Back on March 24th, we took a detailed look at the process involved in administering prescribed fire as a tool for managing grassland and early successional habitat. Today we’re going turn back the hands of time to give you a glimpse of how the treated site fared during the five months since the controlled burn. Let’s go back to Middle Creek Wildlife Management Area for a photo tour to see how things have come along…
Pennsylvania Game Commission crews administering prescribed fire on a grassland demonstration site back on March 16.By late May, native herbaceous perennial wildflowers including Joe-pye Weed had re-greened the site. One of the goals of the burn was to kill fire-sensitive woody plants, thus preventing the process of succession from reforesting the site.The scorched, lifeless remains of small trees and shrubs indicate that that goal was met.Because prescribed fire is administered in a mosaic pattern that permits some early successional growth to remain until the next burn, birds including this thicket-nesting Yellow-breasted Chat are able to take advantage of the mixed habitat during their breeding season in May and June.By August, the site is a haven for native plants and animals.The burn has promoted the growth and late-summer bloom of fire-tolerant native wildflowers and warm-season grasses……including Indiangrass,……Big Bluestem……Thin-leaved Coneflower (Rudbeckia triloba), a plant also known as Brown-eyed Susan,……and Joe-pye Weed, a plant butterflies find irresistible.Eastern Tiger Swallowtail collecting nectar from Joe-pye Weed.A black-morph Eastern Tiger Swallowtail collecting nectar from Spotted Knapweed (Centaurea steobe micranthos), a non-native invasive plant found growing in an area of the burn site missed by this year’s fire. While many non-native plants are unable to survive the flames and heat produced by prescribed fire, it isn’t an absolute cure-all. It doesn’t eliminate all invasive plants, it just keeps them from dominating a landscape by out-competing native species. Left unmanaged, Spotted Knapweed is a tough perennial invasive that can easily become one the species able to overtake a vulnerable grassland. It can be a stubborn survivor of some prescribed burns. On the plus side, butterflies really like it.By August, native grassland plants in the prescribed fire area were already providing an abundance of seeds for birds including this American Goldfinch.For larger birds like turkeys and pheasants, an abundance of Carolina Grasshoppers are providing a protein-dense food source in managed grasslands.And tiny flying insects, a nuisance to us as we take a stroll alongside the grasslands, are a meal taken on the wing by dragonflies including this Black Saddlebags.
Elsewhere around the refuge at Middle Creek, prescribed fire and other management techniques are providing high-quality grassland habitat for numerous species of nesting birds…
Bobolinks nested both in areas subjected to controlled burns……and in hay fields where mowing was delayed until the nesting season, including the fledging process, was completed earlier this month.As advertised, Grasshopper Sparrows nested in these fields as well.
We hope you enjoyed this short photo tour of grassland management practices. Now, we’d like to leave you with one last set of pictures—a set you may find as interesting as we found them. Each is of a different Eastern Cottontail, a species we found to be particularly common on prescribed fire sites when we took these images in late May. The first two are of the individuals we happened to be able to photograph in areas subjected to fire two months earlier in March. The latter two are of cottontails we happened to photograph elsewhere on the refuge in areas not in proximity to ground treated with a prescribed burn or exposed to accidental fire in recent years.
Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.
These first two rabbits are living the good life in a warm-season grass wonderland.
Eastern Cottontail at a site not subjected to any recent fire activity.Eastern Cottontail at a site not subjected to any recent fire activity.
Oh Deer! Oh Deer! These last two rabbits have no clock to track the time; they have only ticks. Better not go for a stroll with them Alice—that’s no wonderland! I know, I know, it’s time to go. See ya later.
Homo sapiens owes much of its success as a species to an acquired knowledge of how to make, control, and utilize fire. Using fire to convert the energy stored in combustible materials into light and heat has enabled humankind to expand its range throughout the globe. Indeed, humans in their furless incomplete mammalian state may have never been able to expand their populations outside of tropical latitudes without mastery of fire. It is fire that has enabled man to exploit more of the earth’s resources than any other species. From cooking otherwise unpalatable foods to powering the modern industrial society, fire has set man apart from the rest of the natural world.
In our modern civilizations, we generally look at the unplanned outbreak of fire as a catastrophe requiring our immediate intercession. A building fire, for example, is extinguished as quickly as possible to save lives and property. And fires detected in fields, brush, and woodlands are promptly controlled to prevent their exponential growth. But has fire gone to our heads? Do we have an anthropocentric view of fire? Aren’t there naturally occurring fires that are essential to the health of some of the world’s ecosystems? And to our own safety? Indeed there are. And many species and the ecosystems they inhabit rely on the periodic occurrence of fire to maintain their health and vigor.
For the war effort- The campaign to reduce the frequency of forest fires got its start during World War II with distribution of this poster in 1942. The goal was to protect the nation’s timber resources from accidental or malicious loss due to fire caused by man-made ignition sources. The release of the Walt Disney film “Bambi” during the same year and the adoption of the Smokey the Bear mascot in 1944 softened the message’s delivery, but the public relations outreach continued to be a key element of a no-fire policy to save trees for lumber. Protection and management of healthy forest ecosystems in their entirety has only recently become a priority. (National Archives image)
Man has been availed of the direct benefits of fire for possibly 40,000 years or more. Here in the Lower Susquehanna River Watershed, the earliest humans arrived as early as 12,000 years ago—already possessing skills for using fire. Native plants and animals on the other hand, have been part of the ever-changing mix of ecosystems found here for a much longer period of time—millions to tens of millions of years. Many terrestrial native species are adapted to the periodic occurrence of fire. Some, in fact, require it. Most upland ecosystems need an occasional dose of fire, usually ignited by lightning (though volcanism and incoming cosmic projectiles are rare possibilities), to regenerate vegetation, release nutrients, and maintain certain non-climax habitat types.
