LIFE IN THE LOWER SUSQUEHANNA RIVER WATERSHED
A Natural History of Conewago Falls—The Waters of Three Mile Island
To take advantage of this unusually mild late-winter day, observers arrived by the thousands to have a look at an even greater number of migratory birds gathered at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area. Here are some highlights…
Trying to get a favorable place to nest before others arrive, the “early birds” are presently racing north through the lower Susquehanna valley. Check out these sightings from earlier today…
Time to get outside and have a look. The spectacle of spring migration passes quickly. You don’t want to miss it!
Flies? Cabbage White butterflies? Can it really be a late-February day? It certainly is. Here are a few more signs of an early spring.
In late March and early April, a rainy night and fog at daybreak can lead to an ideal morning for spotting migratory waterfowl and seabirds during their layover on the lower Susquehanna. Visibility was just good enough to spot these birds at Harrisburg, Pennsylvania, most of them feeding at midriver.
Inside the doorway that leads to your editor’s 3,500 square foot garden hangs a small chalkboard upon which he records the common names of the species of birds that are seen there—or from there—during the year. If he remembers to, he records the date when the species was first seen during that particular year. On New Year’s Day, the results from the freshly ended year are transcribed onto a sheet of notebook paper. On the reverse, the names of butterflies, mammals, and other animals that visited the garden are copied from a second chalkboard that hangs nearby. The piece of paper is then inserted into a folder to join those from previous New Year’s Days. The folder then gets placed back into the editor’s desk drawer beneath a circular saw blade and an old scratched up set of sunglasses—so that he knows exactly where to find it if he wishes to.
A quick glance at this year’s list calls to mind a few recollections.
Before putting the folder back into the drawer for another year, the editor decided to count up the species totals on each of the sheets and load them into the chart maker in the computer.
Despite the habitat improvements in the garden, the trend is apparent. Bird diversity has not cracked the 50 species mark in 6 years. Despite native host plants and nectar species in abundance, butterfly diversity has not exceeded 10 species in 6 years.
It appears that, at the very least, the garden habitat has been disconnected from the home ranges of many species by fragmentation. His little oasis is now isolated in a landscape that becomes increasingly hostile to native wildlife with each passing year. The paving of more parking areas, the elimination of trees, shrubs, and herbaceous growth from the large number of rental properties in the area, the alteration of the biology of the nearby stream by hand-fed domestic ducks, light pollution, and the outdoor use of pesticides have all contributed to the separation of the editor’s tiny sanctuary from the travel lanes and core habitats of many of the species that formerly visited, fed, or bred there. In 2019, migrants, particularly “fly-overs”, were nearly the only sightings aside from several woodpeckers, invasive House Sparrows (Passer domesticus), and hardy Mourning Doves. Even rascally European Starlings became sporadic in occurrence—imagine that! It was the most lackluster year in memory.
If habitat fragmentation were the sole cause for the downward trend in numbers and species, it would be disappointing, but comprehensible. There would be no cause for greater alarm. It would be a matter of cause and effect. But the problem is more widespread.
Although the editor spent a great deal of time in the garden this year, he was also out and about, traveling hundreds of miles per week through lands on both the east and the west shores of the lower Susquehanna. And on each journey, the number of birds seen could be counted on fingers and toes. A decade earlier, there were thousands of birds in these same locations, particularly during the late summer.
In the lower Susquehanna valley, something has drastically reduced the population of birds during breeding season, post-breeding dispersal, and the staging period preceding autumn migration. In much of the region, their late-spring through summer absence was, in 2019, conspicuous. What happened to the tens of thousands of swallows that used to gather on wires along rural roads in August and September before moving south? The groups of dozens of Eastern Kingbirds (Tyrannus tyrannus) that did their fly-catching from perches in willows alongside meadows and shorelines—where are they?
