To pass the afternoon, we sat quietly along the edge of a pond created recently by North American Beavers (Castor canadensis). They first constructed their dam on this small stream about five years ago. Since then, a flourishing wetland has become established. Have a look.
Vegetation surrounding the inundated floodplain helps sequester nutrients and sediments to purify the water while also providing excellent wildlife habitat.The beaver lodge was built among shrubs growing in shallow water in the middle of the pond.Woolgrass (Scirpus cyperinus) is a bulrush that thrives as an emergent and as a terrestrial plant in moist soils bordering the pond.A male Common Whitetail dragonfly keeping watch over his territory.A Twelve-spotted Skimmer perched on Soft Rush.A Blue Dasher dragonfly seizing a Fall Field Cricket (Gryllus pennsylvanicus).A Spicebush Swallowtail visiting a Cardinal Flower.A Green Heron looking for small fish, crayfish, frogs, and tadpoles.The Green Heron stalking potential prey.A Wood Duck feeding on the tiny floating plant known as Lesser Duckweed (Lemna minor).A Least Sandpiper poking at small invertebrates along the muddy edge of the beaver pond.A Solitary Sandpiper.A Solitary Sandpiper testing the waters for proper feeding depth.A Pectoral Sandpiper searches for its next morsel of sustenance.The Sora (Porzana carolina) is a seldom seen rail of marshlands including those created by North American Beavers. Common Cattails, sedges, and rushes provide these chicken-shaped wetland birds with nesting and loafing cover.
Isn’t that amazing? North American Beavers build and maintain what human engineers struggle to master—dams and ponds that reduce pollution, allow fish passage, and support self-sustaining ecosystems. Want to clean up the streams and floodplains of your local watershed? Let the beavers do the job!
At Lake Redman just to the south of York, Pennsylvania, a draw down to provide drinking water to the city while maintenance is being performed on the dam at neighboring Lake Williams, York’s primary water source, has fortuitously coincided with autumn shorebird migration. Here’s a sample of the numerous sandpipers and plovers seen today on the mudflats that have been exposed at the southeast end of the lake…
One of a hundred or more Least Sandpipers seen on mudflats at Lake Redman today.A Semipalmated Plover and a Least SandpiperPectoral Sandpipers.A Pectoral Sandpiper and two Least Sandpipers.A Semipalmated Sandpiper.A Stilt Sandpiper feeding.Stilt Sandpiper consuming an edible.Stilt Sandpiper at rest.A Solitary SandpiperA Lesser Yellowlegs.A Greater Yellowlegs.Stirring up the shorebird crowd every now and then were several Ospreys, but all would soon be back to the business of feeding in the mud.An Osprey hovers above shallow water near the mudflats as it searches for fish.
Not photographed but present at Lake Redman were at least two additional species of shorebirds, Killdeer and Spotted Sandpiper—bringing the day’s tally to ten. Not bad for an inland location! It’s clearly evident that these waders overfly the lower Susquehanna valley in great numbers during migration and are in urgent need of undisturbed habitat for making stopovers to feed and rest so that they might improve their chances of surviving the long journey ahead of them. Mud is indeed a much needed refuge.
Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge. At Bombay Hook, they don’t waste your money mowing grass. Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools. Birds there can often be observed at close range. The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools. Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.Two more Short-billed Dowitchers on the way in.Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed. Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge. Mallards, American Avocets, and egrets are gathered on the shore.More Short-billed Dowitchers arriving to feed in Raymond Pool.Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool. A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.American Avocets probe the muddy bottom of Raymond Pool.Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early. Many migrating Dunlin linger at Bombay Hook into October and even November.This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp PoolA Greater Yellowlegs at Bear Swamp Pool.A Caspian Tern patrolling Raymond Pool.The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass. High tide licks at the roots of the cordgrass supporting the temporary domicile.By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice). It is our only dragonfly able to breed in saltwater. Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.Look up! A migrating Bobolink passes over the dike at Shearness Pool.Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool. Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river. The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century. An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers. Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.A Great Egret prowling Shearness Pool.A Snowy Egret in Bear Swamp Pool.Wood Ducks in Bear Swamp Pool.A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.
As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.
Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay. We found just this one during our visit to Bombay Hook. Man has certainly decimated populations of this ancient crustacean during recent decades.As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.A male Atlantic Marsh Fiddler Crab peers from its den.Herons and egrets including this Great Egret are quite fond of fiddler crabs. As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.An Osprey patrols the vast tidal areas opposite Shearness Pool.
No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook. This one was in shrubby growth along the dike at the north end of Shearness Pool.These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries. The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware. The seed balls are a favorite winter food of goldfinches and siskins.Finis Pool has no frontage on the tidal marsh but is still worth a visit. It lies along a spur road on the tour route and is located within a deciduous coastal plain forest. Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.The White-tailed Deity is common along the road to Finis Pool.Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well. This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.
We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.
It may look like just a puddle in the woods, but this is a very specialized wetland habitat, a habitat that is quickly disappearing from the Lower Susquehanna River Watershed. It’s a vernal pool—also known as a vernal pond or an ephemeral (lasting a short time) pool or pond.
Viable vernal pools have several traits in common…
They contain water in the spring (hence the name vernal).
They have no permanent inflow or outflow of water.
They typically dry up during part of the year—usually in late summer.
They are fish-free.
They provide breeding habitat for certain indicator species of forest-dwelling amphibians and other animals.
They are surrounded by forest habitat that supports the amphibians and other vernal pool species during the terrestrial portion of their life cycle.
To have a closer look at what is presently living in this “black leaf” vernal pool, we’re calling on the crew of the S. S. Haldeman to go down under and investigate.
Along the surface of the pool we’re seeing clusters of amphibian eggs, a sign that this pond has been visited by breeding adult frogs and/or salamanders during recent weeks.Amphibian eggs and the white tail filaments of an invertebrate of interest, Springtime Fairy Shrimp (Eubranchipus vernalis), an endemic of vernal ponds.
Let’s take it down for a better look. Dive, all dive!
Algae provides food for the shrimp and other inhabitants of the pool. Leaf litter furnishes hiding places for the pool’s many inhabitants.These loose clusters of eggs appears to be those of Wood Frogs, a vernal pool indicator species.Clusters of Wood Frog eggs, the embryos within those in the center of the image less developed than those to the left.More Wood Frog eggs. Hatching can take anywhere from two weeks to two months, depending on temperature.Wood Frog eggs with developing larvae (tadpoles) plainly visible. The green color of the eggs is created by a symbiotic algae, Oophila amblystomatis, a species unique to vernal pools. The algae utilizes the waste produced by the developing embryos to fuel its growth and in return releases oxygen into the water during photosynthesis. Upon hatching, the tadpoles rely upon the algae as one of their principle food sources.A zoomed-in view showing development of the larvae and what appears to be a tiny invertebrate clinging on the white egg in the upper right. White eggs don’t hatch and may be infected by a fungus.Wood Frog eggs and Springtime Fairy Shrimp.Wood Frog eggs and Springtime Fairy Shrimp.Springtime Fairy Shrimp swim upside-down. Note the small, bluish clusters of eggs attached to the abdomens of these females. Springtime Fairy Shrimp live their entire lives in the vernal pool. After being deposited in the debris at the bottom of the pool, the eggs will dry out during the summer, then freeze and re-hydrate before hatching during the late winter.A damselfly larva consuming fairy shrimp. (Visible in the margin between the uppermost lobes of the dark-colored oak leaf to the right.)Getting in close we see A) the damselfly larva eating a Springtime Fairy Shrimp and B) one of several discarded exoskeletons of consumed shrimp near this predator.A fishfly larva (Chauliodinae). Mosquito numbers are kept in check by the abundance of predators in these pools.Springtime Fairy Shrimp and a Marbled Salamander (Ambystoma opacum) larva. The presence of these species confirms this small body of water is a fully-functioning vernal pool.Springtime Fairy Shrimp and two more larval Marbled Salamanders. The salamanders’ enlarged gills are necessary to extract sufficient oxygen from the still waters of the pool.The Marbled Salamander is one of three species of mole salamanders found in the Lower Susquehanna River Watershed. All breed in vernal pools and live their air-breathing adult lives under the leaves of the forest in subterranean tunnels where they feed on worms and other invertebrates. Photos of an adult Marbled Salamander and the other two species, Spotted Salamander (Ambystoma maculatum) and Jefferson Salamander (Ambystoma jeffersonianum), can be found by clicking the “Amphibians” tab at the top of this page.Marbled Salamanders lay their eggs during the fall. If the bed of the pool is dry at breeding time, the adult female will remain to guard the eggs until rain floods the pool. The eggs hatch upon inundation, sometimes during the winter.Marbled Salamanders, like all amphibians that develop in vernal pools, must complete transformation into their air-breathing terrestrial life stage before the pool dries up in the summer heat.A larval Marbled Salamander explores the bottom of the pool.A larval Marbled Salamander, Wood Frog eggs, and Springtime Fairy Shrimp, it’s an abundance of life in what at first glance may appear to be just a mud puddle.
