To address the habitat ailments caused by a lack of natural fires, federal, state, and local conservation agencies are adopting the practice of “prescribed fire” as a treatment to restore ecosystem health. A prescribed fire is a controlled burn specifically planned to correct one or more vegetative management problems on a given parcel of land. In the Lower Susquehanna River Watershed, prescribed fire is used to…
Eliminate dangerous accumulations of combustible fuels in woodlands.
Reduce accumulations of dead plant material that may harbor disease.
Provide top kill to promote oak regeneration.
Regenerate other targeted species of trees, wildflowers, grasses, and vegetation.
Kill non-native plants and promote growth of native plants.
Prevent succession.
Remove woody growth and thatch from grasslands.
Promote fire tolerant species of plants and animals.
Improve habitat for rare species (Regal Fritillary, etc.)
Recycle nutrients and minerals contained in dead plant material.
Let’s look at some examples of prescribed fire being implemented right here in our own neighborhood…
Prescribed fires are typically planned for the dormant season extending from late fall into early spring with burns best conducted on days when the relative humidity is low.Prescribed fire is used regularly at Fort Indiantown Gap Military Reservation in Lebanon County, Pennsylvania, to keep accumulations of woody and herbaceous fuels from accumulating on and around the training range areas where live ordinance and other sources of ignition could otherwise spark large, hard-to-control wildfires.Prescribed fires replace the periodic natural burns that would normally reduce the fuel load in forested areas. Where these fuels are allowed to accumulate, south-facing slopes are particularly susceptible to extreme fires due to their exposure to the drying effects of intense sunlight for much of the year. The majority of small oaks subjected to treatment by the prescribed fire shown here will have the chance to regenerate without immediate competition from other species including invasive plants. The larger trees are mostly unaffected by the quick exposure to the flames. Note too that these fires don’t completely burn everything on the forest floor, they burn that which is most combustible. There are still plenty of fallen logs for salamanders, skinks, and other animals to live beneath and within.
A prescribed fire in late winter prevents this grassland consisting of Big Bluestem and native wildflowers from being overtaken by woody growth and invasive species. Fires such as this that are intended to interrupt the process of succession are repeated at least every three to five years.In its wildlife food plots, prescribed fire is used by the Pennsylvania Game Commission to prevent succession and control invasive species such as Multiflora Rose, instead promoting the growth of native plants.An example of a woodlot understory choked with combustible fuels and dense tangles of invasive Multiflora Rose. A forester has the option of prescribing a dose of dormant-season fire for a site like this to reduce the fuel load, top kill non-native vegetation, and regenerate native plants.A dose of prescribed fire was administered on this grassland to kill the woody growth of small trees beginning to overtake the habitat by succession.The Pennsylvania Game Commission employs prescribed fire at Middle Creek Wildlife Management Area and on many of their other holdings to maintain grasslands.Prescribed fire is used to eliminate invasive species including Multiflora Rose from grasslands at Middle Creek W.M.A. Annual burns on the property are conducted in a mosaic pattern so that each individual area of the grassland is exposed to the effects of fire only once every two to five years. Without fire or some type of mechanical or chemical intervention, succession by woody trees and shrubs would take hold after about seven years.Prescribed fire is planned for a fraction of total grassland acreage at Middle Creek W.M.A. each year. Another section of the mosaic is targeted in the following year and yet another in the year that follows that. Because burns are conducted in the spring, grassland cover is available for wildlife throughout the winter. And because each year’s fire burns only a portion of the total grassland acreage, wildlife still has plenty of standing grass in which to take shelter during and after the prescribed fire.Prescribed fire at Middle Creek W.M.A. provides grassland habitat for dozens of species of birds and mammals including the not-so-common Grasshopper Sparrow……and stocked Ring-necked Pheasants that do nest and raise young there.On a few sites in the Lower Susquehanna River Watershed , prescribed fire is being used to establish and maintain savanna-like grasslands. This one, located on a dry, south-facing slope near numerous man-made sources of ignition, can easily be dosed with periodic prescribed burns to both prevent succession and reduce fuel accumulations that may lead to a devastating extreme fire.One year following a prescribed burn, this is the autumn appearance of a savanna-like habitat with fire-tolerant Pitch Pine (Pinus rigida), Bear Oak, warm-season grasses, and a variety of nectar-producing wildflowers for pollinators. These ecosystems are magnets for wildlife and may prove to be a manageable fit on sun-drenched sites adjacent to man-made land disturbances and their sources of ignition.Savanna-like grasslands with oaks and other scattered large trees, some of them dead, make attractive nesting habitat for the uncommon Red-headed Woodpecker.Prescribed fire can benefit hungry Wild Turkeys by maintaining savanna-like grasslands for an abundance of grasshoppers and other insects in summer and improving the success of mast-producing oaks for winter.In the Lower Susquehanna River Watershed, the caterpillar of the rare Eastern Buck Moth feeds on the foliage of the Bear Oak, also known as the Scrub Oak, a shrubby species that relies upon periodic fire to eliminate competition from larger trees in its early successional habitat.Leaves of the Bear Oak in fall. The Bear Oak regenerates readily from top kill caused by fire.Reed Canary Grass (Phalaris arundinacea) is a native cool-season grass with a colorful inflorescence in spring. But given the right situation, it can aggressively overtake other species to create a pure stand lacking biodiversity. It is one of the few native species which is sometimes labelled “invasive”.Prescribed fire can be used to reduce an overabundance of Reed Canary Grass and its thatch in wetlands. Periodic burning can help restore species diversity in these habitats for plants and animals including rare species such as the endangered Bog Turtle (Glyptemys muhlenbergii).On the range areas at Fort Indiantown Gap in Lebanon County, Pennsylvania, disturbances by armored vehicles mimic the effects of large mammals such as the American Bison which periodically trampled grasses to prevent succession and the establishment of woody plants on its prairie habitat. To supplement the activity of the heavy vehicles and to provide suitable habitat for the very rare Regal Fritillary (Speyeria idalia) butterflies found there, prescribed fire is periodically employed to maintain the grasslands on the range. These burns are planned to encourage the growth of “Fort Indiantown Gap Little Bluestem” grass as well as the violets used as host plants by the Regal Fritillary caterpillars. These fires also promote growth of a variety of native summer-blooming wildflowers to provide nectar for the adults butterflies.A last record of a wild American Bison killed in Pennsylvania was an animal taken in the Susquehanna watershed in Union County in 1801. The species is thereafter considered extirpated from the state. Since that time, natural disturbances needed to regenerate warm-season grasses have been limited primarily to fires and riverine ice scour. The waning occurrence of both has reduced the range of these grasses and their prairie-like ecosystems in the commonwealth. (Exhibit: State Museum of Pennsylvania, Harrisburg)A male Regal Fritillary on the range at Fort Indiantown Gap, where armored vehicles and prescribed fire provide suitable prairie-like habitat for this vulnerable species.Prescribed fires return the nutrients and minerals contained in dead plant material to the soil. Following these controlled burns, insects like this Honey Bee can often be seen collecting minerals from the ashes.A Greenbottle Fly gathering minerals from the ash following a prescribed burn.
In Pennsylvania, state law provides landowners and crews conducting prescribed fire burns with reduced legal liability when the latter meet certain educational, planning, and operational requirements. This law may help encourage more widespread application of prescribed fire in the state’s forests and other ecosystems where essential periodic fire has been absent for so very long. Currently in the Lower Susquehanna River Watershed, prescribed fire is most frequently being employed by state agencies on state lands—in particular, the Department of Conservation and Natural Resources on State Forests and the Pennsylvania Game Commission on State Game Lands. Prescribed fire is also part of the vegetation management plan at Fort Indiantown Gap Military Reservation and on the land holdings of the Hershey Trust. Visitors to the nearby Gettysburg National Military Park will also notice prescribed fire being used to maintain the grassland restorations there.