But much of our region has been deprived of natural-type fires since the time of the clearcutting of the virgin forests during the eighteenth and nineteenth centuries. This absence of a natural fire cycle has contributed to degradation and/or elimination of many forest and non-forest habitats. Without fire, a dangerous stockpile of combustible debris has been collecting, season after season, in some areas for a hundred years or more. Lacking periodic fires or sufficient moisture to sustain prompt decomposition of dead material, wildlands can accumulate enough leaf litter, thatch, dry brush, tinder, and fallen wood to fuel monumentally large forest fires—fires similar to those recently engulfing some areas of the American west. So elimination of natural fire isn’t just a problem for native plants and animals, its a potential problem for humans as well.
Indiangrass (seen here), Switchgrass, Big Bluestem, and Little Bluestem are native species requiring periodic forms of disturbance to eliminate competition by woody plants. These warm-season grasses develop roots that penetrate deep into the soil, sometimes to depths of six feet or more, allowing them to survive severe drought and flash fire events. In the tall grass prairies, these extensive root systems allow these grasses to return following heavy grazing by roaming herds of American Bison (Bison bison). Without these habitat disturbances, warm season grasslands succumb to succession in about seven years. With their periodic occurrence, the plants thrive and provide excellent wildlife habitat, erosion control, and grazing forage.
To address the habitat ailments caused by a lack of natural fires, federal, state, and local conservation agencies are adopting the practice of “prescribed fire” as a treatment to restore ecosystem health. A prescribed fire is a controlled burn specifically planned to correct one or more vegetative management problems on a given parcel of land. In the Lower Susquehanna River Watershed, prescribed fire is used to…
Eliminate dangerous accumulations of combustible fuels in woodlands.
Reduce accumulations of dead plant material that may harbor disease.
Provide top kill to promote oak regeneration.
Regenerate other targeted species of trees, wildflowers, grasses, and vegetation.
Kill non-native plants and promote growth of native plants.
Prevent succession.
Remove woody growth and thatch from grasslands.
Promote fire tolerant species of plants and animals.
Improve habitat for rare species (Regal Fritillary, etc.)
Recycle nutrients and minerals contained in dead plant material.
Let’s look at some examples of prescribed fire being implemented right here in our own neighborhood…
Prescribed fires are typically planned for the dormant season extending from late fall into early spring with burns best conducted on days when the relative humidity is low.Prescribed fire is used regularly at Fort Indiantown Gap Military Reservation in Lebanon County, Pennsylvania, to keep accumulations of woody and herbaceous fuels from accumulating on and around the training range areas where live ordinance and other sources of ignition could otherwise spark large, hard-to-control wildfires.Prescribed fires replace the periodic natural burns that would normally reduce the fuel load in forested areas. Where these fuels are allowed to accumulate, south-facing slopes are particularly susceptible to extreme fires due to their exposure to the drying effects of intense sunlight for much of the year. The majority of small oaks subjected to treatment by the prescribed fire shown here will have the chance to regenerate without immediate competition from other species including invasive plants. The larger trees are mostly unaffected by the quick exposure to the flames. Note too that these fires don’t completely burn everything on the forest floor, they burn that which is most combustible. There are still plenty of fallen logs for salamanders, skinks, and other animals to live beneath and within.
A prescribed fire in late winter prevents this grassland consisting of Big Bluestem and native wildflowers from being overtaken by woody growth and invasive species. Fires such as this that are intended to interrupt the process of succession are repeated at least every three to five years.In its wildlife food plots, prescribed fire is used by the Pennsylvania Game Commission to prevent succession and control invasive species such as Multiflora Rose, instead promoting the growth of native plants.An example of a woodlot understory choked with combustible fuels and dense tangles of invasive Multiflora Rose. A forester has the option of prescribing a dose of dormant-season fire for a site like this to reduce the fuel load, top kill non-native vegetation, and regenerate native plants.A dose of prescribed fire was administered on this grassland to kill the woody growth of small trees beginning to overtake the habitat by succession.The Pennsylvania Game Commission employs prescribed fire at Middle Creek Wildlife Management Area and on many of their other holdings to maintain grasslands.Prescribed fire is used to eliminate invasive species including Multiflora Rose from grasslands at Middle Creek W.M.A. Annual burns on the property are conducted in a mosaic pattern so that each individual area of the grassland is exposed to the effects of fire only once every two to five years. Without fire or some type of mechanical or chemical intervention, succession by woody trees and shrubs would take hold after about seven years.Prescribed fire is planned for a fraction