Scenes from a Managed Grassland Ecosystem

Pretty pictures…

Veery
Photographing Neotropical migrants as they arrive to nest in our local forests can be frustrating under closed canopies and diffused light conditions.  We instead decided to focus our efforts and lenses on the grassland species at Middle Creek Wildlife Management Area where, earlier this week, several prescribed fire burns were wrapping up the spring treatments for 2026.
Warm-season grassland
Managed warm-season grassland habitat at Middle Creek Wildlife Management Area.
Tree Swallow
Already tending nests containing eggs and young, Tree Swallows are scattered around the refuge’s grasslands to welcome newly arriving migrants, and human visitors!
Grasshopper Sparrow
Middle Creek is probably the best place in the lower Susquehanna valley to see ground-nesting Grasshopper Sparrows.
Grasshopper Sparrow
To attract a mate and establish a defendable territory, the males are singing their buzzy, insect-like songs from atop roadside fence posts.
Grasshopper Sparrow
The songs persist as the male moves from perch to perch around a potential breeding site.
Grasshopper Sparrow and Tree Swallow
It’s a behavior that occasionally puts the newcomer Grasshopper Sparrow in conflict with a well-ensconced Tree Swallow guarding its nest box.
Eastern Bluebird
Eastern Bluebirds squeeze in among the Tree Swallows to find a vacant nest box within which they can raise their young.
Song Sparrow
Song Sparrows may be present year-round in grasslands with interspersed growth of early successional shrubs and briars, however the populations at a given location changes as birds migrate in and out of the area seasonally.  Those individuals observed nesting here in the spring and summer are not usually the same birds seen during the late fall and winter.
Savannah Sparrow
As a species, the Savannah Sparrow, like the Song Sparrow, may be represented in the grasslands throughout the year, though the individuals vary with the changing seasons. Unlike the Song Sparrow which nests in shrubs, the Savannah Sparrow nests on the ground among thick grasses.
American Goldfinch
Still munching on the seeds of last year’s wildflowers, American Goldfinches are pairing up around the grasslands waiting for thistle down and other fibers to be available for nest construction.  They will then build a small cup within the upper limbs of a small to medium-sized tree to shelter their eggs and nestlings.
Common Yellowthroat
The Common Yellowthroat, one of our Neotropical warblers, will reproduce in grasslands with scattered early successional thorny shrubs to afford a safe place to construct a nest. The males sing almost incessantly from the time they arrive in late April and early May until sometime in July, or even later.
Eastern Kingbird
Eastern Kingbirds are arriving from tropical wintering grounds to nest in grasslands and pastures throughout the lower Susquehanna region.  They are a tyrant flycatcher that will ambush insects from a perch atop a tree, shrub, or tuft of tall grass.
Northern Mockingbird
Northern Mockingbirds are considered a resident species, though there may be some southward movement during severe winters when foods such as berries become scarce. At Middle Creek, some of these evacuees have returned to nest along the interface zone between the grassland and early successional shrub habitats.
Horned Lark
Nesting Horned Larks apparently take advantage of the interface between the managed grasslands and nearly bare soils in spring-planted croplands at Middle Creek. They may be one of the few, possibly the only, species of bird life to take advantage of no-till farming for nesting on the ground.
Red-winged Blackbird
Red-winged Blackbirds not only construct nests in cattails and other marsh growth, but in dense grasses and shrubs in and near grasslands and pastures. The males are easily seen displaying their plumage as they sing from a small tree, shrub, or a cattail seed spike.
Female Red-winged Blackbird
Less frequently seen is the female Red-winged Blackbird.  She spends the majority of her time in the nest, but will periodically come out to join the ruckus when an intruder is being scolded into leaving the premises.
Eastern Meadowlark
Another member of the blackbird family is the Eastern Meadowlark.  It too is an energetic singer during the breeding season.  Meadowlarks build their nests on the ground, often in pastures of cool-season grasses where harvests prior to August are fatal to eggs and young.  Middle Creek’s managers delay harvests to allow the birds enough time to complete their reproduction cycle.
Bobolink
Another blackbird is the Bobolink, an obligate grassland specialist and a Neotropical migrant.  One of its few uses for trees and shrubs is as a place to burst into display and song, though it will also perform these rituals on the wing and among the grass.
Bobolink
A male Bobolink singing and displaying from a perch.
Bobolinks
As the males arrive, competition for suitable nesting sites among the grasses becomes intense, even before a female is anywhere in sight.
Bobolinks
We saw as many as five males at a time clustered into this one tuft of thorny twigs to take turns chattering and showing their wares to one another.
Bobolink
A territorial male Bobolink at Middle Creek Wildlife Management Area.  Delayed mowing allows cool-season grasslands to provide nesting sites for this species too.
Baltimore Oriole
Did you know the Baltimore Oriole is a blackbird?  Well, it is.
Baltimore Oriole
When surrounded by acres of grasslands for collecting insects, groves of mature shade trees including oaks, walnuts, sycamores, and elms provide excellent nesting sites for Baltimore Orioles.
Baltimore Orioles
Competition for these ideal sites can get quite animated.
Baltimore Oriole
Fights like this one between two second-year males can become vicious,…
Baltimore Orioles
…leaving combatants rolling on the ground in their fury,…
Baltimore Orioles
…at least until the more-experienced adult males who’ve been fighting over the place have had enough…
Baltimore Orioles
…and decide to send the clumsy, younger males packing.
Western Cattle Egrets
During our visit to the Middle Creek grasslands earlier this week, we were afforded an encounter with some unusual grassland birds, Western Cattle Egrets.  To the delight of birders who never got to enjoy the abundance of this species on the lower Susquehanna during the 1970s and early 1980s, these birds have been a popular attraction as they’ve lingered around the refuge for a couple of weeks now.
Western Cattle Egrets and Geese
It’s an unusual experience, watching Western Cattle Egrets feeding in a field of cultivated cool-season grasses alongside Snow Geese, though they certainly aren’t looking for the same thing.