For crews administering prescribed fire burns, late March and early April are a busy time. The relative humidity is often at its lowest level of the year, so the probability of ignition of previous years’ growth is generally at its best. We visited with a crew administering a prescribed fire at Middle Creek Wildlife Management Area last week. Have a look…
Members of a Pennsylvania Game Commission burn crew provide visitors to Middle Creek Wildlife Management Area with an overview of prescribed fire and the equipment and techniques they use to conduct a burn.Pennsylvania Game Commission Southeast Region Forester Andy Weaver will fulfill the role of Burn Boss for administering this day’s dose of fire. His responsibilities include assessing the weather before the burn and calculating a probability of ignition.The Burn Boss briefs personnel with information on site layout, water supply location(s), places of refuge, emergency procedures, the event’s goals and plan of action, crew assignments, and the results of the weather check: wind from the northwest at 5 miles per hour, temperature 48 degrees, and the relative humidity 63%. Today’s patient is a parcel of warm-season grasses receiving a dose of fire to eliminate invasive non-native plants, woody growth, and thatch. The probability of ignition is 20%, but improving by the minute.To begin the burn, a test fire is started in the downwind corner of the parcel, which also happens to be the bottom of the slope. Fuel ignition is good. The burn can proceed.Crews proceed uphill from the location of the test fire while igniting combustibles along both flanks of the area being treated.A drip torch is used to ignite the dried stems and leaves of warm-season grasses and wildflowers. Each member of the burn crew wears Nomex fire-resistant clothing and carries safety equipment including a two-way radio, a hydration pack, and a cocoon-like emergency fire shelter.An all-terrain vehicle equipped with various tools, a fire pump, hose, and a small water tank accompanies the crew on each flank of the fire.A mowed strip of cool-season grasses along the perimeter of the burn area is already green and functions as an ideal fire break. While the drip torch is perfect for lighting combustibles along the fire’s perimeter, the paintball gun-looking device is an effective tool used to lob incendiaries into the center areas of the burn zone for ignition.With green cool-season grasses already growing on the trails surrounding the burn zone, very little water was used to contain this prescribed fire. Where such convenient fire breaks don’t already exist, crews carry tools including chain saws, shovels, and leaf blowers to create their own. They also carry flame swatters, backpack water pumps, shovels, and other tools to extinguish fires if necessary. None of these items were needed to control this particular fire.This fast-burning fire provides enough heat to damage the cambium layer of the woody tree and shrub saplings in this parcel being maintained as a grassland/wildflower plot, thus the process of succession is forestalled. Burns conducted during previous years on this and adjacent fields have also controlled aggressive growth of invasive Multiflora Rose and Olives (Elaeagnus species).Crews proceed up the slope while maintaining the perimeter by igniting dry plant material along the flanks of the burn zone.Ignition complete, the crews monitor the fire.The Burn Boss surveys the final stages of a safe and successful prescribed fire. The fire has left behind a mosaic of burned and unburned areas, just as a naturally occurring event may have done. Wildlife dodging the flames may be taking refuge in the standing grasses, so there is no remedial attempt to go back and ignite these areas. They’ll be burned during prescribed fires in coming years.By June, this grassland will again be lush and green with warm-season grasses and blooming wildflowers like this Common Milkweed being visited by a Great Spangled Fritillary.And later in the summer, Eastern Tiger Swallowtails on Joe-pye Weed.Indiangrass in flower in mid-summer.Bobolinks glow in the late August sun while taking flight from a stand of warm-season grasses maintained using springtime prescribed fire. The small dots on the dark background at the top of the image are multitudes of flying insects, many of them pollinators. The vegetation is predominately Indiangrass, excellent winter cover for birds, mammals, and other wildlife.
Prescribed burns aren’t a cure-all for what ails a troubled forest or other ecosystem, but they can be an effective remedy for deficiencies caused by a lack of periodic episodes of naturally occurring fire. They are an important option for modern foresters, wildlife managers, and other conservationists.
It’s that time of year. Your local county conservation district is taking orders for their annual tree sale and it’s a deal that can’t be beat. Order now for pickup in April.
The prices are a bargain and the selection includes the varieties you need to improve wildlife habitat and water quality on your property. For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue). And don’t forget to order packs of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local native birds. They’re only $12.00 for a bundle of 10.
It’s the most desirable block in town, not because the houses are any different from others built during the post-war years of the mid-twentieth century, but because the first owners of these domiciles had the good taste and foresight to plant long-lived trees on their lots, the majority of them native species. Pin Oak, Northern Red Oak, Yellow Poplar, Flowering Dogwood (Cornus florida), Eastern Red Cedar, Eastern White Pine, Eastern Hemlock, Norway Spruce, and American Holly dominate the landscape and create excellent habitat for birds and other wildlife. These 75-year-old plantings provide an abundance of shade in summer and thermal stability in winter, making it a “cool” place to live or take a stroll at any time of the year.
Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024
Cumberland County Conservation District is taking orders for Common Winterberry, the ideal small shrub for wet soil anywhere on your property. To get berries, you’ll need both males and females, so buy a bunch and plant them in a clump or scattered group.To live for a century or more like this towering giant, a Pin Oak needs to grow in well-drained soils with adequate moisture. These sturdy shade providers do well along streams and on low ground receiving clean runoff from hillsides, roofs, streets, and parking areas. As they age, Pin Oaks can fail to thrive and may become vulnerable to disease in locations where rainfall is not adequately infiltrated into the soil. Therefore, in drier areas such as raised ground or slopes, avoid the Pin Oak and select the more durable Northern Red Oak for planting. This year, Pin Oaks are available from the Cumberland and Lancaster County Conservation Districts, while Dauphin, Lancaster, Lebanon, and York Counties are taking orders for Northern Red Oaks.The Cumberland County Conservation District is again offering a “Showy Northeast Native Wildflower and Grass Mix” for seeding your own pollinator meadow or garden. It consists of more than twenty species including this perennial favorite, Purple Coneflower.
Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024
The Eastern Redbud is small tree native to our forest edges, particularly in areas of the Piedmont Province with Triassic geology (Furnace Hills, Conewago Hills, Gettysburg/Hammer Creek Formations, etc.) Also known as the Judas Tree, the redbud’s brilliant flowers are followed by heart-shaped leaves. As seen here, it is suitable for planting near houses and other buildings. Eastern Redbud seedlings are being offered through tree sales in Dauphin, Cumberland, and Lancaster Counties.
The Yellow Poplar, often called Tuliptree or Tulip Poplar for its showy flowers, is a sturdy, fast-growing deciduous tree native to forests throughout the Lower Susquehanna River Watershed. Its pole-straight growth habit in shady woodlands becomes more spreading and picturesque when the plant is grown as a specimen or shade tree in an urban or suburban setting. The Yellow Poplar can live for hundreds of years and is a host plant for the Eastern Tiger Swallowtail butterfly. It is available this year from the Lancaster County Conservation District.The American Sweetgum, also known as Sweet Gum, is a large, long-lived tree adorned with a mix of vibrant colors in autumn.Ever wonder where all the American Goldfinches and particularly the Pine Siskins go after passing through our region in fall? Well, many are headed to the lowland forests of the Atlantic Coastal Plain where they feed on an abundance of seeds contained in spiky American Sweetgum fruits. In the Piedmont and Ridge and Valley Provinces of the Lower Susquehanna River Watershed, American Sweetgum transplants can provide enough sustenance to sometimes lure our friendly finches into lingering through the winter.The American Sweetgum is a versatile tree. It can be planted on upland sites as well as in wet ground along streams, lakes, and rivers. In the beaver pond seen here it is the dominate tree species. This year, you can buy the American Sweetgum from the Lancaster County Conservation District.“Red-twig Dogwood” is a group of similar native shrubs that, in our region, includes Silky Dogwood and the more northerly Red-Osier Dogwood (Cornus sericea). Both have clusters of white flowers in spring and showy red twigs in winter. They are an excellent choice for wet soils. Landscapers often ruin these plants by shearing them off horizontally a foot or two from the ground each year. To produce flowers and fruit, and to preserve winter attractiveness, trim them during dormancy by removing three-year-old and older canes at ground level, letting younger growth untouched.“Red-twig Dogwoods” make ideal mass plantings for streamside buffers and remain showy through winter, even on a gloomy day. They not only mitigate nutrient and sediment pollution, they provide excellent food and cover for birds and other wildlife. Both Silky and Red-osier Dogwoods are available for sale through the Lancaster County Conservation District as part of their special multi-species offers, the former is included in its “Beauty Pack” and the latter in its “Wildlife Pack”. The similar Gray Dogwood (Cornus racemosa) is being offered for sale by the York County Conservation District.