of total grassland acreage at Middle Creek W.M.A. each year. Another section of the mosaic is targeted in the following year and yet another in the year that follows that. Because burns are conducted in the spring, grassland cover is available for wildlife throughout the winter. And because each year’s fire burns only a portion of the total grassland acreage, wildlife still has plenty of standing grass in which to take shelter during and after the prescribed fire.Prescribed fire at Middle Creek W.M.A. provides grassland habitat for dozens of species of birds and mammals including the not-so-common Grasshopper Sparrow……and stocked Ring-necked Pheasants that do nest and raise young there.On a few sites in the Lower Susquehanna River Watershed , prescribed fire is being used to establish and maintain savanna-like grasslands. This one, located on a dry, south-facing slope near numerous man-made sources of ignition, can easily be dosed with periodic prescribed burns to both prevent succession and reduce fuel accumulations that may lead to a devastating extreme fire.One year following a prescribed burn, this is the autumn appearance of a savanna-like habitat with fire-tolerant Pitch Pine (Pinus rigida), Bear Oak, warm-season grasses, and a variety of nectar-producing wildflowers for pollinators. These ecosystems are magnets for wildlife and may prove to be a manageable fit on sun-drenched sites adjacent to man-made land disturbances and their sources of ignition.Savanna-like grasslands with oaks and other scattered large trees, some of them dead, make attractive nesting habitat for the uncommon Red-headed Woodpecker.Prescribed fire can benefit hungry Wild Turkeys by maintaining savanna-like grasslands for an abundance of grasshoppers and other insects in summer and improving the success of mast-producing oaks for winter.In the Lower Susquehanna River Watershed, the caterpillar of the rare Eastern Buck Moth feeds on the foliage of the Bear Oak, also known as the Scrub Oak, a shrubby species that relies upon periodic fire to eliminate competition from larger trees in its early successional habitat.Leaves of the Bear Oak in fall. The Bear Oak regenerates readily from top kill caused by fire.Reed Canary Grass (Phalaris arundinacea) is a native cool-season grass with a colorful inflorescence in spring. But given the right situation, it can aggressively overtake other species to create a pure stand lacking biodiversity. It is one of the few native species which is sometimes labelled “invasive”.Prescribed fire can be used to reduce an overabundance of Reed Canary Grass and its thatch in wetlands. Periodic burning can help restore species diversity in these habitats for plants and animals including rare species such as the endangered Bog Turtle (Glyptemys muhlenbergii).On the range areas at Fort Indiantown Gap in Lebanon County, Pennsylvania, disturbances by armored vehicles mimic the effects of large mammals such as the American Bison which periodically trampled grasses to prevent succession and the establishment of woody plants on its prairie habitat. To supplement the activity of the heavy vehicles and to provide suitable habitat for the very rare Regal Fritillary (Speyeria idalia) butterflies found there, prescribed fire is periodically employed to maintain the grasslands on the range. These burns are planned to encourage the growth of “Fort Indiantown Gap Little Bluestem” grass as well as the violets used as host plants by the Regal Fritillary caterpillars. These fires also promote growth of a variety of native summer-blooming wildflowers to provide nectar for the adults butterflies.A last record of a wild American Bison killed in Pennsylvania was an animal taken in the Susquehanna watershed in Union County in 1801. The species is thereafter considered extirpated from the state. Since that time, natural disturbances needed to regenerate warm-season grasses have been limited primarily to fires and riverine ice scour. The waning occurrence of both has reduced the range of these grasses and their prairie-like ecosystems in the commonwealth. (Exhibit: State Museum of Pennsylvania, Harrisburg)A male Regal Fritillary on the range at Fort Indiantown Gap, where armored vehicles and prescribed fire provide suitable prairie-like habitat for this vulnerable species.Prescribed fires return the nutrients and minerals contained in dead plant material to the soil. Following these controlled burns, insects like this Honey Bee can often be seen collecting minerals from the ashes.A Greenbottle Fly gathering minerals from the ash following a prescribed burn.
In Pennsylvania, state law provides landowners and crews conducting prescribed fire burns with reduced legal liability when the latter meet certain educational, planning, and operational requirements. This law may help encourage more widespread application of prescribed fire in the state’s forests and other ecosystems where essential periodic fire has been absent for so very long. Currently in the Lower Susquehanna River Watershed, prescribed fire is most frequently being employed by state agencies on state lands—in particular, the Department of Conservation and Natural Resources on State Forests and the Pennsylvania Game Commission on State Game Lands. Prescribed fire is also part of the vegetation management plan at Fort Indiantown Gap Military Reservation and on the land holdings of the Hershey Trust. Visitors to the nearby Gettysburg National Military Park will also notice prescribed fire being used to maintain the grassland restorations there.