We’ll have more on the Western Cattle Egrets and other interesting migrants at Middle Creek in an upcoming post.  Check back soon!

Managing a Grassland Ecosystem

If you’ve visited Middle Creek Wildlife Management Area anytime during the past month, you may have noticed quite a bit of activity around the large pole-mounted nest boxes placed out in the open fields.

American Kestrels at nest box
American Kestrels, a male and female, at a nest box at Middle Creek Wildlife Management Area.
Kestrels competing for nesting sites and territory.
Within the past week, we noticed that kestrels are still competing for nesting sites and territory, driving away unwanted trespassers.  A pair of kestrels seemed to be occupying each of the four box sites we observed.  That’s good news, but the presence of these man-made nest cavities is in no way wholly responsible for this positive response from these declining birds.  It’s a matter of habitat, much needed grassland habitat.
Cool Season Grasses
Cool-season grasses including fescue (Lolium species) and non-native Orchard Grass (Dactylis glomerata) mature by late spring.  If left standing, they continue to provide indispensable habitat for grassland wildlife through the summer.  Mowing early and often for hay harvest has rendered most cool-season meadows death traps for nesting birds.  By delaying cuttings until at least early August, ground-nesting birds are given adequate time to fledge their young and get the juvenile birds strong enough to fly away from a set of spinning blades.
Grasshopper Sparrow in Timothy
A male Grasshopper Sparrow sings to demarcate its nesting territory in a stand of Timothy, an introduced species cultivated for harvest as hay.  An insectivore during the breeding season, he and his mate will attempt to raise their brood exclusively within the cover of these cool-season grasses.
Eastern Meadowlark
Eastern Meadowlarks arrive in cool-season grasslands by late-winter to begin their breeding cycle which typically extends into the hot summer days of July.
Orchard Grass with Black-eyed Susans
Prior to being mowed, Orchard Grass provides the short, dense cover meadowlarks and other ground-nesting grassland birds need to successfully reproduce.  Growing a cool season grassland can be as easy as delaying the mowing of a pasture, field, or oversize lawn until August, then have a farmer friend come and take off a cutting or two of hay or straw to prevent woody plants from becoming established.
Warm-season Grasses
More durable stands of native warm-season grasses including Big Bluestem, Indiangrass, Switchgrass, and Little Bluestem thrive in the summer heat and provide wildlife habitat throughout the year.  These perennial “prairie grasses”, fed by root systems five to eight feet deep, are especially drought tolerant .  From these entrenched anchors, the plants would quickly bounce back after the once great herds of hungry bison had grazed the landscape bare of surface vegetation before moving on.  This adaptation also assured “prairie grass” regeneration following naturally occurring seasonal fires, events mimicked in eastern North America by its pre-Columbian human residents.  They used recurrent fire to perpetuate early successional habitat for wildlife propagation, foraging, and agriculture.  Today in the lower Susquehanna watershed, establishing warm-season grassland meadows requires soil prep and seeding to get things going again.
Precribed Burn
To prevent succession, warm-season grassland parcels are most commonly maintained using applications of prescribed fire every 3 to 5 years.  Among their benefits, these burns invigorate native vegetation while inhibiting the invasive tendencies of many non-native plants.  Well-planned periodic fire can significantly reduce fuel accumulations, particularly in tinderbox woodland tracts managed as fire-free zones for the past century or more.  Many forest trees including oaks rely on sporadic fire events for regeneration.

LEARN HOW LAND MANAGERS UTILIZE PRESCRIBED FIRE

This coming Saturday, April 18, 2026, beginning at 10 A.M. (rain date April 25), the Pennsylvania Game Commission is hosting a “Prescribed Fire Festival” at Middle Creek Wildlife Management Area.  Be certain to come out to the visitor’s center at 100 Museum Road, Stevens, PA, for this event.  Land managers will be there to answer questions and to explain the planning and preparations involved in overseeing a prescribed burn.  There will be guided walks of habitats preserved using fire of varying intensities.  You’ll see the equipment and protective clothing used by certified personnel to administer a live prescribed fire burn right before your eyes.  Then you can have lunch—food trucks will be available on site.

Prescribed Fire Demonstration
Visitors witness a prescribed fire demonstration at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area in 2025.