The unique maroon flowers of the Common Pawpaw produce banana-like fruits in summer. These small native trees grow best in damp, well-drained soils on slopes along waterways, where they often form clonal understory patches. To get fruit, plant a small grove to increase the probability of pollination. The Common Pawpaw is a host plant for the Zebra Swallowtail butterfly. It is available through both the Lebanon and Lancaster County sales.The Eastern Red Cedar provides excellent food, cover, and nesting sites for numerous songbirds. Planted in clumps of dozens or groves of hundreds of trees, they can provide winter shelter for larger animals including deer and owls. The Eastern Red Cedar is being offered for purchase through both the Lebanon and Lancaster County Conservation Districts.Care to try your hand at raising some chestnuts? Lebanon County Conservation District has hybrid American Chestnut seedlings for sale.Lebanon County Conservation District is offering Common Winterberry and Eastern White Pine during their 2024 Tree and Plant Sale. Plant them both for striking color during the colder months. Eastern White Pine is also available from the Cumberland, Dauphin, Lancaster, and York County sales.
In addition to a selection of trees and shrubs, the Perry County Conservation District is again selling wildflower seed mixes for starting your own pollinator meadow or garden. For 2024, they have both a “Northeast Perennials and Annuals Mix” and a “Butterfly and Hummingbird Seed Mix” available. Give them a try so you can give up the mower!
Again this year, Perry County is offering bluebird nest boxes for sale. The price?—just $12.00.
Wait, what?,…twelve bucks,…that’s cheaper than renting!
The Buttonbush, a shrub of wet soils, produces a cosmic-looking flower. It grows well in wetlands, along streams, and in rain gardens. Buttonbush seedlings are for sale from both the York and Lancaster County Conservation Districts.
To get your deciduous trees like gums, maples, oaks, birches, and poplars off to a safe start, conservation district tree sales in Cumberland, Dauphin, Lancaster, and Perry Counties are offering protective tree shelters. Consider purchasing these plastic tubes and supporting stakes for each of your hardwoods, especially if you have hungry deer in your neighborhood.
Tree shelters protect newly transplanted seedlings from browsing deer, klutzy hikers, visually impaired mower operators, and other hazards.
There you have it. Be sure to check out each tree sale’s web page to find the selections you like, then get your order placed. The deadlines will be here before you know it and you wouldn’t want to miss values like these!
Since Tuesday’s snow storm, the susquehannawiildlife.net headquarters garden continues to bustle with bird activity.
Our Northern Mockingbird remains ever vigilant in its attempts to discourage American Robins and Eastern Bluebirds from feeding on the berry crop. Slowly, the latter species are winning the contest.A Carolina Chickadee carefully dissects a sunflower seed to snack on the nutritious kernel.This beauty shows us that yes, Red-bellied Woodpeckers do indeed have red bellies.Getting energized for a big move north, the robins keep on gulping berries.
Today, there arrived three species of birds we haven’t seen here since autumn. These birds are, at the very least, beginning to wander in search of food. Then too, these may be individuals creeping slowly north to secure an advantage over later migrants by being the first to establish territories on the most favorable nesting grounds.
This Song Sparrow is the first we’ve seen in the garden since sometime last fall. Is it working its way north or did it just come to town in search of food?Northern Flickers regularly spend the winter in small numbers in the Lower Susquehanna River Watershed. This is the first one we’ve had visit the garden since late last autumn.Fish Crows, seen here feeding on the fruits adorning an Eastern Red Cedar, have returned after being absent in our neighborhood since November. In coming weeks, both they and the more numerous American Crows that remained through winter will begin constructing nests in nearby trees.
They say the early bird gets the worm. More importantly, it gets the most favorable nesting spot. What does the early birder get? He or she gets out of the house and enjoys the action as winter dissolves into the miracle of spring. Do make time to go afield and marvel a bit, won’t you? See you there!
In mid-February each year, large numbers of American Robins descend upon the susquehannawildlife.net headquarters garden to feast on the ripe fruits that adorn several species of our native shrubs and trees. This morning’s wet snowfall provided the needed motivation for these birds and others to make today the big day for the annual feeding frenzy.
Early this morning, branches and limbs in the headquarters garden were loaded with clinging snow and more than one hundred American Robins.To have first grabs at suitable nesting sites, early American Robins are currently beginning to edge their way north. Spring migration is underway.The fruits of Common Winterberry are always a favorite of visiting robins.After selfishly guarding the garden’s berries through the entire season, our Northern Mockingbird finds chasing more than one hundred robins away from its food supply an impossible task.This and other visiting robins will strip the winterberry, cedar, American Holly, and other fruit-producing shrubs and trees within a day or two. To survive what remains of the season, our resident mockingbird will have to look elsewhere for provisions.Another American Robin devouring winterberry fruit.In addition to robins, there were, of course, other guests in the garden refuge on this snowy day.This Red-bellied Woodpecker tries to make sense of all the commotion.A pair of Carolina Chickadees established a family in the garden during the spring of 2023. At least five of the birds still stop by on a daily basis.As spring nears, our American Goldfinches are beginning to show a hint of their bright breeding colors.A Blue Jay peeks out from the cover of the Eastern Hemlocks.Our Carolina Wrens sing throughout the winter,……but today we noticed that this Mourning Dove has begun softly cooing to charm a mate……and the male House Finches are warbling away with the sounds of spring.With the local mockingbird busily harassing robins, our Eastern Bluebirds went unmolested long enough to stop by……for some raisins from their enclosed feeder.A showy male Eastern Bluebird on a snowy day in the garden. Spring must be just around the corner!
With the earth at perihelion (its closest approach to the sun) and with our home star just 27 degrees above the horizon at midday, bright low-angle light offered the perfect opportunity for doing some wildlife photography today. We visited a couple of grasslands managed by the Pennsylvania Game Commission to see what we could find…
On this State Game Lands parcel, prescribed fire is used to maintain a mix of grasslands and brushy early successional growth. In nearby areas, both controlled fire and mechanical cutting are used to remove invasive species from hedgerows and the understory of woodlots. Fire tolerant native species then have an opportunity to recolonize the forest and improve wildlife habitat. This management method also reduces the fuel load in areas with the potential for uncontrolled wildfires.The sun-dried fruits of a native Common Persimmon tree found growing in a hedgerow.Just one year ago, mechanical removal of invasive trees and shrubs (including Multiflora Rose) on this State Game Land was followed by a prescribed fire to create this savanna-like grassland.Hundreds of Song Sparrows were found in the grasses and thickets at both locations.White-throated Sparrows were also abundant, but prefer the tangles and shrubs of the thickets.Northern Mockingbirds were vigilantly guarding winter supplies of berries in the woodlots and hedgerows.In grasses and tangles on wetter ground, about a dozen Swamp Sparrows were discovered.The adult White-crowned Sparrow is always a welcome find.And seeing plenty of juvenile White-crowned Sparrows provides some assurance that there will be a steady stream of handsome adult birds arriving to spend the winter during the years to come.Dark-eyed Juncos were encountered only in the vicinity of trees and large shrubs.Several Savannah Sparrows were observed. Though they’re mostly found in treeless country, this particular one happened to pose atop a clump of shrubs located within, you guessed it, the new savanna-like grasslands.A tiny bird, even when compared to a sparrow, the Winter Wren often provides the observer with just a brief glimpse before darting away into the cover of a thicket.Within grasslands, scattered stands of live and dead timber can provide valuable habitat for many species of animals.Woodpeckers and other cavity-nesting birds rely upon an abundance of “snags” (standing dead trees) for breeding sites.This Red-bellied Woodpecker and about a dozen others were found in trees left standing in the project areas.A Yellow-bellied Sapsucker soaks up some sun.This very cooperative Pileated Woodpecker seemed to be preoccupied by insect activity on the sun-drenched bark of the trees. This denizen of mature forests will oft times wander into open country where larger lumber is left intact.
Just as things were really getting fun, some late afternoon clouds arrived to dim the already fading daylight. Just then, this Northern Harrier made a couple of low passes in search of mice and voles hidden in the grasses.It was a fitting end to a very short, but marvelously sunny, early winter day.
