The Stewards of Inland Shorebird Habitat

As they travel between coastal wintering ranges and breeding territories in Canada and Alaska, vast numbers of shorebirds pass through the lower Susquehanna region each spring and fall—though few stop here to rest, feed, and provide us with an opportunity to observe them.  Prior to the construction of man-made dams and other alterations on our lotic (flowing) waterways including the river, shorebirds took advantage of lateral bars, stream deltas, and other alluvial deposits as places to loaf and re-energize.  Before they were drained and filled, some of the valley’s wetlands probably included sparsely vegetated flats where shorebirds could drop in for a brief visit.  Previous to their extirpation from our region, North American Beavers were the primary providers of quality habitat for shorebirds and other migrating waders on our lotic waters.  Their widespread network of dams, pools, and marshes maximized floodplain function by keeping streams thoroughly connected to their wetlands, nurturing plant communities that not only provided food and shelter for the beavers and other wildlife, but provided superb buffering against erosion while protecting against sediment and nutrient imbalances in lower Susquehanna waterways.

Beaver Dam and Lodge
Beaver dams need not be large, particularly on low-gradient streams where a structure like this is sufficient to create a pool with depth adequate for building and maintaining a lodge and transporting leafy branches and other food items by water.
A beaver lodge assembled in a pool with less than three feet of water, deep enough to provide the family with a measure of protection against terrestrial predators.
Beaver Pool
Beaver pool ecosystems provide homes for hundreds of species of plants and animals, including migrating shorebirds and other waders.
Solitary Sandpiper
Mud flats in the margins between emergent shrubs and herbaceous plant growth attract migrating shorebirds like this Solitary Sandpiper to the abundance of invertebrate life.  Seasonal movements of migrating shorebirds regularly coincide with the reductions of water levels in beaver pools which typically occur between May and September each year.
Least Sandpiper
During its southbound migration in September, a Least Sandpiper searches for arthropods and annelids as it visits a food-rich puddle along the periphery of a beaver pool.
Pectoral Sandpiper
A Pectoral Sandpiper feeds in shallow water near a stand of Common Cattails in a beaver pool.
Sora
During migration, rails including the Sora are attracted to dense emergent vegetation in beaver pools.  Some will nest in these rodent-managed refugia.
Green Heron
Green Herons visit beaver pools during migration and, due to the reliability of the food supply, will often nest in their vicinity.

So how did this happen?  How did the North American Beaver become a keystone species—an animal upon which the majority of other life forms within its ecosystem are so reliant?  Well, it’s largely due to the fact that our beavers aren’t particularly fond of a constant stream of noise.  More specifically, they don’t like the sound of running water in places where they intend to build and maintain a lodge.  And so, as they begin to place sticks, mud, limbs, stones, and other materials within a noisy riffle on a stream, they create a dam, and behind it a pool—a pool that is particularly advantageous for protecting their home and providing a means of conveyance for their construction materials and food supplies.