We hope you enjoyed this quick look at life in a vernal pool. While the crew of the S. S. Haldeman decontaminates the vessel (we always scrub and disinfect the ship before moving between bodies of water) and prepares for its next voyage, you can learn more about vernal pools and the forest ecosystems of which they are such a vital component. Be sure to check out…
If you are a landowner or a land manager, you can find materials specifically providing guidance for protecting, restoring, and re-establishing vernal pool habitats at…
Those who happen to come upon it might think this football-sized gelatinous blob is a sure sign of pollution. A freshwater bryozoan (Pectinatella magnifica) colony is composed of a single microscopic founder and its many clones. Despite its bizarre appearance, the “moss animal” is an indicator of good water quality. Pectinatella magnifica is found in clear lentic (still) waters of streams, lakes, and ponds where each individual in the colony feeds by extending a disk of sticky tentacles, called a lophophore, from within its protective sheath to capture single-celled algae (e.g., diatoms) and other plankton. From now through autumn, these bryozoans are reproducing by means of cell-filled statoblasts, durable little seedlike pods which can survive the harsh conditions of both winter and drought and sometimes be transported by animals, wind, or water currents to new areas. Spring weather and/or rehydration of a dried-up lentic pool stimulates a statoblast to open, the cells contained therein then develop into a zooid that attempts to start a new colony by cloning itself.
Along the lower Susquehanna, an unseasonably mild day in early spring can provide an observer with the opportunity to witness an annual spectacle seldom seen by the average visitor to the river—concentrations of dozens, sometimes hundreds, of turtles as they emerge from their winter slumber to bathe in the year’s first surge of warm air and sunshine.
Snapping Turtles (Chelydra serpentina) spend the winter buried in mud along the river shoreline and in nearby Alluvial Terrace Wetlands. We photographed this one just as it was digging its way out.A cold and stiff Snapping Turtle crawls away from the shade toward sun-drenched shallows where it will have a chance to limber up.A cruise in open water loosens up the muscles and gets rid of some of the accumulations of sticky mud and muck.Freshly emerged Painted Turtles clamber onto a log to bask in the cloud-filtered sun.A Painted Turtle looking for a place to get out of the chilly water soon discovered the obvious solution.It’s catching on, more Painted Turtles atop a Snapping Turtle in an Alluvial Terrace Wetland.The Common Map Turtle (right) is the turtle most frequently observed basking on rocks and logs along the main stem of the Susquehanna. To the left is a Red-eared Slider (Trachemys scripta elegans), an increasingly numerous invasive species. The first Red-eared Sliders arrived in the river as, you guessed it, unwanted pets. Editor’s Note: Special thanks to the local North American Beaver (Castor canadensis) for trimming the trees and providing a clear shot for this photograph!And now, a quick quiz. Name the things that don’t belong in this picture? Here’s a hint: a non-native Red-eared Slider (left) joins indigenous Painted Turtles atop a discarded tire in an Alluvial Terrace Wetland in Dauphin County, Pennsylvania.
Let’s take a quiet stroll through the forest to have a look around. The spring awakening is underway and it’s a marvelous thing to behold. You may think it a bit odd, but during this walk we’re not going to spend all of our time gazing up into the trees. Instead, we’re going to investigate the happenings at ground level—life on the forest floor.
