For the past several weeks, we’ve given you a look at the fallout of Neotropical songbirds on mornings following significant nocturnal flights of southbound migrants. Now let’s examine the diurnal (daytime) flights that have developed in recent days over parts of the lower Susquehanna basin and adjacent regions.
For those observing diurnal migrants, particularly at raptor-counting stations, the third week of September is prime time for large flights of tropics-bound Broad-winged Hawks. Winds from easterly directions during this year’s movements kept the greatest concentrations of these birds in the Ridge and Valley Province and upper areas of the Piedmont as they transited our region along a southwesterly heading. Counts topped 1,000 birds or more on at least one day at each of these lookouts during the past seven days. Meanwhile at hawk watches along the Piedmont/Atlantic Coastal Plain border, where flights topped 10,000 or more birds on the best days last fall, observers struggled to see 100 Broad-winged Hawks in a single day.
Dozens of migrating Broad-winged Hawks during a morning liftoff to the north of Blue Mountain.
Among the challenges counters faced while enumerating migratory Broad-winged Hawks this week was the often clear blue skies, a glaring sun, and the high altitude at which these raptors fly during the mid-day hours.
Large numbers of Broad-winged Hawks climbing on a thermal updraft to gain altitude can become increasingly difficult to spot in a clear blue sky.Higher and higher they go until they practically disappear from view.Cloud cover can help make locating really high-flying birds a bit easier, but they can sometimes climb into the lower portion of the cloud and vanish.
Another factor complicating the hawk counters’ tasks this week, particularly west of the Susquehanna River, was the widespread presence of another group of diurnal fliers.
Was it the migratory Blue Jays that are beginning to move through the area? No.Was it the increasing numbers of Sharp-shinned Hawks and other raptors? No.So what was it that complicated the hawkwatchers’ efforts to find high-flying Broad-winged Hawks during this week’s flights?Is it a bird? Is it a plane? No. And it’s nobody’s superhero either. It’s the non-native Spotted Lanternfly and it is now very numerous west of the Susquehanna and common enough to the east to make even experienced observers take a second look while scanning the skies for Broad-winged Hawks and other migrating raptors. Just how many do raptor counters have to contend with? Even where these invasive insects have become less populous in the lower Susquehanna valley, hundreds still fill the skies above forest clearings around hawk-counting stations including Waggoner’s Gap, Second Mountain, and Rocky Ridge County Park.Spotted Lanternfly populations explode with great vigor as they colonize areas with plenty of tender new growth on native, non-native, and cultivated shrubs, trees, and vines. Earlier this week, National Weather Service Doppler Radar in Pittsburgh, Pennsylvania, captured enormous mid-afternoon flights of Spotted Lanternflies in recently invaded areas of southwestern Pennsylvania and eastern Ohio. (NOAA/National Weather Service Doppler Radar image)A base velocity image consisting primarily of Spotted Lanternflies in southwestern Pennsylvania and eastern Ohio. (NOAA/National Weather Service Doppler Radar image)
As the Broad-winged Hawk migration draws to a close during the coming ten days, the occurrence of the one dozen other migratory diurnal raptors will be on the upswing. They generally fly at lower altitudes, often relying upon wind updrafts on the ridges for lift instead of high-rising thermal updrafts. They therefore present better observation opportunities for visitors at hawk watch lookouts and are less frequently confused with high-flying Spotted Lanternflies.
A Merlin seen earlier this week migrating through the Lower Susquehanna River Watershed. The numbers of migrating Merlins and other falcons will peak sometime in coming weeks.
What’s all this buzz about bees? And what’s a hymanopteran? Well, let’s see.
Hymanoptera—our bees, wasps, hornets and ants—are generally considered to be our most evolved insects. Some form complex social colonies. Others lead solitary lives. Many are essential pollinators of flowering plants, including cultivars that provide food for people around the world. There are those with stingers for disabling prey and defending themselves and their nests. And then there are those without stingers. The predatory species are frequently regarded to be the most significant biological controls of the insects that might otherwise become destructive pests. The vast majority of the Hymanoptera show no aggression toward humans, a demeanor that is seldom reciprocated.
Late summer and early autumn is a critical time for the Hymanoptera. Most species are at their peak of abundance during this time of year, but many of the adult insects face certain death with the coming of freezing weather. Those that will perish are busy, either individually or as members of a colony, creating shelter and gathering food to nourish the larvae that will repopulate the environs with a new generation of adults next year. Without abundant sources of protein and carbohydrates, these efforts can quickly fail. Protein is stored for use by the larval insects upon hatching from their eggs. Because the eggs are typically deposited in a cell directly upon the cache of protein, the larvae can begin feeding and growing immediately. To provide energy for collecting protein and nesting materials, and in some cases excavating nest chambers, Hymanoptera seek out sources of carbohydrates. Species that remain active during cold weather must store up enough of a carbohydrate reserve to make it through the winter. Honey Bees make honey for this purpose. As you are about to see, members of this suborder rely predominately upon pollen or insect prey for protein, and upon nectar and/or honeydew for carbohydrates.