Cascading Series of Beaver Pools
On a high-gradient segment of stream, beavers will create a cascading series of pools. Because water filters through a beaver dam instead of spilling over it, the work of these meticulous rodents soon silences the sounds of water changing altitude.  No more sonorous riffle.
Beaver Dam
Quiet please.  High seasonal stream flow and damage from storms may create areas where water begins to erode the structure of the dam.  Where this condition persists, an adult beaver will soon mend the breach, just to quiet things down.  Why would beavers demand such a hush upon their domain?  Well, they have poor eyesight, but their hearing is excellent, and they rely upon it to detect danger.
Louisiana Waterthrush
The Louisiana Waterthrush (Parkesia motacilla), one of our earliest-arriving warblers, nests in forests along clear, high-gradient streams.  In mid-April, we found this individual and three others squabbling over a breeding territory adjacent to the series of cascading beaver pools shown in the previous images.
Brook Trout and Blacknose Dace
Native denizens of coldwater streams, neither the Brook Trout nor the Eastern Blacknose Dace has any difficulty finding its way through the voids in beaver dams to ascend and descend the sequence of pools.
Beaver Dam and Pool on Low-gradient Stream
On a low-gradient segment of stream, a dam just over a foot in height may be sufficient to create a beaver pool of considerable size.  Resembling the water-logged muskeg of the far north, well-established beaver pools form boggy habitats familiar to migrating shorebirds.
Water levels in the pools usually drop along with the stream’s base flow as soon as the effects of the spring thaw and rain showers subside.  Feeding areas in shallow water and on muddy ground are often revealed just in time for northbound shorebirds and other waders to stop by in late April and May.  These conditions often persist through the growing season and the fall migration of these birds which begins as early as late June and sometimes extends into October.
Tree Swallow at Natural Cavity
Where the pool inundates standing timber such as Red Maple and other species not used by beavers, the dead snags provide vital feeding areas for birds and other wildlife.  Cavity nesters like this Tree Swallow seldom find suitable natural housing elsewhere.
Eastern Newts Among Alderfly Exuvia
The mosaic of marshlands and braided stream channels within the beaver pool complex supports an abundance of aquatic life including these breeding Eastern Newts, seen here surrounded by the exuvia left behind following a massive hatch of alderflies (Sialinae).  Alderflies are a stream inhabitant during their larval stage and are indicator of clean water conditions.
A modern-day example of the way fully functional stream floodplains used to look in the Lower Susquehanna River Watershed.  Though the pools may appear pond-like during the cooler and wetter months of the year, by summer the water levels behind the beaver dams recede as the base flow of the stream wanes.  Now the true nature of the marsh, shallow pool, and braided stream complex is revealed.  Unlike most man-made dams that set a fixed pool elevation regardless of flow by discharging water over the top of the structure or through a spillway or gate, beaver dams merely throttle the flow through their porous construction.  Unless the beaver begins plugging small leaks as fast as the stream flow ebbs, the water level in the pools will drop.  So they’re not left high and dry, lodges are located in the deepest pools, usually in close proximity to the dam and/or one of the stream courses.  Though not inundated during the dry season, the soil in the pool complex is almost always damp and plants grow vigorously there, sequestering nutrients and retaining sediments in beneficial deposition patterns that actually inhibit erosion of the riparian landscape.  These streams and floodplains retain their hyporheic zone and freely exchange water with the underlying water table and aquifer.  It’s the ultimate floodplain management system, and the beavers don’t even know their doing all the work.  (Google Maps base image)
Where another high-gradient segment of stream enters the main pool complex, beavers have created an additional series of cascading pools.  The impoundments created by these dams help diffuse stormwater energy and process more nutrients and sediments within the floodplain’s wetland vegetation system.  (Google Maps base image)
Sandhill Cranes in Beaver Pool
Beaver pools often refill as stream flows increase following autumn rains.  The stockpiles of vegetative foods that grew within the beaver’s domain through the summer then become flooded and are a prime source of nutrition for not only the beavers, but waterfowl during both their autumn and spring migrations.  Gleaning and probing Sandhill Cranes often find these habitats to their liking as well.

While North American Beavers have returned to the region, most live as “bank beavers”, residing in the river and larger creeks of the valley where they excavate shelters among the roots of Black Willows and other shoreline trees and shrubs.  Floodplain encroachment, legacy sediment deposits, and just plain human intolerance have all conspired to prohibit North American Beavers from performing their magic on smaller local streams.  For migrating shorebirds, this continued absence of beaver dam ecosystems has turned much of the lower Susquehanna valley into “flyover country”.  Those travelers that do stop to rest and feed concentrate at the few favorable locations such as the lateral bars and the hydroelectric dam-created delta at the Conejohela Flats on the river in Lancaster County.  But centralization has its drawbacks.  Migrants spending time at concentration points may have a greater chance of contracting and spreading disease.  Protracted heavy foraging can degrade these habitats.  And over time, features such the lateral bars and delta deposits, including those on the Conejohela Flats, transition into other habitats—riparian forests.  A more widespread selection of favorable stopover points for shorebirds, waders, waterfowl, and other migrants is certainly desirable.

IMITATION IS THE SINCEREST FORM OF FLATTERY

Until public sentiment sways in favor of the North American Beaver, wildlife managers are mimicking some of the attributes of their sound-inspired installations.

New Impoundment at Middle Creek Wildlife Management Area
Created by excavating a depression in heavy clay soils, this new impoundment at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area collects water directly from rainfall and from surface runoff.  Its depth at no point is greater than about two or three feet.  During the driest times of the year, this space will be a mudflat, and a haven for migrating shorebirds.

