The Stewards of Inland Shorebird Habitat

As they travel between coastal wintering ranges and breeding territories in Canada and Alaska, vast numbers of shorebirds pass through the lower Susquehanna region each spring and fall—though few stop here to rest, feed, and provide us with an opportunity to observe them.  Prior to the construction of man-made dams and other alterations on our lotic (flowing) waterways including the river, shorebirds took advantage of lateral bars, stream deltas, and other alluvial deposits as places to loaf and re-energize.  Before they were drained and filled, some of the valley’s wetlands probably included sparsely vegetated flats where shorebirds could drop in for a brief visit.  Previous to their extirpation from our region, North American Beavers were the primary providers of quality habitat for shorebirds and other migrating waders on our lotic waters.  Their widespread network of dams, pools, and marshes maximized floodplain function by keeping streams thoroughly connected to their wetlands, nurturing plant communities that not only provided food and shelter for the beavers and other wildlife, but provided superb buffering against erosion while protecting against sediment and nutrient imbalances in lower Susquehanna waterways.

Beaver Dam and Lodge
Beaver dams need not be large, particularly on low-gradient streams where a structure like this is sufficient to create a pool with depth adequate for building and maintaining a lodge and transporting leafy branches and other food items by water.
A beaver lodge assembled in a pool with less than three feet of water, deep enough to provide the family with a measure of protection against terrestrial predators.
Beaver Pool
Beaver pool ecosystems provide homes for hundreds of species of plants and animals, including migrating shorebirds and other waders.
Solitary Sandpiper
Mud flats in the margins between emergent shrubs and herbaceous plant growth attract migrating shorebirds like this Solitary Sandpiper to the abundance of invertebrate life.  Seasonal movements of migrating shorebirds regularly coincide with the reductions of water levels in beaver pools which typically occur between May and September each year.
Least Sandpiper
During its southbound migration in September, a Least Sandpiper searches for arthropods and annelids as it visits a food-rich puddle along the periphery of a beaver pool.
Pectoral Sandpiper
A Pectoral Sandpiper feeds in shallow water near a stand of Common Cattails in a beaver pool.
Sora
During migration, rails including the Sora are attracted to dense emergent vegetation in beaver pools.  Some will nest in these rodent-managed refugia.
Green Heron
Green Herons visit beaver pools during migration and, due to the reliability of the food supply, will often nest in their vicinity.

So how did this happen?  How did the North American Beaver become a keystone species—an animal upon which the majority of other life forms within its ecosystem are so reliant?  Well, it’s largely due to the fact that our beavers aren’t particularly fond of a constant stream of noise.  More specifically, they don’t like the sound of running water in places where they intend to build and maintain a lodge.  And so, as they begin to place sticks, mud, limbs, stones, and other materials within a noisy riffle on a stream, they create a dam, and behind it a pool—a pool that is particularly advantageous for protecting their home and providing a means of conveyance for their construction materials and food supplies.

