At last, the bird migration is picking up where it left off after the third week of September when rain, fog, and gloom arrived for a two week stay in the lower Susquehanna valley. Now that this persistent meteorological interluder has departed the stage, our stalled avians can resume the autumn spectacle.
Southbound Flights: On Their Way Both Night and Day
It’s hard to believe, but for almost two months now, sandpipers, plovers, and terns have been filtering south through the Lower Susquehanna River Watershed on their way to the Atlantic coastline as they complete the first leg of their long autumn migration—a journey that will take some species all the way to the far reaches of the South American continent for winter.
As August draws to a close, these early birds are being joined by widespread nocturnal flights of Neotropical migrants—those species, primarily songbirds, on their way to wintering grounds which lie exclusively south of the continental United States.
To catch a glimpse of these night-flying avians, your best bet may be to position yourself on the crest of a ridge or along a linear break in the forest such as a utility right-of-way where waves of warblers, vireos, flycatchers, and other Neotropical passerines sometimes feed on invertebrates after making landfall at daybreak. Pick a place where the trees are bathed in the warm light of the rising sun and be there by 7 A.M. E.D.T. The activity can be tremendous, but it usually ends between 8 and 9.
Diurnal migrants, birds that make their movements during the daylight hours, are ramping up their flights now as well. Broad-winged Hawks, Bald Eagles, and falcons are currently being tallied at hawk-counting stations throughout the northeast. Many of those lookouts are seeing Ruby-throated Hummingbirds, swallows, and other daytime migrants too.
Autumn migration flights are an ever-changing process, with different species peaking at different times throughout the season. In these months just after the nesting season, each of these species is more numerous than at any other time of the year. And of course, the more often we as observers get out and have a look, the more of them we’ll see.
Be certain to click the “Birds” tab at the top of this page for a photo guide to the species you’re likely to see passing through the lower Susquehanna valley this fall. Nearly four months of autumn hawk migration flights lie ahead, so don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab to find a hawk-counting station near you, then stop by for a visit or two. See you there!
This Week at Regional Hawk Watches
With nearly all of the Neotropical migrants including Broad-winged Hawks gone for the year, observers and counters at eastern hawk watches are busy tallying numbers of the more hardy species of diurnal raptors and other birds. The majority of species now coming through will spend the winter months in temperate and sub-tropical areas of the southern United States and Mexico.
Here is a quick look at the raptors seen this week at two regional counting stations: Kiptopeke Hawk Watch near Cape Charles, Virginia, and Second Mountain Hawk Watch at Fort Indiantown Gap, Pennsylvania.
During coming days, fewer and fewer of these birds will be counted at our local hawk watches. Soon, the larger raptors—Red-tailed Hawks, Red-shouldered Hawks, and Golden Eagles—will be thrilling observers. Cooler weather will bring several flights of these spectacular species. Why not plan a visit to a lookout near you? Click on the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab at the top of this page for site information and a photo guide to identification. See you at the hawk watch!
Shorebirds on the Mud in York County, Pennsylvania
At Lake Redman just to the south of York, Pennsylvania, a draw down to provide drinking water to the city while maintenance is being performed on the dam at neighboring Lake Williams, York’s primary water source, has fortuitously coincided with autumn shorebird migration. Here’s a sample of the numerous sandpipers and plovers seen today on the mudflats that have been exposed at the southeast end of the lake…
Not photographed but present at Lake Redman were at least two additional species of shorebirds, Killdeer and Spotted Sandpiper—bringing the day’s tally to ten. Not bad for an inland location! It’s clearly evident that these waders overfly the lower Susquehanna valley in great numbers during migration and are in urgent need of undisturbed habitat for making stopovers to feed and rest so that they might improve their chances of surviving the long journey ahead of them. Mud is indeed a much needed refuge.
Shorebirds and More at Bombay Hook National Wildlife Refuge
Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.
Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.
As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.
No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.
We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.
Photo of the Day
Photo of the Day
Migration Update
Can it be that time already? Most Neotropical birds have passed through the Lower Susquehanna River Watershed on their way south and the hardier species that will spend our winter in the more temperate climes of the eastern United States are beginning to arrive.