Shallow-water conservation impoundments designed, constructed, and managed for migratory waterfowl and waders including shorebirds are not what we typically refer to as ponds—though they are lentic (still) waters.  Similar shallow freshwater impoundments at our National Wildlife Refuges are referred to as pools by the United States Fish and Wildlife Service, but smaller versions like this example at Middle Creek very closely resemble the prairie potholes created by glacial scour in the north-central United States and adjacent portions of Canada.  Many populations of migratory birds are familiar with pothole ecosystems and, like the beaver pools and marshes, have relied upon them as waypoints along their journeys for centuries.

Pond and Lake Zonation
Impoundments most beneficial to migrating waterfowl and waders including shorebirds are shallow in depth.  They lack the deeper waters of a pond or lake and thus have no limnetic (open water) or aphotic zones.  Managed throughout as a littoral zone, impoundments grow plants in shallow water or on damp soil through the summer months to chew up nutrients accumulated in waste deposited by visiting birds.  This vegetation is then flooded from late fall through early spring as forage habitat for migratory waterfowl.  The timing of the fluctuations in water levels approximates those of the beaver pools and marshes on lotic (flowing) waters.
Impoundment Gate
A gate assembly is a water control structure installed to provide seasonal management of the water levels in a conservation impoundment.  More than once, beavers have heard the sounds of tumbling waters inside these types of devices and tried to dam them up!

You’ve heard the line, “If you build it, they will come.”  Well, it’s true.  Here is a sample of the activity witnessed during the past two weeks at the new impoundment completed just several months ago at Middle Creek Wildlife Management Area along the tour route just beyond the “Stop 3” overlook.

Snow Geese
The ubiquitous Snow Geese were among the first migratory occupants of the new impoundment.  A few are currently lingering in the vicinity.
Blue-winged Teal
Ducks soon followed.  These Blue-winged Teal were among the last to pass through earlier this month.
Wilson's Snipe
Wilson’s Snipe were among the first species of migratory shorebirds to visit the new impoundment at Middle Creek…
Killdeer
…as were Killdeer, a species which nests nearby.
Least Sandpipers
Then, earlier this month, flocks of shorebirds including these Least Sandpipers were arriving to feed and rest.
Least Sandpipers
Least Sandpipers search for small invertebrates in shallow water and exposed soil.
Least Sandpiper and a Semipalmated Sandpiper
A Least Sandpiper (left) and a Semipalmated Sandpiper (right).
Solitary Sandpiper
Not all shorebirds seen by themselves are alone, and that includes the Solitary Sandpiper.
White-rumped, Least, and Solitary Sandpipers
Here’s a Solitary Sandpiper (right) feeding alongside a Least Sandpiper (center) and a White-rumped Sandpiper (left).
Least Sandpiper and a White-rumped Sandpiper
A Least Sandpiper (left) and a White-rumped Sandpiper (right).
White-rumped Sandpiper
Rare in our area during spring, the White-rumped Sandpiper (Calidris fuscicollis) flies north using the central flyway, then heads south along the Atlantic flyway in the fall, when it tends to be more regular here.
Spotted Sandpiper
Rocking and teetering along as it looks for food, the Spotted Sandpiper may be one of the easiest of shorebirds to identify.
Dunlin
Chunky little Dunlin (Calidris alpina) with their conspicuously down-curved bills are another easy-to-identify species, particularly in the spring when their breeding (alternate) plumage includes a black belly.
Black-bellied Plovers
Black-bellied Plovers acquire a handsome set of plumage in the spring as well.
Killdeer
Killdeer too are plovers and this pair appears to have taken up residence on barren ground along the periphery of the new impoundment.
Semipalmated Plover
The Semipalmated Plover doesn’t have nearly the flair for ornament its close relatives the Killdeer do; these little shorebirds wear only one ring around their neck instead of two.  Think this plover is cute?,…
Killdeer
…then check out this newly hatched Killdeer.  It starts life with just one necklace too, but acquires a second as it grows.  Look at those legs!  (If you visit the new impoundment at Middle Creek, drive slowly and please watch where you’re going.  Baby Killdeer and other young birds, as well as mammals and turtles, are commonly crossing the paved surfaces right now.)
Lesser Yellowlegs
Speaking of legs, here’s one of dozens of Lesser Yellowlegs that visited the new impoundment during their recent northbound travels.
Greater Yellowlegs
Though less numerous than the smaller Lesser Yellowlegs, a Greater Yellowlegs seldom goes unnoticed when dropping by the man-made pothole habitat.
Greater Yellowlegs
Have trouble telling a Greater Yellowlegs like this one from a Lesser Yellowlegs?  Look for the heavier, longer bill on the former, as well as dark barring along its flanks below the wings while the bird is in breeding (alternate) plumage during the spring migration.
Wilson's Phalarope
Everyone likes to see something unusual every now and then, and this impoundment delivers.  Just yesterday, we photographed this migrating Wilson’s Phalarope (top center) among two Least Sandpipers (top left and right), a Lesser Yellowlegs (left and slightly forward of the phalarope), and a White-rumped Sandpiper (foreground).
Wilson's Phalarope
Renowned for spinning in circles as they feed in shallow water, Wilson’s Phalaropes (Phalaropus tricolor) passing through the lower Susquehanna basin are headed toward nesting areas in the prairie wetlands, including potholes, of the northern United States and adjacent sections of southern Canada.