We’ve assembled here a collection of images and some short commentary describing nearly two dozen kinds of Hymanoptera found in the Lower Susquehanna River Watershed, the majority photographed as they busily collected provisions during recent weeks. Let’s see what some of these fascinating hymanopterans are up to…
SOLITARY WASPS
A Great Black Wasp on goldenrod (Solidago species). Like other solitary wasps, a female Great Black Wasp will sting and paralyze a host insect upon which she’ll deposit her eggs. After hatching, the larvae will begin consuming the host’s body as a source of protein. The parasitized insects are often katydids or grasshoppers.A Great Black Wasp feeding on nectar, a source of carbohydrates. Unlike social bees and wasps, solitary wasps are equipped with a stinger solely used for immobilizing prey, not defending a nest. They are therefore quite docile and pose little threat to humans.A Great Black Wasp powdered with pollen. Hymanopterans that gather nectar and/or pollen are tremendously important pollinators of hundreds of species of plants.A female Thread-waisted wasp (Ammophilia species, probably A. nigricans) drags a paralyzed caterpillar to her excavated nest where she’ll deposit an egg on the body. After hatching, the larval wasp will feed on the disabled caterpillar. The protein will enable the larvae to grow, pupate, and later emerge as an adult wasp.The female Eastern Cicada Killer (Sphecius speciosus) excavates an underground nest with branch tunnels connecting a dozen chambers or more. As the common name suggests, the female wasp paralyzes a cicada, then makes a strenuous effort to fly and drag it back to the nest for placement in a cell. Each male wasp egg is deposited upon just one immobilized cicada, but a female egg is provided with a cache of several cicadas to provide adequate protein for growth to a larger size. Nest cells are sealed with soil, then the larvae hatch in just a couple of days. Within about two weeks, they have consumed the cicada protein and are fully grown. Wrapped in a cocoon, they spend the winter in the nest, then pupate in the spring before emerging as a new generation of adults.The Black-and-yellow Mud Dauber (Sceliphron caementarium) builds a mud-ball nest within which it packs paralyzed spiders to function as a source of protein for its larvae.A Black-and-yellow Mud Dauber at nest.The Pipe Organ Mud Dauber builds this elaborate nest in which their eggs and paralyzed spiders are deposited in cells sealed with mud partitions. After consuming the spiders, the larvae pupate, overwinter, then emerge from their cells as adults during the following spring. To escape the protection of the nest, the new generation of adults bore through the mud walls. Adult Pipe Organ Mud Daubers resemble the Great Black Wasp, but have a white or yellow distal segment on their rear legs resembling a pair of light-colored socks.A closeup of the previous image with the lengths of the nest tubes compressed to show four scavenger flies (Miltogramminae), possibly two species, that have invaded this Pipe Organ Mud Dauber nest. Scavenger flies are kleptoparasites that victimize various solitary bees and wasps, depositing larvae directly into the host species’ nest cells to consume the protein cache stored therein.
CUCKOO WASPS
Cuckoo Wasps (Chrysididae), also known as Emerald Wasps, parasitize the nests of other species of wasps. Females lay their eggs inside the host’s nest, then flee the scene. Upon hatching, larval Cuckoo Wasps feed on stockpiles of prey intended for the host species’ offspring. Like the adult mud daubers that have already matured and departed this nest by digging a hole through the wall of the cell within which they were hatched, the metallic green Cuckoo Wasp in the upper left has just emerged in much the same way.
SWEAT BEES
A Sweat Bee (Lasioglossum species) collecting nectar and pollen on White Snakeroot (Ageratina altissima).Sweat Bees (Lasioglossum species) visit human skin to lick up the electrolytes left behind by evaporating perspiration.Sweat Bees in the genus Lasioglossum demonstrate various social behaviors ranging from species that are solitary nesters to those that create colonies with work forces ranging in size from as few as four to as many as hundreds of bees. Some Lasioglossum practice kleptoparasitism, while others are quite accomplished foragers.A female Augochlorine Green Sweat Bee (Augochlorini) collecting nectar on White Snakeroot. Notice the pollen “baskets” on the rear leg.An Augochlorine Green Sweat Bee (Augochlorini). Sweat bees nest in subterranean cavities and in hollowed out sections of trees.A copper-colored Augochlorine Green Sweat Bee (Augochlorini) collecting nectar and dusted with pollen.
LEAFCUTTER AND MASON BEES
A Leafcutter Bee (Megachile species). Like Mason Bees, female Mason Bees deposit each of their eggs on a “pollen loaf” within an individual cell inside a preexisting tunnel-like cavity in wood, stone, or in the ground. Unlike Mason Bees, female Leafcutter Bees cut a circular piece of leaf to create each of the cells in their nest. After hatching, the larval bee feeds on the pollen loaf, pupates, then emerges from the shelter of the nest to start a new generation, usually during the following year.A Leafcutter Bee (Megachile species) visiting Wild Bergamot. Female Leafcutter and Mason Bees lack pollen “baskets” on their rear legs but instead have pollen “brushes” on the underside of the abdomen to gather the protein they need to create a “pollen loaf” for each nest cell.A Leafcutter Bee (Megachile species) collecting nectar from White Snakeroot.A Mason Bee (Osmia species) emerging from a nest in spring. Mason Bees create nesting cells within preexisting cavities in wood, stone, and other other supporting structures. Within the nest cavity, each egg is deposited atop a cache of pollen and nectar, a pollen loaf, then enclosed behind a partition of mud. The female Mason Bee will usually repeat this process until an entire cavity is filled with cells. During the following spring, a new generation of adult Mason Bees digs its way through the cell walls to emerge and repeat the process. These bees readily use paper straws or holes drilled in blocks of wood for nesting.A mason bee nest box with holes drilled into blocks of wood.Mason Bees seal each cell and the outer end of their nest cavity with mud. These outer nest cells can been parasitized by a variety of wasps. Here, the outer cell of a Mason Bee nest has been victimized by a tiny chalcid wasp (looks like another one to the lower left). Several species of female chalcid wasps (native Monodontomerus species or non-native Pteromalus venustus) enlarge weak points in the outer partition of a mason bee nest, then sting and paralyze the larval bee inside before depositing their eggs. Within the cell. the wasp larvae consume the larval Mason Bee and the “pollen loaf” provided for its growth. These same parasitic wasps prey upon Leafcutter Bees as well.