Here’s a gallery of sightings from recent days…
Be sure to click on these tabs at the top of this page to find image guides to help you identify the dragonflies, birds, and raptors you see in the Lower Susquehanna River Watershed…
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- Damselflies and Dragonflies
- Birds of Conewago Falls
- Hawkwatcher’s Helper: Identifying Bald Eagles and other Raptors
See you next time!
Smoky Skies in the Lower Susquehanna Region
During the coming two weeks, peak numbers of migrating Neotropical birds will be passing through the northeastern United States including the lower Susquehanna valley. Hawk watches are staffed and observers are awaiting big flights of Broad-winged Hawks—hoping to see a thousand birds or more in a single day.
Broad-winged hawks feed on rodents, amphibians, and a variety of large insects while on their breeding grounds in the forests of the northern United States and Canada. They depart early, journeying to wintering areas in Central and South America before frost robs them of a reliable food supply.
While migrating, Broad-winged Hawks climb to great altitudes on thermal updrafts and are notoriously difficult to see from ground level. Bright sunny skies with no clouds to serve as a backdrop further complicate a hawk counter’s ability to spot passing birds. Throughout the Lower Susquehanna River Watershed, the coming week promises to be especially challenging for those trying to observe and census the passage of high-flying Broad-winged Hawks. The forecast of hot and humid weather is not so unusual, but the addition of smoke from fires in the western states promises to intensify the haze and create an especially irritating glare for those searching the skies for raptors.
It may seem gloomy for the mid-September flights in 2021, but hawk watchers are hardy types. They know that the birds won’t wait. So if you want to see migrating “Broad-wings” and other species, you’ve got to get out there and look up while they’re passing through.
These hawk watches in the Lower Susquehanna River Watershed are currently staffed by official counters and all welcome visitors:
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- Rocky Ridge County Park Hawk Watch—3699 Deininger Road off Mount Zion Road (Route 24) northeast of York, Pennsylvania.
- Second Mountain Hawk Watch—off Cold Spring Road on the grounds of Fort Indiantown Gap in Lebanon County, Pennsylvania.
- Waggoner’s Gap Hawk Watch—where Route 74 crosses Blue Mountain north of Carlisle, Pennsylvania.
—or you can just keep an eye on the sky from wherever you happen to be. And don’t forget to check the trees and shrubs because warbler numbers are peaking too! During recent days…
Shorebirds at Middle Creek
Late August and early September is prime time to see migrating shorebirds as they pass through the lower Susquehanna valley during their autumn migration, which, believe it or not, can begin as early as late June. These species that are often assumed to spend their lives only near the seashore are regular visitors each fall as they make their way from breeding grounds in the interior of Canada to wintering sites in seacoast wetlands—many traveling as far south as Central and South America.
Low water levels on the Susquehanna River often coincide with the shorebird migration each year, exposing gravel and sand bars as well as vast expanses of muddy shorelines as feeding and resting areas for these traveling birds. This week though, rain from the remnants of Tropical Storm Fred arrived to increase the flow in the Susquehanna and inundate most of the natural habitat for shorebirds. Those on the move must either continue through the area without stopping or find alternate locations to loaf and find food.
The draining and filling of wetlands along the river and elsewhere in the region has left few naturally-occurring options. The Conejohela Flats south of Columbia offer refuge to many migrating sandpipers and their allies, the river level there being controlled by releases from the Safe Harbor Dam during all but the severest of floods. Shorebirds will sometimes visit flooded fields, but wide-open puddles and farmland resembling mudflats is more of springtime occurrence—preceding the planting and growth of crops. Well-designed stormwater holding facilities can function as habitat for sandpipers and other wildlife. They are worth checking on a regular basis—you never know what might drop in.
