Columbia-Wrightsville Bridge – LIFE IN THE LOWER SUSQUEHANNA RIVER WATERSHED

March 22, 2022March 22, 2022

Seawatch on the Susquehanna

Birds that one might expect to see wintering among the surf and in tidal waters along the Atlantic coast are currently making their way up the Susquehanna on a route that will ultimately lead most to nesting sites in Canada. To see them as pass, one needs simply to find a good vantage point along the river from which to begin watching. Here are some of today’s sightings from the Veteran’s Memorial Bridge (Route 462) at Columbia/Wrightsville, Pennsylvania.

Birds/Waterfowl of Conewago Falls in the Lower Susquehanna River Watershed: Red-breasted Mergansers — Red-breasted Mergansers spend the winter primarily on saltwater bays. They are regular springtime migrants on the lower Susquehanna in late March and early April each year.

Birds/Waterfowl of Conewago Falls in the Lower Susquehanna River Watershed: Red-breasted Merganser — A male Red-breasted Merganser.

Double-crested Cormorants spend the winter in a variety of salt and brackish water habitats. Some birds breed on the lower Susquehanna, but the vast majority nest to the north of the Great Lakes.

Birds of Conewago Falls in the Lower Susquehanna River Watershed: Ring-billed Gull — Ring-billed Gulls winter throughout the Atlantic Coastal Plain and nest as far south as the Great Lakes region.

Birds of Conewago Falls in the Lower Susquehanna River Watershed: Bonaparte's Gulls — Bonaparte’s Gulls winter on the Atlantic from the surf zone to several miles offshore. In the southern states, some pass the colder months on inland lakes.

Birds of Conewago Falls in the Lower Susquehanna River Watershed: Bonaparte's and ring-billed Gulls — Bonaparte’s Gulls and a few Ring-billed Gulls swarm over the lower Susquehanna River at the Route 30 bridge.

April 25, 2021April 22, 2022

Conowingo Dam: Cormorants, Eagles, Snakeheads and a Run of Hickory Shad

Meet the Double-crested Cormorant, a strangely handsome bird with a special talent for catching fish. You see, cormorants are superb swimmers when under water—using their webbed feet to propel and maneuver themselves with exceptional speed in pursuit of prey.

Like many species of birds that dive for their food, Double-crested Cormorants run across the surface of the water to gain speed for a takeoff. Smaller wings may make it more difficult to get airborne, but when folded, they provide improved streamlining for submerged swimming.

Double-crested Cormorants, hundreds of them, are presently gathered along with several other species of piscivorous (fish-eating) birds on the lower Susquehanna River below Conowingo Dam near Rising Sun, Maryland. Fish are coming up the river and these birds are taking advantage of their concentrations on the downstream side of the impoundment to provide food to fuel their migration or, in some cases, to feed their young.

Double-crested Cormorants, mostly adult birds migrating toward breeding grounds to the north, are gathered on the rocks on the east side of the river channel below Conowingo Dam. A Great Blue Heron from a nearby rookery can be seen at the center of the image.

Bald Eagles normally gather in large numbers at Conowingo Dam in the late fall and early winter. Presently there are more than 50 there, and the majority of them are breeding age adults. Presumably they are still on their way north to nest. Meanwhile, local pairs are already feeding young, so it seems these transient birds are running a bit late. Many of them can be seen on the rocks along the east side of the river channel,…

…on the powerline trestles on the island below the dam…

…and in the trees surrounding Fisherman’s Park on the west shore.

In addition to the birds, the movements of fish attract larger fish, and even larger fishermen.

Anglers gather to fish the placid waters below the dam’s hydroelectric powerhouse . Only a few of the generating turbines are operating, so the flow through the dam is minimal.

Some water is being released along the west shoreline to attract migratory river herring to the west fish lift for sorting and retention as breeding stock for a propagation program. The east lift, the passage that hoists American Shad (Alosa sapidissima) to a trough that allows them to swim over the top of the dam to waters upriver, will begin operating as soon as these larger migratory fish begin arriving.

The excitement starts when the sirens start to wail and the red lights begin flashing. Yes friends, it’s showtime.