BUMBLE BEES, CARPENTER BEES, HONEY BEES, AND DIGGER BEES
A Common Eastern Bumble Bee (Bombus impatiens) collecting nectar and pollen on goldenrod. Bumble bees are our sole native group of social bees. Their wax nests are built in a burrow or other shelter. The eggs are deposited in cells along with a supply of pollen for nourishing the larvae upon hatching. Honey is stored in “honey pots” within the nest. New queens are produced along with male bees during the late-summer and fall. Only the new generation of fertilized queens survive the winter to lay eggs and produce workers to construct a new nest.A pair of Common Eastern Bumble Bees collecting nectar and becoming dusted with pollen. Their fuzzy coats and semi-warm-blooded metabolism allows them to be active in cooler weather than is tolerated by other bees.Flowering plants including the Great Rhododendron find success attracting pollinators to their reproductive blossoms by offering carbohydrate-rich nectar to insects like this Eastern Bumble Bee. The yellow spots on the flower’s upper petal help to guide visitors toward their sweet treat.An Eastern Carpenter Bee feeding on goldenrod nectar. Compare the almost hairless abdomen to that of the bumble bees. Carpenter bees are semi-social insects. Females lay their eggs in cells within galleries bored into wood. These nests are completed with great precision, avoiding creation of any second entrance by mistakenly breaching the outer surface of the excavated wood. Each egg/larvae is provided with a supply of protein-rich pollen. Males often hover outside their mate’s nest to prevent competing males from entering the area.A worker Honey Bee, a female member of a sisterhood of foragers from a nearby hive, visits goldenrod alongside Common Eastern Bumble Bees. Honey Bees were brought to North America during the 1620s, the earliest years of the trans-Atlantic migration of European colonists, to pollinate cultivated plants and to provide a reliable source of honey and beeswax. Within the Honey Bee’s social structure, the queen of each hive lays the eggs to produce the female worker bees. Once each year, male drones are produced along with a new generation of queens.In nature, Honey Bees build hives in tree cavities. Recently, this colony constructed a hive in a screech owl nest box at susquehannawildlife.net headquarters. To provide protein for the hatching larvae, worker bees collect pollen and deposit it within the hexagonal cells of the vertically aligned beeswax combs. After an egg is deposited upon the pollen cache, each cell is sealed with more beeswax. Young females tend these nest combs before maturing and becoming foraging worker bees.In apiculture, Honey Bees are raised in man-made hives. This Pennsylvania Association of Beekeepers display gives visitors to the Pennsylvania Farm Show in Harrisburg a look at the inner workings of a live bee hive. Nectar collected by worker bees is turned into honey to provide the supply of carbohydrates needed to fuel the colony through the winter. Note the honeycombs on the glass.A possible Small Carpenter Bee (Ceratina species) visiting White Snakeroot. Small Carpenter Bees nest inside hollow stems and twigs. Some species are eusocial, with a queen’s daughters and sisters sharing responsibility for finding food and rearing the young. Females overwinter inside a one of the excavated stems and begin a new nest there in the spring.A Digger Bee (possibly Melissodes species) with “pollen baskets” full of pollen collected from nearby flowers. Digger Bees in the genus Melissodes are often known as the Long-horned Bees. These social insects excavate underground nests and many species practice communal living.
SCOLIID WASPS
The Two-spotted Scoliid Wasp (Scolia dubia), also known as the Blue-winged Scoliid Wasp, is most frequently observed feeding on nectar. Scoliid wasps are solitary nesters, though they may assemble into groups while visiting flowers. They often ignore the presence of humans and are seldom disturbed by their presence. Females seek out the burrowing grubs of beetles including the Green June Bug (Cotinis nitida) and possibly the Japanese Beetle. After stinging a grub to paralyze it, the wasp will deposit her egg on its body, then bury it. Upon hatching, the larval wasp will feed on the grub for nourishment as it grows.Don’t like having your watermelon overrun by Green June Bugs while you’re eating? Then you ought to go out of your way to be nice to the Two-spotted Scoliid Wasp.The Double-banded Scoliid (Scolia bicincta) parasitizes beetle larvae as hosts for its larvae. For carbohydrates it relishes flower nectar.
PAPER WASPS
A Northern Paper Wasp (Polistes fuscatus). Paper wasps prey upon numerous garden pests, particularly caterpillars, to collect protein. Though they are social insects equipped with stingers to subdue their victims and defend their nests, paper wasps are surprisingly docile.A Northern Paper Wasp (Polistes fuscatus) feeding on nectar from a goldenrod flower.A Northern Paper Wasp harvesting wood pulp from the side of a mason bee nest box at susquehannawildlife.net headquarters. The pulp is chewed in the wasp’s saliva to create the paper used to construct the colony’s open-cell nest.Common Paper Wasps (Polistes exclamans), also known as Guinea Paper Wasps, at their open-cell nest. This and the nests of most other paper wasps are suspended on a filament or a pedicle. Many paper wasps can excrete an ant repellent on this section of the nest in an effort to prevent invasion. Like many other social hymenopterans, a defending wasp can secrete a pheromone venom during the stinging process to warn the colony of danger at the nest. In winter, Common Paper Wasps seek shelter in stumps and other locations to hibernate.The European Paper Wasp (Polistes dominula) is a non-native species which builds nests in man-made structures including bird houses. To collect protein, they prey on a wide selection of insects and other invertebrates. As such, European Paper Wasps are widespread and successful here in North America.