Right now, there is a new shorebird hot spot in the Lower Susquehanna River Watershed—Middle Creek Wildlife Management Area. The water level in the main impoundment there has been drawn down during recent weeks to expose mudflats along the periphery of nearly the entire lake. Viewing from “Stop 1” (the roadside section of the lake in front of the refuge museum) is best. The variety of species and their numbers can change throughout the day as birds filter in and out—at times traveling to other mudflats around the lake where they are hidden from view. The birds at “Stop 1” are backlit in the morning with favorable illumination developing in the afternoon.
Have a look at a few of the shorebirds currently being seen at Middle Creek…
The aquatic environs at Middle Creek attract other species as well. Here are some of the most photogenic…
A Visit to Rocky Ridge
Early October is prime time for hawk watching, particularly if you want to have the chance to see the maximum variety of migratory species. In coming days, a few Broad-winged Hawks and Ospreys will still be trickling through while numbers of Sharp-shinned Hawks, Cooper’s Hawks, Northern Harriers, and falcons swell to reach their seasonal peak. Numbers of migrating Red-tailed and Red-shouldered Hawks are increasing during this time and late-season specialties including Golden Eagles can certainly make a surprise early visit.
If you enjoy the outdoors and live in the southernmost portion of the lower Susquehanna valley, Rocky Ridge County Park in the Hellam Hills just northwest of York, Pennsylvania, is a must see. The park consists of oak forest and is owned and managed by the York County Parks Department. It features an official hawk watch site staffed by volunteers and park naturalists. Have a look.
If you’re a nature photographer, you might be interested to know that there are still hundreds of active butterflies in Rocky Ridge’s utility right-of-way. Here are a few.
To see the daily totals for the raptor count at Rocky Ridge Hawk Watch and other hawk watches in North America, and to learn more about each site, be certain to visit hawkcount.org
Bird Migration Highlights
The southbound bird migration of 2020 is well underway. With passage of a cold front coming within the next 48 hours, the days ahead should provide an abundance of viewing opportunities.
Here are some of the species moving through the lower Susquehanna valley right now.
Spotted Lanternfly in the Lower Susquehanna River Watershed
Second Mountain Hawk Watch is located on a ridge top along the northern edge of the Fort Indiantown Gap Military Reservation and the southern edge of State Game Lands 211 in Lebanon County, Pennsylvania. The valley on the north side of the ridge, also known as St. Anthony’s Wilderness, is drained to the Susquehanna by Stony Creek. The valley to the south is drained toward the river by Indiantown Run, a tributary of Swatara Creek.
The hawk watch is able to operate at this prime location for observing the autumn migration of birds, butterflies, dragonflies, and bats through the courtesy of the Pennsylvania Game Commission and the Garrison Commander at Fort Indiantown Gap. The Second Mountain Hawk Watch Association is a non-profit organization that staffs the count site daily throughout the season and reports data to the North American Hawk Watch Association (posted daily at hawkcount.org).
Today, Second Mountain Hawk Watch was populated by observers who enjoyed today’s break in the rainy weather with a visit to the lookout to see what birds might be on the move. All were anxiously awaiting a big flight of Broad-winged Hawks, a forest-dwelling Neotropical species that often travels back to its wintering grounds in groups exceeding one hundred birds. Each autumn, many inland hawk watches in the northeast experience at least one day in mid-September with a Broad-winged Hawk count exceeding 1,000 birds. They are an early-season migrant and today’s southeast winds ahead of the remnants of Hurricane Florence (currently in the Carolinas) could push southwest-heading “Broad-wings” out of the Piedmont Province and into the Ridge and Valley Province for a pass by the Second Mountain lookout.
The flight turned out to be steady through the day with over three hundred Broad-winged Hawks sighted. The largest group consisted of several dozen birds. We would hope there are probably many more yet to come after the Florence rains pass through the northeast and out to sea by mid-week. Also seen today were Bald Eagles, Ospreys, American Kestrels, and a migrating Red-headed Woodpecker.
Migrating insects included Monarch butterflies, and the three commonest species of migratory dragonflies: Wandering Glider, Black Saddlebags, and Common Green Darner. The Common Green Darners swarmed the lookout by the dozens late in the afternoon and attracted a couple of American Kestrels, which had apparently set down from a day of migration. American Kestrels and Broad-winged Hawks feed upon dragonflies and often migrate in tandem with them for at least a portion of their journey.