A glimpse of the rowdy guests crowding the Thanksgiving Day dinner table at susquehannawildlife.net headquarters…
A male White-breasted Nuthatch visits a peanut feeder……soon to be joined by a female White-breasted Nuthatch.A male Downy Woodpecker gets a bill full of suet.A Carolina Wren nibbles at a peanut.An Eastern Gray Squirrel stuffs itself on peanuts dropped by the birds.A territorial Northern Mockingbird stands guard over its supply of Common Winterberry fruit.To avoid the mockingbird’s aggression, the Eastern Bluebirds opted out of fresh fruit in favor of raisins offered at the feeders.This persistent American Robin has made an art of repeatedly sneaking in to quickly devour a few berries before being chased away by the vigilant mockingbird.After everyone has had their fill, Dark-eyed Juncos clean up the leftovers.
Among the hardy wildflowers still in bloom in the forests of the lower Susquehanna valley is the Blue Wood Aster (Symphyotrichum cordifolium), a variable species also known as the Heart-leaved Aster.
Where should you go this weekend to see vibrantly colored foliage in our region? Where are there eye-popping displays of reds, oranges, yellows, and greens without so much brown and gray? The answer is Michaux State Forest on South Mountain in Adams, Cumberland, and Franklin Counties.
South Mountain is the northern extension of the Blue Ridge Section of the Ridge and Valley Province in Pennsylvania. Michaux State Forest includes much of the wooded land on South Mountain. Within or adjacent to its borders are located four state parks: King’s Gap Environmental Education Center, Pine Grove Furnace State Park, Caledonia State Park, and Mont Alto State Park. The vast network of trails on these state lands includes the Appalachian Trail, which remains in the mountainous Blue Ridge Section all the way to its southern terminus in Georgia.
In Pennsylvania, the forested highlands of the Blue Ridge Section of the Ridge and Valley Province are known as South Mountain. Much of South Mountain lies within the boundaries of Michaux State Forest. Stars indicate the locations of 1) King’s Gap Environmental Education Center, 2) Pine Grove Furnace State Park, 3) Caledonia State Park, and 4) Mont Alto State Park. A drive on US 30 between Gettysburg and Chambersburg will take you right through Michaux State Forest along an east to west axis while a scenic northbound or southbound trip along PA 233 will bring you in proximity to each of the state parks located therein. (Base image from NASA Earth Observatory Collection)
If you want a closeup look at the many species of trees found in Michaux State Forest, and you want them to be labeled so you know what they are, a stop at the Pennsylvania State University’s Mont Alto arboretum is a must. Located next door to Mont Alto State Park along PA 233, the Arboretum at Penn State Mont Alto covers the entire campus. Planting began on Arbor Day in 1905 shortly after establishment of the Pennsylvania State Forest Academy at the site in 1903. Back then, the state’s “forests” were in the process of regeneration after nineteenth-century clear cutting. These harvests balded the landscape and left behind the combustible waste which fueled the frequent wildfires that plagued reforestation efforts for more than half a century. The academy educated future foresters on the skills needed to regrow and manage the state’s woodlands.
Online resources can help you plan your visit to the Arboretum at Penn State Mont Alto. More than 800 trees on the campus are numbered with small blue tags. The “List of arboretum trees by Tag Number” can be downloaded to tell you the species or variety of each. The interactive map provides the locations of individual trees plotted by tag number while the Grove Map displays the locations of groups of trees on the campus categorized by region of origin. A Founder’s Tree Map will help you find some of the oldest specimens in the collection and a Commemorative Tree Map will help you find dedicated trees. There is also a species list of the common and scientific tree names.
The Yellow Buckeye (Aesculus flava) is a tree found in the forests of the Blue Ridge Section of the Ridge and Valley Province in West Virginia, Virginia, North Carolina and Georgia. You can see it in Pennsylvania by visiting the collection of trees in the Arboretum at Penn State Mont Alto.The American Chestnut can be difficult to find due to the impact of chestnut blight, but you can see it in the Arboretum at Penn State Mont Alto.Shagbark Hickory is a common tree in the forests of South Mountain.The Sweet Cherry (Prunus avium), a native of Europe, is naturalized throughout eastern and south-central Pennsylvania and is one of the more than 150 species of trees in the arboretum’s collection.Sweet Birch (Betula lenta) foliage is particularly bright yellow on South Mountain this autumn. It really “pops” against the backdrop of the evergreen Eastern White Pines and Eastern Hemlocks. During less-than-ideal years, Sweet Birch leaves can subtly transition from green to drab brown without much fanfare before falling.You might have a difficult time finding a Common Persimmon (Diospyros virginiana) growing wild in Pennsylvania, but you can find it in the Arboretum at Penn State Mont Alto.
The autumn leaves will be falling fast, so make it a point this weekend to check out the show on South Mountain.
Last October 3rd, a late-season Ruby-throated Hummingbird stopped by the garden at susquehannawildlife.net headquarters to take shelter from a rainy autumn storm. It was so raw and chilly that we felt compelled to do something we don’t normally do—put out the sugar water feeder to supplement the nectar produced by our fall-flowering plants. After several days of constant visits to the feeder and the flowers, our lingering hummer resumed its southbound journey on October 7th.
Fast forward to this afternoon and what do you know, at least two migrating hummingbirds have stopped by to visit the flowers in our garden. This year, we have an exceptional abundance of blooms on some of their favorite plants. In the ponds, aquatic Pickerelweed is topped with purple spikes and we still have bright orange tubular flowers on one of our Trumpet Vines—a full two to three months later than usual.
We checked each of our late-season visitors carefully to be reasonably certain that none was a stray western species of hummingbird. All appear to be female or juvenile “Ruby-throats”. If you have an abundance of flowering plants and/or you’re going to maintain your hummingbird feeders through the coming weeks, be on the lookout for western species. Most are more hardy than our Ruby-throats and some have remained in the lower Susquehanna valley through the winter.As is typically the case, Ruby-throated Hummingbirds quickly gravitate toward the tubular flowers of our Cuphea ignea, the Mexican Cigar. They find these showy plants to be absolutely irresistible.Mexican Cigar grows wild in parts of the Ruby-throated Hummingbird’s winter range. To them, it’s comfort food.We’re trying some new cultivars of Cuphea to see how they do. As this composite image shows, the hummingbirds won’t let our Bat-faced Cuphea (Cuphea llavea) alone. It’s another plant native to Mexico and Central America, right where some of our hummingbirds spend the winter.
Remember, keep those feeders clean and the provisions fresh! You’ll be glad you did.
What’s all this buzz about bees? And what’s a hymanopteran? Well, let’s see.
Hymanoptera—our bees, wasps, hornets and ants—are generally considered to be our most evolved insects. Some form complex social colonies. Others lead solitary lives. Many are essential pollinators of flowering plants, including cultivars that provide food for people around the world. There are those with stingers for disabling prey and defending themselves and their nests. And then there are those without stingers. The predatory species are frequently regarded to be the most significant biological controls of the insects that might otherwise become destructive pests. The vast majority of the Hymanoptera show no aggression toward humans, a demeanor that is seldom reciprocated.
Late summer and early autumn is a critical time for the Hymanoptera. Most species are at their peak of abundance during this time of year, but many of the adult insects face certain death with the coming of freezing weather. Those that will perish are busy, either individually or as members of a colony, creating shelter and gathering food to nourish the larvae that will repopulate the environs with a new generation of adults next year. Without abundant sources of protein and carbohydrates, these efforts can quickly fail. Protein is stored for use by the larval insects upon hatching from their eggs. Because the eggs are typically deposited in a cell directly upon the cache of protein, the larvae can begin feeding and growing immediately. To provide energy for collecting protein and nesting materials, and in some cases excavating nest chambers, Hymanoptera seek out sources of carbohydrates. Species that remain active during cold weather must store up enough of a carbohydrate reserve to make it through the winter. Honey Bees make honey for this purpose. As you are about to see, members of this suborder rely predominately upon pollen or insect prey for protein, and upon nectar and/or honeydew for carbohydrates.