For crews administering prescribed fire burns, late March and early April are a busy time. The relative humidity is often at its lowest level of the year, so the probability of ignition of previous years’ growth is generally at its best. We visited with a crew administering a prescribed fire at Middle Creek Wildlife Management Area last week. Have a look…
Members of a Pennsylvania Game Commission burn crew provide visitors to Middle Creek Wildlife Management Area with an overview of prescribed fire and the equipment and techniques they use to conduct a burn.Pennsylvania Game Commission Southeast Region Forester Andy Weaver will fulfill the role of Burn Boss for administering this day’s dose of fire. His responsibilities include assessing the weather before the burn and calculating a probability of ignition.The Burn Boss briefs personnel with information on site layout, water supply location(s), places of refuge, emergency procedures, the event’s goals and plan of action, crew assignments, and the results of the weather check: wind from the northwest at 5 miles per hour, temperature 48 degrees, and the relative humidity 63%. Today’s patient is a parcel of warm-season grasses receiving a dose of fire to eliminate invasive non-native plants, woody growth, and thatch. The probability of ignition is 20%, but improving by the minute.To begin the burn, a test fire is started in the downwind corner of the parcel, which also happens to be the bottom of the slope. Fuel ignition is good. The burn can proceed.Crews proceed uphill from the location of the test fire while igniting combustibles along both flanks of the area being treated.A drip torch is used to ignite the dried stems and leaves of warm-season grasses and wildflowers. Each member of the burn crew wears Nomex fire-resistant clothing and carries safety equipment including a two-way radio, a hydration pack, and a cocoon-like emergency fire shelter.An all-terrain vehicle equipped with various tools, a fire pump, hose, and a small water tank accompanies the crew on each flank of the fire.A mowed strip of cool-season grasses along the perimeter of the burn area is already green and functions as an ideal fire break. While the drip torch is perfect for lighting combustibles along the fire’s perimeter, the paintball gun-looking device is an effective tool used to lob incendiaries into the center areas of the burn zone for ignition.With green cool-season grasses already growing on the trails surrounding the burn zone, very little water was used to contain this prescribed fire. Where such convenient fire breaks don’t already exist, crews carry tools including chain saws, shovels, and leaf blowers to create their own. They also carry flame swatters, backpack water pumps, shovels, and other tools to extinguish fires if necessary. None of these items were needed to control this particular fire.This fast-burning fire provides enough heat to damage the cambium layer of the woody tree and shrub saplings in this parcel being maintained as a grassland/wildflower plot, thus the process of succession is forestalled. Burns conducted during previous years on this and adjacent fields have also controlled aggressive growth of invasive Multiflora Rose and Olives (Elaeagnus species).Crews proceed up the slope while maintaining the perimeter by igniting dry plant material along the flanks of the burn zone.Ignition complete, the crews monitor the fire.The Burn Boss surveys the final stages of a safe and successful prescribed fire. The fire has left behind a mosaic of burned and unburned areas, just as a naturally occurring event may have done. Wildlife dodging the flames may be taking refuge in the standing grasses, so there is no remedial attempt to go back and ignite these areas. They’ll be burned during prescribed fires in coming years.By June, this grassland will again be lush and green with warm-season grasses and blooming wildflowers like this Common Milkweed being visited by a Great Spangled Fritillary.And later in the summer, Eastern Tiger Swallowtails on Joe-pye Weed.Indiangrass in flower in mid-summer.Bobolinks glow in the late August sun while taking flight from a stand of warm-season grasses maintained using springtime prescribed fire. The small dots on the dark background at the top of the image are multitudes of flying insects, many of them pollinators. The vegetation is predominately Indiangrass, excellent winter cover for birds, mammals, and other wildlife.
Prescribed burns aren’t a cure-all for what ails a troubled forest or other ecosystem, but they can be an effective remedy for deficiencies caused by a lack of periodic episodes of naturally occurring fire. They are an important option for modern foresters, wildlife managers, and other conservationists.
We’ve got the summertime blues for you, right here at susquehannawildlife.net…
In warm-season grass meadows, Big Bluestem is now in flower. This and other species of native prairie grasses provide excellent habitat for birds, mammals, and insects including butterflies. To survive drought and fire, their roots run much deeper than cool season grasses, creeping down four to six feet or more. This adaptation allowed warm season grasses to recover from heavy grazing by large Pre-Anthropocene mammals. Today, it makes them ideal plants for soil stabilization.A male Indigo Bunting has already found ripe seeds among the heads of flowering Big Bluestem.Look closely and you’ll see our Indigo Bunting is beginning a pre-migration molt out of its bright-blue breeding (alternate) plumage and into a gray-brown winter (basic) plumage. The berries of the American Pokeweed upon which it is perched will soon ripen into a dark blue, almost black, color. Though toxic to humans, these fruits find favor with many species of birds and mammals.Another great wildlife food is Silky Dogwood (Cornus amomum), a deciduous shrub that sports blue-colored berries in summer and showy, bright-red twigs in winter. It grows well in wet ground along streams and ponds, as well as in rain gardens.A Great Blue Heron searches the shallows for small fish. This species is also a good mouser, at times seen hunting in grassy meadows. Right now is prime time to see it and a variety of other herons and egrets throughout the Lower Susquehanna River Watershed.
…so don’t let the summertime blues get you down. Grab a pair of binoculars and/or a camera and go for a stroll!
This tallgrass prairie wildflower planting on a health campus in Hershey, Pennsylvania, enhances stormwater management and benefits butterflies and other wildlife. Reducing the acreage maintained as manicured lawn has helped disperse the large flocks of resident Canada Geese (a population of invasive native transplants) that frequented the property and posed a serious hazard to medevac helicopters flying in and out of the facility.Big Bluestem, a warm-season grass, dominates the site and is complemented by Indiangrass and tall wildflowers including Common Milkweed, Wild Bergamot, Oxeye, Black-eyed Susan, Prairie Coneflower, and Purple Coneflower.
It’s surprising how many millions of people travel the busy coastal routes of Delaware each year to leave the traffic congestion and hectic life of the northeast corridor behind to visit congested hectic shore towns like Rehobeth Beach, Bethany Beach, and Ocean City, Maryland. They call it a vacation, or a holiday, or a weekend, and it’s exhausting. What’s amazing is how many of them drive right by a breathtaking national treasure located along Delaware Bay just east of the city of Dover—and never know it. A short detour on your route will take you there. It’s Bombay Hook National Wildlife Refuge, a quiet but spectacular place that draws few crowds of tourists, but lots of birds and other wildlife.