YELLOWJACKETS AND HORNETS
An Eastern Yellowjacket feeding on lanternfly honeydew. Eastern Yellowjackets derive much of their success from being generalists, collecting carbohydrates from nearly any sweet source, natural or man made. They are quite fond of ripe fruits, flower nectar, and sugary snacks and drinks, especially soda. Protein for nourishing their larvae is derived from the wide variety invertebrates upon which they prey and from carrion. These foods are chewed into a paste form in preparation for placement into the brood cells.A subterranean colony of Eastern Yellowjackets is started anew each spring by a young queen that has survived winter hibernation in diapause, a state of interrupted development. She constructs the new nest’s first cells using pulp made by chewing rotting wood. The first brood of workers scales up construction while the queen continues producing eggs. At the nest, these social insects will viciously attack anyone or anything perceived to be a threat, so give them their space and leave them alone. Many yellowjacket infestations of homes and other buildings are the work of non-native German Yellowjacket (Vespula germanica) [not shown], an invasive species that constructs paper nests in void spaces including walls and attics.Yellowjackets may be moody and aggressive, but they do fall victim to a number of predators. A Robber Fly (Promachus species) has taken down and is devouring this Eastern Yellowjacket.A Bald-faced Hornet (Dolichovespula maculata) feeding on Spotted Lanternfly honeydew on a Tree-of-heaven (Ailanthus altissima). In the absence of nectar-producing flowers, many bees, yellowjackets, and hornets have turned to the invasive lanternfly and Ailanthus combo to turn the sun’s energy into the carbohydrates they need. For protein, they prey upon spiders, flies, caterpillars, and a variety of other insects.To create paper for nest construction, this Bald-faced Hornet is collecting wood pulp from the surface of a weathered picnic table. Away from the nest, these hornets demonstrate a calm, carefree demeanor and can be closely observed.A Bald-faced Hornet nest in a pine tree. These hives are strictly temporary. Within the nest, a generation of drones (males) and new queens are produced late each year. These wasps leave the colony to mate. With the arrival of freezing weather, all inhabitants within the nest, including the old queen, perish, as do the drones that departed to breed. Only the new queens survive winter hibernation to propagate the next generation of wasps, starting with the workers needed to construct a fresh nest and reestablish the colony.Did you ever get the feeling you’re being watched? Don’t go messing around with Bald-faced Hornet nests. The occupants therein, like other social bees, wasps, and hornets, are equipped with stingers and venom for defending their colony. This is an adaptation that has developed over time to assure the survival of populations of these insects. Think about it this way, a solitary wasp that loses a nest loses only their individual brood of offspring. There is minimal impact on the wider local population of such insects. Conversely, a social wasp or hornet that loses a nest loses an entire colony, possibly negating the benefits of their cooperative behavior and threatening the survival of the species. Insects that cooperate to build societies for survival can be more vulnerable to the catastrophic impacts of certain circumstances like disease, weather, and invasion of their colonies. Therefore, natural selection has provided them with contingencies for these dangers, for example, the instinct to construct protective shelters and the adaptation of stingers and venom for defense against intruders and would-be predators. Oh, and by the way, the Bald-faced Hornet can spray venom, often aiming for the eyes, so keep your distance.European Hornets (Vespa crabro), an introduced species, are predatory on a variety of flying insects for protein. For carbohydrates they are attracted to sweets like this lanternfly honeydew on Tree-of-heaven.European Hornets constructing a paper nest in a tree cavity.
POTTER WASPS
A potter wasp (Eumenes species), probably a Fraternal Potter Wasp (E. fraternus), hovering near a European Paper Wasp on Partridge Pea. The female potter wasp builds a small mud nest resembling a tiny clay pot. One of her eggs is inserted and left hanging on a thin thread. Then a paralyzed caterpillar is deposited as a source of protein to nourish the larva upon hatching. Lastly, the pot is sealed with a lid made of wet mud. Upon maturing, the new generation of adult wasps perform a pottery breaking to emerge and take flight.
ANTS
Field Ants (Formica species, possibly Formica pallidefulva) clear the entrance to their underground nest. Field ants are eusocial insects, they work in concert to build, maintain, and defend the nest, rear young, and find food. There is no social caste system. Field Ants are predators and scavengers when collecting protein. For carbohydrates they often rely on the honeydew produced by aphids. As a method of improving and sustaining the production of honeydew, some ant species will tend colonies of aphids by moving the younger individuals from depleted portions of plants to more healthy tissue. Field Ant nests contain chambers used for a variety of functions including raising young and storing food. Some nests include multiple queens and some colonies consist of more than one nest. Ants in the genus Formica are weaponized; they can spray formic acid to repel intruders and defend their colony.
We hope this brief but fascinating look at some of our more common bees, wasps, hornets, and ants has provided the reader with an appreciation for the complexity with which their food webs and ecology have developed over time. It should be no great mystery why bees and other insects, particularly native species, are becoming scarce or absent in areas of the Lower Susquehanna River Watershed where the landscape is paved, hyper-cultivated, sprayed, mowed, and devoid of native vegetation, particularly nectar-producing plants. Late-summer and autumn can be an especially difficult time for hymanopterans seeking the sources of proteins and carbohydrates needed to complete preparations for next year’s generations of these valuable insects. An absence of these staples during this critical time of year quickly diminishes the diversity of species and begins to tear at the fabric of the food web. This degradation of a regional ecosystem can have unforeseen impacts that become increasingly widespread and in many cases permanent.
How can anyone be surprised by the absence of bees and other pollinators in farmland? Manicured and cultivated ground offers little in the way of year-round shelter and food sources for insects and other wildlife.This savanna-like habitat on a south-facing slope provides the abundance of nectar-producing, pollen-rich wildflowers needed to nourish a diverse population of insects including bees, wasps, hornets, and ants. Goldenrods, asters, and White Snakeroot are some of their late-season favorites.
Editor’s Note: No bees, wasp, hornets, or ants were harmed during this production. Neither was the editor swarmed, attacked, or stung. Remember, don’t panic, just observe.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York, NY.
Grasshoppers are perhaps best known for the occasions throughout history when an enormous congregation of these insects—a “plague of locusts”—would assemble and rove a region to feed. These swarms, which sometimes covered tens of thousands of square miles or more, often decimated crops, darkened the sky, and, on occasion, resulted in catastrophic famine among human settlements in various parts of the world.