Rotting logs and leaf litter create the moisture retaining detritus in which mesic forest plants grow and thrive. Note the presence of mosses and a vernal pool in this damp section of forest.The earliest green leaves in the forest are often those of the Skunk Cabbage (Simplocarpus foetidus). This member of the arum family gets a head start by growing in the warm waters of a spring seep or in a stream-fed wetland. Like many native wildflowers of the forest, Skunk Cabbage takes advantage of early-springtime sun to flower and grow prior to the time in late April when deciduous trees grow foliage and cast shade beneath their canopy.Among the bark of dead and downed trees, the Mourning Cloak butterfly (Nymphalis antiopa) hibernates for the winter. It emerges to alight on sun-drenched surfaces in late winter and early spring.Another hibernating forest butterfly that emerges on sunny early-spring days is the Eastern Comma (Polygonia comma), also known as the Hop Merchant.In a small forest brook, a water strider (Gerridae) chases its shadow using the surface tension of the water to provide buoyancy. Forests are essential for the protection of headwaters areas where our streams get their start.Often flooded only in the springtime, fish-free pools of water known as vernal ponds are essential breeding habitat for many forest-dwelling amphibians. Unfortunately, these ephemeral wetland sites often fall prey to collecting, dumping, filling, and vandalism by motorized and non-motorized off-roaders, sometimes resulting in the elimination of the populations of frogs, toads, and salamanders that use them.Wood Frogs (Lithobates sylvaticus) emerge from hiding places among downed timber and leaf litter to journey to a nearby vernal pond where they begin calling still more Wood Frogs to the breeding site.Wood Frog eggs must hatch and tadpoles must transform into terrestrial frogs before the pond dries up in the summertime. It’s a risky means of reproduction, but it effectively evades the enormous appetites of fish.When the egg laying is complete, adult Wood Frogs return to the forest and are seldom seen during the rest of the year.In early spring, Painted Turtles emerge from hideouts in larger forest pools, particularly those in wooded swamps, to bask in sunny locations.Dead standing trees, often called snags, are essential habitats for many species of forest wildlife. There is an entire biological process, a micro-ecosystem, involved in the decay of a dead tree. It includes fungi, bacteria, and various invertebrate animals that reduce wood into the detritus that nourishes and hydrates new forest growth.Birds like this Red-headed Woodpecker feed on insects found in large snags and nest almost exclusively in them. Many species of wildlife rely on dead trees, both standing and fallen, during all or part of their lives.
There certainly is more to a forest than the living trees. If you’re hiking through a grove of timber getting snared in a maze of prickly Multiflora Rose (Rosa multiflora) and seeing little else but maybe a wild ungulate or two, then you’re in a has-been forest. Logging, firewood collection, fragmentation, and other man-made disturbances inside and near forests take a collective toll on their composition, eventually turning them to mere woodlots. Go enjoy the forests of the lower Susquehanna valley while you still can. And remember to do it gently; we’re losing quality as well as quantity right now—so tread softly.
The White-tailed Deity in a woodlot infested by invasive tangles of Multiflora Rose.
It’s been an atypical summer. The lower Susquehanna River valley has been in a cycle of heavy rains for over a month and stream flooding has been a recurring event. At Conewago Falls, the Pothole Rocks have been inundated for weeks. The location used as a lookout for the Autumn Migration Count last fall is at the moment submerged in ten feet of roaring water. Any attempt to tally the migrants which are passing thru in 2018 will thus be delayed indefinitely. Of greater import, the flooding at Conewago Falls is impacting many of the animals and plants there at a critical time in their annual life cycle. Having been displaced from its usual breeding sites on the river, one insect species in particular seems to be omnipresent in upland areas right now, and few people have ever heard of it.
So, you take a cruise in the motorcar to your favorite store and arrive at the sprawling parking lot. Not wishing to have your doors dented or paint chipped because you settled for a space tightly packed among other shopper’s conveyances, you park out there in the “boondocks”. You know the place, the lightly-used portion of the lot where sometimes brush grows from cracks in the asphalt and you must be on alert for impatient consumers who throttle-up to high speeds and dash diagonally across the carefully painted grids on the pavement to reach their favorite parking destination in the front row. Coming to a stop, you take the car out of gear, set the brake, disengage the safety belt, and gather your shopping list. You grasp the door handle and, not wanting to be flattened by one of the aforementioned motorists, you have a look around before exiting.
It was then that you saw the thing, hovering above your shiny bright hood. For a brief moment, it seemed to be peering right through the windshield at you with big reddish-brown eyes. In just a second or two, it turned its whole bronze body ninety degrees to the left and darted away on its cellophane wings. Maybe you didn’t really get a good look at it. It was so fast. But it certainly was odd. Oh well, time to walk inside a grab a few provisions. Away you go.