After the burn demonstration, why not go for a walk or drive around the refuge.  You can take note of how grassland and early successional plant communities are responding to previous doses of prescribed fire…

Wingstem
Greening up a week or two after the burn, Wingstem is blooming by June in areas treated using prescribed fire earlier in the spring.
Joe-Pye-Weed
Another scene from June: Four-foot-tall Joe-Pye-Weed amidst lush growth of goldenrod in early successional habitat where a prescribed burn eliminated accumulating wildfire fuel and turned back the growth of invasive plant species in March.

And how the grassland animals respond as well…

Ring-necked Pheasants
Ring-necked Pheasants in early successional habitat maintained by the periodic application of prescribed fire.
Eastern Cottontail
Eastern Cottontails prosper in the mosaic of warm-season grasses and early successional thickets on lands sustained using prescribed fire.  Rabbits are herbivores, primary consumers eating mostly legumes and other plants, the producers that through the process of photosynthesis convert the energy of the the sun into food energy.
Northern Short-tailed Shrew
Other small vegetation-eating rodents including mice and voles thrive in managed grasslands.  Similar in appearance are the shrews.  Though seldom noticed, Northern Short-tailed Shrews spend day and night foraging for food, even in the shallow waters of wet meadows and thickets.  Unlike the aforementioned herbivorous rodents, shrews are insectivores, secondary consumers feeding mostly on primary consumers including a variety of insects and other arthropods.
Masked Shrew
Some years ago, we found this tiny Masked Shrew (Sorex cinereus) in a grassland area being preserved using prescribed fire.  Like other shrews, Masked Shrews are secretive, but always on the go.  They feed constantly to fuel their vast energy requirements, sometimes consuming three times their body weight in a single day.  But their voracious appetite can get them into trouble, causing these incessant eaters to encounter numerous potentially infective parasites during their non-stop foraging missions.
Female American Kestrel
Patrolling the grasslands is the female American Kestrel, a secondary consumer.  She’s on the lookout for primary consumers including large insects like grasshoppers or crickets.  But perhaps more likely is a small rodent whose less-than-ideal vigor may cause it to slip up, creating an easy target.  If she selects a careless shrew as her prey, she may be assuming the role of a tertiary consumer, eating a secondary consumer (the shrew) that fed on insects (the primary consumers) that derived their energy from photosynthesized plant matter.
American Barn Owl
The strictly nocturnal American Barn Owl (Tyto furcata), another secondary and sometimes tertiary consumer, takes the night shift, hunting unwary voles, mice, and shrews, often by sensing the sounds they make among runways in the grass.  As a native predator in its favored habitat, the owl’s selection of each victim actually helps to keep the prey species’ population healthy, eliminating the weak and vulnerable to provide a qualitative service to the surviving wildlife of the grasslands.  While we may not think of the barn owl as a direct consumer of insects, its positive influence on insectivorous shrew populations makes it an important functionary in maintaining balance in the ecosystems it calls home.
Male American Kestrel
As food becomes increasing plentiful in the grasslands, a female kestrel will remain mostly out of sight, performing the majority of the egg incubation duties while her colorful mate stands guard nearby.
Male American Kestrel
The male not only keeps watch, but also continues the hunt for insects, small mammals, and other prey to feed not only the newborn nestlings, but also his mate while she tends the nest.  As the young grow and no longer need brooding to stay warm, the female will join the male in a joint effort to snatch up enough food to keep their three to seven offspring nourished.  Like other raptors, populations of these predators are abruptly regulated in the nest.  The first hatched of these falcon’s young will receive the most food, giving them, particularly the oldest individual, the best chances of survival.  The later-hatched and thus smaller offspring may have trouble competing for the available provisions brought to the nest.  If food is plentiful, there may be enough for all of the birds to grow and survive.  If food is scarce, only the oldest (which also happen to be the biggest, strongest, and most aggressive) baby falcon(s) will live to fledge and leave the nest.  If hunting becomes really poor, the adults will sustain themselves at the expense of their young.

The fate of an avian predator such as a kestrel lies at the mercy of the fate of its quarry.  Because, you see, the sun’s energy, after being converted to chemical energy by photosynthesizing plants, flows upward through the trophic levels of the food chain—herbivores (primary consumers) such as rodents and insects to carnivores (secondary and tertiary consumers) including kestrels.  Grasslands, when abundant and diverse, are correspondingly abundant and diverse with small mammals and insects and will therefore support thriving populations of American Kestrels and other predators.  These secondary and sometimes tertiary consumers fulfill a role in cultivating healthier populations of their prey—the primary and secondary consumers in the food web—as a balanced component of a flourishing grassland ecosystem.  Sparse and fragmented grasslands, on the other hand, beget negligible small mammal and insect populations, are stricken with broken food webs, and champion few if any American Kestrels or other predators.  If the land it occupies is neat, tidy, manicured, exploited, or sprayed sterile and dead, the energy flow cycle of the ecosystem is dead as well.  There’s nothing animal introductions, reintroductions, rescues, culling, stocking, or harvesting can do about it, because in the end, it’s all about the habitat.