We’ve assembled here a collection of images and some short commentary describing nearly two dozen kinds of Hymanoptera found in the Lower Susquehanna River Watershed, the majority photographed as they busily collected provisions during recent weeks. Let’s see what some of these fascinating hymanopterans are up to…
SOLITARY WASPS
A Great Black Wasp on goldenrod (Solidago species). Like other solitary wasps, a female Great Black Wasp will sting and paralyze a host insect upon which she’ll deposit her eggs. After hatching, the larvae will begin consuming the host’s body as a source of protein. The parasitized insects are often katydids or grasshoppers.A Great Black Wasp feeding on nectar, a source of carbohydrates. Unlike social bees and wasps, solitary wasps are equipped with a stinger solely used for immobilizing prey, not defending a nest. They are therefore quite docile and pose little threat to humans.A Great Black Wasp powdered with pollen. Hymanopterans that gather nectar and/or pollen are tremendously important pollinators of hundreds of species of plants.A female Thread-waisted wasp (Ammophilia species, probably A. nigricans) drags a paralyzed caterpillar to her excavated nest where she’ll deposit an egg on the body. After hatching, the larval wasp will feed on the disabled caterpillar. The protein will enable the larvae to grow, pupate, and later emerge as an adult wasp.The female Eastern Cicada Killer (Sphecius speciosus) excavates an underground nest with branch tunnels connecting a dozen chambers or more. As the common name suggests, the female wasp paralyzes a cicada, then makes a strenuous effort to fly and drag it back to the nest for placement in a cell. Each male wasp egg is deposited upon just one immobilized cicada, but a female egg is provided with a cache of several cicadas to provide adequate protein for growth to a larger size. Nest cells are sealed with soil, then the larvae hatch in just a couple of days. Within about two weeks, they have consumed the cicada protein and are fully grown. Wrapped in a cocoon, they spend the winter in the nest, then pupate in the spring before emerging as a new generation of adults.The Black-and-yellow Mud Dauber (Sceliphron caementarium) builds a mud-ball nest within which it packs paralyzed spiders to function as a source of protein for its larvae.A Black-and-yellow Mud Dauber at nest.The Pipe Organ Mud Dauber builds this elaborate nest in which their eggs and paralyzed spiders are deposited in cells sealed with mud partitions. After consuming the spiders, the larvae pupate, overwinter, then emerge from their cells as adults during the following spring. To escape the protection of the nest, the new generation of adults bore through the mud walls. Adult Pipe Organ Mud Daubers resemble the Great Black Wasp, but have a white or yellow distal segment on their rear legs resembling a pair of light-colored socks.A closeup of the previous image with the lengths of the nest tubes compressed to show four scavenger flies (Miltogramminae), possibly two species, that have invaded this Pipe Organ Mud Dauber nest. Scavenger flies are kleptoparasites that victimize various solitary bees and wasps, depositing larvae directly into the host species’ nest cells to consume the protein cache stored therein.
CUCKOO WASPS
Cuckoo Wasps (Chrysididae), also known as Emerald Wasps, parasitize the nests of other species of wasps. Females lay their eggs inside the host’s nest, then flee the scene. Upon hatching, larval Cuckoo Wasps feed on stockpiles of prey intended for the host species’ offspring. Like the adult mud daubers that have already matured and departed this nest by digging a hole through the wall of the cell within which they were hatched, the metallic green Cuckoo Wasp in the upper left has just emerged in much the same way.
SWEAT BEES
A Sweat Bee (Lasioglossum species) collecting nectar and pollen on White Snakeroot (Ageratina altissima).Sweat Bees (Lasioglossum species) visit human skin to lick up the electrolytes left behind by evaporating perspiration.Sweat Bees in the genus Lasioglossum demonstrate various social behaviors ranging from species that are solitary nesters to those that create colonies with work forces ranging in size from as few as four to as many as hundreds of bees. Some Lasioglossum practice kleptoparasitism, while others are quite accomplished foragers.A female Augochlorine Green Sweat Bee (Augochlorini) collecting nectar on White Snakeroot. Notice the pollen “baskets” on the rear leg.An Augochlorine Green Sweat Bee (Augochlorini). Sweat bees nest in subterranean cavities and in hollowed out sections of trees.A copper-colored Augochlorine Green Sweat Bee (Augochlorini) collecting nectar and dusted with pollen.
LEAFCUTTER AND MASON BEES
A Leafcutter Bee (Megachile species). Like Mason Bees, female Mason Bees deposit each of their eggs on a “pollen loaf” within an individual cell inside a preexisting tunnel-like cavity in wood, stone, or in the ground. Unlike Mason Bees, female Leafcutter Bees cut a circular piece of leaf to create each of the cells in their nest. After hatching, the larval bee feeds on the pollen loaf, pupates, then emerges from the shelter of the nest to start a new generation, usually during the following year.A Leafcutter Bee (Megachile species) visiting Wild Bergamot. Female Leafcutter and Mason Bees lack pollen “baskets” on their rear legs but instead have pollen “brushes” on the underside of the abdomen to gather the protein they need to create a “pollen loaf” for each nest cell.A Leafcutter Bee (Megachile species) collecting nectar from White Snakeroot.A Mason Bee (Osmia species) emerging from a nest in spring. Mason Bees create nesting cells within preexisting cavities in wood, stone, and other other supporting structures. Within the nest cavity, each egg is deposited atop a cache of pollen and nectar, a pollen loaf, then enclosed behind a partition of mud. The female Mason Bee will usually repeat this process until an entire cavity is filled with cells. During the following spring, a new generation of adult Mason Bees digs its way through the cell walls to emerge and repeat the process. These bees readily use paper straws or holes drilled in blocks of wood for nesting.A mason bee nest box with holes drilled into blocks of wood.Mason Bees seal each cell and the outer end of their nest cavity with mud. These outer nest cells can been parasitized by a variety of wasps. Here, the outer cell of a Mason Bee nest has been victimized by a tiny chalcid wasp (looks like another one to the lower left). Several species of female chalcid wasps (native Monodontomerus species or non-native Pteromalus venustus) enlarge weak points in the outer partition of a mason bee nest, then sting and paralyze the larval bee inside before depositing their eggs. Within the cell. the wasp larvae consume the larval Mason Bee and the “pollen loaf” provided for its growth. These same parasitic wasps prey upon Leafcutter Bees as well.
BUMBLE BEES, CARPENTER BEES, HONEY BEES, AND DIGGER BEES
A Common Eastern Bumble Bee (Bombus impatiens) collecting nectar and pollen on goldenrod. Bumble bees are our sole native group of social bees. Their wax nests are built in a burrow or other shelter. The eggs are deposited in cells along with a supply of pollen for nourishing the larvae upon hatching. Honey is stored in “honey pots” within the nest. New queens are produced along with male bees during the late-summer and fall. Only the new generation of fertilized queens survive the winter to lay eggs and produce workers to construct a new nest.A pair of Common Eastern Bumble Bees collecting nectar and becoming dusted with pollen. Their fuzzy coats and semi-warm-blooded metabolism allows them to be active in cooler weather than is tolerated by other bees.Flowering plants including the Great Rhododendron find success attracting pollinators to their reproductive blossoms by offering carbohydrate-rich nectar to insects like this Eastern Bumble Bee. The yellow spots on the flower’s upper petal help to guide visitors toward their sweet treat.An Eastern Carpenter Bee feeding on goldenrod nectar. Compare the almost hairless abdomen to that of the bumble bees. Carpenter bees are semi-social insects. Females lay their eggs in cells within galleries bored into wood. These nests are completed with great precision, avoiding creation of any second entrance by mistakenly breaching the outer surface of the excavated wood. Each egg/larvae is provided with a supply of protein-rich pollen. Males often hover outside their mate’s nest to prevent competing males from entering the area.A worker Honey Bee, a female member of a sisterhood of foragers from a nearby hive, visits goldenrod alongside Common Eastern Bumble Bees. Honey Bees were brought to North America during the 1620s, the earliest years of the trans-Atlantic migration of European colonists, to pollinate cultivated plants and to provide a reliable source of honey and beeswax. Within the Honey Bee’s social structure, the queen of each hive lays the eggs to produce the female worker bees. Once each year, male drones are produced along with a new generation of queens.In nature, Honey Bees build hives in tree cavities. Recently, this colony constructed a hive in a screech owl nest box at susquehannawildlife.net headquarters. To provide protein for the hatching larvae, worker bees collect pollen and deposit it within the hexagonal cells of the vertically aligned beeswax combs. After an egg is deposited upon the pollen cache, each cell is sealed with more beeswax. Young females tend these nest combs before maturing and becoming foraging worker bees.In apiculture, Honey Bees are raised in man-made hives. This Pennsylvania Association of Beekeepers display gives visitors to the Pennsylvania Farm Show in Harrisburg a look at the inner workings of a live bee hive. Nectar collected by worker bees is turned into honey to provide the supply of carbohydrates needed to fuel the colony through the winter. Note the honeycombs on the glass.A possible Small Carpenter Bee (Ceratina species) visiting White Snakeroot. Small Carpenter Bees nest inside hollow stems and twigs. Some species are eusocial, with a queen’s daughters and sisters sharing responsibility for finding food and rearing the young. Females overwinter inside a one of the excavated stems and begin a new nest there in the spring.A Digger Bee (possibly Melissodes species) with “pollen baskets” full of pollen collected from nearby flowers. Digger Bees in the genus Melissodes are often known as the Long-horned Bees. These social insects excavate underground nests and many species practice communal living.