Cascading Series of Beaver Pools
On a high-gradient segment of stream, beavers will create a cascading series of pools. Because water filters through a beaver dam instead of spilling over it, the work of these meticulous rodents soon silences the sounds of water changing altitude.  No more sonorous riffle.
Beaver Dam
Quiet please.  High seasonal stream flow and damage from storms may create areas where water begins to erode the structure of the dam.  Where this condition persists, an adult beaver will soon mend the breach, just to quiet things down.  Why would beavers demand such a hush upon their domain?  Well, they have poor eyesight, but their hearing is excellent, and they rely upon it to detect danger.
Louisiana Waterthrush
The Louisiana Waterthrush (Parkesia motacilla), one of our earliest-arriving warblers, nests in forests along clear, high-gradient streams.  In mid-April, we found this individual and three others squabbling over a breeding territory adjacent to the series of cascading beaver pools shown in the previous images.
Brook Trout and Blacknose Dace
Native denizens of coldwater streams, neither the Brook Trout nor the Eastern Blacknose Dace has any difficulty finding its way through the voids in beaver dams to ascend and descend the sequence of pools.
Beaver Dam and Pool on Low-gradient Stream
On a low-gradient segment of stream, a dam just over a foot in height may be sufficient to create a beaver pool of considerable size.  Resembling the water-logged muskeg of the far north, well-established beaver pools form boggy habitats familiar to migrating shorebirds.
Water levels in the pools usually drop along with the stream’s base flow as soon as the effects of the spring thaw and rain showers subside.  Feeding areas in shallow water and on muddy ground are often revealed just in time for northbound shorebirds and other waders to stop by in late April and May.  These conditions often persist through the growing season and the fall migration of these birds which begins as early as late June and sometimes extends into October.
Tree Swallow at Natural Cavity
Where the pool inundates standing timber such as Red Maple and other species not used by beavers, the dead snags provide vital feeding areas for birds and other wildlife.  Cavity nesters like this Tree Swallow seldom find suitable natural housing elsewhere.
Eastern Newts Among Alderfly Exuvia
The mosaic of marshlands and braided stream channels within the beaver pool complex supports an abundance of aquatic life including these breeding Eastern Newts, seen here surrounded by the exuvia left behind following a massive hatch of alderflies (Sialinae).  Alderflies are a stream inhabitant during their larval stage and are indicator of clean water conditions.
A modern-day example of the way fully functional stream floodplains used to look in the Lower Susquehanna River Watershed.  Though the pools may appear pond-like during the cooler and wetter months of the year, by summer the water levels behind the beaver dams recede as the base flow of the stream wanes.  Now the true nature of the marsh, shallow pool, and braided stream complex is revealed.  Unlike most man-made dams that set a fixed pool elevation regardless of flow by discharging water over the top of the structure or through a spillway or gate, beaver dams merely throttle the flow through their porous construction.  Unless the beaver begins plugging small leaks as fast as the stream flow ebbs, the water level in the pools will drop.  So they’re not left high and dry, lodges are located in the deepest pools, usually in close proximity to the dam and/or one of the stream courses.  Though not inundated during the dry season, the soil in the pool complex is almost always damp and plants grow vigorously there, sequestering nutrients and retaining sediments in beneficial deposition patterns that actually inhibit erosion of the riparian landscape.  These streams and floodplains retain their hyporheic zone and freely exchange water with the underlying water table and aquifer.  It’s the ultimate floodplain management system, and the beavers don’t even know their doing all the work.  (Google Maps base image)
Where another high-gradient segment of stream enters the main pool complex, beavers have created an additional series of cascading pools.  The impoundments created by these dams help diffuse stormwater energy and process more nutrients and sediments within the floodplain’s wetland vegetation system.  (Google Maps base image)
Sandhill Cranes in Beaver Pool
Beaver pools often refill as stream flows increase following autumn rains.  The stockpiles of vegetative foods that grew within the beaver’s domain through the summer then become flooded and are a prime source of nutrition for not only the beavers, but waterfowl during both their autumn and spring migrations.  Gleaning and probing Sandhill Cranes often find these habitats to their liking as well.

While North American Beavers have returned to the region, most live as “bank beavers”, residing in the river and larger creeks of the valley where they excavate shelters among the roots of Black Willows and other shoreline trees and shrubs.  Floodplain encroachment, legacy sediment deposits, and just plain human intolerance have all conspired to prohibit North American Beavers from performing their magic on smaller local streams.  For migrating shorebirds, this continued absence of beaver dam ecosystems has turned much of the lower Susquehanna valley into “flyover country”.  Those travelers that do stop to rest and feed concentrate at the few favorable locations such as the lateral bars and the hydroelectric dam-created delta at the Conejohela Flats on the river in Lancaster County.  But centralization has its drawbacks.  Migrants spending time at concentration points may have a greater chance of contracting and spreading disease.  Protracted heavy foraging can degrade these habitats.  And over time, features such the lateral bars and delta deposits, including those on the Conejohela Flats, transition into other habitats—riparian forests.  A more widespread selection of favorable stopover points for shorebirds, waders, waterfowl, and other migrants is certainly desirable.

IMITATION IS THE SINCEREST FORM OF FLATTERY

Until public sentiment sways in favor of the North American Beaver, wildlife managers are mimicking some of the attributes of their sound-inspired installations.

New Impoundment at Middle Creek Wildlife Management Area
Created by excavating a depression in heavy clay soils, this new impoundment at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area collects water directly from rainfall and from surface runoff.  Its depth at no point is greater than about two or three feet.  During the driest times of the year, this space will be a mudflat, and a haven for migrating shorebirds.

Shallow-water conservation impoundments designed, constructed, and managed for migratory waterfowl and waders including shorebirds are not what we typically refer to as ponds—though they are lentic (still) waters.  Similar shallow freshwater impoundments at our National Wildlife Refuges are referred to as pools by the United States Fish and Wildlife Service, but smaller versions like this example at Middle Creek very closely resemble the prairie potholes created by glacial scour in the north-central United States and adjacent portions of Canada.  Many populations of migratory birds are familiar with pothole ecosystems and, like the beaver pools and marshes, have relied upon them as waypoints along their journeys for centuries.