Several studies published during the autumn of 2019 have documented and/or predicted losses in bird populations in the eastern half of the United States and elsewhere. These studies looked at data samples collected during recent decades to either arrive at conclusions or project future trends. They cite climate change, the feline infestation, and habitat loss/degradation among the factors contributing to alterations in range, migration, and overall numbers.
There’s not much need for analysis to determine if bird numbers have plummeted in certain Lower Susquehanna Watershed habitats during the aforementioned seasons—the birds are gone. None of these studies documented or forecast such an abrupt decline. Is there a mysterious cause for the loss of the valley’s birds? Did they die off? Is there a disease or chemical killing them or inhibiting their reproduction? Is it global warming? Is it Three Mile Island? Is it plastic straws, wind turbines, or vehicle traffic?
The answer might not be so cryptic. It might be right before our eyes. And we’ll explore it during 2020.
In the meantime, Uncle Ty and I going to the Pennsylvania Farm Show in Harrisburg. You should go too. They have lots of food there.
There was a hint of what was to come. If you were out and about before dawn this morning, you may have been lucky enough to hear them passing by high overhead. It was 5:30 A.M. when I opened the door and was greeted by that distinctive nasal whistle. Stepping through the threshold and into the cold, I peered into the starry sky and saw them, their feathers glowing orange in the diffused light from the streets and parking lots below. Their size and snow-white plumage make Tundra Swans one of the few species of migrating birds you’ll ever get to visibly discern in a dark moonless nighttime sky.
The calm air at daybreak and through the morning transitioned to a steady breeze from the south in the afternoon. Could this be it? Would this be that one day in late February or the first half of March each year when waterfowl (and other birds too) seem to take advantage of the favorable wind to initiate an “exodus” and move in conspicuous numbers up the lower Susquehanna valley on their way to breeding grounds in the north? Well, indeed it would be. And with the wind speeding up the parade, an observer at a fixed point on the ground gets to see more birds fly by.
In the late afternoon, an observation location in the Gettysburg Basin about five miles east of Conewago Falls in Lancaster County seemed to be well-aligned with a northwesterly flight path for migrating Tundra Swans. At about 5:30 P.M., the clear sky began clouding over, possibly pushing high-flying birds more readily into view. During the next several hours, over three thousand Tundra Swans passed overhead, flocks continuing to pass for a short time after nightfall. There were more than one thousand Canada Geese, the most numerous species on similar days in previous years. Sometimes on such a day there are numerous ducks. Not today. The timing, location, and conditions put Tundra Swans in the spotlight for this year’s show.
Other migrants moving concurrently with the waterfowl included Ring-billed Gulls, American Herring Gulls (6+), American Robins (50+), Red-winged Blackbirds (500+), and Common Grackles (100+).
Though I’ve only seen such a spectacle only once during a season in recent years, there certainly could be another large flight of ducks, geese, or swans yet to come. The breeze is forecast to continue from southerly directions for at least another day. Keep you eyes skyward, no matter where you might happen to be in the lower Susquehanna valley. These or other migratory species may put on another show, a “big day”, just for you.
You remember the signs of an early spring, don’t you? It was a mild, almost balmy, February. The earliest of the spring migrants such as robins and blackbirds were moving north through the Lower Susquehanna River Watershed. The snow had melted and ice on the river had passed. Everyone was outdoors once again. At last, winter was over and only the warmer months lie ahead…beginning with March.
Ah yes, March, the cold windy month of March. We remember February fondly, but this March has startled us out of our vernal daydreams to wrestle with the reality of the season. And if you’re anywhere near the Mid-Atlantic states on this first full day of spring, you know that a long winter’s nap and visions of sugar peas would be time better spent than a stroll outdoors. Presently it’s dusk, and the snow from the 4th “Nor’easter” in a month is a foot deep and still falling.
In honor of “The Spring That Was”, here then is a sampling of some of the migratory waterfowl that have found their way to the Lower Susquehanna River Watershed during March. Some are probably lingering and feeding for a while. All will move along to their breeding grounds within a couple of weeks, regardless of the weather.