Upon completion of your shopping, you’re taking the long stroll back to your car and you notice more of these peculiar creatures. Two are coupled together and are hovering above someone’s automobile hood, then they drop down, and the lower of the two taps its abdomen on the paint. You ask yourself, “What are these bizarre things?”
Meet the Wandering Glider (Pantala flavescens), also known as the Globe Wanderer or Globe Skimmer, a wide-ranging dragonfly known to occur on every continent with the exception of Antarctica.
Wandering Gliders sometimes arrive in the lower Susquehanna River valley in large numbers after catching a ride on sustained winds from southerly directions and will often fly and migrate in storm systems. Conditions for such movements have been optimal in our region since mid-July. These dragonflies will often hover above motor vehicle hoods and, after mating, females will deposit eggs upon them, apparently mistaking their glossy surface for small pools of water.
Wandering Gliders travel the globe, and as such are accomplished fliers. Adults spend most of the day on the wing, feeding upon a variety of flying insects. Days ago, I watched several intercepting a swarm of flying ants. As fast as ants left the ground they were grabbed and devoured by the gliders. Wandering Gliders are adept at taking day-flying mosquitos, often zipping stealthily past a person’s head or shoulders to grab one of the little pests—the would-be skeeter victim usually unaware of the whole affair.
Due to their nomadic life history, Wandering Gliders are opportunists when breeding and will lay eggs in most any body of freshwater. Their larvae do not overwinter prior to maturity; adults can be expected in a little more than one to two months. Repetitive flooding in the Lower Susquehanna River Watershed this summer may be reducing the availability of the best local breeding sites for this species—riverine, stream, and floodplain pools of standing water with prey. This may explain why thousands of Wandering Gliders are patrolling parking lots, farmlands, and urban areas this summer. And it’s the likely reason for their use of puddles on asphalt pavement, on rubber roofs, and in fields as places to try to deposit eggs. Unfortunately, they may be as likely to succeed there as they are on your motor vehicle hood.
At this time a year ago, the airspace above the Diabase Pothole Rocks at Conewago Falls was jammed with territorial male Wandering Gliders. Each male hovered at various locations around his breeding territory consisting of pools and water-filled potholes. Intruders would quickly be dispatched from the area, then the male would resume his patrols from a set of repetitively-used hovering positions about six feet above the rocks. Mating and egg-laying continued into late September. The larvae, also called nymphs or naiads, were readily observed in many pools and potholes in early October and the emergence of juveniles was noted in mid-October. The absence of flooding, the mild autumn weather, and the moderation of water temperatures in the pools and potholes courtesy of the sun-drenched diabase boulders helped to extend the 2017 breeding season for Wandering Gliders in Conewago Falls. They aren’t likely to experience the same favor this year, but their great ability to travel and adapt should overcome this momentary misfortune.
A male Wandering Glider aggressively patrols his territory in the Diabase Pothole Rocks Microhabitat at Conewago Falls. August 20, 2017.A mating pair of Wandering Gliders continue flying non-stop above one of thousands of suitable breeding pools among the Diabase Pothole Rocks at Conewago Falls. September 23, 2017.A female (bottom)Wandering Glider has deposited eggs in a pool while flying in tandem with a male (top). They’ll do the same thing on your automobile hood! Conewago Falls Diabase Pothole Rocks Microhabitat. September 23, 2017.Wandering Glider larvae are at the top of the food chain in flooded potholes. As they grew, these dragonfly larvae decimated the mosquito larvae which were abundant there earlier in the summer. October 7, 2017.A juvenile male Wandering Glider emerges from the pool where it fed and grew as a larva. It remained at water’s edge on the surface of a sun-warmed diabase rock for several hours to dry its wings. It soon flew away to parts unknown, possibly traveling hundreds or thousands of miles. Look carefully at the wings for the beige dash marks on the forward edge near the terminal end. Females lack this marking. Conewago Falls Diabase Pothole Rocks Microhabitat. October 14, 2017.A Wandering Glider exuviae, the shed exoskeleton of a creature gone, but not forgotten. October 14, 2017.