Pond and Lake Zonation
Impoundments most beneficial to migrating waterfowl and waders including shorebirds are shallow in depth.  They lack the deeper waters of a pond or lake and thus have no limnetic (open water) or aphotic zones.  Managed throughout as a littoral zone, impoundments grow plants in shallow water or on damp soil through the summer months to chew up nutrients accumulated in waste deposited by visiting birds.  This vegetation is then flooded from late fall through early spring as forage habitat for migratory waterfowl.  The timing of the fluctuations in water levels approximates those of the beaver pools and marshes on lotic (flowing) waters.
Impoundment Gate
A gate assembly is a water control structure installed to provide seasonal management of the water levels in a conservation impoundment.  More than once, beavers have heard the sounds of tumbling waters inside these types of devices and tried to dam them up!

You’ve heard the line, “If you build it, they will come.”  Well, it’s true.  Here is a sample of the activity witnessed during the past two weeks at the new impoundment completed just several months ago at Middle Creek Wildlife Management Area along the tour route just beyond the “Stop 3” overlook.

Snow Geese
The ubiquitous Snow Geese were among the first migratory occupants of the new impoundment.  A few are currently lingering in the vicinity.
Blue-winged Teal
Ducks soon followed.  These Blue-winged Teal were among the last to pass through earlier this month.
Wilson's Snipe
Wilson’s Snipe were among the first species of migratory shorebirds to visit the new impoundment at Middle Creek…
Killdeer
…as were Killdeer, a species which nests nearby.
Least Sandpipers
Then, earlier this month, flocks of shorebirds including these Least Sandpipers were arriving to feed and rest.
Least Sandpipers
Least Sandpipers search for small invertebrates in shallow water and exposed soil.
Least Sandpiper and a Semipalmated Sandpiper
A Least Sandpiper (left) and a Semipalmated Sandpiper (right).
Solitary Sandpiper
Not all shorebirds seen by themselves are alone, and that includes the Solitary Sandpiper.
White-rumped, Least, and Solitary Sandpipers
Here’s a Solitary Sandpiper (right) feeding alongside a Least Sandpiper (center) and a White-rumped Sandpiper (left).
Least Sandpiper and a White-rumped Sandpiper
A Least Sandpiper (left) and a White-rumped Sandpiper (right).
White-rumped Sandpiper
Rare in our area during spring, the White-rumped Sandpiper (Calidris fuscicollis) flies north using the central flyway, then heads south along the Atlantic flyway in the fall, when it tends to be more regular here.
Spotted Sandpiper
Rocking and teetering along as it looks for food, the Spotted Sandpiper may be one of the easiest of shorebirds to identify.
Dunlin
Chunky little Dunlin (Calidris alpina) with their conspicuously down-curved bills are another easy-to-identify species, particularly in the spring when their breeding (alternate) plumage includes a black belly.
Black-bellied Plovers
Black-bellied Plovers acquire a handsome set of plumage in the spring as well.
Killdeer
Killdeer too are plovers and this pair appears to have taken up residence on barren ground along the periphery of the new impoundment.
Semipalmated Plover
The Semipalmated Plover doesn’t have nearly the flair for ornament its close relatives the Killdeer do; these little shorebirds wear only one ring around their neck instead of two.  Think this plover is cute?,…
Killdeer
…then check out this newly hatched Killdeer.  It starts life with just one necklace too, but acquires a second as it grows.  Look at those legs!  (If you visit the new impoundment at Middle Creek, drive slowly and please watch where you’re going.  Baby Killdeer and other young birds, as well as mammals and turtles, are commonly crossing the paved surfaces right now.)
Lesser Yellowlegs
Speaking of legs, here’s one of dozens of Lesser Yellowlegs that visited the new impoundment during their recent northbound travels.
Greater Yellowlegs
Though less numerous than the smaller Lesser Yellowlegs, a Greater Yellowlegs seldom goes unnoticed when dropping by the man-made pothole habitat.
Greater Yellowlegs
Have trouble telling a Greater Yellowlegs like this one from a Lesser Yellowlegs?  Look for the heavier, longer bill on the former, as well as dark barring along its flanks below the wings while the bird is in breeding (alternate) plumage during the spring migration.
Wilson's Phalarope
Everyone likes to see something unusual every now and then, and this impoundment delivers.  Just yesterday, we photographed this migrating Wilson’s Phalarope (top center) among two Least Sandpipers (top left and right), a Lesser Yellowlegs (left and slightly forward of the phalarope), and a White-rumped Sandpiper (foreground).
Wilson's Phalarope
Renowned for spinning in circles as they feed in shallow water, Wilson’s Phalaropes (Phalaropus tricolor) passing through the lower Susquehanna basin are headed toward nesting areas in the prairie wetlands, including potholes, of the northern United States and adjacent sections of southern Canada.

