This Week’s Enormous Diurnal Flights

For the past several weeks, we’ve given you a look at the fallout of Neotropical songbirds on mornings following significant nocturnal flights of southbound migrants.  Now let’s examine the diurnal (daytime) flights that have developed in recent days over parts of the lower Susquehanna basin and adjacent regions.

For those observing diurnal migrants, particularly at raptor-counting stations, the third week of September is prime time for large flights of tropics-bound Broad-winged Hawks.  Winds from easterly directions during this year’s movements kept the greatest concentrations of these birds in the Ridge and Valley Province and upper areas of the Piedmont as they transited our region along a southwesterly heading.  Counts topped 1,000 birds or more on at least one day at each of these lookouts during the past seven days.  Meanwhile at hawk watches along the Piedmont/Atlantic Coastal Plain border, where flights topped 10,000 or more birds on the best days last fall, observers struggled to see 100 Broad-winged Hawks in a single day.

Broad-winged Hawks
Dozens of migrating Broad-winged Hawks during a morning liftoff to the north of Blue Mountain.

Among the challenges counters faced while enumerating migratory Broad-winged Hawks this week was the often clear blue skies, a glaring sun, and the high altitude at which these raptors fly during the mid-day hours.

"Kettling" Broad-winged Hawks
Large numbers of Broad-winged Hawks climbing on a thermal updraft to gain altitude can become increasingly difficult to spot in a clear blue sky.
"Kettling" Broad-winged Hawks
Higher and higher they go until they practically disappear from view.
Broad-winged Hawks
Cloud cover can help make locating really high-flying birds a bit easier, but they can sometimes climb into the lower portion of the cloud and vanish.

Another factor complicating the hawk counters’ tasks this week, particularly west of the Susquehanna River, was the widespread presence of another group of diurnal fliers.

Blue Jay
Was it the migratory Blue Jays that are beginning to move through the area?   No.
Sharp-shinned Hawk
Was it the increasing numbers of Sharp-shinned Hawks and other raptors?  No.
Hawkwatchers Scanning the Skies
So what was it that complicated the hawkwatchers’ efforts to find high-flying Broad-winged Hawks during this week’s flights?
Spotted Lanternfly
Is it a bird?  Is it a plane?  No.  And it’s nobody’s superhero either.  It’s the non-native Spotted Lanternfly and it is now very numerous west of the Susquehanna and common enough to the east to make even experienced observers take a second look while scanning the skies for Broad-winged Hawks and other migrating raptors.  Just how many do raptor counters have to contend with?  Even where these invasive insects have become less populous in the lower Susquehanna valley, hundreds still fill the skies above forest clearings around hawk-counting stations including Waggoner’s Gap, Second Mountain, and Rocky Ridge County Park.
Spotted Lanternflies on Radar
Spotted Lanternfly populations explode with great vigor as they colonize areas with plenty of tender new growth on native, non-native, and cultivated shrubs, trees, and vines.  Earlier this week, National Weather Service Doppler Radar in Pittsburgh, Pennsylvania, captured enormous mid-afternoon flights of Spotted Lanternflies in recently invaded areas of southwestern Pennsylvania and eastern Ohio.  (NOAA/National Weather Service Doppler Radar image)
Spotted Lanternflies on Radar
A base velocity image consisting primarily of Spotted Lanternflies in southwestern Pennsylvania and eastern Ohio.  (NOAA/National Weather Service Doppler Radar image)

As the Broad-winged Hawk migration draws to a close during the coming ten days, the occurrence of the one dozen other migratory diurnal raptors will be on the upswing.  They generally fly at lower altitudes, often relying upon wind updrafts on the ridges for lift instead of high-rising thermal updrafts.  They therefore present better observation opportunities for visitors at hawk watch lookouts and are less frequently confused with high-flying Spotted Lanternflies.

Merlin
A Merlin seen earlier this week migrating through the Lower Susquehanna River Watershed.  The numbers of migrating Merlins and other falcons will peak sometime in coming weeks.

Pot o’ Gold on the Ridgetops

Bathed in glowing sunshine, a very large fallout of migrating Neotropical songbirds enlivened the forest edge atop Second Mountain in northern Lebanon County, Pennsylvania, this morning.  While last night’s flight was widespread beneath a dome of atmospheric high pressure covering the Mid-Atlantic States, this is a look at some of the 500 -1,000 migrants observed feeding on insects and other natural foods at just this single location.