Bombay Hook National Wildlife Refuge is located just off Route 9, a lightly-traveled coastal road east of Dover, Delaware. Note the Big Bluestem and other warm season grasses in the background. Bombay Hook, like other refuges in the system, is managed for the benefit of the wildlife that relies upon it to survive. Within recent years, most of the mowed grass and tilled ground that once occurred here has been replaced by prairie grasses or successional growth, much to the delight of Northern Bobwhite (Colinus virginianus) and other species.
Let’s join Uncle Tyler Dyer and have a look around Bombay Hook. He’s got his duck stamp and he’s ready to go.
Uncle Ty’s current United States Fish and Wildlife Service Duck Stamp displayed on his dashboard is free admission to the tour road at Bombay Hook and other National Wildlife Refuges.The refuge at Bombay Hook includes woodlands, grasslands, and man-made freshwater pools, but it is predominately a protectorate of thousands of acres of tidal salt marsh bordering and purifying the waters of Delaware Bay. These marshes are renowned wintering areas for an Atlantic population of Snow Goose known as the “Greater Snow Goose” (Anser caerulescens atlanticus). Witnessing thousands of these birds rising over the marsh and glowing in the amber light of a setting sun is an unforgettable experience.Trails at various stops along the auto tour route lead to observation towers and other features. This boardwalk meanders into the salt marsh grasses and includes a viewing area alongside a tidal creek. Our visit coincided with a very high tide induced by east winds and a new moon.During high tide, an Eastern Cottontail (Sylvilagus floridanus) seeks higher ground near the boardwalk and the wooded edge of the salt marsh.As the tide rises, fast-flying shorebirds scramble from flooded mudflats in the salt marsh on the east side of the tour road.When high tide arrives in the salt marshes, shorebirds and waterfowl often concentrate in the man-made freshwater pools on the west side of the tour road. Glaring afternoon sun is not the best for viewing birds located west of the road. For ideal light conditions, time your visit for a day when high tide occurs in the morning and recedes to low tide in the afternoon.A view looking west into Shearness Pool, largest of the freshwater impoundments at Bombay Hook.Bombay Hook has many secretive birds hiding in its wetlands, but they can often be located by the patient observer. Here, two Pied-billed Grebes feed in an opening among the vegetation in a freshwater pool.One of Bombay Hook’s resident Bald Eagles patrols the wetlands.American Avocets (Recurvirostra americana) gather by the hundreds at Bombay Hook during the fall. A passing eagle will stir them into flight.An American Avocet, a delicate wader with a peculiar upturned bill.As soon as the tide begins receding, shorebirds and waterfowl like these Green-winged Teal begin dispersing into the salt marshes to feed on the exposed mudflats.The woodlands and forested areas of the refuge host resident songbirds and can be attractive to migrating species like this Yellow-rumped Warbler.For much of its course, the tour road at Bombay Hook is located atop the dike that creates the man-made freshwater pools on the western edge of the tidal salt marsh. If you drive slowly and make frequent stops to look and listen, you’ll notice an abundance of birds and other wildlife living along this border between two habitats. Here, a Swamp Sparrow has a look around.Savannah Sparrows are common along the tour road where native grasses grow wild.Bombay Hook is renowned for its rarities. One of the attractions during the late summer and autumn of 2021 was a group of Roseate Spoonbills (Platalea ajaja), vagrants from the southern states, seen here with Great Egrets and Snowy Egrets (Egretta thula).Roseate Spoonbills and Great Egrets at Bombay Hook.
Remember to go the Post Office and get your duck stamp. You’ll be supporting habitat acquisition and improvements for the wildlife we cherish. And if you get the chance, visit a National Wildlife Refuge. November can be a great time to go, it’s bug-free! Just take along your warmest clothing and plan to spend the day. You won’t regret it.
The southbound bird migration of 2020 is well underway. With passage of a cold front coming within the next 48 hours, the days ahead should provide an abundance of viewing opportunities.
Here are some of the species moving through the lower Susquehanna valley right now.
Blue-winged Teal are among the earliest of the waterfowl to begin southward migration.Sandpipers and plovers have been on the move since July. The bird in the foreground with these Killdeer is not one of their offspring, but rather a Semipalmated Plover (Charadrius semipalmatus), a regular late-summer migrant in the Lower Susquehanna River Watershed.Hawk watch sites all over North America are counting birds right now. The Osprey is an early-season delight as it glides past the lookouts. Look for them moving down the Susquehanna as well.Bald Eagles will be on the move through December. To see these huge raptors in numbers, visit a hawk watch on a day following passage of a cold front when northwest winds are gusting.Merlins were seen during this past week in areas with good concentrations of dragonflies. This particular one at Middle Creek Wildlife Management Area in Lancaster and Lebanon Counties……was soon visited by another.Check the forest canopy for Yellow-billed Cuckoos. Some local birds are still on breeding territories while others from farther north are beginning to move through.Ruby-throated Hummingbirds are darting through the lower Susquehanna valley on their way to the tropics. This one has no trouble keeping pace with a passing Tree Swallow.Nocturnal flights can bring new songbirds to good habitat each morning. It’s the best time of year to see numbers of Empidonax flycatchers. But, because they’re often silent during fall migration, it’s not the best time of year to easily identify them. This one lacking a prominent eye ring is a Willow Flycatcher (Empidonax traillii).During the past two weeks, Red-eyed Vireos have been numerous in many Susquehanna valley woodlands. Many are migrants while others are breeding pairs tending late-season broods.During mornings that follow heavy overnight flights, Blackburnian Warblers have been common among waves of feeding songbirds.Chestnut-sided Warblers are regular among flocks of nocturnal migrants seen foraging among foliage at sunrise.Scarlet Tanagers, minus the brilliant red breeding plumage of the males, are on their way back to the tropics for winter.While passing overhead on their way south, Bobolinks can be seen or heard from almost anywhere in the lower Susquehanna valley. Their movements peak in late August and early September.During recent evenings, Bobolinks have been gathering by the hundreds in fields of warm-season grasses at Middle Creek Wildlife Management Area.If you go to see the Bobolinks there, visit Stop 3 on the tour route late in the afternoon and listen for their call. You’ll soon notice their wings glistening in the light of the setting sun as they take short flights from point to point while they feed. Note the abundance of flying insects above the Big Bluestem and Indiangrass in this image. Grasslands like these are essential habitats for many of our least common resident and migratory birds.