Species of wildlife in addition to shorebirds and waterfowl have already found the new impoundment favorable…

Spot-winged Gliders
Pairs of breeding Spot-winged Gliders (Pantala hymenaea), seen here in tandem while ovipositing, were swarming the impoundment after arriving earlier this week.  These dragonflies are ofttimes unpredictable nomads and are similar in appearance to the usually more numerous Wandering Gliders.  To recolonize seasonal portions of their range, both are famous for hitching a ride en masse on storm systems.  They share the behavior of finding ephemeral and new bodies of water favorable for egg laying due to their low density of aquatic predators.
Snapping Turtle
We watched this Snapping Turtle arrive to apparently find the newly created waters to its liking.  Snapping Turtles are important consumers in a wetland ecosystem.  Larger specimens may fill the role of an upper trophic level or apex predator, eliminating vulnerable mid-level consumers including other Snapping Turtles.
American Pipit
Because it bounces its tail up and down like a Spotted Sandpiper, you may at first glance mistake an American Pipit for a shorebird.  Long known as the Water Pipit, these songbirds have been visiting the impoundment while migrating north and stopping to feed in nearby croplands.
Glossy Ibis
Seen here with Least Sandpipers is a visiting Glossy Ibis.
Glossy Ibis
Sometimes twenty or more of these mostly coastal waders have made a pit stop at the “new Middle Creek pothole”, though none have thus far chosen to remain long.  Apparently, food sources sufficient to sustain a bird of their size have yet to develop in its benthos.
Western Cattle Egrets
Western Cattle Egrets visiting Middle Creek this spring have been frequenting the new impoundment.
Western Cattle Egret
With still little in the way of insects such as grasshoppers available in the surrounding landscape,…
Green Frog
…cattle egrets are looking to find prey like this Green Frog.
Western Cattle Egret
Many have been observed hunting the adjacent grasslands..
Western Cattle Egret
Where small mammals, mostly Meadow Voles, are being taken in abundance.

Managing saturation levels in shallow-water impoundments to resemble the seasonal variations in beaver pool and marsh systems can create lush growth and wildlife-rich environments.  Take a look at some images from a project in a headwaters area of a tributary to Conewago Creek (west)…