Blackpoll and Black-throated Green Warblers
As the composition of the nightly flights begins to transition to species that start their migration later in the season, numbers of birds like the Blackpoll Warbler (top) are beginning to rival those of the heretofore very common Black-throated Green Warbler (bottom).
Blackpoll Warbler
A Blackpoll Warbler.
Bay-breasted Warbler
The Bay-breasted Warbler, another common species found among the fallout flock.
Tennessee Warbler
The Tennessee Warbler was today’s most abundant migrant with at least 200 birds seen moving among the other species.
Tennessee Warbler
A newly arrived Tennessee Warbler at sunrise.
Tennessee Warbler
A Tennessee Warbler snatches breakfast.
Magnolia Warbler
The Magnolia Warbler was another common find among today’s migrants.
Magnolia Warbler
A Magnolia Warbler.
Nashville Warbler
A Nashville Warbler among tangles of Mile-a-minute Weed.
Nashville Warbler
A Nashville Warbler.
Black-throated Blue Warbler
Black-throated Blue Warbler.
Adult Male Black-throated Blue Warbler
An adult male Black-throated Blue Warbler.
Northern Parula
An acrobatic Northern Parula.
Blackburnian Warbler
Blackburnian Warblers continue their southerly trek.
American Redstart
An American Redstart.
 Chestnut-sided Warbler
A Chestnut-sided Warbler feeding on Mile-a-minute Weed berries.
 Cape May Warbler
A Cape May Warbler.
Adult Male Cape May Warbler
A handsome adult male Cape May Warbler.
Swainson's Thrush
A Swainson’s Thrush.
Scarlet Tanager
Scarlet Tanagers continue to work their way south through the Lower Susquehanna River Watershed.
Olive-sided Flycatcher
A composite image of an uncommon find, a large-headed Olive-sided Flycatcher seen perched atop a dead tree snag.
Philadelphia Vireo
A brief encounter with a Philadelphia Vireo, another uncommon species.
Blue-headed Vireo
A Blue-headed Vireo, typically the last of its genus to move south in the fall, put in a welcome appearance among the hundreds of migrants.
Ruby-throated Hummingbird
A favorite diurnal migrant, the Ruby-throated Hummingbird, was seen feeding on flowers that grew from the remains of garden waste dumped illegally along the mountain road last autumn.

Many of the observers at the Second Mountain Hawk Watch commented that this morning’s fallout was by far the best they had seen anywhere in the region during recent years.  Others believed it to be the best they had ever seen.  It was indeed less like a “wave” of migrants and more like a “tsunami”.  Choosing a good viewing location and being there at the right time can improve your chances of seeing a spectacle like this.  The good news is, it looks like another big flight is currently underway, so finding a forest edge on a ridgetop or along a utility right-of-way just might pay off for you early tomorrow morning.

Nocturnal migrating birds, a southwest-bound flight down the ridges of central Pennsylvania as indicated by Doppler radar between 10:30 and 11:30 PM EDT on September 11, 2024.  (NOAA/National Weather Service image)

This Week’s Review of the Morning Fallouts

Clear, cool nights have provided ideal flight conditions for nocturnal Neotropical migrants and other southbound birds throughout the week.  Fix yourself a drink and a little snack, then sit down and enjoy this set of photographs that includes just some of the species we found during sunrise feeding frenzies atop several of the Lower Susquehanna River Watershed’s ridges.  Hurry up, because here they come…

Black-throated Green Warbler
A Black-throated Green Warbler.
Black-throated Green Warbler
The Black-throated Green Warbler was perhaps the most frequently identified treetop warbler during the most recent four mornings.
Black-throated Green Warbler
A Black-throated Green Warbler with a unique variation in the crown plumage.
Blackburnian Warbler
The Blackburnian Warbler was another plentiful species.
Cape May Warblers
Cape May Warblers have an affinity for conifers like this Eastern White Pine.
Cape May Warbler
But when traveling in mixed flocks with other migrants, Cape May Warblers can also be found feeding in the crown foliage of deciduous trees.
Tennessee Warbler
This adult Tennessee Warbler appears to be adorned in a very worn set of plumage…
Tennessee Warbler
…and its traveling companion looks like it’s overdue for a new set of feathers as well.
Nashville Warbler
Like the Tennessee Warbler, the Nashville Warbler was common among mixed flocks.
Nashville Warbler
A Nashville Warbler atop a Black Cherry.
Chestnut-sided Warbler
This Chestnut-sided Warbler was one of several found among the more common species of migrants.
Chestnut-sided Warbler
A Chestnut-sided Warbler.
Chestnut-sided Warbler
We were lucky enough to spot this male Chestnut-sided Warbler sporting his namesake flank feathers.
 Black-and-white Warbler
A Black-and-white Warbler uses its nuthatch-like feeding behavior to search the tree bark for edible invertebrates.
Common Yellowthroat
To see the Common Yellowthroat, one must cease looking upward into the high canopy and instead give the aching neck a rest by peering into the low vegetation at the forest edge.
Magnolia Warbler
While checking the low growth, keep an eye open for other migrants among the shrubs and tangles. This Magnolia Warbler glows in the rays of a rising sun as it searches for a meal after a long night of travel.
House Wren
Here we found a perky little House Wren.
Eastern Wood-Pewee
Back in the middle and upper reaches of the trees, we find what has been by far the most numerous of the flycatchers seen during our visits to fallout sunrises.  Eastern Wood-Pewees are appearing in very good numbers and can be seen quarreling and battling for hunting perches from which they are ambushing flying insects.
 Eastern Wood-Pewee
An Eastern Wood-Pewee fiercely defending its hunting perch.
Eastern Wood-Pewee
An Eastern Wood-Pewee.
Least Flycatcher
The numbers of migrating Least Flycatchers and other members of the genus Empidonax may be reaching their seasonal peak this week.
Scarlet Tanagers
Scarlet Tanagers are currently a common find following nocturnal flights.
Scarlet Tanager
A Scarlet Tanager peers down from the top of a Red Maple.
Rose-breasted Grosbeak
Did you hear a loud squeak in the treetops?  It could be a southbound Rose-breasted Grosbeak stopping by for the day.
Rose-breasted Grosbeak
A Rose-breasted Grosbeak in a dead tree snag.
Swainson's Thrush
The Neotropical thrushes are beginning to move south now as well.  We found this newly arrived Swainson’s Thrush at Rocky Ridge County Park in York County during sunrise this morning.
Red-eyed Vireo
Not surprisingly, the Red-eyed Vireo is one of the most numerous of the migrants seen feeding in the deciduous canopy following a nocturnal flight event.  It’s not at all unusual to see dozens filing the trees around a ridgetop overlook or along a forest edge.  Be certain to check these congregations carefully, especially the groups of birds feeding in the lower branches of tall timber or in the tops of smaller trees.  This week we found…
Yellow-throated Vireo
…several hungry Yellow-throated Vireos arriving after nocturnal flights,…
Philadelphia Vireo
…and a Philadelphia Vireo (Vireo philadelphicus) at the hawkwatch at Rocky Ridge County Park in York County.
Red-breasted Nuthatch
Though not a Neotropical migrant, the easier-heard-than-seen Red-breasted Nuthatch is beginning to wander south into the lower Susquehanna region.  Most of these birds will eventually continue on to the pine forests of the southern United States for winter, but a few could remain to become seasonal visitors at feeding stations.
Ruby-throated Hummingbird
Ruby-throated Hummingbirds are on the move; their migration to the tropics is well underway and nearing its peak.  Ruby-throats are diurnal migrants that do a majority of their flying during the hours of daylight.  The notable exception: the nighttime portion of the long southbound flight some of the birds make to cross the Gulf of Mexico.
Red-headed Woodpecker
The Red-headed Woodpecker is another diurnal migrant.   This denizen of temperate climates is currently beginning to move to its wintering grounds, an area that extends from the latitudes of the lower Susquehanna south to the Gulf of Mexico and central Texas.
Juvenile Red-headed Woodpecker
A juvenile Red-headed Woodpecker during a brief pit stop.
Cedar Waxwings
The flights of roving bands of masked Cedar Waxwings continue.  Their numbers appear to be an improvement over those of 2023.
Broad-winged Hawk
At regional hawk-counting stations, observers are seeing more Broad-winged Hawks and other species beginning to move through.
"Kettling" Broad-winged Hawks
The frequency of Broad-winged Hawks passing the lookouts one at a time is giving way to the occurrence of larger and larger “kettling” groups that search out thermal updrafts to save energy while migrating.  By mid-September each of these “kettles” can include one hundred birds or more.  On the peak days, the daily Broad-winged Hawk totals can reach one thousand or more.
Broad-winged Hawk
A Broad-winged Hawk soaring to gain lift from a thermal updraft above a hawkwatch lookout.