It’s sprayed with herbicides. It’s mowed and mangled. It’s ground to shreds with noisy weed-trimmers. It’s scorned and maligned. It’s been targeted for elimination by some governments because it’s undesirable and “noxious”. And it has that four letter word in its name which dooms the fate of any plant that possesses it. It’s the Common Milkweed, and it’s the center of activity in our garden at this time of year. Yep, we said milk-WEED.
Now, you need to understand that our garden is small—less than 2,500 square feet. There is no lawn, and there will be no lawn. We’ll have nothing to do with the lawn nonsense. Those of you who know us, know that the lawn, or anything that looks like lawn, are through.
Anyway, most of the plants in the garden are native species. There are trees, numerous shrubs, some water features with aquatic plants, and filling the sunny margins is a mix of native grassland plants including Common Milkweed. The unusually wet growing season in 2018 has been very kind to these plants. They are still very green and lush. And the animals that rely on them are having a banner year. Have a look…
The flowers of the Common Milkweed were exceptionally fragrant this year. At their peak in early June, their hyacinth-or lilac-like aroma was so prevalent, it drifted into our building and overwhelmed the stink of the neighbor’s filthy dumpster that he had placed 12 feet away from our walls (100 feet from his).Common Milkweed attracts a pollinating Greenbottle Fly (Lucilia species). The dumpster attracts the invasive House Fly (Musca domestica), carrier of dysentery, typhoid, and other wonderful diseases. Are you following this? Remember as we proceed, milkweed is “noxious”.Busy Eastern Carpenter Bees (Xylocopa virginica) load up with pollen from the flowers of the Common Milkweed.A Red Milkweed Beetle (Tetraopes tetrophthalmus) munches on a tender fresh Common Milkweed leaf in mid-June.Following the pollination of the flowers, seed pods will begin to grow. We trim these off the plants. The removal of the extra weight allows most of the stems to remain erect through stormy weather. You’ll still get new plants from underground runners. As you may have guessed, we’re trying to keep these plants upright and strong to host Monarch butterfly larvae.
We’ve planted a variety of native grassland species to help support the milkweed structurally and to provide a more complete habitat for Monarch butterflies and other native insects. This year, these plants are exceptionally colorful for late-August due to the abundance of rain. The warm season grasses shown below are the four primary species found in the American tall-grass prairies and elsewhere.
Big Bluestem, a native warm-season grass in flower.Little Bluestem (Schizachyrium scoparium “Fort Indiantown Gap”) in flower. This variety grows on the tank range at the military base where the armored vehicles and prescribed burns substitute for the herd animals and fires of the prairie to prevent succession and allow it to thrive.Partridge Pea can tolerate sandy soil and is a host plant for vagrant Cloudless Sulphur butterflies.Black-eyed Susan (Rudbeckia hirta) is a popular native grassland wildflower.Indiangrass (Sorghastrum nutans) in flower. This and the other native plants shown here are available as seed from Ernst Seed Company in Meadville (PA). They have an unbelievably large selection of indigenous species. You can plant a small plot or acres and acres using really good mixes blended for purposes ranging from reclaiming pipeline right-of-ways and strip mines to naturalizing backyard gardens.A Red Admiral (Vanessa atalanta) butterfly, a migratory species like the Monarch, on Purple Coneflower (Echinacea purpurea). Yes, it is that Echinacea, the same one used as a supplement and home remedy.
There was Monarch activity in the garden today like we’ve never seen before—and it revolved around milkweed and the companion plants.