Species of wildlife in addition to shorebirds and waterfowl have already found the new impoundment favorable…

Spot-winged Gliders
Pairs of breeding Spot-winged Gliders (Pantala hymenaea), seen here in tandem while ovipositing, were swarming the impoundment after arriving earlier this week.  These dragonflies are ofttimes unpredictable nomads and are similar in appearance to the usually more numerous Wandering Gliders.  To recolonize seasonal portions of their range, both are famous for hitching a ride en masse on storm systems.  They share the behavior of finding ephemeral and new bodies of water favorable for egg laying due to their low density of aquatic predators.
Snapping Turtle
We watched this Snapping Turtle arrive to apparently find the newly created waters to its liking.  Snapping Turtles are important consumers in a wetland ecosystem.  Larger specimens may fill the role of an upper trophic level or apex predator, eliminating vulnerable mid-level consumers including other Snapping Turtles.
American Pipit
Because it bounces its tail up and down like a Spotted Sandpiper, you may at first glance mistake an American Pipit for a shorebird.  Long known as the Water Pipit, these songbirds have been visiting the impoundment while migrating north and stopping to feed in nearby croplands.
Glossy Ibis
Seen here with Least Sandpipers is a visiting Glossy Ibis.
Glossy Ibis
Sometimes twenty or more of these mostly coastal waders have made a pit stop at the “new Middle Creek pothole”, though none have thus far chosen to remain long.  Apparently, food sources sufficient to sustain a bird of their size have yet to develop in its benthos.
Western Cattle Egrets
Western Cattle Egrets visiting Middle Creek this spring have been frequenting the new impoundment.
Western Cattle Egret
With still little in the way of insects such as grasshoppers available in the surrounding landscape,…
Green Frog
…cattle egrets are looking to find prey like this Green Frog.
Western Cattle Egret
Many have been observed hunting the adjacent grasslands..
Western Cattle Egret
Where small mammals, mostly Meadow Voles, are being taken in abundance.

Managing saturation levels in shallow-water impoundments to resemble the seasonal variations in beaver pool and marsh systems can create lush growth and wildlife-rich environments.  Take a look at some images from a project in a headwaters area of a tributary to Conewago Creek (west)…

Well-established Shallow-water Impoundment
By late July, southbound shorebirds were already using these mudflats to feed and rest.  Other sections of the impoundment were dense with emergent and aquatic plants, the latter kept hydrated in deeper pools of the project by the inflow from several captured springs that supplement direct rainfall and sheet runoff to supply its water.  During a seasonal drawdown, the exposure of the impoundment’s soils to direct sunlight can provide a measure of disinfection to reduce the chances of disease transmission among its populations of visiting birds and other animals.
Water Lily Growth
In the deeper pools of the impoundment, water lilies and other aquatic plants grow in lush mats to provide cover and feeding areas for resident populations of breeding reptiles and amphibians.
Green Heron
An abundance of foods are available for waders including this Green Heron…
Immature Little Blue Heron
…and this immature Little Blue Heron, a wanderer typical of more southern latitudes.
Sandhill Cranes
While walking the road among tall grasses in the supporting landscape surrounding this impoundment, we were at first startled when these Sandhill Cranes strode by going the other direction.  We quietly kept moving,…
Sandhill Cranes
…then spotted them again as we looked across the impoundment to realize they weren’t alone, but were escorting a colt.
Sandhill Cranes with Colt
The hatching of this colt is testimony to the vital role wetland ecosystems play in the lives of hundreds of species.  Whether they be beaver pools and marshes on lotic waters or man-made shallow lentic waters, each of these habitats is filling a void that left floodplains and other critical lowland biomes faltering.  While they can’t replace the full-function floodplain management provided by an active beaver colony, shallow-water impoundments can provide relief for habitat-starved populations of the animals and plants that rely upon them.  A constellation of these projects on lands public or private across the lower Susquehanna watershed could help provide refuge for many of our flora and fauna with the most desperately fragmented of ranges.

So that you can relax while observing the comings and goings at a pair of the lower Susquehanna valley’s man-made impoundments, the Pennsylvania Game Commission has erected two viewing pavilions for public use on its lands…

The Haldeman Island observation pavilion is located on State Game Lands 290  just upstream from the Juniata River’s mouth on the Susquehanna at Clark’s Ferry in Dauphin County.
Haldeman Island Pavilion
It overlooks not only the island’s man-made shallow-water impoundments and neighboring grasslands, but the tower used in the 1970s to reintroduce Bald Eagles from Saskatchewan to the lower Susquehanna.  Interpretive signs explain the conservation stories of habitats and the eagle reintroduction program.
Middle Creek W.M.A. Observation Pavilion
The observation pavilion at Middle Creek Wildlife Management Area is of similar construction to the one at Haldeman Island.  It is accessed from the parking pull-off along the tour road at its intersection with Chapel Road, just before the right turn and incline that leads to the “Stop 3” grassland overlook.
It too includes numerous interpretive signs to help visitors understand impoundment management.