Red lights and sirens are a warning that additional flow is about to be released from the dam. Boaters should anticipate rough water and persons in and along the river need to seek higher ground immediately.

Gates are opened at mid-river to release a surge of water through the dam.

The wake from the release quickly reaches the shoreline, raising the water level in moments.

Experienced anglers know that the flow through the dam gets fish moving and can improve the catch significantly, especially in spring when many species are ascending the river.

Within minutes of the renewed flow, birds are catching fish.

A Double-crested Cormorant with a young Channel Catfish (Ictalurus punctatus).

A Double-crested Cormorant fleeing others trying to steal its Channel Catfish.

Another Double-crested Cormorant eating a Channel Catfish. Did you realize that Channel Catfish were an introduced species in the Susquehanna River system?

An Osprey with a stick, it’s too busy building a nest right now to fish.

Great Blue Herons swallow their prey at the spot of capture, then fly back to the nest to regurgitate a sort of “minced congealed fish product” to their young.

Then the anglers along the wave-washed shoreline began catching fish too.

This young man led off a flurry of catches that would last for the remainder of the afternoon.

Though Gizzard Shad are filter feeders that don’t readily take baits and lures, they are regularly foul-hooked and reeled in from the large schools that ascend the river in spring.

Gizzard Shad are very abundant in the lower Susquehanna, providing year-round forage for many species of predatory animals including Bald Eagles.

A Double-crested Cormorant swallowing a Gizzard Shad.

This angler soon helped another fisherman by landing his large catch, a Northern Snakehead (Channa argus).

The teeth of a Northern Snakehead are razor sharp. It is an aggressive non-native invasive species currently overtaking much of the Lower Susquehanna River Watershed. Anglers are encouraged to fish for them, catch them, keep them, and kill them at the site of capture. **Never transport a live Northern Snakehead anywhere at any time. It is illegal in both Maryland and Pennsylvania to possess a live snakehead.**

Northern Snakehead advisory sign posted at Exelon Energy’s Conowingo Fishermen’s Park.

A stringer of Northern Snakeheads. This species was imported from Asia as a food fish, so it has excellent culinary possibilities. It’s better suited for a broiler or frying pan than a river or stream.

Another stringer of Northern Snakeheads. It’s pretty safe to say that they have quickly become one of the most abundant predatory fish in the river. Their impact on native species won’t be good, so catch and eat as many as you can. Remember, snakeheads swim better in butter and garlic than in waters with native fish.

This foul-hooked Shorthead Redhorse (Moxostoma macrolepidotum), a native species of sucker, was promptly released.

Striped Bass are anadromous fish that leave the sea in spring to spawn in fresh water. They ascend the Susquehanna in small numbers, relying upon the operation of the fish passages at the Conowingo, Holtwood, Safe Harbor, and York Haven Dams to continue their journey upstream. During spring spawning, Striped Bass in the Susquehanna River and on the Susquehanna Flats portion of the upper Chesapeake Bay are not in season and may not be targeted, even for catch-and-release. This accidental catch was immediately turned loose.

After removal from the hook, this hefty Smallmouth Bass was returned to the river. Many anglers are surprised to learn that Smallmouth Bass are not native to the Susquehanna basin.

This angler’s creel contains a Northern Snakehead (left) and a Walleye (right). Did you know that the Walleye (Sander vitreus) is an introduced species in the Susquehanna watershed?

By late afternoon, anglers using shad darts began hooking into migrating Hickory Shad (Alosa mediocris), a catch-and-release species in Maryland.

Hickory Shad are recognized by their lengthy lower jaw. They are anadromous herring that leave the sea to spawn in freshwater streams. Hickory Shad ascend the Susquehanna as far as Conowingo Dam each year, but shy away from the fish lifts. Downriver from the dam, they do ascend Deer Creek along the river’s west shore and Octoraro Creek on the east side. In Pennsylvania, the Hickory Shad is an endangered species.

A Hickory Shad angled on a dual shad dart rig. During the spring spawning run, they feed mostly on small fish, and are the most likely of the Susquehanna’s herring to take the hook.