Photo of the Day

Tree Swallow
With southerly breezes to push them along, insectivorous songbirds are beginning to arrive throughout the lower Susquehanna valley.  By the first week of March, Tree Swallows started their push up the river corridor, relying upon the late-winter hatches of stoneflies and other cold-hardy flying insects for sustenance.  As temperatures warmed during mid-month, greater numbers of these colorful cavity nesters ventured out across upland areas in search of suitable breeding sites in grasslands and pastures near water.  Upon finding a good home in a dead tree or nest box, they’ll defend it vigorously to discourage encroachment.

Back in the Day: Down on the Farm

Let us travel through time for just a little while to recall those sunny, late-spring days down on the farm—back when the rural landscape was a quiet, semi-secluded realm with little in the way of traffic, housing projects, or industrialized agriculture.  Those among us who grew up on one of these family homesteads, or had friends who did, remember the joy of exploring the meadows, thickets, soggy springs, and woodlots they protected.

Low-intensity Farming
During much of the twentieth century, low-intensity agriculture provided a haven for wildlife.  Periodic disturbances helped maintain cool-season grassland and early successional habitat for a number of species we currently find in decline.

For many of us, farmland was the first place we encountered and began to understand wildlife.  Vast acreage provided an abundance of space to explore.  And the discovery of each new creature provided an exciting experience.

Distributed by the Pennsylvania Game Commission, artist Ned Smith’s wildlife posters introduced many residents of the lower Susquehanna region to its birds and mammals.  This poster of “Birds of Field and Garden” helped us learn what to expect and search for during our forays to the farm.

Today, high-intensity agriculture, relentless mowing, urban sprawl, and the increasing costs and demand for land have all conspired to seriously deplete habitat quality and quantity for many of the species we used to see on the local farm.  Unfortunately for them, farm wildlife has largely been the victim of modern economics.

For old time’s sake, we recently passed a nostalgic afternoon at Middle Creek Wildlife Management Area examining what maintenance of traditional farm habitat has done and can do for breeding birds.  Join us for a quick tour to remember how it used to be at the farm next door…

Barn Swallow
Always found nesting under the forebay of the barn, the Barn Swallow relentlessly pursued flying insects over the pond and meadow.
Eastern Meadowlark
Eastern Meadowlarks arrived during March and April to begin nesting in their namesake.  Their song, “spring-of-the-year”, heralded the new season.
Eastern Kingbird
Arriving in meadows and pastures during early May, the Eastern Kingbird provided for its nestlings by ambushing a variety of flying insects.  By August, congregations of these birds could be found gathering along ponds and streams ahead of their fall migration.
Orchard Oriole
In the cherry grove down by the creek, the Orchard Oriole would be singing incessantly to defend its territory.
American Goldfinches
Normally seed eaters through the colder months, American Goldfinches would regularly find a source of protein in the occupants of Eastern Tent Caterpillar nests.
Yellow Warbler
Along the wet margins of the creek, Yellow Warblers would nest in the shrubs and small trees.
Willow Flycatcher
The “Traill’s Flycatcher” was a familiar find in low-lying areas of successional shrubs and small trees.  Today, “Traill’s Flycatcher” is recognized as two distinct species, the Alder Flycatcher (Empidonax alnorum) and the Willow Flycatcher.  In the lower Susquehanna valley, the latter (seen here) is by far the most common of the two.
Eastern Bluebirds
During the nineteenth century, Eastern Bluebirds became a rarity on lower Susquehanna farms due to a combination of factors: pesticide (DDT) use, habitat loss, and competition with other birds for nest sites.  The species saw a resurgence beginning in the 1970s with discontinuation of DDT applications and widespread provision of nesting boxes.  Around human habitations, competition with invasive House Sparrows continues to be detrimental to their success.
Purple Martins and Tree Swallows
Purple Martins suffered a similar fate to the bluebirds.  The potential for their recovery remains dubious and they continue to be very local breeders, fussy about selection of suitable man-made provisions for nesting.  After considerable effort, Purple Martins have at last been attracted to nest in the condos placed for their use at Middle Creek Wildlife Management Area.  In the artificial gourds, there are nesting Tree Swallows, a species which also benefits from the placement of boxes intended for bluebirds.
Field Sparrow
Abandoned fields and other successional habitats were and continue to be favored homes for Field Sparrows.
Cedar Waxwing
At almost any time of year, roving bands of Cedar Waxwings would suddenly visit old field habitat looking for berries among the shrubs and other pioneering woody growth.  In early summer, after most species have already hatched their young, nesting would commence and these fruit eaters would transform into accomplished fly catchers.
Ring-necked Pheasant
During the twentieth century prior to the 1980s, Ring-necked Pheasant populations in the Lower Susquehanna River Watershed were composed of breeding descendants of introduced birds supplemented by additional releases to maintain numbers sufficient for hunting.  Year-round populations did and can reside in mosaic landscapes of early successional and grassland habitats, the latter including hay fields left unmowed through the nesting season.
Red-winged Blackbird
Red-winged Blackbirds have always been a fixture of hay fields and meadows on farms.  While the increase in mowing frequency has reduced their nesting success, they have persevered as a species by nesting earlier than other birds and by utilizing other landscape features such as densely vegetated stormwater basins for breeding sites.
Bobolink
Do you recall the last time you saw a Bobolink nesting in a hay field near you?  Arriving in early May as a Neotropical migrant, the Bobolink requires a cool-season grassland such as hay field through at least July to complete its nesting cycle.  Even earlier this century, we remember nesting Bobolinks being more widespread on farms throughout the region.  Now, you almost have to go to Middle Creek if you want to see them.
Grasshopper Sparrow
Formerly more widespread in hay fields throughout the lower Susquehanna valley, the native Grasshopper Sparrow is yet another species falling victim to early mowing and intensive farming.
Grasshopper Sparrow
The solution to their dilemma is as advertised.  Instead of cutting the grass, why not take heed of the example set here and cut back on the tens of thousands of acres that are excessively or needlessly mowed in the Lower Susquehanna River Watershed?  How ’bout letting a significant percentage of your property regenerate as successional habitat as well?  It can and does make a difference!
Cool-season Grasses
Beautiful cool-season grasses waving in the spring breeze.  Meadows and hay fields can be managed to function as cool-season grasslands to provide nesting opportunities for many of the species we used to find down on the farm.