Well-established Shallow-water Impoundment
By late July, southbound shorebirds were already using these mudflats to feed and rest.  Other sections of the impoundment were dense with emergent and aquatic plants, the latter kept hydrated in deeper pools of the project by the inflow from several captured springs that supplement direct rainfall and sheet runoff to supply its water.  During a seasonal drawdown, the exposure of the impoundment’s soils to direct sunlight can provide a measure of disinfection to reduce the chances of disease transmission among its populations of visiting birds and other animals.
Water Lily Growth
In the deeper pools of the impoundment, water lilies and other aquatic plants grow in lush mats to provide cover and feeding areas for resident populations of breeding reptiles and amphibians.
Green Heron
An abundance of foods are available for waders including this Green Heron…
Immature Little Blue Heron
…and this immature Little Blue Heron, a wanderer typical of more southern latitudes.
Sandhill Cranes
While walking the road among tall grasses in the supporting landscape surrounding this impoundment, we were at first startled when these Sandhill Cranes strode by going the other direction.  We quietly kept moving,…
Sandhill Cranes
…then spotted them again as we looked across the impoundment to realize they weren’t alone, but were escorting a colt.
Sandhill Cranes with Colt
The hatching of this colt is testimony to the vital role wetland ecosystems play in the lives of hundreds of species.  Whether they be beaver pools and marshes on lotic waters or man-made shallow lentic waters, each of these habitats is filling a void that left floodplains and other critical lowland biomes faltering.  While they can’t replace the full-function floodplain management provided by an active beaver colony, shallow-water impoundments can provide relief for habitat-starved populations of the animals and plants that rely upon them.  A constellation of these projects on lands public or private across the lower Susquehanna watershed could help provide refuge for many of our flora and fauna with the most desperately fragmented of ranges.

So that you can relax while observing the comings and goings at a pair of the lower Susquehanna valley’s man-made impoundments, the Pennsylvania Game Commission has erected two viewing pavilions for public use on its lands…

The Haldeman Island observation pavilion is located on State Game Lands 290  just upstream from the Juniata River’s mouth on the Susquehanna at Clark’s Ferry in Dauphin County.
Haldeman Island Pavilion
It overlooks not only the island’s man-made shallow-water impoundments and neighboring grasslands, but the tower used in the 1970s to reintroduce Bald Eagles from Saskatchewan to the lower Susquehanna.  Interpretive signs explain the conservation stories of habitats and the eagle reintroduction program.
Middle Creek W.M.A. Observation Pavilion
The observation pavilion at Middle Creek Wildlife Management Area is of similar construction to the one at Haldeman Island.  It is accessed from the parking pull-off along the tour road at its intersection with Chapel Road, just before the right turn and incline that leads to the “Stop 3” grassland overlook.
It too includes numerous interpretive signs to help visitors understand impoundment management.

During the next two weeks, the exodus of migrating shorebirds now staged and feeding upon Atlantic Horseshoe Crab eggs on Delaware Bay will commence.  During the evening of their departure from the bay, many of these birds cross portions of the Lower Susquehanna River Watershed, particularly east of the river.  Stormy weather and other climatic conditions may force some of them to seek a place to put down temporarily, so keeping a close eye on the new pothole-like impoundment at Middle Creek may be a prudent move.  After that, waders known as “post-breeding wanderers” can show up at any time.  Then, beginning as early as late June, shorebirds begin moving south on a migration that can provide us with viewing opportunities into September and beyond.  See you out there!

Shorebirds on the Mud in York County, Pennsylvania

At Lake Redman just to the south of York, Pennsylvania, a draw down to provide drinking water to the city while maintenance is being performed on the dam at neighboring Lake Williams, York’s primary water source, has fortuitously coincided with autumn shorebird migration.  Here’s a sample of the numerous sandpipers and plovers seen today on the mudflats that have been exposed at the southeast end of the lake…

Least Sandpiper at Lake Redman, York County, Pennsylvania.
One of a hundred or more Least Sandpipers seen on mudflats at Lake Redman today.
 A Semipalmated Plover and a Least Sandpiper at Lake Redman, York County, Pennsylvania.
A Semipalmated Plover and a Least Sandpiper
Pectoral Sandpipers at Lake Redman, York County, Pennsylvania.
Pectoral Sandpipers.
A Pectoral Sandpiper and Least Sandpipers at Lake Redman, York County, Pennsylvania.
A Pectoral Sandpiper and two Least Sandpipers.
A Semipalmated Sandpiper at Lake Redman, York County, Pennsylvania.
A Semipalmated Sandpiper.
A Stilt Sandpiper at Lake Redman, York County, Pennsylvania.
A Stilt Sandpiper feeding.
Stilt Sandpiper consuming an edible at Lake Redman, York County, Pennsylvania.
Stilt Sandpiper consuming an edible.
Stilt Sandpiper at rest at Lake Redman, York County, Pennsylvania.
Stilt Sandpiper at rest.
A Solitary Sandpiper at Lake Redman, York County, Pennsylvania.
A Solitary Sandpiper
A Lesser Yellowlegs at Lake Redman, York County, Pennsylvania.
A Lesser Yellowlegs.
A Greater Yellowlegs at Lake Redman, York County, Pennsylvania.
A Greater Yellowlegs.
Osprey at Lake Redman, York County, Pennsylvania.
Stirring up the shorebird crowd every now and then were several Ospreys, but all would soon be back to the business of feeding in the mud.
An Osprey hovers above shallow water near the mudflats as it searches for fish.
An Osprey hovers above shallow water near the mudflats as it searches for fish.