Simultaneous hook-ups became common after fours hours worth of release water from the dam worked its way toward the mouth of the river and got the schools moving. Water temperatures in the mid-to-upper-fifties trigger the ascent of Hickory Shad. On the Susquehanna, those temperatures were slow to materialize in the spring of 2021, so the Hickory Shad migration is a bit late.

Catch-and-release fishing for Hickory Shad appears to be in full swing not only at the dam, but along the downstream shoreline to at least the mouth of Deer Creek at Susquehanna State Park too.

Many Hickory Shad could be seen feeding on some of the millions of caddisflies (Trichoptera) swarming on the river. These insects, along with earlier hatches of Winter Stoneflies (Taeniopterygidae), not only provide forage for many species of fish, but are a vital source of natural food for birds that migrate up the river in March and April each year. Swallows, Ring-billed Gulls, and Bonaparte’s Gulls are particularly fond of snatching them from the surface of the water.

A Winter Stonefly (Taeniopterygidae) from an early-season hatch on the Susquehanna River at the Veteran’s Memorial Bridge at Columbia/Wrightsville, Pennsylvania. (March 3, 2021)

Just below Conowingo Dam, a lone fly fisherman was doing a good job mimicking the late-April caddisfly hatch, successfully reeling in numerous surface-feeding Hickory Shad.

You may have noticed the extraordinary number of introduced fish species listed in this account of a visit to Conowingo Dam. Sorry to say that there are two more: the Flathead Catfish (Pylodictis olivaris) and the Blue Catfish (Ictalurus furcatus). Like the Northern Snakehead, each has become a plentiful invasive species during recent years. Unlike the Northern Snakehead, these catfish are “native transplants”, species introduced from populations in the Mississippi River and Gulf Slope drainages of the United States. So if you visit the area, consider getting a fishing license and catching a few. Like the snakeheads, they too are quite palatable.

The arrival of migrating Hickory Shad heralds the start of a movement that will soon include White Perch, anadromous American Shad, and dozens of other fish species that swim upstream during the springtime. Do visit Fisherman’s Park at Conowingo Dam to see this spectacle before it’s gone. The fish and birds have no time to waste, they’ll soon be moving on.

To reach Exelon’s Conowingo Fisherman’s Park from Rising Sun, Maryland, follow U.S. Route 1 south across the Conowingo Dam, then turn left onto Shuresville Road, then make a sharp left onto Shureslanding Road. Drive down the hill to the parking area along the river. The park’s address is 2569 Shureslanding Road, Darlington, Maryland.

A water release schedule for the Conowingo Dam can be obtained by calling Exelon Energy’s Conowingo Generation Hotline at 888-457-4076. The recording is updated daily at 5 P.M. to provide information for the following day.

And remember, the park can get crowded during the weekends, so consider a weekday visit.

March 3, 2021March 4, 2021

Common Goldeneyes, Buffleheads, and Migrating Canada Geese

Spring migration is underway and waterfowl are on the move along the lower Susquehanna River. Here is a sample of sightings collected during a walk across the Veteran’s Memorial Bridge at Columbia-Wrightsville this morning.

At the Veteran’s Memorial Bridge, the Susquehanna was cresting this morning after recent rains and accompanying snow melt.

An overnight breeze from the southwest and calm winds during the morning hours created ideal conditions for flocks of Canada Geese to begin migrating north from Chesapeake Bay through the lower Susquehanna valley.

These three Common Mergansers and a Common Goldeneye (right) are some of the hundreds of diving ducks presently gathered on the river in the vicinity of the Veteran’s Memorial Bridge.

Common Goldeneyes and a first-winter male Bufflehead (upper right). All of the ducks seen today in the waters surrounding the bridge are benthic feeders, diving to the river bottom to pluck invertebrates from the substrate.

A male Common Goldeneye (Bucephala clangula).

Common Goldeneyes repositioning for another series of feeding dives.

During the morning flight, thousands of Canada Geese were seen moving north in flocks numbering about 50 to 100 birds each. The bird in the lower right was one of thousands of Ring-billed Gulls seen headed upriver as well.