At the moment there is a heavy snow falling, not an unusual occurrence for mid-February, nevertheless, it is a change in weather. Forty-eight hours ago we were in the midst of a steady rain and temperatures were in the sixties. The snow and ice had melted away and a touch of spring was in the air.
Anyone casually looking about while outdoors during these last several days may have noticed that birds are indeed beginning to migrate north in the lower Susquehanna valley. Killdeer, American Robins, Eastern Bluebirds, Red-winged Blackbirds, and Common Grackles are easily seen or heard in most of the area now.
Just hours ago, between nine o’clock this morning and one o’clock this afternoon, there was a spectacular flight of birds following the river north, their spring migration well underway. In the blue skies above Conewago Falls, a steady parade of Ring-billed Gulls was utilizing thermals and riding a tailwind from the south-southeast to cruise high overhead on a course toward their breeding range.
The swirling hoards of Ring-billed Gulls attracted other migrants to take advantage of the thermals and glide paths on the breeze. Right among them were 44 American Herring Gulls, 3 Great Black-backed Gulls, 12 Tundra Swans (Cygnus columbianus), 10 Canada Geese, 3 Northern Pintails (Anas acuta), 6 Common Mergansers, 3 Red-tailed Hawks, a Red-shouldered Hawk, 6 Bald Eagles (non-adults), 8 Black Vultures, and 5 Turkey Vultures.
In the afternoon, the clouds closed in quickly, the flight ended, and by dusk more than an inch of snow was on the ground. Looks like spring to me.
Temperatures plummeted to well below freezing during the past two nights, but there was little sign of it in Conewago Falls this morning. The fast current in the rapids and swirling waters in flooded Pothole Rocks did not freeze. Ice coated the standing water in potholes only in those rocks lacking a favorable orientation to the sun for collecting solar heat during the day to conduct into the water during the cold nights.
On the shoreline, the cold snap has left its mark. Ice covers the still waters of the wetlands. Frost on exposed vegetation lasted until nearly noontime in shady areas. Insect activity is now grounded and out of sight. The leaves of the trees tumble and fall to cover the evidence of a lively summer.
The nocturnal bird flight is narrowing down to just a few species. White-throated Sparrows, a Swamp Sparrow (Melospiza georgiana), and Song Sparrows are still on the move. Though their numbers are not included in the migration count, hundreds of the latter are along the shoreline and in edge habitat around the falls right now. Song Sparrows are present year-round, migrate at night, and are not seen far from cover in daylight, so migratory movements are difficult to detect. It is certain that many, if not all of the Song Sparrows here today have migrated and arrived here recently. The breeding population from spring and summer has probably moved further south. And many of the birds here now may remain for the winter. Defining the moment of this dynamic, yet discrete, population change and logging it in a count would certainly require different methods.
Diurnal migration was foiled today by winds from southerly directions and moderating temperatures. The only highlight was an American Robin flight that extended into the morning for a couple of hours after daybreak and totaled over 800 birds. This flight was peppered with an occasional flock of blackbirds. Then too, there were the villains.
They’re dastardly, devious, selfish, opportunistic, and abundant. Today, they were the most numerous diurnal migrant. Their numbers made this one of the biggest migration days of the season, but they are not recorded on the count sheet. It’s no landmark day. They excite no one. For the most part, they are not recognized as migrants because of their nearly complete occupation of North America south of the taiga. If people build on it or alter it, these birds will be there. They’re everywhere people are. If the rotten attributes of man were wrapped up into one bird, an “anthropoavian”, this would be it.
Meet the European Starling (Sturnus vulgaris). Introduced into North America in 1890, the species has spread across the entire continent. It nests in cavities in buildings and in trees. Starlings are aggressive, particularly when nesting, and have had detrimental impacts on the populations of native cavity nesting birds, particularly Red-headed Woodpeckers, Purple Martins (Progne subis), and Eastern Bluebirds. They commonly terrorize these and other native species to evict them from their nest sites. European Starlings are one of the earlier of the scores of introduced plants and animals we have come to call invasive species.