A couple of inches of rain this week caused a small increase in the flow of the river, just a burp, nothing major. This higher water coincided with some breezy days that kicked up some chop on the open waters of the Susquehanna upstream of Conewago Falls. Apparently it was just enough turbulence to uproot some aquatic plants and send them floating into the falls.
Piled against and upon the upstream side of many of the Pothole Rocks were thousands of two to three feet-long flat ribbon-like opaque green leaves of Tapegrass, also called Wild Celery, but better known as American Eelgrass (Vallisneria americana). Some leaves were still attached to a short set of clustered roots. It appears that most of the plants broke free from creeping rootstock along the edge of one of this species’ spreading masses which happened to thrive during the second half of the summer. You’ll recall that persistent high water through much of the growing season kept aquatic plants beneath a blanket of muddy current. The American Eelgrass colonies from which these specimens originated must have grown vigorously during the favorable conditions in the month of August. A few plants bore the long thread-like pistillate flower stems with a fruit cluster still intact. During the recent few weeks, there have been mats of American Eelgrass visible, the tops of their leaves floating on the shallow river surface, near the east and west shorelines of the Susquehanna where it begins its pass through the Gettysburg Basin near the Pennsylvania Turnpike bridge at Highspire. This location is a probable source of the plants found in the falls today.
Uprooted American Eelgrass floating into the Pothole Rocks under the power of a north wind. Note the white thread-like pistillate flower stem to the left and the small rooted specimen to the upper right. The latter is likely a plant from the creeping rootstock on the edge of a colony. As a native aquatic species, American Eelgrass is a critical link in the Susquehanna River and Chesapeake Bay food chain. Its decimation by pollution during the twentieth century led to migration pattern alterations and severe population losses for the Canvasback (Aythya valisineria) duck.American Eelgrass, a very small specimen, found growing in a low-lying Pothole Rock alongside the accumulations of freshly arriving material from upstream. Note that the creeping rootstock has leaves growing from at least three nodes on this plant. Eelgrass dislocations are regular occurrences which sometimes begin new colonies, like the small one seen here in this Diabase Pothole Rock Microhabitat.
The cool breeze from the north was a perfect fit for today’s migration count. Nocturnal migrants settling down for the day in the Riparian Woodlands at sunrise included more than a dozen warblers and some Gray Catbirds (Dumetella carolinensis). Diurnal migration was underway shortly thereafter.
A moderate flight of nocturnal migrants is indicated around NEXRAD sites in the northeastern states at 3:18 AM EDT. The outer rain bands of Hurricane Irma can be seen approaching the Florida Keys as the storm closes in on the peninsula. (NOAA/National Weather Service image)
Four Bald Eagles were counted as migrants this morning. Based on plumage, two were first-year eagles (Juvenile) seen up high and flying the river downstream, one was a second-year bird (Basic I) with a jagged-looking wing molt, and a third was probably a fourth year (Basic III) eagle looking much like an adult with the exception of a black terminal band on the tail. These birds were the only ones which could safely be differentiated from the seven or more Bald Eagles of varying ages found within the past few weeks to be lingering at Conewago Falls. There were as many as a dozen eagles which appeared to be moving through the falls area that may have been migrating, but the four counted were the only ones readily separable from the locals.
Red-tailed Hawks (Buteo jamaicensis) were observed riding the wind to journey not on a course following the river, but flying across it and riding the updraft on the York Haven Diabase ridge from northeast to southwest.
Bank Swallows (Riparia riparia) seem to have moved on. None were discovered among the swarms of other species today.
Ruby-throated Hummingbirds, Caspian Terns, Cedar Waxwings (Bombycilla cedrorum), and Chimney Swifts (Chaetura pelagica) were migrating today, as were Monarch butterflies.
Not migrating, but always fun to have around, all four wise guys were here today. I’m referring to the four members of the Corvid family regularly found in the Mid-Atlantic states: Blue Jay (Cyanocitta cristata), American Crow (Corvus brachyrhynchos), Fish Crow (Corvus ossifragus), and Common Raven (Corvus corax).
It looks like a big Blue Jay, but it’s not. This Belted Kingfisher (Megaceryle alcyon) takes a break after flying around the falls trying to shake a marauding Ruby-throated Hummingbird off its tail.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
SOURCES
Klots, Elsie B. 1966. The New Field Book of Freshwater Life. G. P. Putnam’s Sons. New York, NY.