Still later, as the last of the Broad-winged Hawks descended from great heights and began passing by just above the trees looking for a place to settle down, a most unwelcome visitor arrived at the lookout. It glided in from the St. Anthony’s Wilderness side of the ridge on showy crimson-red wings, then became nearly indiscernible from gray tree bark when it landed on a limb. It was the dreaded and potentially invasive Spotted Lanternfly (Lycorma delicatula). This large leafhopper is native to Asia and was first discovered in North America in the Oley Valley of eastern Berks County, Pennsylvania in 2014. The larval stage is exceptionally damaging to cultivated grape and orchard crops. It poses a threat to forest trees as well. Despite efforts to contain the species through quarantine and other methods, it’s obviously spreading quickly. Here on the Second Mountain lookout, we know that wind has a huge influence on the movement of birds and insects. The east and southeast winds we’ve experienced for nearly a week may be carrying Spotted Lanternflies well out of their most recent range and into the forests of the Ridge and Valley Province. We do know for certain that the Spotted Lanternfly has found its way into the Lower Susquehanna River Watershed.
Winter Won’t Let Go…the Birds Don’t Care
It seems as though the birds have grown impatient for typical spring weather to arrive. The increase in hours of daylight has signaled them that breeding time is here. No further delays can be entertained. They’ve got a schedule to keep.
Thursday, March 29: Winds began blowing from the southwest, breaking a cold spell which had persisted since last week’s snowfall. Birds were on the move ahead of an approaching rainy cold front.
Friday, March 30: Temperatures reached 60 degrees at last. Birds were again moving north through the day, despite rain showers and a change in wind direction—from the northwest and cooler following the passage of the front in the late morning.
Saturday, March 31: It was cooler, but birds were still on the wing headed north.
Sunday, April 1: The morning was pleasant, but conditions became cooler and breezy in the afternoon. Migratory and resident birds began feeding ahead of another storm.
Monday, April 2: Snow fell again, overnight and through the morning—a couple of inches. Most of the snow had melted away by late afternoon.
A Quick Getaway
It was a placid morning on Conewago Falls with blue skies dotted every now and then by a small flock of migrating robins or blackbirds. The jumbled notes of a singing Winter Wren (Troglodytes hiemalis) in the Riparian Woodland softly mixed with the sounds of water spilling over the dam. The season’s first Wood Ducks (Aix sponsa), Blue-winged Teal (Spatula discors), Herring Gull (Larus argentatus), Horned Larks (Eremophila alpestris), and White-throated Sparrows (Zonotrichia albicollis) were seen.
There was a small ruckus when one of the adult Bald Eagles from a local pair spotted an Osprey passing through carrying a fish. This eagle’s effort to steal the Osprey’s catch was soon interrupted when an adult eagle from a second pair that has been lingering in the area joined the pursuit. Two eagles are certainly better than one when it’s time to hustle a skinny little Osprey, don’t you think?
But you see, this just won’t do. It’s a breach of eagle etiquette, don’t you know? Soon both pairs of adult eagles were engaged in a noisy dogfight. It was fussing and cackling and the four eagles going in every direction overhead. Things calmed down after about five minutes, then a staring match commenced on the crest of the dam with the two pairs of eagles, the “home team” and the “visiting team”, perched about 100 feet from each other. Soon the pair which seems to be visiting gave up and moved out of the falls for the remainder of the day. The Osprey, in the meantime, was able to slip away.
In recent weeks, the “home team” pair of Bald Eagles, seen regularly defending territory at Conewago Falls, has been hanging sticks and branched tree limbs on the cross members of the power line tower where they often perch. They seem only to collect and display these would-be nest materials when the “visiting team” pair is perched in the nearby tower just several hundred yards away…an attempt to intimidate by homesteading. It appears that with winter and breeding time approaching, territorial behavior is on the increase.