Today, thousands of European Starlings were on the move, working their way down the river shoreline and raiding berries from the vines and trees of the Riparian Woodlands. My estimate is between three and five thousand migrated through during the morning. But don’t worry, thousands more will be around for the winter.
The NOAA National Weather Service radar images from last evening provided an indication that there may be a good fallout of birds at daybreak in the lower Susquehanna valley. The moon was bright, nearly full, and there was a gentle breeze from the north to move the nocturnal migrants along. The conditions were ideal.
The Riparian Woodlands at Conewago Falls were alive with migrants this morning. American Robins and White-throated Sparrows were joined by new arrivals for the season: Brown Creeper (Certhia americana), Ruby-crowned Kinglets (Regulus calendula), Golden-crowned Kinglets (Regulus satrapa), Dark-eyed Junco (Junco hyemalis), and Yellow-rumped Warbler (Setophaga coronata). These are the perching birds one would expect to have comprised the overnight flight. While the individuals that will remain may not yet be among them, these are the species we will see wintering in the Mid-Atlantic states. No trip to the tropics for these hardy passerines.
It was a placid morning on Conewago Falls with blue skies dotted every now and then by a small flock of migrating robins or blackbirds. The jumbled notes of a singing Winter Wren (Troglodytes hiemalis) in the Riparian Woodland softly mixed with the sounds of water spilling over the dam. The season’s first Wood Ducks (Aix sponsa), Blue-winged Teal (Spatula discors), American Herring Gull (Larus smithsonianus), Horned Larks (Eremophila alpestris), and White-throated Sparrows (Zonotrichia albicollis) were seen.
There was a small ruckus when one of the adult Bald Eagles from a local pair spotted an Osprey passing through carrying a fish. This eagle’s effort to steal the Osprey’s catch was soon interrupted when an adult eagle from a second pair that has been lingering in the area joined the pursuit. Two eagles are certainly better than one when it’s time to hustle a skinny little Osprey, don’t you think?
But you see, this just won’t do. It’s a breach of eagle etiquette, don’t you know? Soon both pairs of adult eagles were engaged in a noisy dogfight. It was fussing and cackling and the four eagles going in every direction overhead. Things calmed down after about five minutes, then a staring match commenced on the crest of the dam with the two pairs of eagles, the “home team” and the “visiting team”, perched about 100 feet from each other. Soon the pair which seems to be visiting gave up and moved out of the falls for the remainder of the day. The Osprey, in the meantime, was able to slip away.
In recent weeks, the “home team” pair of Bald Eagles, seen regularly defending territory at Conewago Falls, has been hanging sticks and branched tree limbs on the cross members of the power line tower where they often perch. They seem only to collect and display these would-be nest materials when the “visiting team” pair is perched in the nearby tower just several hundred yards away…an attempt to intimidate by homesteading. It appears that with winter and breeding time approaching, territorial behavior is on the increase.
In the afternoon, a fresh breeze from the south sent ripples across the waters among the Pothole Rocks. The updraft on the south face of the diabase ridge on the east shore was like a highway for some migrating hawks, falcons, and vultures. Black Vultures (Coragyps atratus) and Turkey Vultures streamed off to the south headlong into the wind after leaving the ridge and crossing the river. A male and female Northern Harrier (Circus hudsonius), ten Red-tailed Hawks, two Red-shouldered Hawks (Buteo lineatus), six Sharp-shinned Hawks, and two Merlins crossed the river and continued along the diabase ridge on the west shore, accessing a strong updraft along its slope to propel their journey further to the southwest. Four high-flying Bald Eagles migrated through, each following the east river shore downstream and making little use of the ridge except to gain a little altitude while passing by.