Photo of the Day

Ring-necked Pheasant
A Ring-necked Pheasant forages among the fresh green growth two weeks to the day after prescribed fire was administered to this parcel at Middle Creek Wildlife Management Area to turn back plant succession and maintain grassland habitat.

Wildlife in the Burn Zone

During Saturday’s Prescribed Fire Demonstration at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area, we noticed just how fast some species of wildlife return to areas subjected to burns administered to maintain grassland habitat and reduce the risk of high-intensity blazes.

Back Fire Ignition during prescribed burn demonstration.
Pennsylvania Game Commission crews ignite a back fire to contain a prescribed burn along its downwind/upslope perimeter during a demonstration at Middle Creek Wildlife Management Area on Saturday.
Prescribed Fire Demonstration
Visitors observe a fire planned to maintain this section of the refuge as warm-season grassland.  A species with roots several feet deep, the light-colored vegetation is Indiangrass, a plant adapted to thrive following periodic episodes of wildfire.  Prescribed fire can be used to replace naturally occurring infernos with much safer controlled burns that eliminate successional and invasive plants to promote the establishment of Indiangrass and other native warm-season species including Big Bluestem, Little Bluestem, Switchgrass, and a variety of wildflowers as well.
Prescribed Fire Demonstration
Even as the fire reached its brief peak of intensity, we noticed birds already attracted to the site…
Tree Swallow
Dozens of recently arrived Tree Swallows swept in to patrol for flying insects as the burn was in progress.
Tree Swallow
One even stopped by to have a look inside the kestrel nest box as fire approached the dry stand of goldenrod on the slope behind.
Red-tailed Hawk
Red-tailed Hawks and other raptors, including nocturnal owls, are frequently the first visitors attracted to the scene of a prescribed burn or wildfire.  In grassland and successional habitats, they come looking for any vulnerable voles or mice that may be moving about looking for cover.

Eastern Meadowlarks

Eastern Meadowlarks

Eastern Meadowlarks
These three Eastern Meadowlarks spent the morning in the grassland areas adjacent to the prescribed fire site, mostly where a burn had been conducted one week prior.  During the demonstration, one even perched and sang from the oak trees in the museum/visitor’s center parking lot.

Following the Prescribed Fire Demonstration, we decided to pay a visit to some of the parcels where burns had been administered one week earlier on the north side of Middle Creek’s main impoundment.  We found a surprising amount of activity.

White-crowned and White-throated Sparrows
Apparently feeding upon slightly heat-treated seeds, sparrows were found by the dozens. White-crowned (left), White-throated (right), Song, and Savannah Sparrows were identified.
Downy Woodpecker
This Downy Woodpecker was finding something to its liking among the scorched leaves, stems, and twigs.
American Robins seem to find areas with lightly burnt vegetation and ash-dusted soil advantageous for finding invertebrates following a fire.
Mixed Flock of Blackbirds
We found this flock of Red-winged Blackbirds, Browned-headed Cowbirds, and a few European Starlings feeding throughout a grassland field cleared of early-successional growth by a prescribed fire administered one week ago.
Mixed Flock of Blackbirds
They seemed to favor gleaning seeds from among the lightly burned areas of the plot.
Eastern Meadowlark
Nearby, in an island of unburned grass in the same field, we found yet another Eastern Meadowlark, our fourth of the day.  High-intensity agriculture, particularly early hay mowing and pesticide treatments, have mostly eliminated this and other grassland species from modern farms.  Management practices like prescribed fire and delayed mowing (no spinning blades until at least early August) can maintain ideal grassland habitat for stunningly colorful blackbirds including nesting Eastern Meadowlarks, Bobolinks, and many other species as well.
American Kestrel at Nest Box
A male American Kestrel at a nest box located among Middle Creek’s warm-season and cool-season grassland habitats, the former maintained by prescribed fire, the latter by delayed mowing.