The migration is by no means over; it has only just begun.  So plan to visit a local hawkwatch or other suitable ridgetop in coming weeks.  Arrive early (between 7 and 8 AM) to catch a glimpse of a nocturnal migrant fallout, then stay through the day to see the hundreds, maybe even thousands, of Broad-winged Hawks and other diurnal raptors that will pass by.  It’s an experience you won’t forget.

Broad-winged Hawk Gliding Away to the Southwest
A Broad-winged Hawk gliding away to the southwest.

Be certain to click the “Birds” tab at the top of this page for a photo guide to the species you’re likely to see passing south through the lower Susquehanna valley in coming months.  And don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab to find a hawk-counting station near you.

More birds are on the way.  Here’s a look at this evening’s liftoff of nocturnal migrants detected by National Weather Service Radar in State College, Pennsylvania.  (NOAA/National Weather Service Doppler Radar image)

Southbound Flights: On Their Way Both Night and Day

It’s hard to believe, but for almost two months now, sandpipers, plovers, and terns have been filtering south through the Lower Susquehanna River Watershed on their way to the Atlantic coastline as they complete the first leg of their long autumn migration—a journey that will take some species all the way to the far reaches of the South American continent for winter.

Caspian Terns and Osprey
Migrating Caspian Terns and an Osprey take a break on a lower Susquehanna valley gravel bar.

As August draws to a close, these early birds are being joined by widespread nocturnal flights of Neotropical migrants—those species, primarily songbirds, on their way to wintering grounds which lie exclusively south of the continental United States.

To catch a glimpse of these night-flying avians, your best bet may be to position yourself on the crest of a ridge or along a linear break in the forest such as a utility right-of-way where waves of warblers, vireos, flycatchers, and other Neotropical passerines sometimes feed on invertebrates after making landfall at daybreak.  Pick a place where the trees are bathed in the warm light of the rising sun and be there by 7 A.M. E.D.T.  The activity can be tremendous, but it usually ends between 8 and 9.

Red-eyed Vireo
One of more than half a dozen Red-eyed Vireos seen during a daybreak fallout at Second Mountain Hawkwatch in Lebanon County earlier this week.
Blackburnian Warbler
During sunrise, a just-arrived Blackburnian Warbler checks the foliage of a ridgetop Red Maple for insects.
Least Flycatcher
A Least Flycatcher quietly searches the shrubby growth along the forest edge for a morning meal.
Black-and-white Warbler
A Black-and-white Warbler in the first hour of sunlight after a big nocturnal flight.
Scarlet Tanager
A Scarlet Tanager having a look around.  This individual and other migrating birds may be here for a few days before moving on.
Black-throated Green Warbler
A Black-throated Green Warbler in a treetop glowing with the light of sunrise.
Chestnut-sided Warbler
A hungry Chestnut-sided Warbler inspects limbs, leaves, and twigs looking for nourishment after an all-night flight.