The largest “plague of locusts” in the United States occurred during the mid-1870s in the Great Plains. The Rocky Mountain Locust (Melanoplus spretus), a grasshopper of prairies in the American west, had a range that extended east into New England, possibly settling there on lands cleared for farming. Rocky Mountain Locusts, aside from their native habitat on grasslands, apparently thrived on fields planted with warm-season crops. Like most grasshoppers, they fed and developed most vigorously during periods of dry, hot weather. With plenty of vegetative matter to consume during periods of scorching temperatures, the stage was set for populations of these insects to explode in agricultural areas, then take wing in search of more forage. Plagues struck parts of northern New England as early as the mid-1700s and were numerous in various states in the Great Plains through the middle of the 1800s. The big ones hit between 1873 and 1877 when swarms numbering as many as trillions of grasshoppers did $200 million in crop damage and caused a famine so severe that many farmers abandoned the westward migration. To prevent recurrent outbreaks of locust plagues and famine, experts suggested planting more cool-season grains like winter wheat, a crop which could mature and be harvested before the grasshoppers had a chance to cause any significant damage. In the years that followed, and as prairies gave way to the expansive agricultural lands that presently cover most of the Rocky Mountain Locust’s former range, the grasshopper began to disappear. By the early years of the twentieth century, the species was extinct. No one was quite certain why, and the precise cause is still a topic of debate to this day. Conversion of nearly all of its native habitat to cropland and grazing acreage seems to be the most likely culprit.
The critically endangered Eskimo Curlew (Numenius borealis), a species not photographed since 1962 and not confirmed since 1963, fed on Rocky Mountain Locusts during its spring migration through the Great Plains. Excessive hunting and conversion of grasslands to agriculture are believed responsible for the bird’s demise. (United States Fish and Wildlife Service image by Christina Nelson)
In the Mid-Atlantic States, the mosaic of the landscape—farmland interspersed with a mix of forest and disturbed urban/suburban lots—prevents grasshoppers from reaching the densities from which swarms arise. In the years since the implementation of “Green Revolution” farming practices, numbers of grasshoppers in our region have declined. Systemic insecticides including neonicotinoids keep grasshoppers and other insects from munching on warm-season crops like corn and soybeans. And herbicides including 2,4-D (2,4-Dichlorophenoxyacetic acid) have, in effect, become the equivalent of insecticides, eliminating broadleaf food plants from the pasturelands and hayfields where grasshoppers once fed and reproduced in abundance. As a result, few of the approximately three dozen species of grasshoppers with ranges that include the Lower Susquehanna River Watershed are common here. Those that still thrive are largely adapted to roadsides, waste ground, and small clearings where native and some non-native plants make up their diet.
Here’s a look at four species of grasshoppers you’re likely to find in disturbed habitats throughout our region. Each remains common in relatively pesticide-free spaces with stands of dense grasses and broadleaf plants nearby.
CAROLINA GRASSHOPPER
Dissosteira carolina
The Carolina Grasshopper, also known as the Carolina Locust or Quaker, is one of the band-winged grasshoppers. It is commonly found along roadsides and on other bare ground near stands of tall grass and broadleaf plants.The Carolina Grasshopper is variable in color, ranging from very dark brown……to a rich tan or khaki shade. These earth-tone colors provide the insect with effective camouflage while spending time on the ground.The Carolina Grasshopper is most readily detected and identified when it flies. The colors of the wings resemble those of the Mourning Cloak butterfly.Carolina Grasshoppers are among the preferred victims of Great Black Wasps (Sphex pensylvanicus). A female wasp stings the grasshopper to paralyze it, then drags it away to one of numerous cells in an underground burrow where she lays an egg on it. The body of the disabled grasshopper then provides nourishment for the larval wasp.
DIFFERENTIAL GRASSHOPPER
Melanoplus differentialis
Differential Grasshopper nymph with small “fairy wings”.An adult female Differential Grasshopper with fully developed wings.An adult female Differential Grasshopper
TWO-STRIPED GRASSHOPPER
Melanoplus bivittatus
An early-stage Two-striped Grasshopper nymph.A Two-striped Grasshopper nymph in a later stage.An adult female Two-striped Grasshopper.An adult female Two-striped Grasshopper. Note the pale stripe originating at each eye and joining near the posterior end of the wings to form a V-shaped pattern.An adult female Two-striped Grasshopper.
RED-LEGGED GRASSHOPPER
Melanoplus femurrubrum
An adult male Red-legged Grasshopper hiding in dense urban vegetation.The Red-legged Grasshopper may currently be our most abundant and widespread species.An adult male Red-legged Grasshopper.An adult female Red-legged Grasshopper.
Protein-rich grasshoppers are an important late-summer, early-fall food source for birds. The absence of these insects has forced many species of breeding birds to abandon farmland or, in some cases, disappear altogether.
Beginning in the early 1930s, the Western Cattle Egret (Bubulcus ibis), a notoriously nomadic species, transited the Atlantic from Africa to colonize the Americas…and they did it without any direct assistance from humans. During the 1970s and early 1980s, a nesting population of Western Cattle Egrets on river islands adjacent to the Susquehanna’s Conejohela Flats off Washington Boro was the largest inland rookery in the northeastern United States. The Lancaster County Bird Club censused the birds each August and found peak numbers in 1981 (7,580). During their years of abundance, V-shaped flocks of cattle egrets from the rookery islands ventured into grazing lands throughout portions of Lancaster, York, Dauphin, and Lebanon Counties to hunt grasshoppers. These daily flights were a familiar summertime sight for nearly two decades. Then, in the early 1980s, reductions in pastureland acreage and plummeting grasshopper numbers quickly took their toll. By 1988, the rookery was abandoned. The cattle egrets had moved on. (Vintage 33 mm image)During the summer and early fall, juvenile and adult Ring-necked Pheasants feed heavily on grasshoppers. Earlier and more frequent mowing along with declining numbers of grasshoppers on farmlands due to an increase in pesticide use were factors contributing to the crash of the pheasant population in the early 1980s.To the delight of Wild Turkeys, each of the four species of grasshoppers shown above frequents clearings and roadsides adjacent to forest areas. While changes in grasshopper distribution have been detrimental to populations of birds like pheasants, they’ve created a feeding bonanza for turkeys.Wild Turkeys feeding on an abundance of grasshoppers along a forest road.An American Kestrel feeds on a grasshopper while ignoring the abundance of Spotted Lanternflies swarming the adjacent utility pole. In Susquehanna valley farmlands, grasshopper and kestrel numbers are down. Lanternflies, on the other hand, have got it made.Maintaining areas bordering roads, forests, wetlands, farmlands, and human development in a state of early succession can provide and ideal mix of mature grasses and broadleaf plants for grasshoppers, pollinators, birds, and other wildlife.
Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge. At Bombay Hook, they don’t waste your money mowing grass. Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools. Birds there can often be observed at close range. The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools. Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.Two more Short-billed Dowitchers on the way in.Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed. Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge. Mallards, American Avocets, and egrets are gathered on the shore.More Short-billed Dowitchers arriving to feed in Raymond Pool.Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool. A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.American Avocets probe the muddy bottom of Raymond Pool.Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early. Many migrating Dunlin linger at Bombay Hook into October and even November.This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp PoolA Greater Yellowlegs at Bear Swamp Pool.A Caspian Tern patrolling Raymond Pool.The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass. High tide licks at the roots of the cordgrass supporting the temporary domicile.By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice). It is our only dragonfly able to breed in saltwater. Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.Look up! A migrating Bobolink passes over the dike at Shearness Pool.Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool. Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river. The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century. An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers. Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.A Great Egret prowling Shearness Pool.A Snowy Egret in Bear Swamp Pool.Wood Ducks in Bear Swamp Pool.A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.
As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.
Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay. We found just this one during our visit to Bombay Hook. Man has certainly decimated populations of this ancient crustacean during recent decades.As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.A male Atlantic Marsh Fiddler Crab peers from its den.Herons and egrets including this Great Egret are quite fond of fiddler crabs. As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.An Osprey patrols the vast tidal areas opposite Shearness Pool.
No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook. This one was in shrubby growth along the dike at the north end of Shearness Pool.These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries. The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware. The seed balls are a favorite winter food of goldfinches and siskins.Finis Pool has no frontage on the tidal marsh but is still worth a visit. It lies along a spur road on the tour route and is located within a deciduous coastal plain forest. Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.The White-tailed Deity is common along the road to Finis Pool.Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well. This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.
We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.
Here’s wishing you and yours a Happy Halloween. It’s a much-anticipated day of excitement capped by surprise visits from strange-looking hideous creatures you’ve never seen before. They don’t stop by for a chat. Nope, not a word. Just a little bit of nearly imperceptible buzzing when the move around. You see, the little sneaks have hatched a plan. They want to eat your stuff and maybe trash the place before they go. And when you finally get rid of them, more start showing up—dozens and dozens, then hundreds. The more you have, the more you attract. You’ll be shocked that there are that many living in your neighborhood. It’s like a scene from “Nightmare on Maple Street”. The invasion drives some people mad, but you’re just going to love it. So, get ready, because here they come. Trick or Treat!
Q: What’s scariest thing about having Brown Marmorated Stink Bugs over for Halloween?A: They bring along their friends.
Once upon a time, a great panic spread throughout the lower Susquehanna region. A destructive mob of invaders was overtaking our verdant land and was sure to decimate all in its path. Clad in gray and butternut, they came by the thousands. Flashing their crimson banners, they signaled their arrival at each new waypoint along their route. The Pennsylvania government called upon the populace to heed the call and turn out in defense of the state. Small bands of well-intentioned citizens tried in vain to turn back the progress of the hostiles—none succeeded. But for a cadre of civic-minded elites and some small groups of college professors and their students, few responded to a call to confine the invasion along designated lines of containment. Word spread quickly throughout the valley that farms had been overrun by waves of the merciless intruders. Agrarians reported that their orchards had been stripped; they had lost all of the fruits of their labor. Stories exaggerating the hideous appearance of the approaching aliens struck fear into the faint-of-heart. The growing sentiment among the terror-stricken residents: this horde must be stopped before pestilence is visited upon everyone in the state!
And so, on the evening of June 28, 1863, just three days before the Battle of Gettysburg, the wooden Susquehanna River bridge at Columbia-Wrightsville was set ablaze just as Brigadier General John B. Gordon’s brigade of the invading Army of Northern Virginia approached the span’s west entrance preparing to cross to the eastern shore. Thus, the rebel tide was turned away from the Susquehanna at the point some contend to be the authentic “High-water Mark of the Confederacy”.
Just one among an army of Spotted Lanternflies marching westbound across the Susquehanna River on the present-day Columbia-Wrightsville Bridge. Same story, different ending.
During the coming two weeks, peak numbers of migrating Neotropical birds will be passing through the northeastern United States including the lower Susquehanna valley. Hawk watches are staffed and observers are awaiting big flights of Broad-winged Hawks—hoping to see a thousand birds or more in a single day.
During its passage through the Lower Susquehanna River Watershed, an adult Broad-winged Hawk sails over Second Mountain Hawk Watch in Lebanon County, Pennsylvania.A hatch-year/juvenile Broad-winged Hawk gazes toward hawk watchers on the ground.
Broad-winged hawks feed on rodents, amphibians, and a variety of large insects while on their breeding grounds in the forests of the northern United States and Canada. They depart early, journeying to wintering areas in Central and South America before frost robs them of a reliable food supply.
The Carolina Saddlebags (Tramea carolina), this one photographed at Second Mountain Hawk Watch on September 8th, is the rarest of the lower Susquehanna region’s migratory dragonflies. Autumn Broad-winged Hawk movements coincide with southbound flights of the Carolina Saddlebags and the more numerous migratory dragonfly species: Common Green Darner, Wandering Glider, Twelve-spotted Skimmer, and Black Saddlebags. “Broad-wings” will often eat these and other dragonflies during migration and can sometimes be seen catching and feeding upon them while still soaring high overhead.