A Prescribed Fire Site After the Burn

Back on March 24th, we took a detailed look at the process involved in administering prescribed fire as a tool for managing grassland and early successional habitat.  Today we’re going turn back the hands of time to give you a glimpse of how the treated site fared during the five months since the controlled burn.  Let’s go back to Middle Creek Wildlife Management Area for a photo tour to see how things have come along…

Prescribed Fire application in March
Pennsylvania Game Commission crews administering prescribed fire on a grassland demonstration site back on March 16.
Prescribed Fire Site in May
By late May, native herbaceous perennial wildflowers including Joe-pye Weed had re-greened the site.  One of the goals of the burn was to kill fire-sensitive woody plants, thus preventing the process of succession from reforesting the site.
Prescribed Fire Site in May
The scorched, lifeless remains of small trees and shrubs indicate that that goal was met.
Yellow-breasted Chat in May
Because prescribed fire is administered in a mosaic pattern that permits some early successional growth to remain until the next burn, birds including this thicket-nesting Yellow-breasted Chat are able to take advantage of the mixed habitat during their breeding season in May and June.
Prescribed Fire Site in August
By August, the site is a haven for native plants and animals.
Native Warm-season Grasses and Wildlflowers in August
The burn has promoted the growth and late-summer bloom of fire-tolerant native wildflowers and warm-season grasses…
Indiangrass in Flower
…including Indiangrass,…
Big Bluestem
…Big Bluestem…
Thin-leaved Coneflower, a plant also known as Brown-eyed Susan.
…Thin-leaved Coneflower (Rudbeckia triloba), a plant also known as Brown-eyed Susan,…
Joe-pye Weed
…and Joe-pye Weed, a plant butterflies find irresistible.
Eastern Tiger Swallowtail collecting nectar from Joe-pye Weed.
Eastern Tiger Swallowtail collecting nectar from Joe-pye Weed.
Black-morph Eastern Tiger Swallowtail
A black-morph Eastern Tiger Swallowtail collecting nectar from Spotted Knapweed (Centaurea steobe micranthos), a non-native invasive plant found growing in an area of the burn site missed by this year’s fire.  While many non-native plants are unable to survive the flames and heat produced by prescribed fire, it isn’t an absolute cure-all.  It doesn’t eliminate all invasive plants, it just keeps them from dominating a landscape by out-competing native species.  Left unmanaged, Spotted Knapweed is a tough perennial invasive that can easily become one the species able to overtake a vulnerable grassland.  It can be a stubborn survivor of some prescribed burns.  On the plus side, butterflies really like it.
American Goldfinch
By August, native grassland plants in the prescribed fire area were already providing an abundance of seeds for birds including this American Goldfinch.
Carolina Grasshopper
For larger birds like turkeys and pheasants, an abundance of Carolina Grasshoppers are providing a protein-dense food source in managed grasslands.
Black Saddlebags
And tiny flying insects, a nuisance to us as we take a stroll alongside the grasslands, are a meal taken on the wing by dragonflies including this Black Saddlebags.

Elsewhere around the refuge at Middle Creek, prescribed fire and other management techniques are providing high-quality grassland habitat for numerous species of nesting birds…

Bobolink
Bobolinks nested both in areas subjected to controlled burns…
Bobolink in Hay Field
…and in hay fields where mowing was delayed until the nesting season, including the fledging process, was completed earlier this month.
Grasshopper Sparrow
As advertised, Grasshopper Sparrows nested in these fields as well.

We hope you enjoyed this short photo tour of grassland management practices.  Now, we’d like to leave you with one last set of pictures—a set you may find as interesting as we found them.  Each is of a different Eastern Cottontail, a species we found to be particularly common on prescribed fire sites when we took these images in late May.  The first two are of the individuals we happened to be able to photograph in areas subjected to fire two months earlier in March.  The latter two are of cottontails we happened to photograph elsewhere on the refuge in areas not in proximity to ground treated with a prescribed burn or exposed to accidental fire in recent years.

Eastern Cottontail
Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.
Eastern Cottontail
Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.

These first two rabbits are living the good life in a warm-season grass wonderland.

Eastern Cottontail with ticks.
Eastern Cottontail at a site not subjected to any recent fire activity.
Eastern Cottontail with ticks.
Eastern Cottontail at a site not subjected to any recent fire activity.

Oh Deer!  Oh Deer!  These last two rabbits have no clock to track the time; they have only ticks.  Better not go for a stroll with them Alice—that’s no wonderland!  I know, I know, it’s time to go.  See ya later.

Birds of the Sunny Grasslands

With the earth at perihelion (its closest approach to the sun) and with our home star just 27 degrees above the horizon at midday, bright low-angle light offered the perfect opportunity for doing some wildlife photography today.  We visited a couple of grasslands managed by the Pennsylvania Game Commission to see what we could find…