During the next two weeks, the exodus of migrating shorebirds now staged and feeding upon Atlantic Horseshoe Crab eggs on Delaware Bay will commence.  During the evening of their departure from the bay, many of these birds cross portions of the Lower Susquehanna River Watershed, particularly east of the river.  Stormy weather and other climatic conditions may force some of them to seek a place to put down temporarily, so keeping a close eye on the new pothole-like impoundment at Middle Creek may be a prudent move.  After that, waders known as “post-breeding wanderers” can show up at any time.  Then, beginning as early as late June, shorebirds begin moving south on a migration that can provide us with viewing opportunities into September and beyond.  See you out there!

Early Season Wildflowers

Here are five common forest flowers that the average visitor to these environs may easily overlook during an early April visit.

Skunk Cabbage
The flowers of the Skunk Cabbage, a native member of the arum family, are contained within a spadix which is partially hidden inside the mottled maroon spathe at the base of the plant.  In late winter, the closed spathe generates its own heat to melt through frozen water and soils to make its appearance in streamside and spring-fed wooded wetlands, just ahead of the emergence of the large green leaves.
Common Spicebush
Common Spicebush is a native flowering shrub of damp woodland understories.  Later in the season, its foliage provides food for Spicebush Swallowtail caterpillars.  Pollinated blossoms yield bright-red oblong berries relished by a variety of birds in fall and winter.
Red Maple
The flowers of the indigenous Red Maple will soon generate the familiar helicopter-like winged seeds which readily distribute this native tree into new ground ranging from lowlands to the crests of our highest ridges.
Coltsfoot
Coltsfoot (Tussilago farfara) is non-native wildflower of Eurasian origin.  Reminding one at first glance of a dandelion, it is commonly seen blooming in disturbed areas of woodlands.  Coltsfoot often grows where it has little competition from other plants, such as among the debris left behind due to snow plowing and grading along forest roads.
Sweet Cherry
Though not particularly abundant, the non-native Sweet Cherry, an escape from cultivation, is widespread in forests and woodlots throughout the lower Susquehanna valley.  Sweet Cherry is believed to be an ancestor of the Sour Cherry (Prunus cerasus) and is frequently used as rootstock for orchard and garden varieties of this popular fruit.  Sweet Cherry is often called “Wild Cherry”, a name also applied to the Black Cherry, a native tree which blooms later in the spring.

Be certain to get out and enjoy this year’s blooming seasons of our hundreds of varieties of flowering plants.  But, particularly when it comes to native species,…

Early-spring Maples: Clouds of Color Across the Forest Canopy

By late winter, sunshine and warming temperatures awaken the sleeping maples of the lower Susquehanna basin and the sap begins to flow.  The new growing season is evident by the first days of spring when their swelling buds and flowers paint the drab gray canopy of deciduous woodlands with an overlying coat of red, orange, and maroon.

Red Maples in Early Spring
Red Maples, renowned for their vibrant autumn foliage, are just as conspicuous in the early spring landscape.
Eastern Gray Squirrel Feeding on Red Maple Buds
For birds and mammals like this Eastern Gray Squirrel, the buds and flowers of maples mean relief from the diminished natural food supplies that helped them survive winter’s worst.  The season of abundance has arrived and these animals instantly recognize this source of sweet, fresh energy, just in time for the breeding season.  The flowers of maples, which later in the spring yield those familiar helicopter-like seeds, are an important early season source of energy for many species of bees and other pollinators.

This Week’s Review of the Morning Fallouts

Clear, cool nights have provided ideal flight conditions for nocturnal Neotropical migrants and other southbound birds throughout the week.  Fix yourself a drink and a little snack, then sit down and enjoy this set of photographs that includes just some of the species we found during sunrise feeding frenzies atop several of the Lower Susquehanna River Watershed’s ridges.  Hurry up, because here they come…