Not photographed but present at Lake Redman were at least two additional species of shorebirds, Killdeer and Spotted Sandpiper—bringing the day’s tally to ten.  Not bad for an inland location!  It’s clearly evident that these waders overfly the lower Susquehanna valley in great numbers during migration and are in urgent need of undisturbed habitat for making stopovers to feed and rest so that they might improve their chances of surviving the long journey ahead of them.  Mud is indeed a much needed refuge.

The Colorful Birds Are Here

You need to get outside and go for a walk.  You’ll be sorry if you don’t.  It’s prime time to see wildlife in all its glory.  The songs and colors of spring are upon us!

Flooding that resulted from mid-week rains is subsiding.  The muddy torrents of Conewago Falls are seen here racing by the powerhouse at the York Haven Dam.
Receding waters will soon leave the parking area at Falmouth and other access points along the river high and dry.
Migrating Yellow-rumped Warblers are currently very common in the riparian woodlands near Conewago Falls.  They and all the Neotropical warblers, thrushes, vireos, flycatchers are moving through the Susquehanna watershed right now.
A Baltimore Oriole feeds in a riverside maple tree.
Ruby-crowned Kinglets are migrating through the Susquehanna valley.  These tiny birds may be encountered among the foliage of trees and shrubs as they feed upon insects .
Gray Catbirds are arriving.  Many will stay to nest in shrubby thickets and in suburban gardens.
American Robins and other birds take advantage of rising flood waters to feed upon earthworms and other invertebrates that are forced to the soil’s surface along the inundated river shoreline.
Spotted Sandpipers are a familiar sight as they feed along water’s edge.
The Yellow Warbler (Setophaga petechia) is a Neotropical migrant that nests locally in wet shrubby thickets.  Let your streamside vegetation grow and in a few years you just might have these “wild canaries” singing their chorus of “sweet-sweet-sweet-I’m-so-sweet” on your property.

If you’re not up to a walk and you just want to go for a slow drive, why not take a trip to Middle Creek Wildlife Management Area and visit the managed grasslands on the north side of the refuge.  To those of us over fifty, it’s a reminder of how Susquehanna valley farmlands were before the advent of high-intensity agriculture.  Take a look at the birds found there right now.

Red-winged Blackbirds commonly nest in cattail marshes, but are very fond of untreated hayfields, lightly-grazed pastures, and fallow ground too.  These habitats are becoming increasingly rare in the lower Susquehanna region.  Farmers have little choice, they either engage in intensive agriculture or go broke.
Nest boxes are provided for Tree Swallows at the refuge.
Numbers of American Kestrels have tumbled with the loss of grassy agricultural habitats that provide large insects and small rodents for them to feed upon.
White-crowned Sparrows (Zonotrichia leucophrys) are a migrant and winter resident species that favors small clumps of shrubby cover in pastures and fallow land.
When was the last time you saw an Eastern Meadowlark (Sturnella magna) singing “spring-of-the-year” in a pasture near your home?
And yes, the grasslands at Middle Creek do support nesting Ring-necked Pheasants (Phasianus colcichus).  If you stop for a while and listen, you’ll hear the calls of “kowk-kuk” and a whir of wings.  Go check it out.

And remember, if you happen to own land and aren’t growing crops on it, put it to good use.  Mow less, live more.  Mow less, more lives.

Birds of a Feather-The Basics

When we look at birds, we are fascinated by the unique structure and appearance of their feathers.  They set birds apart from all other life forms on the planet.  Feathers enable most birds to achieve a feat long envied by humans…flight.  Birds on the wing awaken a curiosity in man.  They are generally the largest animals one will see in the air.  People want to know the name of a bird they see flying by, and want to know more about it.  The method and style of bird flight can aid an observer who attempts to determine which of the world’s 10,000 bird species he or she is studying.  Body shape and bird sounds often tell us a lot about the birds we encounter.  But most often, we rely on the unique colors, patterns, and shapes of the feathers, the plumage, to identify the bird we are seeing.