SCOLIID WASPS
The Two-spotted Scoliid Wasp (Scolia dubia), also known as the Blue-winged Scoliid Wasp, is most frequently observed feeding on nectar. Scoliid wasps are solitary nesters, though they may assemble into groups while visiting flowers. They often ignore the presence of humans and are seldom disturbed by their presence. Females seek out the burrowing grubs of beetles including the Green June Bug (Cotinis nitida) and possibly the Japanese Beetle. After stinging a grub to paralyze it, the wasp will deposit her egg on its body, then bury it. Upon hatching, the larval wasp will feed on the grub for nourishment as it grows.Don’t like having your watermelon overrun by Green June Bugs while you’re eating? Then you ought to go out of your way to be nice to the Two-spotted Scoliid Wasp.The Double-banded Scoliid (Scolia bicincta) parasitizes beetle larvae as hosts for its larvae. For carbohydrates it relishes flower nectar.
PAPER WASPS
A Northern Paper Wasp (Polistes fuscatus). Paper wasps prey upon numerous garden pests, particularly caterpillars, to collect protein. Though they are social insects equipped with stingers to subdue their victims and defend their nests, paper wasps are surprisingly docile.A Northern Paper Wasp (Polistes fuscatus) feeding on nectar from a goldenrod flower.A Northern Paper Wasp harvesting wood pulp from the side of a mason bee nest box at susquehannawildlife.net headquarters. The pulp is chewed in the wasp’s saliva to create the paper used to construct the colony’s open-cell nest.Common Paper Wasps (Polistes exclamans), also known as Guinea Paper Wasps, at their open-cell nest. This and the nests of most other paper wasps are suspended on a filament or a pedicle. Many paper wasps can excrete an ant repellent on this section of the nest in an effort to prevent invasion. Like many other social hymenopterans, a defending wasp can secrete a pheromone venom during the stinging process to warn the colony of danger at the nest. In winter, Common Paper Wasps seek shelter in stumps and other locations to hibernate.The European Paper Wasp (Polistes dominula) is a non-native species which builds nests in man-made structures including bird houses. To collect protein, they prey on a wide selection of insects and other invertebrates. As such, European Paper Wasps are widespread and successful here in North America.
YELLOWJACKETS AND HORNETS
An Eastern Yellowjacket feeding on lanternfly honeydew. Eastern Yellowjackets derive much of their success from being generalists, collecting carbohydrates from nearly any sweet source, natural or man made. They are quite fond of ripe fruits, flower nectar, and sugary snacks and drinks, especially soda. Protein for nourishing their larvae is derived from the wide variety invertebrates upon which they prey and from carrion. These foods are chewed into a paste form in preparation for placement into the brood cells.A subterranean colony of Eastern Yellowjackets is started anew each spring by a young queen that has survived winter hibernation in diapause, a state of interrupted development. She constructs the new nest’s first cells using pulp made by chewing rotting wood. The first brood of workers scales up construction while the queen continues producing eggs. At the nest, these social insects will viciously attack anyone or anything perceived to be a threat, so give them their space and leave them alone. Many yellowjacket infestations of homes and other buildings are the work of non-native German Yellowjacket (Vespula germanica) [not shown], an invasive species that constructs paper nests in void spaces including walls and attics.Yellowjackets may be moody and aggressive, but they do fall victim to a number of predators. A Robber Fly (Promachus species) has taken down and is devouring this Eastern Yellowjacket.A Bald-faced Hornet (Dolichovespula maculata) feeding on Spotted Lanternfly honeydew on a Tree-of-heaven (Ailanthus altissima). In the absence of nectar-producing flowers, many bees, yellowjackets, and hornets have turned to the invasive lanternfly and Ailanthus combo to turn the sun’s energy into the carbohydrates they need. For protein, they prey upon spiders, flies, caterpillars, and a variety of other insects.To create paper for nest construction, this Bald-faced Hornet is collecting wood pulp from the surface of a weathered picnic table. Away from the nest, these hornets demonstrate a calm, carefree demeanor and can be closely observed.A Bald-faced Hornet nest in a pine tree. These hives are strictly temporary. Within the nest, a generation of drones (males) and new queens are produced late each year. These wasps leave the colony to mate. With the arrival of freezing weather, all inhabitants within the nest, including the old queen, perish, as do the drones that departed to breed. Only the new queens survive winter hibernation to propagate the next generation of wasps, starting with the workers needed to construct a fresh nest and reestablish the colony.Did you ever get the feeling you’re being watched? Don’t go messing around with Bald-faced Hornet nests. The occupants therein, like other social bees, wasps, and hornets, are equipped with stingers and venom for defending their colony. This is an adaptation that has developed over time to assure the survival of populations of these insects. Think about it this way, a solitary wasp that loses a nest loses only their individual brood of offspring. There is minimal impact on the wider local population of such insects. Conversely, a social wasp or hornet that loses a nest loses an entire colony, possibly negating the benefits of their cooperative behavior and threatening the survival of the species. Insects that cooperate to build societies for survival can be more vulnerable to the catastrophic impacts of certain circumstances like disease, weather, and invasion of their colonies. Therefore, natural selection has provided them with contingencies for these dangers, for example, the instinct to construct protective shelters and the adaptation of stingers and venom for defense against intruders and would-be predators. Oh, and by the way, the Bald-faced Hornet can spray venom, often aiming for the eyes, so keep your distance.European Hornets (Vespa crabro), an introduced species, are predatory on a variety of flying insects for protein. For carbohydrates they are attracted to sweets like this lanternfly honeydew on Tree-of-heaven.European Hornets constructing a paper nest in a tree cavity.
POTTER WASPS
A potter wasp (Eumenes species), probably a Fraternal Potter Wasp (E. fraternus), hovering near a European Paper Wasp on Partridge Pea. The female potter wasp builds a small mud nest resembling a tiny clay pot. One of her eggs is inserted and left hanging on a thin thread. Then a paralyzed caterpillar is deposited as a source of protein to nourish the larva upon hatching. Lastly, the pot is sealed with a lid made of wet mud. Upon maturing, the new generation of adult wasps perform a pottery breaking to emerge and take flight.
ANTS
Field Ants (Formica species, possibly Formica pallidefulva) clear the entrance to their underground nest. Field ants are eusocial insects, they work in concert to build, maintain, and defend the nest, rear young, and find food. There is no social caste system. Field Ants are predators and scavengers when collecting protein. For carbohydrates they often rely on the honeydew produced by aphids. As a method of improving and sustaining the production of honeydew, some ant species will tend colonies of aphids by moving the younger individuals from depleted portions of plants to more healthy tissue. Field Ant nests contain chambers used for a variety of functions including raising young and storing food. Some nests include multiple queens and some colonies consist of more than one nest. Ants in the genus Formica are weaponized; they can spray formic acid to repel intruders and defend their colony.
We hope this brief but fascinating look at some of our more common bees, wasps, hornets, and ants has provided the reader with an appreciation for the complexity with which their food webs and ecology have developed over time. It should be no great mystery why bees and other insects, particularly native species, are becoming scarce or absent in areas of the Lower Susquehanna River Watershed where the landscape is paved, hyper-cultivated, sprayed, mowed, and devoid of native vegetation, particularly nectar-producing plants. Late-summer and autumn can be an especially difficult time for hymanopterans seeking the sources of proteins and carbohydrates needed to complete preparations for next year’s generations of these valuable insects. An absence of these staples during this critical time of year quickly diminishes the diversity of species and begins to tear at the fabric of the food web. This degradation of a regional ecosystem can have unforeseen impacts that become increasingly widespread and in many cases permanent.