Black-throated Green Warbler
A Black-throated Green Warbler.
Black-throated Green Warbler
The Black-throated Green Warbler was perhaps the most frequently identified treetop warbler during the most recent four mornings.
Black-throated Green Warbler
A Black-throated Green Warbler with a unique variation in the crown plumage.
Blackburnian Warbler
The Blackburnian Warbler was another plentiful species.
Cape May Warblers
Cape May Warblers have an affinity for conifers like this Eastern White Pine.
Cape May Warbler
But when traveling in mixed flocks with other migrants, Cape May Warblers can also be found feeding in the crown foliage of deciduous trees.
Tennessee Warbler
This adult Tennessee Warbler appears to be adorned in a very worn set of plumage…
Tennessee Warbler
…and its traveling companion looks like it’s overdue for a new set of feathers as well.
Nashville Warbler
Like the Tennessee Warbler, the Nashville Warbler was common among mixed flocks.
Nashville Warbler
A Nashville Warbler atop a Black Cherry.
Chestnut-sided Warbler
This Chestnut-sided Warbler was one of several found among the more common species of migrants.
Chestnut-sided Warbler
A Chestnut-sided Warbler.
Chestnut-sided Warbler
We were lucky enough to spot this male Chestnut-sided Warbler sporting his namesake flank feathers.
 Black-and-white Warbler
A Black-and-white Warbler uses its nuthatch-like feeding behavior to search the tree bark for edible invertebrates.
Common Yellowthroat
To see the Common Yellowthroat, one must cease looking upward into the high canopy and instead give the aching neck a rest by peering into the low vegetation at the forest edge.
Magnolia Warbler
While checking the low growth, keep an eye open for other migrants among the shrubs and tangles. This Magnolia Warbler glows in the rays of a rising sun as it searches for a meal after a long night of travel.
House Wren
Here we found a perky little House Wren.
Eastern Wood-Pewee
Back in the middle and upper reaches of the trees, we find what has been by far the most numerous of the flycatchers seen during our visits to fallout sunrises.  Eastern Wood-Pewees are appearing in very good numbers and can be seen quarreling and battling for hunting perches from which they are ambushing flying insects.
 Eastern Wood-Pewee
An Eastern Wood-Pewee fiercely defending its hunting perch.
Eastern Wood-Pewee
An Eastern Wood-Pewee.
Least Flycatcher
The numbers of migrating Least Flycatchers and other members of the genus Empidonax may be reaching their seasonal peak this week.
Scarlet Tanagers
Scarlet Tanagers are currently a common find following nocturnal flights.
Scarlet Tanager
A Scarlet Tanager peers down from the top of a Red Maple.
Rose-breasted Grosbeak
Did you hear a loud squeak in the treetops?  It could be a southbound Rose-breasted Grosbeak stopping by for the day.
Rose-breasted Grosbeak
A Rose-breasted Grosbeak in a dead tree snag.
Swainson's Thrush
The Neotropical thrushes are beginning to move south now as well.  We found this newly arrived Swainson’s Thrush at Rocky Ridge County Park in York County during sunrise this morning.
Red-eyed Vireo
Not surprisingly, the Red-eyed Vireo is one of the most numerous of the migrants seen feeding in the deciduous canopy following a nocturnal flight event.  It’s not at all unusual to see dozens filing the trees around a ridgetop overlook or along a forest edge.  Be certain to check these congregations carefully, especially the groups of birds feeding in the lower branches of tall timber or in the tops of smaller trees.  This week we found…
Yellow-throated Vireo
…several hungry Yellow-throated Vireos arriving after nocturnal flights,…
Philadelphia Vireo
…and a Philadelphia Vireo (Vireo philadelphicus) at the hawkwatch at Rocky Ridge County Park in York County.
Red-breasted Nuthatch
Though not a Neotropical migrant, the easier-heard-than-seen Red-breasted Nuthatch is beginning to wander south into the lower Susquehanna region.  Most of these birds will eventually continue on to the pine forests of the southern United States for winter, but a few could remain to become seasonal visitors at feeding stations.
Ruby-throated Hummingbird
Ruby-throated Hummingbirds are on the move; their migration to the tropics is well underway and nearing its peak.  Ruby-throats are diurnal migrants that do a majority of their flying during the hours of daylight.  The notable exception: the nighttime portion of the long southbound flight some of the birds make to cross the Gulf of Mexico.
Red-headed Woodpecker
The Red-headed Woodpecker is another diurnal migrant.   This denizen of temperate climates is currently beginning to move to its wintering grounds, an area that extends from the latitudes of the lower Susquehanna south to the Gulf of Mexico and central Texas.
Juvenile Red-headed Woodpecker
A juvenile Red-headed Woodpecker during a brief pit stop.
Cedar Waxwings
The flights of roving bands of masked Cedar Waxwings continue.  Their numbers appear to be an improvement over those of 2023.
Broad-winged Hawk
At regional hawk-counting stations, observers are seeing more Broad-winged Hawks and other species beginning to move through.
"Kettling" Broad-winged Hawks
The frequency of Broad-winged Hawks passing the lookouts one at a time is giving way to the occurrence of larger and larger “kettling” groups that search out thermal updrafts to save energy while migrating.  By mid-September each of these “kettles” can include one hundred birds or more.  On the peak days, the daily Broad-winged Hawk totals can reach one thousand or more.
Broad-winged Hawk
A Broad-winged Hawk soaring to gain lift from a thermal updraft above a hawkwatch lookout.