As they pass over the lower Susquehanna region, migrating Canada Geese are typically observed flying much higher than flocks from the local resident populations, often reaching the cruising altitudes of aircraft. Aviators are always alert for flights of resident geese around airfields. But to prevent bird strikes during days like today when thousands of migratory geese traverse the airspace, air traffic controllers can become extra busy relaying the location and altitude of potential targets to pilots flying aircraft entering areas where birds have been reported.

This is, of course, just the beginning of the great spring migration. Do make a point of getting out to observe the spectacle. And remember, keep looking up—you wouldn’t want to miss anything.

July 15, 2019July 16, 2019

Friendly Neighborhood Spider, Man

Within the last few years, the early-summer emergence of vast waves of mayflies has caused great consternation among residents of riverside towns and motorists who cross the bridges over the lower Susquehanna. Fishermen and others who frequent the river are familiar with the phenomenon. Mayflies rise from their benthic environs where they live for a year or more as an aquatic larval stage (nymph) to take flight as a short-lived adult (imago), having just one night to complete the business of mating before perishing by the following afternoon.

In 2015, an emergence on a massive scale prompted the temporary closure of the mile-long Columbia-Wrightsville bridge while a blizzard-like flight of huge mayflies reduced visibility and caused road conditions to deteriorate to the point of causing accidents. The slimy smelly bodies of dead mayflies, probably millions of them, were removed like snow from the normally busy Lincoln Highway. Since then, to prevent attraction of the breeding insects, lights on the bridge have been shut down from about mid-June through mid-July to cover the ten to fourteen day peak of the flight period of Hexagenia bilineata, sometimes known as the Great Brown Drake, the species that swarms the bridge.

An adult (imago) male Great Brown Drake (Hexagenia bilineata) burrowing mayfly. Adult mayflies are also known as spinners.

A sub-adult (based on the translucence of the wings) female burrowing mayfly (Hexagenia species). The sub-adult (subimago or dun) stage lasts less than a day. Normally within 18 hours of leaving the water and beginning flight, it will molt into an adult, ready to breed during its final night of life.

After so many years, why did the swarms of these mayflies suddenly produce the enormous concentrations seen on this particular bridge across the lower Susquehanna? Let’s have a look.

Following the 2015 flight, conservation organizations were quick to point out that the enormous numbers of mayflies were a positive thing—an indicator that the waters of the river were getting cleaner. Generally, assessments of aquatic invertebrate populations are considered to be among the more reliable gauges of stream health. But some caution is in order in this case.

Prior to the occurrence of large flights several years ago, Hexagenia bilineata was not well known among the species in the mayfly communities of the lower Susquehanna and its tributaries. The native range of the species includes the southeastern United States and the Mississippi River watershed. Along segments of the Mississippi, swarms such as occurred at Columbia-Wrightsville in 2015 are an annual event, sometimes showing up on local weather radar images. These flights have been determined to be heaviest along sections of the river with muddy bottoms—the favored habitat of the burrowing Hexagenia bilineata nymph. This preferred substrate can be found widely in the Susquehanna due to siltation, particularly behind dams, and is the exclusive bottom habitat in Lake Clarke just downstream of the Columbia-Wrightsville bridge.

Native mayflies in the Susquehanna and its tributaries generally favor clean water in cobble-bottomed streams. Hexagenia bilineata, on the other hand, appears to have colonized the river (presumably by air) and has found a niche in segments with accumulated silt, the benthic habitats too impaired to support the native taxa formerly found there. Large flights of burrowing mayflies do indicate that the substrate didn’t become severely polluted or eutrophic during the preceding year. And big flights tell us that the Susquehanna ecosystem is, at least in areas with silt bottoms, favorable for colonization by the Great Brown Drake. But large flights of Hexagenia bilineata mayflies don’t necessarily give us an indication of how well the Susquehanna ecosystem is supporting indigenous mayflies and other species of native aquatic life. Only sustained recoveries by populations of the actual native species can tell us that. So, it’s probably prudent to hold off on the celebrations. We’re a long way from cleaning up this river.