Diurnal migrants, birds that make their movements during the daylight hours, are ramping up their flights now as well.  Broad-winged Hawks, Bald Eagles, and falcons are currently being tallied at hawk-counting stations throughout the northeast.  Many of those lookouts are seeing Ruby-throated Hummingbirds, swallows, and other daytime migrants too.

Broad-winged Hawk
While many diurnal migrants gather in flocks as a method of defense against predators, Broad-winged Hawks congregate as a reconnaissance measure.  Not waiting around for strong autumn winds to be deflected upwards by the region’s numerous ridges, these Neotropical birds migrate early enough in the season to rely upon thermal updrafts from sun-heated surfaces to provide lift, gain altitude, and save energy during their long trip.  By traveling in groups, there are collectively able to better locate and utilize rising air columns as they progress southwesterly along their route.  Broad-winged Hawks travel to Central and South America for winter.  Their numbers in the Lower Susquehanna River Watershed will peak during the third week of September.
American Kestrel
Many American Kestrels transit our area in August and early September, a time coinciding with flights of migratory dragonflies, insects upon which they and another falcon, the Merlin, frequently prey.
Chimney Swifts
Chimney Swifts are fast-flying diurnal migrants.  They are beginning to congregate into larger groups in preparation for their departure.  Some are already on the move.
Cedar Waxwing
Cedar Waxwings are currently being seen in roving flocks throughout the region.  Their migration is less of a point-to-point flight and more of a continuous wandering in search of berries and other wild foods.
American Goldfinch
Not everyone is yet ready to go.  American Goldfinches are still in the midst of their nesting cycle and won’t begin leaving until the young are on their own and colorful adult males like this one are beginning to molt into their drab winter plumage.

Autumn migration flights are an ever-changing process, with different species peaking at different times throughout the season.  In these months just after the nesting season, each of these species is more numerous than at any other time of the year.  And of course, the more often we as observers get out and have a look, the more of them we’ll see.

Be certain to click the “Birds” tab at the top of this page for a photo guide to the species you’re likely to see passing through the lower Susquehanna valley this fall.  Nearly four months of autumn hawk migration flights lie ahead, so don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab to find a hawk-counting station near you, then stop by for a visit or two.  See you there!

A large nightfall liftoff of nocturnal migrating birds is shown here on this evening’s State College, Pennsylvania, Doppler Radar base velocity loop.  Returns indicating birds moving toward the radar site (the white dot in the center of the color mass) are shown in green, and those indicating birds moving away are indicated in red.  This is a big flight headed generally in a south-southwest direction.  (NOAA/National Weather Service Doppler Radar image)

A Pre-dawn Thunderstorm and a Fallout of Migrating Birds

In recent days, the peak northbound push of migratory birds that includes the majority of our colorful Neotropical species has been slowed to a trickle by the presence of rain, fog, and low overcast throughout the Mid-Atlantic States.  Following sunset last evening, the nocturnal flight resumed—only to be grounded this morning during the pre-dawn hours by the west-to-east passage of a fast-moving line of strong thundershowers.  The NOAA/National Weather Service images that follow show the thunderstorms as well as returns created by thousands of migrating birds as they pass through the Doppler Radar coverage areas that surround the lower Susquehanna valley.

Sterling, Virginia, Doppler Radar west of Washington, D.C., at 4:00 A.M. E.D.T. indicates a dense flight of northbound migrating birds located just to the south of the approaching line of rain and thunderstorms over the State College, Pennsylvania, radar coverage area.  (NOAA/National Weather Service image)
More northbound birds are indicated at 4 A.M. by the radar station located at Dover, Delaware…  (NOAA/National Weather Service image)
…and by the Mount Holly, New Jersey, radar site.  (NOAA/National Weather Service image)
Many of the migrating birds shown here over the Binghamton, New York, radar station at 4 A.M. probably overflew the lower Susquehanna region earlier in the night.  (NOAA/National Weather Service image)
And these birds over Albany, New York’s, radar station at 4 A.M. are mostly migrants that passed north over New Jersey and easternmost Pennsylvania last evening and during the wee hours of this morning.  (NOAA/National Weather Service image)

Just after 4 A.M., flashes of lightning in rapid succession repeatedly illuminated the sky over susquehannawildlife.net headquarters.  Despite the rumbles of thunder and the din of noises typical for our urban setting, the call notes of nocturnal migrants could be heard as these birds descended in search of a suitable place to make landfall and seek shelter from the storm.  At least one Wood Thrush and a Swainson’s Thrush (Catharus ustulatus) were in the mix of species passing overhead.  A short time later at daybreak, a Great Crested Flycatcher was heard calling from a stand of nearby trees and a White-crowned Sparrow was seen in the garden searching for food.  None of these aforementioned birds is regular here at our little oasis, so it appears that a significant and abrupt fallout has occurred.

White-crowned Sparrow
A White-crowned Sparrow in the headquarters garden at daybreak.  It’s the first visit by this species in a decade or more.

Looks like a good day to take the camera for a walk.  Away we go!