How can anyone be surprised by the absence of bees and other pollinators in farmland? Manicured and cultivated ground offers little in the way of year-round shelter and food sources for insects and other wildlife.This savanna-like habitat on a south-facing slope provides the abundance of nectar-producing, pollen-rich wildflowers needed to nourish a diverse population of insects including bees, wasps, hornets, and ants. Goldenrods, asters, and White Snakeroot are some of their late-season favorites.
Editor’s Note: No bees, wasp, hornets, or ants were harmed during this production. Neither was the editor swarmed, attacked, or stung. Remember, don’t panic, just observe.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York, NY.
To pass the afternoon, we sat quietly along the edge of a pond created recently by North American Beavers (Castor canadensis). They first constructed their dam on this small stream about five years ago. Since then, a flourishing wetland has become established. Have a look.
Vegetation surrounding the inundated floodplain helps sequester nutrients and sediments to purify the water while also providing excellent wildlife habitat.The beaver lodge was built among shrubs growing in shallow water in the middle of the pond.Woolgrass (Scirpus cyperinus) is a bulrush that thrives as an emergent and as a terrestrial plant in moist soils bordering the pond.A male Common Whitetail dragonfly keeping watch over his territory.A Twelve-spotted Skimmer perched on Soft Rush.A Blue Dasher dragonfly seizing a Fall Field Cricket (Gryllus pennsylvanicus).A Spicebush Swallowtail visiting a Cardinal Flower.A Green Heron looking for small fish, crayfish, frogs, and tadpoles.The Green Heron stalking potential prey.A Wood Duck feeding on the tiny floating plant known as Lesser Duckweed (Lemna minor).A Least Sandpiper poking at small invertebrates along the muddy edge of the beaver pond.A Solitary Sandpiper.A Solitary Sandpiper testing the waters for proper feeding depth.A Pectoral Sandpiper searches for its next morsel of sustenance.The Sora (Porzana carolina) is a seldom seen rail of marshlands including those created by North American Beavers. Common Cattails, sedges, and rushes provide these chicken-shaped wetland birds with nesting and loafing cover.
Isn’t that amazing? North American Beavers build and maintain what human engineers struggle to master—dams and ponds that reduce pollution, allow fish passage, and support self-sustaining ecosystems. Want to clean up the streams and floodplains of your local watershed? Let the beavers do the job!
“Waves” of warblers and other Neotropical songbirds continue to roll along the ridgetops of southern Pennsylvania. The majority of these migrants are headed to wintering habitat in the tropics after departing breeding grounds in the forests of southern Canada. At Second Mountain Hawk Watch, today’s early morning flight kicked off at sunrise, then slowed considerably by 8:30 A.M. E.D.T. Once again, in excess of 400 warblers were found moving through the trees and working their way southwest along the spine of the ridge. Each of the 12 species seen yesterday were observed today as well. In addition, there was a Northern Parula and a Canada Warbler. Today’s flight was dominated by Bay-breasted, Blackburnian, Black-throated Green, and Tennessee Warblers.
A Blackburnian Warbler at sunrise on Second Mountain.A hungry Blackburnian Warbler feeding on insects.Black-throated Green Warblers were a plentiful species among both yesterday’s and today’s waves of Neotropical migrants.A juvenile Black-throated Green Warbler.One of the scores of Tennessee Warblers seen on Second Mountain early this morning.Cape May Warblers were still common today, but not moving through in the numbers seen yesterday.A male Black-throated Blue Warbler.Compared to yesterday’s flight, lesser numbers of Magnolia Warblers were seen today.An adult male Wilson’s Warbler was a good find among the hundreds of birds swarming the ridgetop.This Nashville Warbler spent much of the day in the tangles of Mile-a-minute Weed surrounding the lookout.
Other interesting Neotropical migrants joined the “waves” of warblers…
Red-eyed Vireo numbers were higher than yesterday.This Warbling Vireo was found peering from the cover of the shady forest.A minimum of six Rose-breasted Grosbeaks were identified including the juvenile male seen here in first-fall plumage. Other good sightings were Scarlet Tanagers, an adult male Baltimore Oriole, and a dozen or more Ruby-throated Hummingbirds.Three Least Flycatchers were heard calling and seen chasing one another through a stand of dead timber on the south slope below the lookout.After the warbler flight settled, the task of counting migrating raptors commenced. Five Broad-winged Hawks including this one were tallied as they glided away to the southwest for a winter vacation in the tropics of Central and South America.
It may be one of the most treasured plants among native landscape gardeners. The Cardinal Flower (Lobelia cardinalis) blooms in August each year with a startling blaze of red color that, believe it or not, will sometimes be overlooked in the wild.
Cardinal Flower is most often found in wet soil along forested bodies of water. The blooms of this shade-loving species may go unnoticed until rays of sunshine penetrate the canopy to strike their brilliant red petals.
The Cardinal Flower grows in wetlands as well as in a variety of moist soils along streams, rivers, lakes, and ponds. Shady locations with short periods of bright sun each day seem to be favored for an abundance of color.
Cardinal Flower in bloom in a riparian forest along the Susquehanna. To its right is its close relative, Great Lobelia, a plant sometimes known as Great Blue Lobelia or Blue Cardinal Flower.Cardinal Flower in a wet bottomland woods.The Cardinal Flower can find favorable growing conditions along stream, river, and lake shores. Even though they are perennial plants, their presence along such waters often seems temporary. Changing conditions cause them to suddenly disappear from known locations, then sometimes reappear at the same place or elsewhere nearby. Some of this phenomenon may be due to the fact that stressed plants can fail to bloom, so they easily escape notice. When producing flowers during favorable years, the plants seem to mysteriously return.Cardinal Flowers along a wave-swept shoreline light up the greenery of erosion-controlling riparian vegetation with glowing red color.
The Cardinal Flower can be an ideal plant for attracting hummingbirds, bees, butterflies, and other late-summer pollinators. It grows well in damp ground, especially in rain gardens and along the edges streams, garden ponds, and stormwater retention pools. If you’re looking to add Cardinal Flower to your landscape, you need first to…
REMEMBER the CARDINAL RULE…
Cardinal Flower plants are available at many nurseries that carry native species of garden and/or pond plants. Numerous online suppliers offer seed for growing your own Cardinal Flowers. Some sell potted plants as well. A new option is to grow Cardinal Flowers from tissue cultures. Tissue-cultured plants are raised in laboratory media, so the pitfalls of disease and hitchhikers like invasive insects and snails are eliminated. These plants are available through the aquarium trade from most chain pet stores. Though meant to be planted as submerged aquatics in fish tank substrate, we’ve reared the tissue-cultured stock indoors as emergent plants in sandy soil and shallow water through the winter and early spring. When it warms up, we transplant them into the edges of the outdoor ponds to naturalize. As a habit, we always grow some Cardinal Flower plants in the fish tanks to take up the nitrates in the water and to provide a continuous supply of cuttings for starting more emergent stock for outdoor use.
A tissue-cultured Cardinal Flower rooted in sandy substrate and being grown as a submerged aquatic plant in a fish tank. Cuttings from this plant will be used to grow emergent specimens in shallow water for transplanting outdoors around the garden pond.A Cardinal Flower grown from an aquarium store tissue culture blooms in the pond at susquehannawildlife.net headquarters.Grown as an emergent, Cardinal Flower may bloom very late in the season. This tissue-cultured specimen in the headquarters pond was photographed in early November, 2022.
We’ve got the summertime blues for you, right here at susquehannawildlife.net…
In warm-season grass meadows, Big Bluestem is now in flower. This and other species of native prairie grasses provide excellent habitat for birds, mammals, and insects including butterflies. To survive drought and fire, their roots run much deeper than cool season grasses, creeping down four to six feet or more. This adaptation allowed warm season grasses to recover from heavy grazing by large Pre-Anthropocene mammals. Today, it makes them ideal plants for soil stabilization.A male Indigo Bunting has already found ripe seeds among the heads of flowering Big Bluestem.Look closely and you’ll see our Indigo Bunting is beginning a pre-migration molt out of its bright-blue breeding (alternate) plumage and into a gray-brown winter (basic) plumage. The berries of the American Pokeweed upon which it is perched will soon ripen into a dark blue, almost black, color. Though toxic to humans, these fruits find favor with many species of birds and mammals.Another great wildlife food is Silky Dogwood (Cornus amomum), a deciduous shrub that sports blue-colored berries in summer and showy, bright-red twigs in winter. It grows well in wet ground along streams and ponds, as well as in rain gardens.A Great Blue Heron searches the shallows for small fish. This species is also a good mouser, at times seen hunting in grassy meadows. Right now is prime time to see it and a variety of other herons and egrets throughout the Lower Susquehanna River Watershed.