The migration is by no means over; it has only just begun.  So plan to visit a local hawkwatch or other suitable ridgetop in coming weeks.  Arrive early (between 7 and 8 AM) to catch a glimpse of a nocturnal migrant fallout, then stay through the day to see the hundreds, maybe even thousands, of Broad-winged Hawks and other diurnal raptors that will pass by.  It’s an experience you won’t forget.

Broad-winged Hawk Gliding Away to the Southwest
A Broad-winged Hawk gliding away to the southwest.

Be certain to click the “Birds” tab at the top of this page for a photo guide to the species you’re likely to see passing south through the lower Susquehanna valley in coming months.  And don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab to find a hawk-counting station near you.

More birds are on the way.  Here’s a look at this evening’s liftoff of nocturnal migrants detected by National Weather Service Radar in State College, Pennsylvania.  (NOAA/National Weather Service Doppler Radar image)

Southbound Flights: On Their Way Both Night and Day

It’s hard to believe, but for almost two months now, sandpipers, plovers, and terns have been filtering south through the Lower Susquehanna River Watershed on their way to the Atlantic coastline as they complete the first leg of their long autumn migration—a journey that will take some species all the way to the far reaches of the South American continent for winter.

Caspian Terns and Osprey
Migrating Caspian Terns and an Osprey take a break on a lower Susquehanna valley gravel bar.

As August draws to a close, these early birds are being joined by widespread nocturnal flights of Neotropical migrants—those species, primarily songbirds, on their way to wintering grounds which lie exclusively south of the continental United States.

To catch a glimpse of these night-flying avians, your best bet may be to position yourself on the crest of a ridge or along a linear break in the forest such as a utility right-of-way where waves of warblers, vireos, flycatchers, and other Neotropical passerines sometimes feed on invertebrates after making landfall at daybreak.  Pick a place where the trees are bathed in the warm light of the rising sun and be there by 7 A.M. E.D.T.  The activity can be tremendous, but it usually ends between 8 and 9.

Red-eyed Vireo
One of more than half a dozen Red-eyed Vireos seen during a daybreak fallout at Second Mountain Hawkwatch in Lebanon County earlier this week.
Blackburnian Warbler
During sunrise, a just-arrived Blackburnian Warbler checks the foliage of a ridgetop Red Maple for insects.
Least Flycatcher
A Least Flycatcher quietly searches the shrubby growth along the forest edge for a morning meal.
Black-and-white Warbler
A Black-and-white Warbler in the first hour of sunlight after a big nocturnal flight.
Scarlet Tanager
A Scarlet Tanager having a look around.  This individual and other migrating birds may be here for a few days before moving on.
Black-throated Green Warbler
A Black-throated Green Warbler in a treetop glowing with the light of sunrise.
Chestnut-sided Warbler
A hungry Chestnut-sided Warbler inspects limbs, leaves, and twigs looking for nourishment after an all-night flight.

Diurnal migrants, birds that make their movements during the daylight hours, are ramping up their flights now as well.  Broad-winged Hawks, Bald Eagles, and falcons are currently being tallied at hawk-counting stations throughout the northeast.  Many of those lookouts are seeing Ruby-throated Hummingbirds, swallows, and other daytime migrants too.

Broad-winged Hawk
While many diurnal migrants gather in flocks as a method of defense against predators, Broad-winged Hawks congregate as a reconnaissance measure.  Not waiting around for strong autumn winds to be deflected upwards by the region’s numerous ridges, these Neotropical birds migrate early enough in the season to rely upon thermal updrafts from sun-heated surfaces to provide lift, gain altitude, and save energy during their long trip.  By traveling in groups, there are collectively able to better locate and utilize rising air columns as they progress southwesterly along their route.  Broad-winged Hawks travel to Central and South America for winter.  Their numbers in the Lower Susquehanna River Watershed will peak during the third week of September.
American Kestrel
Many American Kestrels transit our area in August and early September, a time coinciding with flights of migratory dragonflies, insects upon which they and another falcon, the Merlin, frequently prey.
Chimney Swifts
Chimney Swifts are fast-flying diurnal migrants.  They are beginning to congregate into larger groups in preparation for their departure.  Some are already on the move.
Cedar Waxwing
Cedar Waxwings are currently being seen in roving flocks throughout the region.  Their migration is less of a point-to-point flight and more of a continuous wandering in search of berries and other wild foods.
American Goldfinch
Not everyone is yet ready to go.  American Goldfinches are still in the midst of their nesting cycle and won’t begin leaving until the young are on their own and colorful adult males like this one are beginning to molt into their drab winter plumage.