While migrating, Broad-winged Hawks climb to great altitudes on thermal updrafts and are notoriously difficult to see from ground level. Bright sunny skies with no clouds to serve as a backdrop further complicate a hawk counter’s ability to spot passing birds. Throughout the Lower Susquehanna River Watershed, the coming week promises to be especially challenging for those trying to observe and census the passage of high-flying Broad-winged Hawks. The forecast of hot and humid weather is not so unusual, but the addition of smoke from fires in the western states promises to intensify the haze and create an especially irritating glare for those searching the skies for raptors.
Smoke from fires along the California coast and in central Utah can be seen streaming east this morning. (NOAA/GOES image)Smoke from western fires and humid air creates a band of haze in Ohio, Pennsylvania, New Jersey, and states to the south this morning. (NOAA/GOES image)
A migrating Broad-winged Hawk in the glare of a hazy sky. In addition to visibility problems, swarms of Spotted Lanternflies above the treetops make distant hawks difficult to discern for hawk watchers scanning the horizon with binoculars.
It may seem gloomy for the mid-September flights in 2021, but hawk watchers are hardy types. They know that the birds won’t wait. So if you want to see migrating “Broad-wings” and other species, you’ve got to get out there and look up while they’re passing through.
Migrating Ospreys typically fly low enough and are large enough to be spotted even during the haziest of conditions.Bald Eagles like this fourth-year bird can ascend to great altitude, but their size usually prevents them from sneaking past a lookout unnoticed.Peregrines escape notice not due to hazy sky conditions, but because they pass by so quickly. They’re being seen at local hawk watches now through October.
These hawk watches in the Lower Susquehanna River Watershed are currently staffed by official counters and all welcome visitors:
Rocky Ridge County Park Hawk Watch—3699 Deininger Road off Mount Zion Road (Route 24) northeast of York, Pennsylvania.
Second Mountain Hawk Watch—off Cold Spring Road on the grounds of Fort Indiantown Gap in Lebanon County, Pennsylvania.
Waggoner’s Gap Hawk Watch—where Route 74 crosses Blue Mountain north of Carlisle, Pennsylvania.
—or you can just keep an eye on the sky from wherever you happen to be. And don’t forget to check the trees and shrubs because warbler numbers are peaking too! During recent days…
Northern Parula at Chiques Rock County Park in Lancaster County.Black-and-white Warbler at Rocky Ridge County Park in York County.Cape May Warbler at Chiques Rock County Park in Lancaster County.Bay-breasted Warbler at Rocky Ridge County Park in York County.
The Magi have arrived. Emanating from the shadows of a nearby forest, you may hear the endless drone of what sounds like an extraterrestrial craft. Then you get your first look at those beady red eyes set against a full suit of black armor—out of this world. The Magicicada are here at last.
This exuvia, the leftover from a cicada’s final molt, tells us they are here.A Brood X Periodical Cicada soon after emergence and final molt.Not to worry, cicadas are harmless and docile when handled. This is Magicicada septendecim, the largest and most common of our three species of Brood X seventeen-year cicadas. They are currently emerging along south-facing borders of forests and wooded parks and lawns.Magicicada septendecim can be recognized by the orange on the thorax behind each eye and in front of the wing insertions. The smaller M. cassinii and M. septendecula have no orange coloration between the eye and wing.Magicicada septendecim (seen here) has broad orange stripes on the abdomen. M. cassinii has an all black abdomen and M. septendecula, the rarest species, has narrow well-defined orange stripes.
If you go out and about to observe Periodical Cicadas, keep an eye open for these species too…
Spotted Lanternflies, one of our most dreaded invasive species, have hatched. These tiny nymphs about 5 millimetres in length were found feeding on a Virginia Creeper (Parthenocissus quinquefolia), a native vine in the grape family (Vitaceae).Deer Ticks, also known as Black-legged Ticks, are hanging around on vegetation of all kinds looking to hitch a ride on a suitable host. Don’t let it be you. This adult female, less than 5 millimetres in length, was washed loose during an after-hike shower.
With autumn coming to a close, let’s have a look at some of the fascinating insects (and a spider) that put on a show during some mild afternoons in the late months of 2019.