Gray Catbird
Along woodland edges, in thickets, and in gardens, Gray Catbirds were everywhere today.  We heard and/or saw hundreds of them.
American Redstart
During our travels, American Redstarts were the most frequently encountered warbler.  Look for them in low-lying forested habitats.
Many early-arriving Baltimore Orioles have already begun building nests.  But widespread territorial fighting today may be an indication that some latecomer orioles became trespassers after dropping in on existing territories during the morning fallout.
Red-eyed Vireo
Red-eyed Vireos are difficult to see but easily heard in forested areas throughout the Lower Susquehanna River Watershed.
Scarlet Tanager
If the oriole isn’t the showiest of the Neotropical migrants, then the Scarlet Tanager is certainly a contender…
Scarlet Tanager
Listen for their burry, robin-like song in the treetops of mature upland forests.
Wood Thrush
No woodland chorus is complete without the flute-like harmony of the Wood Thrush.  Look and listen for them in rich forests with dense understory vegetation.
Eastern Wood-Pewee
The Eastern Wood-Pewee, another forest denizen, has an easy song to learn…a series of ascending “pee-a-wee” phrases interspersed with an occasional descending “pee-urr”.  It was one of the few flycatchers we found today, but more are certainly on the way.  Their numbers should peak in coming days.
Yellow-rumped Warbler
Yellow-rumped Warblers can be especially numerous during migration but tend to peak prior to the arrival of the bulk of the Neotropical species.  This was the only “yellow-rump” we encountered today.  The majority have already passed through on their way to breeding grounds to our north.
Common Yellowthroat
If today you were to visit a streamside thicket or any type of early successional habitat, you would probably find this perky little warbler there, the Common Yellowthroat.
Yellow Warbler
The Yellow Warbler likes streamside thickets too.  You can also find them along lakes, ponds, and wetlands, especially among shrubby willows and alders.
White-crowned Sparrow
While nowhere near the headquarters garden, we ran into another White-crowned Sparrow in less-than-ideal habitat.  This one was in a row of trees in a paved parking lot.
Bobolink
Not all songbirds migrate at night. The Bobolink is an example of a diurnal (day-flying) migrant.  They’re currently arriving in hay fields that are spared the mower until after nesting season.
Blue-gray Gnatcatcher
While looking for Neotropical species and other late-season migrants, we also found numerous early arrivals that had already begun their breeding cycles.  We discovered this Blue-gray Gnatcatcher on its nest in a Black Walnut tree…
Blue-gray Gnatcatcher
…then, later in the day, we found this one in its nest, again in a Black Walnut tree.  Note the freshly emerging set of leaves and flower clusters.  With many tree species already adorned in a full set of foliage, open canopies in stands of walnuts we found growing in reforested areas seemed to be good places to see lots of migrants and other birds today.  It’s hard to say whether birds were more numerous in these sections of woods or were just easier to observe among the sparse leaf cover.  In either case, the nut-burying squirrels that planted these groves did us and the birds a favor.

There’s obviously more spring migration to come, so do make an effort to visit an array of habitats during the coming weeks to see and hear the wide variety of birds, including the spectacular Neotropical species, that visit the Lower Susquehanna River Watershed each May.  You won’t regret it!

Wood Ducks
Wood Ducks arrived in February and March to breed in the lower Susquehanna valley.  Soon after hatching in April or May, the young leave the nest cavity to travel under the watchful gaze of their ever-vigilant mother as they search for food along our local waterways.  If you’re fortunate, you might catch a glimpse of a brood and hen while you’re out looking at the more than one hundred species of birds that occur in our region during the first half of May.  Good luck!

It’s Not Too Late

In the event you found yourself missing the spectacle of the migration of Neotropical birds through the northeastern United States earlier this month, there is good news.  Apparently the cool and wet weather has prolonged the spring flights which by now are normally slowing to a trickle.  Take a look at this composite radar loop and see the masses of birds rising skyward as nightfall progresses from east to west during the past couple of hours.

 

Nocturnal migrating birds are indicated as they ascend into the path of sweeping radar beams at sites throughout the eastern United States.  The outer bands of Tropical Storm Alberto appear over southern Florida and strong thunderstorms are widespread elsewhere.   (NOAA/National Weather Service image)
A zoomed-in view of Texas shows a number of items of interest.   A tornado warning was issued for the thunderstorm seen north of San Antonio (hook-shaped red in lower center).  In the earliest frames of this loop, two well-defined rings expand around radar sites west of San Antonio, followed by a partial ring in the city and another to the north near the thunderstorm.  These rings plot accurately with the locations of large bat roosts.  The bats, millions of them, ascend high enough to be detected by radar as they depart their roosts to feed at dusk.  Finally at nightfall, great masses of Neotropical birds are displayed as they rise into the busy flyway used by northbound migrants after crossing the Rio Grande from Mexico.  (NOAA/National Weather Service image)

Any of the birds presently moving through Texas could travel for another week or more to reach the Susquehanna region—so there’s still time to see the show.  Let’s grab a camera or some binoculars and go have a look.  It’s not too late.

Feathered Fallout

The NOAA National Weather Service radar images from last evening provided an indication that there may be a good fallout of birds at daybreak in the lower Susquehanna valley.  The moon was bright, nearly full, and there was a gentle breeze from the north to move the nocturnal migrants along.  The conditions were ideal.

Rising from daytime roosts in New York and Pennsylvania, then streaming south in moonlit skies, migrating birds are recorded as echoes on this post-sunset composite NEXRAD loop from last evening.  (NOAA/National Weather Service image)

The Riparian Woodlands at Conewago Falls were alive with migrants this morning.  American Robins and White-throated Sparrows were joined by new arrivals for the season: Brown Creeper (Certhia americana), Ruby-crowned Kinglets (Regulus calendula), Golden-crowned Kinglets (Regulus satrapa), Dark-eyed Junco (Junco hyemalis), and Yellow-rumped Warbler (Setophaga coronata).  These are the perching birds one would expect to have comprised the overnight flight.  While the individuals that will remain may not yet be among them, these are the species we will see wintering in the Mid-Atlantic states.  No trip to the tropics for these hardy passerines.