Grasslands and Hedgerows
On this State Game Lands parcel, prescribed fire is used to maintain a mix of grasslands and brushy early successional growth.  In nearby areas, both controlled fire and mechanical cutting are used to remove invasive species from hedgerows and the understory of woodlots.  Fire tolerant native species then have an opportunity to recolonize the forest and improve wildlife habitat.  This management method also reduces the fuel load in areas with the potential for uncontrolled wildfires.
The sun-dried fruits of a Common Persimmon tree found growing in a hedgerow.
The sun-dried fruits of a native Common Persimmon tree found growing in a hedgerow.
Savanna-like Grasslands
Just one year ago, mechanical removal of invasive trees and shrubs (including Multiflora Rose) on this State Game Land was followed by a prescribed fire to create this savanna-like grassland.
Song Sparrow
Hundreds of Song Sparrows were found in the grasses and thickets at both locations.
White-throated Sparrow
White-throated Sparrows were also abundant, but prefer the tangles and shrubs of the thickets.
Northern Mockingbird
Northern Mockingbirds were vigilantly guarding winter supplies of berries in the woodlots and hedgerows.
Swamp Sparrow
In grasses and tangles on wetter ground, about a dozen Swamp Sparrows were discovered.
White-crowned Sparrow
The adult White-crowned Sparrow is always a welcome find.
White-crowned Sparrow
And seeing plenty of juvenile White-crowned Sparrows provides some assurance that there will be a steady stream of handsome adult birds arriving to spend the winter during the years to come.
Dark-eyed Junco
Dark-eyed Juncos were encountered only in the vicinity of trees and large shrubs.
Savannah Sparrow
Several Savannah Sparrows were observed.  Though they’re mostly found in treeless country, this particular one happened to pose atop a clump of shrubs located within, you guessed it, the new savanna-like grasslands.
Winter Wren
A tiny bird, even when compared to a sparrow, the Winter Wren often provides the observer with just a brief glimpse before darting away into the cover of a thicket.
Standing Clump of Timber
Within grasslands, scattered stands of live and dead timber can provide valuable habitat for many species of animals.
A "snag" with an excavated nest cavity.
Woodpeckers and other cavity-nesting birds rely upon an abundance of “snags” (standing dead trees) for breeding sites.
Red-bellied Woodpecker
This Red-bellied Woodpecker and about a dozen others were found in trees left standing in the project areas.
Yellow-bellied Sapsucker
A Yellow-bellied Sapsucker soaks up some sun.
Pileated Woodpecker
This very cooperative Pileated Woodpecker seemed to be preoccupied by insect activity on the sun-drenched bark of the trees.  This denizen of mature forests will oft times wander into open country where larger lumber is left intact.

Pileated Woodpecker

Northern Harrier
Just as things were really getting fun, some late afternoon clouds arrived to dim the already fading daylight.  Just then, this Northern Harrier made a couple of low passes in search of mice and voles hidden in the grasses.
Northern Harrier
It was a fitting end to a very short, but marvelously sunny, early winter day.

Butterflies and More at Boyd Big Tree Preserve Conservation Area

If you’re feeling the need to see summertime butterflies and their numbers just don’t seem to be what they used to be in your garden, then plan an afternoon visit to the Boyd Big Tree Preserve along Fishing Creek Valley Road (PA 443) just east of U.S. 22/322 and the Susquehanna River north of Harrisburg.  The Pennsylvania Department of Conservation and Natural Resources manages the park’s 1,025 acres mostly as forested land with more than ten miles of trails.  While located predominately on the north slope of Blue Mountain, a portion of the preserve straddles the crest of the ridge to include the upper reaches of the southern exposure.

American Chestnut at Boyd Big Tree Preserve
A grove of American Chestnuts (Castanea dentata) planted at Boyd Big Tree Preserve is part of a propagation program working to restore blight-resistant trees to Pennsylvania and other areas of their former range which included the Appalachians and the upper Ohio River watershed.

Fortunately, one need not take a strenuous hike up Blue Mountain to observe butterflies.  Open space along the park’s quarter-mile-long entrance road is maintained as a rolling meadow of wildflowers and cool-season grasses that provide nectar for adult butterflies and host plants for their larvae.

Butterfly Meadow at Boyd Big Tree Preserve
A view looking north at the butterfly meadow and entrance road at Boyd Big Tree Preserve Conservation Area.  Second Mountain is in the background.
Walking a Meadow Path
Mowed paths follow the entrance road and a portion of the perimeter of the meadow allowing visitors a chance to wander among the waist-high growth to see butterflies, birds, and blooming plants at close range without trampling the vegetation or risking exposure to ticks.
A Silver-spotted Skipper feeding on nectar from Dogbane (Apocynum cannabinum) flowers.
A Silver-spotted Skipper feeding on nectar from the flowers of Indian Hemp (Apocynum cannabinum).  Like the milkweeds, Indian Hemp is a member of the dogbane family (Apocynaceae).
An Eastern Tiger Swallowtail feeding on Common Milkweed.
An Eastern Tiger Swallowtail feeding on Common Milkweed.
Great Spangled Fritillary on Common Milkweed.
A Great Spangled Fritillary (Speyeria cybele) on Common Milkweed.
A Black Swallowtail feeding on Common Milkweed nectar.
A Black Swallowtail (Papilio polyxenes) feeding on Common Milkweed nectar.
A Pipevine Swallowtail on Common Milkweed.
A Pipevine Swallowtail (Battus philenor) on Common Milkweed.
A Pipevine Swallowtail on Common Milkweed.
Another Pipevine Swallowtail on Common Milkweed.  Note the hook-shaped row of red-orange spots on the underside of the hindwing.
A Pipevine Swallowtail on visiting Butterfly Weed.
A Pipevine Swallowtail visiting the brilliant blooms of Butterfly Weed, a favorite of a wide variety of pollinators.
A Black Swallowtail on Butterfly Weed
A Black Swallowtail with damaged wings alights atop a Butterfly Weed flower cluster.  Note the pair of parallel rows of red-orange spot on the underside of the hindwing.
A Monarch on Butterfly Weed
A Monarch feeding on nectar from the flowers of Butterfly Weed.
A mating pair of Eastern Tailed Blues.
A mating pair of Eastern Tailed Blues on a Timothy (Phleum pratense) spike.
A female (left) and male Great Spangled Fritillary.
A male Great Spangled Fritillary (right) pursuing a female.
Common Green Darner
Butterflies aren’t the only colorful insects patrolling the meadows at Boyd Big Tree Preserve.  Dragonflies including Common Green Darners are busily pursuing prey, particularly small flying insects like mosquitos, gnats, and flies.
Juvenile Broad-winged Hawk
Dragonflies themselves can become prey and are much sought after by Broad-winged Hawks. This very vocal juvenile gave us several good looks as it ventured from the forest into the skies above the upper meadow during midday.  It wasn’t yet a good enough flier to snag a dragonfly, but it will have plenty of opportunities for practice during its upcoming fall migration which, for these Neotropical raptors, will get underway later this month.