Autumn migration flights are an ever-changing process, with different species peaking at different times throughout the season.  In these months just after the nesting season, each of these species is more numerous than at any other time of the year.  And of course, the more often we as observers get out and have a look, the more of them we’ll see.

Be certain to click the “Birds” tab at the top of this page for a photo guide to the species you’re likely to see passing through the lower Susquehanna valley this fall.  Nearly four months of autumn hawk migration flights lie ahead, so don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab to find a hawk-counting station near you, then stop by for a visit or two.  See you there!

A large nightfall liftoff of nocturnal migrating birds is shown here on this evening’s State College, Pennsylvania, Doppler Radar base velocity loop.  Returns indicating birds moving toward the radar site (the white dot in the center of the color mass) are shown in green, and those indicating birds moving away are indicated in red.  This is a big flight headed generally in a south-southwest direction.  (NOAA/National Weather Service Doppler Radar image)

Plantings for Wet Lowlands

This linear grove of mature trees, many of them nearly one hundred years old, is a planting of native White Oaks (Quercus alba) and Swamp White Oaks (Quercus bicolor).

Imagine the benefit of trees like this along that section of stream you’re mowing or grazing right now.  The Swamp White Oak in particular thrives in wet soils and is available now for just a couple of bucks per tree from several of the lower Susquehanna’s County Conservation District Tree Sales.  These and other trees and shrubs planted along creeks and rivers to create a riparian buffer help reduce sediment and nutrient pollution.  In addition, these vegetated borders protect against soil erosion, they provide shade to otherwise sun-scorched waters, and they provide essential wildlife habitat.  What’s not to love?

Swamp White Oak
Autumn leaf of a Swamp White Oak

The following native species make great companions for Swamp White Oaks in a lowland setting and are available at bargain prices from one or more of the County Conservation District Tree Sales now underway…

Red Maple
The Red Maple is an ideal tree for a stream buffer project. They do so well that you should limit them to 10% or less of the plants in your project so that they don’t overwhelm slower-growing species.
River Birch
The River Birch (Betula nigra) is a multi-trunked tree of lowlands.  Large specimens with arching trunks help shade waterways and provide a source of falling insects for surface-feeding fish.  Its peeling bark is a distinctive feature.
Common Winterberry
The Common Winterberry with its showy red winter-time fruit is a slow-growing shrub of wet soils.  Only female specimens of this deciduous holly produce berries, so you need to plant a bunch to make sure you have both genders for successful pollination.
American Robins feeding on Common Winterberry.
An American Robin feeding on Common Winterberry.
Common Spicebush
Common Spicebush is a shrub of moist lowland soils.  It is the host plant for the Spicebush Swallowtail butterfly and produces small red berries for birds and other wildlife.  Plant it widely among taller trees to provide native vegetation in the understory of your forest.
Common Spicebush foliage and berries.
Common Spicebush foliage and berries in the shade beneath a canopy of tall trees.
Common Pawpaw
The Common Pawpaw a small shade-loving tree of the forest understory.
Common Pawpaw
Common Pawpaw is a colony-forming small tree which produces a fleshy fruit.  It is the host plant for the caterpillars of the Zebra Swallowtail.
Buttonbush
The Buttonbush is a shrub of wet soils.  It produces a round flower cluster, followed by this globular seed cluster.
Eastern Sycamore
And don’t forget the Eastern Sycamore, the giant of the lowlands.  At maturity, the white-and-tan-colored bark on massive specimens makes them a spectacular sight along stream courses and river shores.  Birds ranging from owls, eagles, and herons to smaller species including the Yellow-throated Warbler rely upon them for nesting sites.
Yellow-crowned Night Herons Nesting in an Eastern Sycamore
Yellow-crowned Night Herons, an endangered species in Pennsylvania, nesting in an Eastern Sycamore.

So don’t mow, do something positive and plant a buffer!

Act now to order your plants because deadlines are approaching fast.  For links to the County Conservation District Tree Sales in the Lower Susquehanna River Watershed, see our February 18th post.

Red Maples

Red Maple
Many cultivars of ornamental maples for the home garden are derived from this native species, and it’s easy to see why.  Despite the variation in the foliage of each, every tree in this view is a Red Maple (Acer rubrum).
Red Maple Leaves
The Red Maple is the most common and widespread tree in all of eastern North America.  It is easily recognized by its three-lobed leaves.  Not only do the fall foliage colors vary among a population of trees, but some individual specimens have a well-stocked palette of their own.  This collection came from a single Red Maple.  Visit a forested area today and check them out!