American Robins continued migratory flight into the first hour of daylight this morning.  Their calls are commonly heard at night as migrating individuals pass overhead.
White-throated Sparrows are nocturnal migrants, and are a familiar find on woodland edges and at suburban feeding stations through the winter.
Dark-eyed Juncos, also nocturnal migrants, are common winter residents in the Lower Susquehanna River Watershed, frequently visiting bird feeders.
Heavy rain earlier this week in the Susquehanna River drainage basin has flooded most of the Pothole Rocks; the rapids of Conewago Falls have returned.
CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS

Yellowlegs for Breakfast

A few nocturnal migrants flew through the moonlit night to arrive at Conewago Falls for a sunrise showing this morning.  A dozen warblers were in the treetops and a Wood Thrush (Hylocichla mustelina) chattered away in the understory of the Riparian Woodlands.  Three species of shorebirds were in the falls and on the Pothole Rocks: Least Sandpiper (Calidris minutilla), Lesser Yellowlegs (Tringa flavipes), and Greater Yellowlegs (Tringa melanoleuca).

A Greater Yellowlegs (right) and two Lesser Yellowlegs sandpipers dropped by for breakfast.

The diurnal migration was highlighted by a Merlin (Falco columbarius), an Osprey, and a Bald Eagle, each flying down the river.  Most of the other birds in the falls seemed content to linger and feed.  There’s no need to hurry folks, only trouble lurks down there in paradise at the moment.

A light to moderate flight of nocturnal migrants in the eastern United States is displayed on NOAA National Weather Service NEXRAD radar at 4:58 AM EDT.  The eye of Hurricane Irma can be seen approaching the Florida Keys.  (NOAA/National Weather Service image)
CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS

Piles of Green Tape

A couple of inches of rain this week caused a small increase in the flow of the river, just a burp, nothing major.  This higher water coincided with some breezy days that kicked up some chop on the open waters of the Susquehanna upstream of Conewago Falls.  Apparently it was just enough turbulence to uproot some aquatic plants and send them floating into the falls.

Piled against and upon the upstream side of many of the Pothole Rocks were thousands of two to three feet-long flat ribbon-like opaque green leaves of Tapegrass, also called Wild Celery, but better known as American Eelgrass (Vallisneria americana).  Some leaves were still attached to a short set of clustered roots.  It appears that most of the plants broke free from creeping rootstock along the edge of one of this species’ spreading masses which happened to thrive during the second half of the summer.  You’ll recall that persistent high water through much of the growing season kept aquatic plants beneath a blanket of muddy current.  The American Eelgrass colonies from which these specimens originated must have grown vigorously during the favorable conditions in the month of August.  A few plants bore the long thread-like pistillate flower stems with a fruit cluster still intact.  During the recent few weeks, there have been mats of American Eelgrass visible, the tops of their leaves floating on the shallow river surface, near the east and west shorelines of the Susquehanna where it begins its pass through the Gettysburg Basin near the Pennsylvania Turnpike bridge at Highspire.  This location is a probable source of the plants found in the falls today.

Uprooted American Eelgrass floating into the Pothole Rocks under the power of a north wind.  Note the white thread-like pistillate flower stem to the left and the small rooted specimen to the upper right.  The latter is likely a plant from the creeping rootstock on the edge of a colony.  As a native aquatic species, American Eelgrass is a critical link in the Susquehanna River and Chesapeake Bay food chain.  Its decimation by pollution during the twentieth century led to migration pattern alterations and severe population losses for the Canvasback (Aythya valisineria) duck.
American Eelgrass, a very small specimen, found growing in a low-lying Pothole Rock alongside the accumulations of freshly arriving material from upstream.  Note that the creeping rootstock has leaves growing from at least three nodes on this plant.  Eelgrass dislocations are regular occurrences which sometimes begin new colonies, like the small one seen here in this Diabase Pothole Rock Microhabitat.

The cool breeze from the north was a perfect fit for today’s migration count.  Nocturnal migrants settling down for the day in the Riparian Woodlands at sunrise included more than a dozen warblers and some Gray Catbirds (Dumetella carolinensis).  Diurnal migration was underway shortly thereafter.

A moderate flight of nocturnal migrants is indicated around NEXRAD sites in the northeastern states at 3:18 AM EDT.  The outer rain bands of Hurricane Irma can be seen approaching the Florida Keys as the storm closes in on the peninsula.  (NOAA/National Weather Service image)

Four Bald Eagles were counted as migrants this morning.  Based on plumage, two were first-year eagles (Juvenile) seen up high and flying the river downstream, one was a second-year bird (Basic I) with a jagged-looking wing molt, and a third was probably a fourth year (Basic III) eagle looking much like an adult with the exception of a black terminal band on the tail.  These birds were the only ones which could safely be differentiated from the seven or more Bald Eagles of varying ages found within the past few weeks to be lingering at Conewago Falls.  There were as many as a dozen eagles which appeared to be moving through the falls area that may have been migrating, but the four counted were the only ones readily separable from the locals.