Late in the afternoon, the local Bald Eagles were again airborne and cackling up a storm. This time they intercepted an eagle coming down the ridge toward the river and immediately forced the bird to climb if it intended to pass. It turned out to be the best sighting of the day, and these “home team” eagles found it first. It was a Golden Eagle (Aquila chrysaetos) in crisp juvenile plumage. On its first southward voyage, it seemed to linger after climbing high enough for the Bald Eagles to loose concern, then finally selected the ridge route and crossed the river to head off to the southwest.
The humid rainy remains of Hurricane Nate have long since passed by Pennsylvania, yet mild wet weather lingers to confuse one’s sense of the seasons. This gloomy misty day was less than spectacular for watching migrating birds and insects, but some did pass by. Many resident animals of the falls are availing themselves of the opportunity to continue active behavior before the cold winds of autumn and winter force a change of lifestyle.
Warm drizzle at daybreak prompted several Northern Spring Peepers (Pseudacris crucifer crucifer) to begin calling from the wetlands in the Riparian Woodlands of Conewago Falls. An enormous chorus of these calls normally begins with the first warm rains of early spring to usher in this tiny frog’s mating season. Today, it was just a few “peeps” among anxious friends.
Any additional river flow that resulted from the rains of the previous week is scarcely noticeable among the Pothole Rocks. The water level remains low, the water column is fairly clear, and the water temperatures are in the 60s Fahrenheit.
It’s no real surprise then to see aquatic turtles climbing onto the boulders in the falls to enjoy a little warmth, if not from the sun, then from the stored heat in the rocks. As usual, they’re quick to slide into the depths soon after sensing someone approaching or moving nearby. Seldom found anywhere but on the river, these skilled divers are Common Map Turtles (Graptemys geographica), also known as Northern Map Turtles. Their paddle-like feet are well adapted to swimming in strong current. They are benthic feeders, feasting upon a wide variety of invertebrates found among the stone and substrate of the river bottom.
Adult Common Map Turtles hibernate communally on the river bottom in a location protected from ice scour and turbulent flow, often using boulders, logs, or other structures as shelter from strong current. The oxygenation of waters tumbling through Conewago Falls may be critical to the survival of the turtles overwintering downstream. Dissolved oxygen in the water is absorbed by the nearly inactive turtles as they remain submerged at their hideout through the winter. Though Common Map Turtles, particularly males, may occasionally move about in their hibernation location, they are not seen coming to the surface to breathe.
The Common Map Turtles in the Susquehanna River basin are a population disconnected from that found in the main range of the species in the Great Lakes and upper Mississippi basin. Another isolated population exists in the Delaware River.
SOURCES
Committee on the Status of Endangered Wildlife in Canada. 2002. Status Report of the Northern Map Turtle. Canadian Wildlife Service. Ottawa, Ontario.
A moderate breeze from the south placed a headwind into the face of migrants trying to wing their way to winter quarters. The urge to reach their destination overwhelmed any inclination a bird or insect may have had to stay put and try again another day.
Blue Jays were joined by increasing numbers of American Robins crossing the river in small groups to continue their migratory voyages. Killdeer (Charadrius vociferous) and a handful of sandpipers headed down the river route. Other migrants today included a Cooper’s Hawk (Astur cooperii), Eastern Bluebirds (Sialia sialis), and a few Common Mergansers (Mergus merganser), House Finches (Haemorhous mexicanus), and Common Grackles (Quiscalus quiscula).
The afternoon belonged to the insects. The warm wind blew scores of Monarchs toward the north as they persistently flapped on a southwest heading. Many may have actually lost ground today. Painted Lady (Vanessa cardui) and Cloudless Sulphur butterflies were observed battling their way south as well. All three of the common migrating dragonflies were seen: Common Green Darner (Anax junius), Wandering Glider (Pantala flavescens), and Black Saddlebags (Tramea lacerata).
The warm weather and summer breeze are expected to continue as the rain and wind from Hurricane Nate, today striking coastal Alabama and Mississippi, progresses toward the Susquehanna River watershed during the coming forty-eight hours.