To birds, feathers are survival.  They are lightweight and strong to support the mechanics of flight.  Feathers are superb insulators against the elements, and provide additional buoyancy for birds spending time on the water.  For most birds, feathers provide a coloration and a texture similar to their surroundings, enabling them to hide from predators or to stalk prey.  In the case of some species, extravagant showy plumage is acquired, at least during the breeding season, and often only by males, as a way to attract a mate, intimidate rivals, defend a territory, or lure an intruder away from a nest site.  Because they become worn and damaged, all feathers are periodically molted and replaced by fresh plumage.

The feathers worn by a young bird leaving the nest are called the juvenile plumage.  Typically, this is followed by a molt into a basic (non-breeding) plumage.  The oft times extravagant breeding feathers are the result of a molt into an alternate (breeding) plumage.

While making field observations, the species, subspecies, gender, age, and other vital statistics of a bird can often by discerned easily by noting the plumage.  In the case of some other birds, diligence, experience, research, and an exceptionally good look and/or a photograph may be required to interpret these particulars.  In still other instances, a trained expert with a specimen in the hand is the only method of learning the bird’s identity and background.

This juvenile House Wren (Troglodytes aedon) left the nest wearing plumage very similar to that of its parents.  In lieu of bright colors, the male House Wren relies upon a vigorous bubbly song and a scrappy demeanor to defend its breeding territory.  This species nests in cavities on the edges of the Riparian Woodlands of Conewago Falls.  Males may have more than one mate.  House Wrens probably migrate at night and will winter in the southern border states and further south.

The age at which birds acquire adult breeding and non-breeding plumages varies by species.  Many juvenile birds resemble adults in basic (non-breeding) plumage as soon as they leave the nest.  For these birds, there is little difference between their juvenile plumage and the appearance of the feathers which follow the molt into their first basic (non-breeding) plumage.  Bird species which sexually mature within their first year may acquire their first basic (non-breeding) plumage before arrival of their first winter, followed by an alternate (breeding) plumage by their first spring.  This is particularly true for smaller short-lived birds.  Other species, normally larger long-lived ones, may experience a sequence of molts through multiple basic (non-breeding) plumages over a period of years prior to resembling an adult.  Some of these species, such as eagles, retain their juvenile plumage for as long as a year before extensive molting into a first basic (non-breeding) plumage begins.  Still others, including many gulls, attain a first-winter (formative) plumage prior to molting into their first basic (non-breeding) set of feathers.  Sexual maturity and initiation of an annual molt to alternate (breeding) plumage, if there is one, may take as long as three to five years for these bigger birds.

For nearly all species of birds, the molts which produce basic (non-breeding) plumage occur on at least an annual basis and include a total replacement of feathers.  This process renews worn and missing plumes including the flight feathers of the wings and tail.  Any molt to alternate (breeding) plumage often excludes the replacement of the feathers of the wings and tail.  There are many exceptions to these generalities.