…so don’t let the summertime blues get you down. Grab a pair of binoculars and/or a camera and go for a stroll!
If you’re feeling the need to see summertime butterflies and their numbers just don’t seem to be what they used to be in your garden, then plan an afternoon visit to the Boyd Big Tree Preserve along Fishing Creek Valley Road (PA 443) just east of U.S. 22/322 and the Susquehanna River north of Harrisburg. The Pennsylvania Department of Conservation and Natural Resources manages the park’s 1,025 acres mostly as forested land with more than ten miles of trails. While located predominately on the north slope of Blue Mountain, a portion of the preserve straddles the crest of the ridge to include the upper reaches of the southern exposure.
A grove of American Chestnuts (Castanea dentata) planted at Boyd Big Tree Preserve is part of a propagation program working to restore blight-resistant trees to Pennsylvania and other areas of their former range which included the Appalachians and the upper Ohio River watershed.
Fortunately, one need not take a strenuous hike up Blue Mountain to observe butterflies. Open space along the park’s quarter-mile-long entrance road is maintained as a rolling meadow of wildflowers and cool-season grasses that provide nectar for adult butterflies and host plants for their larvae.
A view looking north at the butterfly meadow and entrance road at Boyd Big Tree Preserve Conservation Area. Second Mountain is in the background.Mowed paths follow the entrance road and a portion of the perimeter of the meadow allowing visitors a chance to wander among the waist-high growth to see butterflies, birds, and blooming plants at close range without trampling the vegetation or risking exposure to ticks.A Silver-spotted Skipper feeding on nectar from the flowers of Indian Hemp (Apocynum cannabinum). Like the milkweeds, Indian Hemp is a member of the dogbane family (Apocynaceae).An Eastern Tiger Swallowtail feeding on Common Milkweed.A Great Spangled Fritillary (Speyeria cybele) on Common Milkweed.A Black Swallowtail (Papilio polyxenes) feeding on Common Milkweed nectar.A Pipevine Swallowtail (Battus philenor) on Common Milkweed.Another Pipevine Swallowtail on Common Milkweed. Note the hook-shaped row of red-orange spots on the underside of the hindwing.A Pipevine Swallowtail visiting the brilliant blooms of Butterfly Weed, a favorite of a wide variety of pollinators.A Black Swallowtail with damaged wings alights atop a Butterfly Weed flower cluster. Note the pair of parallel rows of red-orange spot on the underside of the hindwing.A Monarch feeding on nectar from the flowers of Butterfly Weed.A mating pair of Eastern Tailed Blues on a Timothy (Phleum pratense) spike.A male Great Spangled Fritillary (right) pursuing a female.Butterflies aren’t the only colorful insects patrolling the meadows at Boyd Big Tree Preserve. Dragonflies including Common Green Darners are busily pursuing prey, particularly small flying insects like mosquitos, gnats, and flies.Dragonflies themselves can become prey and are much sought after by Broad-winged Hawks. This very vocal juvenile gave us several good looks as it ventured from the forest into the skies above the upper meadow during midday. It wasn’t yet a good enough flier to snag a dragonfly, but it will have plenty of opportunities for practice during its upcoming fall migration which, for these Neotropical raptors, will get underway later this month.
Do yourself a favor and take a trip to the Boyd Big Tree Preserve Conservation Area. Who knows? It might actually inspire you to convert that lawn or other mowed space into much-needed butterfly/pollinator habitat.
While you’re out, you can identify your sightings using our photographic guide—Butterflies of the Lower Susquehanna River Watershed—by clicking the “Butterflies” tab at the top this page. And while you’re at it, you can brush up on your hawk identification skills ahead of the upcoming migration by clicking the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab. Therein you’ll find a listing and descriptions of hawk watch locations in and around the lower Susquehanna region. Plan to visit one or more this autumn!
Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge. At Bombay Hook, they don’t waste your money mowing grass. Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools. Birds there can often be observed at close range. The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools. Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.Two more Short-billed Dowitchers on the way in.Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed. Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge. Mallards, American Avocets, and egrets are gathered on the shore.More Short-billed Dowitchers arriving to feed in Raymond Pool.Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool. A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.American Avocets probe the muddy bottom of Raymond Pool.Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early. Many migrating Dunlin linger at Bombay Hook into October and even November.This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp PoolA Greater Yellowlegs at Bear Swamp Pool.A Caspian Tern patrolling Raymond Pool.The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass. High tide licks at the roots of the cordgrass supporting the temporary domicile.By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice). It is our only dragonfly able to breed in saltwater. Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.Look up! A migrating Bobolink passes over the dike at Shearness Pool.Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool. Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river. The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century. An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers. Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.A Great Egret prowling Shearness Pool.A Snowy Egret in Bear Swamp Pool.Wood Ducks in Bear Swamp Pool.A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.
As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.
Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay. We found just this one during our visit to Bombay Hook. Man has certainly decimated populations of this ancient crustacean during recent decades.As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.A male Atlantic Marsh Fiddler Crab peers from its den.Herons and egrets including this Great Egret are quite fond of fiddler crabs. As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.An Osprey patrols the vast tidal areas opposite Shearness Pool.
No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook. This one was in shrubby growth along the dike at the north end of Shearness Pool.These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries. The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware. The seed balls are a favorite winter food of goldfinches and siskins.Finis Pool has no frontage on the tidal marsh but is still worth a visit. It lies along a spur road on the tour route and is located within a deciduous coastal plain forest. Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.The White-tailed Deity is common along the road to Finis Pool.Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well. This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.
We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.
Here’s a look at some native plants you can grow in your garden to really help wildlife in late spring and early summer.
The showy bloom of a Larger Blue Flag (Iris versicolor) and the drooping inflorescence of Soft Rush (Juncus effusus). These plants favor moist soils in wetlands and damp meadows where they form essential cover and feeding areas for insects, amphibians, and marsh birds. Each is an excellent choice for helping to absorb nutrients in a rain garden or stream-side planting. They do well in wet soil or shallow water along the edges of garden ponds too.The fruits of Smooth Shadbush (Amelanchier laevis), also known as Allegheny Serviceberry, Smooth Serviceberry, or Smooth Juneberry, ripen in mid-June and are an irresistible treat for catbirds, robins, bluebirds, mockingbirds, and roving flocks of Cedar Waxwings.Also in mid-June, the fragrant blooms of Common Milkweed attract pollinators like Eastern Carpenter Bees,……Honey Bees,……and butterflies including the Banded Hairstreak (Satyrium calanus). In coming weeks, Monarch butterflies will find these Common Milkweed plants and begin laying their eggs on the leaves. You can lend them a hand by planting milkweed species (Asclepias) in your garden. Then watch the show as the eggs hatch and the caterpillars begin devouring the foliage. Soon, they’ll pupate and, if you’re lucky, you’ll be able to watch an adult Monarch emerge from a chrysalis!
Let’s have a look at some of the magnificent native wildflowers blooming on this Mother’s Day in forests and thickets throughout the Lower Susquehanna River Watershed.
Rue Anemone (Thalictrum thalictroides) is a fragile little wildflower of open woods.The Blackhaw (Viburnum prunifolium) is a shrub of thickets and forest borders. The flower clusters mature into an abundance of berries relished by birds and other wildlife.A large Black Cherry tree adorned in white blossoms can be pleasing to the eye and very beneficial for pollinating insects.The Common Pawpaw is a small tree of damp well-drained soils. Where conditions are ideal, it may form clonal colonies in the forest understory. The deep maroon blooms mature into banana-like fruits, hence its alternate common name “Custard Apple”.The Mayapple or Mandrake (Podophyllum peltatum) commonly blankets the forest floor. It, like other ephemeral wildflowers, is vulnerable to invasions by non-native shrubs that leaf out earlier in spring than native species and rob them of sunlight during their already abbreviated growing season.
Aren’t they spectacular?
Why not take a stroll to have a look at these and other plants you can find growing along a nearby woodland trail? You could check out some place new and get a little exercise too!
If you go, please tread lightly, take your camera, mind your manners, and…