A female Monarch laying eggs on a Common Milkweed leaf.A third instar Monarch caterpillar with Oleander Aphids (Aphis nerii) on a Common Milkweed leaf. Both of these insect species absorb toxins from the milkweed which makes them distasteful to predators.Fifth instar (left and center) and fourth instar (right) monarch caterpillars devour a Common Milkweed leaf. There were over thirty of these caterpillars in just a ten by ten feet area this morning. We hope that if you’re keeping a habitat for Monarchs, you’re enjoying the same fortune right now!A slow-moving Monarch stopped for a break after making the circuit to deposit eggs on milkweed throughout the garden.Third instar (top), fourth instar (right), and fifth instar (left) Monarch caterpillars quickly consume the leaf of a Common Milkweed plant. Caterpillars emerging from eggs deposited today may not have sufficient late-season food to complete the larval segment of their life cycle. Need more milkweed!After benefiting from the nourishment of the Common Milkweed plant, a fifth instar Monarch caterpillar begins pupation on Big Bluestem grass.Two hours later, the chrysalis is complete.Another chrysalis, this one on flowering Switchgrass just two feet away from the previous one. An adult Monarch will emerge from this pupa to become part of what we hope will be the most populated southbound exodus for the species in over five years.There it is, soon ready to fly away. And all courtesy of the noxious milkweed.A chrysalis can often be found on man-made objects too. This one is on the rim of a flower pot.Ornamental flowers can attract adult Monarch butterflies seeking nectar. We’re now more careful to select seeds and plants that have not been treated with neonicotinoid insecticides. There’s growing concern over the impact these compounds may be having on pollinating species of animals. Oh…and we don’t mow, whack, cut, mutilate, or spray herbicides on our milkweed, but you probably figured that out already.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York.
At the moment there is a heavy snow falling, not an unusual occurrence for mid-February, nevertheless, it is a change in weather. Forty-eight hours ago we were in the midst of a steady rain and temperatures were in the sixties. The snow and ice had melted away and a touch of spring was in the air.
Big Bluestem in the Riverine Grasslands is inundated by the rising waters of the Susquehanna at Conewago Falls. The river ice has been dispersed by the recent mild temperatures and rains.
Anyone casually looking about while outdoors during these last several days may have noticed that birds are indeed beginning to migrate north in the lower Susquehanna valley. Killdeer, American Robins, Eastern Bluebirds, Red-winged Blackbirds, and Common Grackles are easily seen or heard in most of the area now.
Just hours ago, between nine o’clock this morning and one o’clock this afternoon, there was a spectacular flight of birds following the river north, their spring migration well underway. In the blue skies above Conewago Falls, a steady parade of Ring-billed Gulls was utilizing thermals and riding a tailwind from the south-southeast to cruise high overhead on a course toward their breeding range.
Ring-billed Gulls swarm in a thermal updraft above Conewago Falls to gain altitude prior to streaming off to the north and continuing their journey.Ring-billed Gulls climbing to heights sometimes exceeding 1,000 feet before breaking off and gliding away to the north.
The swirling hoards of Ring-billed Gulls attracted other migrants to take advantage of the thermals and glide paths on the breeze. Right among them were 44 Herring Gulls, 3 Great Black-backed Gulls, 12 Tundra Swans (Cygnuscolumbianus), 10 Canada Geese, 3 Northern Pintails (Anas acuta), 6 Common Mergansers, 3 Red-tailed Hawks, a Red-shouldered Hawk, 6 Bald Eagles (non-adults), 8 Black Vultures, and 5 Turkey Vultures.
A first-year Herring Gull (top center) is a standout in a “kettle” of Ring-billed Gulls.How many Ring-billed Gulls passed by today? More than 18,000…with emphasis on MORE THAN. You see, early this afternoon, the handy-dandy clicker-counter used to tick off and tally the big flights of birds as they pass by quit clicking and counting. Therefore, 18,000 is the absolute minimum number of Ring-billed Gulls seen migrating north today. Hopefully the trusty old oil can will get the clicker working again soon.
In the afternoon, the clouds closed in quickly, the flight ended, and by dusk more than an inch of snow was on the ground. Looks like spring to me.
The tall seed-topped stems swaying in a summer breeze are a pleasant scene. And the colorful autumn shades of blue, orange, purple, red, and, of course, green leaves on these clumping plants are nice. But of the multitude of flowering plants, Big Bluestem, Freshwater Cordgrass, and Switchgrass aren’t much of a draw. No self-respecting bloom addict is going out of their way to have a gander at any grass that hasn’t been subjugated and tamed by a hideous set of spinning steel blades. Grass flowers…are you kidding?
Big Bluestem in flower in the Riverine Grasslands at Conewago Falls.
O.K., so you need something more. Here’s more.
Meet the Partridge Pea (Chamaecrista fasciculata). It’s an annual plant growing in the Riverine Grasslands at Conewago Falls as a companion to Big Bluestem. It has a special niche growing in the sandy and, in summertime, dry soils left behind by earlier flooding and ice scour. The divided leaves close upon contact and also at nightfall. Bees and other pollinators are drawn to the abundance of butter-yellow blossoms. Like the familiar pea of the vegetable garden, the flowers are followed by flat seed pods.
The Partridge Pea can tolerate dry sandy soils.
But wait, here’s more.
In addition to its abundance in Conewago Falls, the Halberd-leaved Rose Mallow (Hibiscus laevis) is the ubiquitous water’s edge plant along the free-flowing Susquehanna River for miles downstream. It grows in large clumps, often defining the border between the emergent zone and shore-rooted plants. It is particularly successful in accumulations of alluvium interspersed with heavier pebbles and stone into which the roots will anchor to endure flooding and scour. Such substrate buildup around the falls, along mid-river islands, and along the shores of the low-lying Riparian Woodlands immediately below the falls are often quite hospitable to the species.