Bush Katydids (Scudderia species) are found in brushy habitats and along rural roadsides. Their green summer color fades to brown, maroon, and gold to match the autumn foliage where they hide. Bush katydids often remain active until a hard freeze finally does them in.The Eastern Buck Moth (Hemileuca maia) is fuzzy, appearing to wear a warm coat for its autumn expeditions. Adults emerge in October and may fly as late as December. Females deposit their eggs on the twigs of Bear Oak (Quercus ilicifolia), Blackjack Oak (Q. marilandica), or Chestnut Oak (Q. montana), trees that, in our region, seem most favorable for the moth’s use when growing on burned barrens and mountain slopes. The spiny caterpillars are known to feed only on the foliage of these few trees. In the lower Susquehanna valley, the Eastern Buck Moth is rare because its specialized habitat is in short supply, and it’s all Smokey The Bear’s fault.The Sachem (Atalopedes campestris) wanders north from the Atlantic Coastal Plain into the Susquehanna valley each summer. In some years they become the most numerous small orange butterfly of all, particularly around home gardens. The larvae will feed on Crabgrass (Digitaria species), but have not found success overwintering this far north. By November, adults begin to look pretty drab.From 1978 through 1982, the Asian Multicolored Lady Beetle (Harmonia axyridis) was introduced into the eastern states by the United States Department of Agriculture. It has become a nuisance in many areas where it swarms, sometimes bites, and often overwinters in large smelly masses within homes and other warm buildings. As you may have guessed, it’s possibly displacing some of the less aggressive native lady beetle species.On a chilly afternoon, a sun-warmed Bold Jumping Spider (Phidippus audax) pounced and dispatched this sluggish worker Honey Bee (Apis mellifera) that was trying to gather pollen from a late-season Purple Coneflower bloom. This spider is bold indeed.Under bridges, inside bird nest boxes, and sometimes beneath porches, the female Pipe Organ Mud Dauber (Trypoxylon politum), a predatory wasp, builds these elaborate nests composed of long rows (pipes) of nursery cells. Into each cell one or more paralyzed spiders is deposited along with one of the female’s eggs. When hatched, each larva will feed upon the paralyzed spider(s) inside its cell, then pupate. The pupae overwinter, then emerge from their cells as adults during the following spring. In the autumn, males often stand guard at an entrance to the nest to prevent parasitic species, including some flies (look at the fifth pipe from the right), from laying eggs on the pupae. These wasps are not aggressive toward humans.A Black-and-yellow Mud Dauber (Sceliphron caementarium) observes a neighboring nest of Common Paper Wasps (Polistes exclamans). The Common Paper Wasp, a species also known as the Guinea Paper Wasp, is a native of the southern United States. It is currently expanding its range into the lower Susquehanna valley from the Atlantic Coastal Plain. These two wasp species and the Pipe Organ Mud Dauber are known to regularly coexist. All three will take advantage of man-made structures for their nest sites. People using the picnic tables beneath this pavilion roof never noticed the hundreds of docile wasps above.Those moody Eastern Yellowjackets (Vespula maculifrons) can get very temperamental during warm autumn days. These wasps may appear to have no enemies, but away from areas impacted by man’s everyday activities, they do. The Robber Fly (Promachus species) hunts like a flycatcher or other woodland bird, waiting on a perch along the forest’s edge for prey to pass by, then ambushing it, yellowjackets included.The invasive Spotted Lanternfly, a native of eastern Asia, continues to spread destruction. It established itself throughout much of the east side of the lower Susquehanna River during the summer and fall of 2019. Their route of travel across the farmlands of the region intersects with plenty of vineyards to obliterate and few, if any, natural enemies. Expect them to begin colonizing the west shore en masse during 2020.In 2020, plan to roll a few Spotted Lanternflies over, enjoy the view, and wait for the crimson tide to pass. With any luck, they’ll peak in a year or two.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York, NY.
Second Mountain Hawk Watch is located on a ridge top along the northern edge of the Fort Indiantown Gap Military Reservation and the southern edge of State Game Lands 211 in Lebanon County, Pennsylvania. The valley on the north side of the ridge, also known as St. Anthony’s Wilderness, is drained to the Susquehanna by Stony Creek. The valley to the south is drained toward the river by Indiantown Run, a tributary of Swatara Creek.
The hawk watch is able to operate at this prime location for observing the autumn migration of birds, butterflies, dragonflies, and bats through the courtesy of the Pennsylvania Game Commission and the Garrison Commander at Fort Indiantown Gap. The Second Mountain Hawk Watch Association is a non-profit organization that staffs the count site daily throughout the season and reports data to the North American Hawk Watch Association (posted daily at hawkcount.org).
Today, Second Mountain Hawk Watch was populated by observers who enjoyed today’s break in the rainy weather with a visit to the lookout to see what birds might be on the move. All were anxiously awaiting a big flight of Broad-winged Hawks, a forest-dwelling Neotropical species that often travels back to its wintering grounds in groups exceeding one hundred birds. Each autumn, many inland hawk watches in the northeast experience at least one day in mid-September with a Broad-winged Hawk count exceeding 1,000 birds. They are an early-season migrant and today’s southeast winds ahead of the remnants of Hurricane Florence (currently in the Carolinas) could push southwest-heading “Broad-wings” out of the Piedmont Province and into the Ridge and Valley Province for a pass by the Second Mountain lookout.
The flight turned out to be steady through the day with over three hundred Broad-winged Hawks sighted. The largest group consisted of several dozen birds. We would hope there are probably many more yet to come after the Florence rains pass through the northeast and out to sea by mid-week. Also seen today were Bald Eagles, Ospreys, American Kestrels, and a migrating Red-headed Woodpecker.
Migrating Broad-winged Hawks circle on a thermal updraft above Second Mountain Hawk Watch to gain altitude before gliding away to the southwest.
Migrating insects included Monarch butterflies, and the three commonest species of migratory dragonflies: Wandering Glider, Black Saddlebags, and Common Green Darner. The Common Green Darners swarmed the lookout by the dozens late in the afternoon and attracted a couple of American Kestrels, which had apparently set down from a day of migration. American Kestrels and Broad-winged Hawks feed upon dragonflies and often migrate in tandem with them for at least a portion of their journey.
Still later, as the last of the Broad-winged Hawks descended from great heights and began passing by just above the trees looking for a place to settle down, a most unwelcome visitor arrived at the lookout. It glided in from the St. Anthony’s Wilderness side of the ridge on showy crimson-red wings, then became nearly indiscernible from gray tree bark when it landed on a limb. It was the dreaded and potentially invasive Spotted Lanternfly (Lycorma delicatula). This large leafhopper is native to Asia and was first discovered in North America in the Oley Valley of eastern Berks County, Pennsylvania in 2014. The larval stage is exceptionally damaging to cultivated grape and orchard crops. It poses a threat to forest trees as well. Despite efforts to contain the species through quarantine and other methods, it’s obviously spreading quickly. Here on the Second Mountain lookout, we know that wind has a huge influence on the movement of birds and insects. The east and southeast winds we’ve experienced for nearly a week may be carrying Spotted Lanternflies well out of their most recent range and into the forests of the Ridge and Valley Province. We do know for certain that the Spotted Lanternfly has found its way into the Lower Susquehanna River Watershed.
This adult Spotted Lanternfly landed in a birch tree behind the observers at the Second Mountain Hawk Watch late this afternoon. It was first recognized by its bright red wings as it glided from treetops on the north side of the lookout.