In the absence of man-made lighting, male Great Brown Drakes congregate over waterways lit often by moonlight alone. The males hover in position within a swarm, often downwind of an object in the water. As females begin flight and pass through the swarm, they are pursued by the males in the vicinity. The male response is apparently sight motivated—anything moving through their field of view in a straight line will trigger a pursuit. That’s why they’re so pesky, landing on your face whenever you approach them. Mating takes place as males rendezvous with airborne females. The female then drops to the water surface to deposit eggs and later die—if not eaten by a fish first. Males return to the swarm and may mate again and again. They die by the following afternoon. After hatching, the larvae (nymphs) burrow in the silt where they’ll grow for the coming year. Feathery gills allow them to absorb oxygen from water passing through the U-shaped refuge they’ve excavated.

Several factors increase the likelihood of large swarms of Great Brown Drakes at bridges. Location is, of course, a primary factor. Bridges spanning suitable habitat will, as a minimum, experience incidental occurrences of the flying forms of the mayflies that live in the waters below. Any extraordinarily large emergence will certainly envelop the bridge in mayflies. Lights, both fixed and those on motor vehicles, enhance the appearance of movement on a bridge deck, thus attracting hovering swarms of male Hexagenia bilineata and other species from a greater distance, leading to larger concentrations. Concrete walls along the road atop the bridge lure the males to try to hover in a position of refuge behind them, despite the vehicles that disturb the still air each time they pass. The walls also function as the ultimate visual attraction as headlamp beams and shadows cast by moving vehicles are projected onto them over the length of the bridge. Vast numbers of dead, dying, and maimed mayflies tend to accumulate along these walls for this reason.

The absence of illumination from fixed lighting on the deck of the bridge reduces the density of Great Brown Drake swarms. Some communities take mayfly countermeasures one step further. Along the Mississippi, some bridges are fitted with lights on the underside of the deck to attract the mayflies to the area directly over the water, concentrating the breeding mayflies and fishermen alike. The illumination below the bridge is intended to draw mayflies away from light created by headlamps on motor vehicles passing by on the otherwise dark deck above. Lights beneath the bridge also help prevent large numbers of mayflies from being drawn away from the water toward lights around businesses and homes in neighborhoods along the shoreline—where they can become a nuisance.

Lights out on the Columbia-Wrightsville bridge. Dousing the lights to eliminate fixed illumination on bridges is an effective method of reducing the density of *Hexagenia bilineata* swarms.

With the bridge lights darkened, male Great Brown Drakes, their cellophane-like wings illuminated by headlamps to appear as white spots on the road, number in the hundreds instead of hundreds of thousands in swarms on the bridge near the east and west shorelines.

Swarms of Great Brown Drake mayflies are still present at the Columbia-Wrightsville bridge, they’re just not concentrated there in enormous numbers. Evidence includes their bodies found in cobwebs along the entire length of the span.

The aptly-named Bridge Orb Weaver (Larinioides sclopetarius) constructs webs along the entire length of the Columbia-Wrightsville bridge, and on many of the buildings at both ends. The abundance of victims tangled in silk must overwhelm their appetite, or maybe they actually consume only the smaller insects. They have their choice. Of the Bridge Orb Weaver, Uncle Ty Dyer says, “When you live along the river, it’s your friendly neighborhood spider, man.”

The native Eastern Dobsonfly (Corydalus cornutus) is among the reliable indicators of stream quality in the Susquehanna at the Columbia-Wrightsville bridge. Winged adults, which live for about a week, are clumsy fliers attracted to lights. The aquatic larvae are known as hellgrammites, which require clean flowing water over rocky or pebbly substrate to thrive. Two adults were found on the bridge last evening. It would be encouraging to find more. Maybe we’ll stop back to have another look when the lights are back on.

SOURCES

Edsall, Thomas A. 2001. “Burrowing Mayflies (Hexagenia) as Indicators of Ecosystem Health.” Aquatic Ecosystem Health and Management. 43:283-292.

Fremling, Calvin R. 1960. Biology of a Large Mayfly, Hexagenia bilineata (Say), of the Upper Mississippi River. Research Bulletin 482. Agricultural and Home Economics Experiment Station, Iowa State University. Ames, Iowa.

McCafferty, W. P. 1994. “Distributional and Classificatory Supplement to the Burrowing Mayflies (Ephemeroptera: Ephimeroidea) of the United States.” Entomological News. 105:1-13.