Do yourself a favor and take a trip to the Boyd Big Tree Preserve Conservation Area.  Who knows?  It might actually inspire you to convert that lawn or other mowed space into much-needed butterfly/pollinator habitat.

While you’re out, you can identify your sightings using our photographic guide—Butterflies of the Lower Susquehanna River Watershed—by clicking the “Butterflies” tab at the top this page.  And while you’re at it, you can brush up on your hawk identification skills ahead of the upcoming migration by clicking the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab.  Therein you’ll find a listing and descriptions of hawk watch locations in and around the lower Susquehanna region.  Plan to visit one or more this autumn!

Some Good Reasons to Postpone Mowing Until Mid-August

Here in a series of photographs are just a handful of the reasons why the land stewards at Middle Creek Wildlife Management Area and other properties where conservation and propagation practices are employed delay the mowing of fields composed of cool-season grasses until after August 15 each year.

Eastern Meadowlark
Eastern Meadowlarks, birds of large pastures, hay lots and other meadows of cool-season grasses, build their nests and raise their young on the ground.  In the years since the early twentieth century, loss in the volume of acreage maintained in the lower Susquehanna Valley as grassland habitat types has dramatically reduced the prevalence and abundance of this and other birds with similar nesting requirements.  During the most recent fifty years, early and frequent mowing and other practices introduced as part of agriculture’s Green Revolution have all but eliminated ground-nesting grassland species from the region.
Grasshopper Sparrow
Like the meadowlarks, Grasshopper Sparrows (Ammodramus savannarum) nest on the ground in fields of cool-season grasses.  Mowing prior to the time the young leave the nest and are able to fly away can obliterate a generation of grassland birds.  Because their life span is short, widespread loss of an entire year of reproduction can quickly impact overall populations of native sparrows and other small birds.  Delayed mowing can improve numbers of Grasshopper Sparrows as well as Savannah Sparrows, Vesper Sparrows (Pooecetes gramineus), and the very rare Henslow’s Sparrow (Centronyx henslowii).
Bobolink
The Bobolink, like the meadowlark, is a member of the blackbird family (Icteridae).  It too requires grasslands free of disturbances like mowing for the duration of the nesting season which, for this particular bird, lasts until mid-August in the lower Susquehanna region.  In places lacking their specific habitat requirements, Bobolinks will seldom be detected except as flyovers during migration.
Ring-necked Pheasant
Ring-necked Pheasants were introduced to the lower Susquehanna basin, and their populations were maintained thereafter, by stocking for the purpose of hunting.  But throughout the middle twentieth century, there was a substantial population of ring-necks breeding in fields of cool-season grasses in farmlands throughout the region.  High-intensity agriculture with frequent mowing eliminated not only nesting habitat in grasslands, but winter cover in areas of early successional growth.  Populations of Ring-necked Pheasants, as well as native Northern Bobwhite, crumbled during the late 1970s and early 1980s due to these changes.  For these resident birds that don’t migrate or routinely travel great distances to find new places to live and breed, widespread habitat loss can be particularly catastrophic.  Not surprisingly, the Northern Bobwhite is no longer found in the Lower Susquehanna River Watershed and has been extirpated from all of Pennsylvania.
Blue Grosbeak
At places like Middle Creek Wildlife Management Area where a mix of grasslands, early successional growth, and even some cropland are maintained, the Blue Grosbeak has extended its range well north of the Mason-Dixon and has become a regular nesting species during recent decades.  Good habitat management does pay dividends.

Right now is a good time to visit Middle Creek Wildlife Management Area to see the effectiveness a delayed mowing schedule can have when applied to fields of cool-season grasses.  If you slowly drive, walk, or bicycle the auto tour route on the north side of the lake, you’ll pass through vast areas maintained as cool-season and warm-season grasses and early successional growth—and you’ll have a chance to see these and other grassland birds raising their young.  It’s like a trip back in time to see farmlands they way they were during the middle years of the twentieth century.