Red-tailed Hawks (Buteo jamaicensis) were observed riding the wind to journey not on a course following the river, but flying across it and riding the updraft on the York Haven Diabase ridge from northeast to southwest.

Bank Swallows (Riparia riparia) seem to have moved on.  None were discovered among the swarms of other species today.

Ruby-throated Hummingbirds, Caspian Terns, Cedar Waxwings (Bombycilla cedrorum), and Chimney Swifts (Chaetura pelagica) were migrating today, as were Monarch butterflies.

Not migrating, but always fun to have around, all four wise guys were here today.  I’m referring to the four members of the Corvid family regularly found in the Mid-Atlantic states: Blue Jay (Cyanocitta cristata), American Crow (Corvus brachyrhynchos), Fish Crow (Corvus ossifragus), and Common Raven (Corvus corax).

It looks like a big Blue Jay, but it’s not.  This Belted Kingfisher (Megaceryle alcyon) takes a break after flying around the falls trying to shake a marauding Ruby-throated Hummingbird off its tail.
CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS

SOURCES

Klots, Elsie B.  1966.  The New Field Book of Freshwater Life.  G. P. Putnam’s Sons.  New York, NY.

Strangers In The Night

We all know that birds (and many other animals) migrate.  It’s a survival phenomenon which, above all, allows them to utilize their mobility to translocate to a climate which provides an advantage for obtaining food, enduring seasonal weather, and raising offspring.

In the northern hemisphere, most migratory birds fly north in the spring to latitudes with progressively greater hours of daylight to breed, nest, and provide for their young.  In the southern hemisphere there are similar movements, these to the south during their spring (our autumn).  The goal is the same, procreation, though the landmass offering sustenance for species other than seabirds is limited “down under”.  Interestingly, there are some seabirds that breed in the southern hemisphere during our winter and spend our summer (their winter) feeding on the abundant food sources of the northern oceans.

Each autumn, migratory breeding birds leave their nesting grounds as the hours of sunlight slowly recede with each passing day.  They fly to lower latitudes where the nights aren’t so long and the climate is less brutal.  There, they pass their winter season.

Food supply, weather, the start/finish of the nesting cycle, and other factors motivate some birds to begin their spring and autumn journeys.  But overall, the hours of daylight and the angle of the sun prompt most species to get going.

But what happens after birds begin their trips to favorable habitats?  Do they follow true north and south routes?  Do they fly continuously, day and night?  Do they ease their way from point to point, stopping to feed along the way?  Do they all migrate in flocks?  Well, the tactics of migration differ widely from bird species to species, from population to population, and sometimes from individual to individual.  The variables encountered when examining the dynamics of bird migration are seemingly endless, but fascinatingly so.  Bird migration is well-studied, but most of its intricacies and details remain a mystery.

Consider for a moment that just 10,000 years ago, an Ice Age was coming to an end, with the southernmost edge of the most recent glaciers already withdrawn into present-day Canada from points as near as the upper Susquehanna River watershed.  Back then, the birds migrating to the lower portion of the drainage basin each spring probably weren’t forest-dwelling tropical warblers, orioles, and other songbirds.  The migratory birds that nested in the lower Susquehanna River valley tens of millennia ago were probably those species found nesting today in taiga and tundra much closer to the Arctic Circle.  And the ancestors of most of the tropical migrants that nest here now surely spent their entire lives much closer to the Equator, finding no advantage by journeying to the frigid Susquehanna valley to nest.  It’s safe to say that since those times, and probably prior to them, migration patterns have been in a state of flux.

During the intervening years since the great ice sheets, birds have been able to adapt to the shifts in their environment on a gradual basis, often using their unmatched mobility to exploit new opportunities.  Migration patterns change slowly, but continuously, resulting in differences that can be substantial over time.  If the natural transformations of habitat and climate have kept bird migration evolving, then man’s impact on the planet shows great potential to expedite future changes, for better or worse.

Now, let’s look at two different bird migration strategies, that of day-fliers or diurnal migrants, and that of night-fliers, the nocturnal migrants.

Diurnal migrants are the most familiar to people who notice birds on the move.  The majority of these species have one thing in common, some form of defense to lessen the threat of becoming the victim of a predator while flying in daylight.  Of course the vultures, hawks, and eagles fly during the day.  Swallows and swifts employ speed and agility on the wing to avoid becoming prey, as do hummingbirds.  Finches have an undulating flight, never flying on a horizontal plane, which makes their capture more difficult.  Other songbirds seen migrating by day, Red-winged Blackbirds for example, congregate into flocks soon after breeding season to avoid being alone.  Defense flocks change shape constantly as birds position themselves toward the center and away from the vulnerable fringes of the swarm.  The larger the flock, the safer the individual.  For a lone bird, large size can be a form of protection against all but the biggest of predators.  Among the more unusual defenses is that of birds like Indigo Buntings and other tropical migrants that fly across the Gulf of Mexico each autumn (often completing a portion of the flight during the day), risking exhaustion at sea to avoid the daylight hazards, including numerous predators, found in the coastal and arid lands of south Texas.  Above all, diurnal migrants capture our attention and provide a spectacle which fascinates us.  Perhaps diurnal migrants attract our favor because we can just stand or sit somewhere and watch them go by.  We can see, identify, and even count them.  It’s fantastic.