In the afternoon, a fresh breeze from the south sent ripples across the waters among the Pothole Rocks. The updraft on the south face of the diabase ridge on the east shore was like a highway for some migrating hawks, falcons, and vultures. Black Vultures (Coragyps atratus) and Turkey Vultures streamed off to the south headlong into the wind after leaving the ridge and crossing the river. A male and female Northern Harrier (Circus hudsonius), ten Red-tailed Hawks, two Red-shouldered Hawks (Buteo lineatus), six Sharp-shinned Hawks, and two Merlins crossed the river and continued along the diabase ridge on the west shore, accessing a strong updraft along its slope to propel their journey further to the southwest. Four high-flying Bald Eagles migrated through, each following the east river shore downstream and making little use of the ridge except to gain a little altitude while passing by.
Late in the afternoon, the local Bald Eagles were again airborne and cackling up a storm. This time they intercepted an eagle coming down the ridge toward the river and immediately forced the bird to climb if it intended to pass. It turned out to be the best sighting of the day, and these “home team” eagles found it first. It was a Golden Eagle (Aquila chrysaetos) in crisp juvenile plumage. On its first southward voyage, it seemed to linger after climbing high enough for the Bald Eagles to loose concern, then finally selected the ridge route and crossed the river to head off to the southwest.
Swallows by the Thousands
A fresh breeze from the north brought cooler air and a reminder that summer is gone and autumn has arrived.
Fast-moving dark clouds provided a perfect backdrop for viewing passing diurnal migrants. Bald Eagles utilized the tail wind to cruise down the Susquehanna toward Chesapeake Bay and points further south. A migrating Merlin began a chase from which a Northern Flicker narrowly escaped by finding shelter among Pothole Rocks and a few small trees. The season’s first American Black Duck (Anas rubripes), Common Loon (Gavia immer), Yellow-bellied Sapsucker (Sphyrapicus varia), and American Pipits (Anthus rubescens) moved through.
Blue Jays continued their hesitant crossings of the river at Conewago Falls. The majority completed the journey by forming groups of a dozen or more birds and following the lead of a lone American Robin, a Northern Flicker, or, odd as it appeared, a small warbler.
By far the most numerous migrants today were swallows. Thousands of Northern Rough-winged Swallows and hundreds of Tree Swallows were on the wing in search of what was suddenly a sparse flying insect supply. To get out of the brisk wind, some of the more resourceful birds landed on the warm rocks. To satisfy their appetite, many were able to pick crawling arthropods from the surface of the boulders. They swallow them whole.
Blue Jay Way
The Neotropical birds that raised their young in Canada and in the northern United States have now logged many miles on their journey to warmer climates for the coming winter. As their density decreases among the masses of migrating birds, a shift to species with a tolerance for the cooler winter weather of the temperate regions will be evident.
Though it is unusually warm for this late in September, the movement of diurnal migrants continues. This morning at Conewago Falls, five Broad-winged Hawks (Buteo platypterus) lifted from the forested hills to the east, then crossed the river to continue a excursion to the southwest which will eventually lead them and thousands of others that passed through Pennsylvania this week to wintering habitat in South America. Broad-winged Hawks often gather in large migrating groups which swarm in the rising air of thermal updrafts, then, after gaining substantial altitude, glide away to continue their trip. These ever-growing assemblages from all over eastern North America funnel into coastal Texas where they make a turn to south around the Gulf of Mexico, then continue on toward the tropics. In the coming weeks, a migration count at Corpus Christi in Texas could tally 100,000 or more Broad-winged Hawks in a single day as a large portion of the continental population passes by. You can track their movement and that of other diurnal raptors as recorded at sites located all over North America by visiting hawkcount.org on the internet. Check it out. You’ll be glad you did.
Nearly all of the other migrants seen today have a much shorter flight ahead of them. Red-bellied Woodpeckers (Melanerpes carolinus), Red-headed Woodpeckers (Melanerpes erythrocephalus), and Northern Flickers (Colaptes auratus) were on the move. Migrating American Robins (Turdus migratorius) crossed the river early in the day, possibly leftovers from an overnight flight of this primarily nocturnal migrant. The season’s first Great Black-backed Gulls (Larus marinus) arrived. American Goldfinches are easily detected by their calls as they pass overhead. Look carefully at the goldfinches visiting your feeder, the birds of summer are probably gone and are being replaced by migrants currently passing through.