A mid-summer nesting species, the American Goldfinch, Spinus tristis, male (left and right) molts into a glamorous alternate plumage for the breeding season.  The adult female’s alternate plumage (top) is a subdued green-yellow and black.  Her feathers resemble the foliage around the nest and offer protection from discovery during incubation.  Juvenile plumage (bottom) is similar to that of the female, but duller with a buffy tone.  By November each of these birds will have molted into a tan-buff basic (non-breeding) plumage, the male’s with a slight yellow hue.  During their first spring, juveniles attain sexual maturity and, like the adults from the previous year, molt into alternate (breeding) plumage.  The breeding birds seen here will probably winter in the southern United States, and birds that nested to our north will arrive to remain as winter residents.  Various stages of molt can be seen simultaneously during spring and autumn migrations as populations of goldfinches from multiple latitudes intermingle as they pass through the Susquehanna River watershed.
This juvenile Spotted Sandpiper (Actitis macularius), lacks the namesake dark markings on its white underside, thus it presently resembles an adult in basic (non-breeding) plumage.  Upon reaching sexual maturity, it will molt into a spotted alternate (breeding) plumage for the nesting season during each remaining year of its life.  Females of this species defend a territory and lay eggs in a nest located among cover near the shoreline.  A female may have as many as five mates.  Males alone incubate the eggs, usually four, for 20-24 days.  The young leave the nest upon hatching and are escorted by the male for about two to three additional weeks.  They are the only sandpiper to nest on the Susquehanna River shoreline.  Spotted Sandpipers migrate to the southern border states and further south for winter.  At Conewago Falls, they arrive in late April to nest, with birds, possibly migrants from further north, remaining until well into October.  In any plumage, you can easily recognize the Spotted Sandpiper by its habit of teetering its body at the hips to pump the tail up and down.
Bald Eagles go through a series of five molts before reaching adult plumage.  The first plumage, Juvenile, is nearly all dark brown with white linings along the forward underside of the wings.  The wing and tail feathers are a bit longer than in later plumages to aid the inexperienced birds during their clumsy first flights.  Due to the additional feather length, Juveniles look larger than older birds, but they are not as heavy as their seniors.  The bird seen here flying above Conewago Falls is probably in its second year.  This plumage, Basic I, also known as “White Belly I”, is characterized by a nearly full set of new flight feathers.  Note that some of the longer Juvenile feathers are still present, giving the wings a jagged sloppy look, particularly near the center of the trailing edge.  Third-year (Basic II) birds often have at least some white belly feathers and are sometimes known as “White Belly II” Bald Eagles.  Basic II birds typically possess a complete set of adult flight feathers, so the trailing edge of the wing has a neater and more uniform appearance.

The Bald Eagle in the two photographs above is in its first year.  This plumage, known as “Juvenile”, is characterized by dark flight feathers which appear uniformly long in length when the bird is airborne.  The eye is dark brown.  The iris of the eye will lighten in the second year (Basic I) and will become cream-colored by the third year (Basic II).  The bill, which is all dark gray when the bird is in the nest, has begun the slow progression to a yellow color that will be complete in the bird’s fifth year (Basic IV).  Third year (Basic II) birds molt to white crown and throat feathers, but have a dark set of feathers through the eye producing an “Osprey face” in most individuals.  In its fourth year (Basic III), this eagle will molt to a white tail with just a thin dark brown terminal band.  The head will become nearly all white except for a few dark spots through the eye, which will have a yellow iris.  A cleaner white head and tail will develop during the fifth year (Basic IV) and will persist through the familiar adult Bald Eagle plumages (Definitive Plumage) for the remainder of its life.

The Juvenile and non-breeding (basic) plumages of late-summer may seem drab and confusing, but learning them is a worthwhile endeavor.  Consider that most of the birds coming south during the migration will be adorned in this fashion.  The birds of North America are in their greatest numerical mass of the year right now, and nearly all are females, juveniles, other non-adults, or molting males.  There are few males in breeding plumage among the autumn waves of migrants.  In the coming months, there will be an abundance of opportunities to enjoy these marvels on wings, so getting to know the birds in non-breeding feathers is time well spent.  Make haste and get ready.  For our feathered friends, it’s autumn and they’re on their way south.

Here come the confusing fall migrants.  Twelve Semipalmated Sandpipers (Calidris pusilla) and sixteen unidentified “peep” sandpipers (Calidris species) were seen feeding in Conewago Falls on August 27.  This Semipalmated Sandpiper is either an adult in worn alternate (breeding) plumage or a Juvenile.  Adults of this species molt into basic plumage on the wintering grounds.  The Semipalmated Sandpiper breeds in the high arctic tundra and winters in the West Indies and northern South America.
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SOURCES

Dunne, Pete.  2006.  Pete Dunne’s Essential Field Guide Companion.  Houghton Mifflin Company.  New York.

Hayman, Peter; John Marchant, and Tony Prater.  1986.  Shorebirds, An Identification Guide to the Waders of the World.  Houghton Mifflin Company.  New York.

Kauman, Kenn.  1996.  Lives of North American Birds.  Houghton Mifflin Company.  New York.

McCullough, Mark A.  1989.  Molting Sequence and Aging Of Bald Eagles.  The Wilson Bulletin.  101:1-10.