What about a bird like the Canada Goose (Branta canadensis)?  It is often seen migrating in flocks during the day (the truly migratory ones flying much higher than the local year-round resident “transplants”), but then, during the big peak movements of spring and fall, they can be heard overhead all through the night.  Perhaps the Canada Goose and related waterfowl bridge the gap between day and night, introducing us to the secretive starlight and moonshine commuters, the nocturnal migrants.

The high-flying migratory Canada Goose can be seen during the daytime and heard at night when passing over the lower Susquehanna River valley.  A large flight exiting from Chesapeake Bay in late February or early March often results in a 12 to 24 hour-long stream of northbound flocks.

The skies are sometimes filled with thousands of them, mostly small perching birds and waders.  These strangers in the night fly inconspicuously in small groups or individually, and most can be detected when passing above us only when heard making short calls to remain in contact with their travel partners.  They need not worry about predators, but instead must have a method of finding their way.  Many, like the Indigo Bunting, can navigate by the stars, a capability which certainly required many generations to refine.  The nocturnal migrants begin moving just after darkness falls and ascend without delay to establish a safe flight path void of obstacles (though lights and tall structures can create a deadly counter to this tactic).  Often, the only clue we have that a big overnight flight has occurred is the sudden appearance of new bird species or individuals, on occasion in great numbers, in a place where we observe regularly.  Just days ago, the arrival of various warbler species at Conewago Falls indicated that there was at least a small to moderate movement of these birds during previous nights.

In recent years, the availability of National Weather Service radar has brought the capability to observe nocturnal migrants into easy reach.  Through the night, you can log on to your local National Oceanic and Atmospheric Administration’s National Weather Service radar page (State College for the Conewago Falls area) and watch on the map as the masses of migrating bird pass through the sweep of the radar beam.  As they lift off just after nightfall, rising birds will create an echo as they enter the sweeping beam close to the radar site.  Then, due to the incline of the transmitted signal and the curvature of the earth, migrants will be displayed as an expanding donut-like ring around the radar’s map location as returns from climbing birds are received from progressively higher altitudes at increasing distances from the center of the site’s coverage area.  On a night with a local or regional flight, several radar locations may show signs of birds in the air.  On nights with a widespread flight, an exodus of sorts, the entire eastern half of the United States may display birds around the sites.  You’ll find the terrain in the east allows it to be well-covered while radars in the west are less effective due to the large mountains.  At daybreak, the donut-shaped displays around each radar site location on the map contract as birds descend out of the transmitted beam and are no longer detected.

Weather systems sometimes seem to motivate some flights and stifle others.  The first example seen below is a northbound spring exodus, the majority of which is probably migrants from the tropics, the Neotropical migrants, including our two dozen species of warblers.  A cold front passing into the northeastern United States appears to have stifled any flight behind it, while favorable winds from the southwest are motivating a heavy concentration ahead of the front.

National Oceanic and Atmospheric Administration/National Weather Service radar image from May 5, 2010, at 11:18 PM EDT, shows rain associated with a cold front moving east from the border of Ontario, Canada, and the United States into New England and the Mid-Atlantic region.  The heavy blue and green reflections surrounding the radar locations ahead of the front are nocturnal migrating birds taking advantage of favorable conditions for flight including a tail wind from the southwest.  Note the lighter migration behind the advancing front.  Heavy radar echoes on the gulf coast, particularly in Texas, indicate dense bird concentrations exiting the tropics to fan out into North American breeding areas.  The westward progression of expanding echoes surrounding individual radar sites indicates birds rising into the radar beam at local nightfall.

The second and third examples seen below are an autumn nocturnal migration movement, probably composed of many of the same tropics-bound species which were on the way north in the previous example.  Note that during autumn, the cold front seems to motivate the flight following its passage.  Ahead of the front, there is a reduced and, in places, undetectable volume of birds.  The two images below are separated by about 42 hours.

National Oceanic and Atmospheric Administration/National Weather Service radar image (still) from September 5, 2017, at 2:38 AM EDT, shows rain in the northeast associated with a slow-moving cold front stretching from Maine southwest to New Mexico.  Heaviest nocturnal bird flights can be seen behind the advancing front where there are favorable tail winds from the north or northwest.
Nearly two full days later, the slow-moving cold front from the previous image has crossed Pennsylvania.  As nightfall progresses from east to west, ascending nocturnal migrants enter NEXRAD radar beams, their echoes creating expanding rings around individual sites.  Concentrations of southbound birds can be seen along the gulf coast.  Many will follow the Texas coastline into Mexico.  The Neotropical migrants that try to cross the Gulf of Mexico this night could be in for a perilous voyage.  Hurricane Katia is churning in the southern gulf and a much stronger storm, Hurricane Irma, is rolling toward the Bahamas and Florida from the southeast.  Masses of birds that follow learned routes or instinct to venture offshore and cross seas under such circumstances could suffer catastrophic losses.  (NOAA/National Weather Service image)

You can easily learn much more about birds (and insects and bats) on radar, including both diurnal and nocturnal migrants, by visiting the Clemson University Radar Ornithology Laboratory (CUROL) website.  There you’ll find information on using the various mode settings on NEXRAD (Next-Generation Radar) to differentiate between birds, other flying animals, and inanimate airborne or grounded objects.  It’s superbly done and you’ll be glad you gave it a try.

SOURCES

Clemson University Radar Ornithology Laboratory (CUROL) website:   http://virtual.clemson.edu/groups/birdrad/    as accessed September 6, 2017.