Halberd-leaved Rose Mallow is a durable inhabitant of the falls. Regular flooding keeps competing species at bay. A taproot helps to safeguard against dislocation, allowing plants to grow in places subjected to turbulent currents.Halberd-leaved Rose Mallow in bloom. The similarity to cultivated members of the Hibiscus genus can readily be seen. It is one of the showiest of perennial wildflowers in the floodplain. Note the lobed, halberd-shaped leaves, source of its former species name militaris.The seeds of Halberd-leaved Rose Mallow are contained in bladders which can float to assist in their distribution. Some of these bladders cling to the dead leafless stems in winter, making it an easy plant to identify in nearly any season.
A second native wildflower species in the genus Hibiscus is found in the Conewago Falls floodplain, this one in wetlands. The Swamp Rose Mallow (H. moscheutos) is similar to Halberd-leaved Rose Mallow, but sports more variable and colorful blooms. The leaves are toothed without the deep halberd-style lobes and, like the stems, are downy. As the common name implies, it requires swampy habitat with ample water and sunlight.
Swamp Rose Mallow in a sunny wetland. This variety with solid-colored flowers (without dark centers) and pale green leaves and stems was formerly known as a separate species of Swamp Rose Mallow, H. palustris. Note that the flowers are terminal on the stems.A few scattered specimens of a more typical variety of Swamp Rose Mallow are found on the shoreline and in the Riverine Grasslands of Conewago Falls. The blooms are bright pink with darker centers and the leaf stems are robust and reddish. This one is seen growing among Halberd-leaved Rose Mallow, with which it shares the characteristic of having flower stems growing from some of the upper leaf axils. A variety with red-centered white flowers is often found throughout the plant’s range.
In summary, we find Partridge Pea in the Riverine Grasslands growing in sandy deposits left by flood and ice scour. We find Halberd-leaved Rose Mallow rooted at the border between shore and the emergent zone. We find Swamp Rose Mallow as an emergent in the wetlands of the floodplain. And finally, we find marshmallows in only one location in the area of Conewago Falls. Bon ap’.
Here’s S’more
SOURCES
Newcomb, Lawrence. 1977. Newcomb’s Wildflower Guide. Little, Brown and Company. Boston, Massachusetts.
It has not been a good summer if you happen to be a submerged plant species in the lower Susquehanna River. Regularly occurring showers and thunderstorms have produced torrents of rain and higher than usual river stages. The high water alone wouldn’t prevent you from growing, colonizing a wider area, and floating several small flowers on the surface, however, the turbidity, the suspended sediment, would. The muddy current casts a dirty shadow on the benthic zone preventing bottom-rooted plants from getting much headway. There will be smaller floating mats of the uppermost leaves of these species. Fish and invertebrates which rely upon this habitat for food and shelter will find sparse accommodation…better luck next year.
Due to the dirty water, fish-eating birds are having a challenging season as they try to catch sufficient quantities of prey to feed themselves and their offspring. A family of Ospreys (Pandion haliaetus) at Conewago Falls, including recently fledged young, were observed throughout this morning and had no successful catches. Of the hundred or more individual piscivores of various species present, none were seen retrieving fish from the river. The visibility in the water column needs to improve before fishing is a viable enterprise again.
Ospreys competing for a suitable fishing perch. Improving water conditions in the coming week should increase their success as predators.Versatile at finding food, adult Bald Eagles are experienced and know to be on the lookout for Ospreys with fish, a meal they can steal through intimidation.
While the submerged plant communities may be stunted by 2017’s extraordinary water levels, there is a very unique habitat in Conewago Falls which endures summer flooding and, in addition, requires the scouring effects of river ice to maintain its mosaic of unique plants. It is known as a Riverine Grassland or scour grassland.
The predominant plants of the Riverine Grasslands are perennial warm-season grasses. The deep root systems of these hardy species have evolved to survive events which prevent the grassland from reverting to woodland through succession. Fire, intense grazing by wild herd animals, poor soils, drought, and other hardships, including flooding and ice scour, will eliminate intolerant plant species and prevent an area from reforesting. In winter and early spring, scraping and grinding by flood-driven chunk ice mechanically removes large woody and poorly rooted herbaceous growth from susceptible portions of the falls. These adverse conditions clear the way for populations of species more often associated with North America’s tall grass prairies to take root. Let’s have a look at some of the common species found in the “Conewago Falls Pothole Rocks Prairie”.
Big Bluestem (Andropogon gerardi), seen heregrowing in the cracks of a pothole rock. High water nourishes the plant by filling the crevices with nutrient-loaded sediment. This species evolved with roots over three feet deep to survive fires, trampling by bison, and drought.Freshwater Cordgrass (Spartina pectinata) does well with its roots in water. It creates exceptional bird habitat and grows in the falls and on ice-scoured small islands in free-flowing segments of the Susquehanna River downstream.Switchgrass (Panicum virgatum), like Big Bluestem, is one of the tall grass prairie species and, like Freshwater Cordgrass, grows in near pure stands on ice-scoured islands. It takes flooding well and its extensive root system prevents erosion.Though not a grass, Water Willow (Justicia americana) is familiar as a flood-enduring emergent plant of river islands, gravel bars, and shorelines where its creeping rhizome root system spreads the plant into large masses. These stands are often known locally as “grass beds”. This member of the acanthus family provides habitat for fish and invertebrates among its flooded leaves and stems. Its presence is critical to aquatic life in a year such as this.
The Conewago Falls Riverine Grassland is home to numerous other very interesting plants. We’ll look at more of them next time.
SOURCES
Brown, Lauren. 1979. Grasses, An Identification Guide. Houghton Mifflin Company. New York, NY.