By far, the most conspicuous migrant today was the Blue Jay. Hundreds were seen as they filtered out of the hardwood forests of the diabase ridge to cautiously cross the river and continue to the southwest. Groups of five to fifty birds would noisily congregate in trees along the river’s edge, then begin flying across the falls. Many wary jays abandoned their small crossing parties and turned back. Soon, they would try the trip again in a larger flock.
A look at this morning’s count reveals few Neotropical migrants. With the exception of the Broad-winged Hawks and warblers, the migratory species seen today will winter in a sub-tropical temperate climate, primarily in the southern United States, but often as far north as the lower Susquehanna River valley. The individual birds observed today will mostly continue to a winter home a bit further south. Those that will winter in the area of Conewago Falls will arrive in October and later.
The long-distance migrating insect so beloved among butterfly enthusiasts shows signs of improving numbers. Today, more than two dozen Monarchs were seen crossing the falls and slowly flapping and gliding their way to Mexico.
Yellowlegs for Breakfast
A few nocturnal migrants flew through the moonlit night to arrive at Conewago Falls for a sunrise showing this morning. A dozen warblers were in the treetops and a Wood Thrush (Hylocichla mustelina) chattered away in the understory of the Riparian Woodlands. Three species of shorebirds were in the falls and on the Pothole Rocks: Least Sandpiper (Calidris minutilla), Lesser Yellowlegs (Tringa flavipes), and Greater Yellowlegs (Tringa melanoleuca).
The diurnal migration was highlighted by a Merlin (Falco columbarius), an Osprey, and a Bald Eagle, each flying down the river. Most of the other birds in the falls seemed content to linger and feed. There’s no need to hurry folks, only trouble lurks down there in paradise at the moment.
Summer Grasses
It has not been a good summer if you happen to be a submerged plant species in the lower Susquehanna River. Regularly occurring showers and thunderstorms have produced torrents of rain and higher than usual river stages. The high water alone wouldn’t prevent you from growing, colonizing a wider area, and floating several small flowers on the surface, however, the turbidity, the suspended sediment, would. The muddy current casts a dirty shadow on the benthic zone preventing bottom-rooted plants from getting much headway. There will be smaller floating mats of the uppermost leaves of these species. Fish and invertebrates which rely upon this habitat for food and shelter will find sparse accommodation…better luck next year.
Due to the dirty water, fish-eating birds are having a challenging season as they try to catch sufficient quantities of prey to feed themselves and their offspring. A family of Ospreys (Pandion haliaetus) at Conewago Falls, including recently fledged young, were observed throughout this morning and had no successful catches. Of the hundred or more individual piscivores of various species present, none were seen retrieving fish from the river. The visibility in the water column needs to improve before fishing is a viable enterprise again.
While the submerged plant communities may be stunted by 2017’s extraordinary water levels, there is a very unique habitat in Conewago Falls which endures summer flooding and, in addition, requires the scouring effects of river ice to maintain its mosaic of unique plants. It is known as a Riverine Grassland or scour grassland.
The predominant plants of the Riverine Grasslands are perennial warm-season grasses. The deep root systems of these hardy species have evolved to survive events which prevent the grassland from reverting to woodland through succession. Fire, intense grazing by wild herd animals, poor soils, drought, and other hardships, including flooding and ice scour, will eliminate intolerant plant species and prevent an area from reforesting. In winter and early spring, scraping and grinding by flood-driven chunk ice mechanically removes large woody and poorly rooted herbaceous growth from susceptible portions of the falls. These adverse conditions clear the way for populations of species more often associated with North America’s tall grass prairies to take root. Let’s have a look at some of the common species found in the “Conewago Falls Pothole Rocks Prairie”.
The Conewago Falls Riverine Grassland is home to numerous other very interesting plants. We’ll look at more of them next time.
SOURCES
Brown, Lauren. 1979. Grasses, An Identification Guide. Houghton Mifflin Company. New York, NY.