Five Flowering Plants for Cleaner Water

Currently in the Lower Susquehanna River Watershed, you can find these five species of herbaceous plants in full bloom. As they grow, they and others like them help to purify waters within their respective ecosystems by taking up nutrients—namely, the nitrogen and phosphorus that can lead to detrimental algal blooms and eutrophication in ponds, lakes, streams, and rivers.

(United States Geological Survey image by Virginia-West Virginia Science Center)

The Marsh Blue Violet (Viola cucullata) is most frequently found growing in the wet soils of forest bottomlands, usually where springs first break the earth’s surface and begin slowly trickling away to form a small brook or join an existing stream. The blooms are recognized by their darker purple centers and their long stems.

The seldom-noticed flowers of the Soft Rush (Juncus effusus), also known as the Common Rush, emerge from the sides of its quill-like stems. This wetland species is found in damp soils, sometimes in standing water, and grows in stiff, erect clumps that persist through winter. When found in pastures, Soft Rush is seldom of interest to cattle or other livestock. It therefore doesn’t lure these animals into muddy, puddle-prone areas. When subjected to heavy grazing in dry weather and flooding during wet spells, these puddle sites may host nearly pure stands of Soft Rush, the only plant able to thrive there. When it comes to nutrient uptake in these soggy sections of the meadow, the soft Rush is the lone ranger. Soft Rush seeds are available from Ernst Conservation Seeds in Meadville, PA, and are included in many of their mixes formulated for stormwater management basins and other wet soil applications.

Larger Blue Flag (Iris versicolor) is a plant of wetlands and shorelines. It can be grown as an emergent in ponds and lakes where it will help to absorb nutrients from both the water and the underlying substrate.

The Yellow Iris (Iris pseudacorus), also known as the Water Flag, is native to Eurasia and Africa. Seen here growing as an emergent among native Common Cattails (a superb water purifier), the Yellow Iris can easily escape cultivation and become invasive. The showy flowers and water-cleansing benefits of this plant make it attractive for use in the garden or farm pond, but considerations must be made for its aggressive growth and proclivity to escape to neighboring habitats. If you’re purchasing irises for transplanting, you’re probably better off sticking with the native Larger Blue Flag; it is far less vigorous and you’ll be able to grow other aquatic species along with it.

In large ponds, lakes, and low-gradient streams, one of the best aquatic plants for sequestering nutrients and clarifying water is Spatterdock, also known as Cow-lily or Yellow Pond Lily. Spatterdock does best as an emergent in shallow water along the shoreline. It grows well in full sunshine and makes excellent habitat for wildlife. Depending on the nutrient load from fish, waterfowl, decaying vegetation, and other sources, plant cover may need to be as high as 30% or more of the surface area to keep algae from overtaking a lake or large pond. Spatterdock can often be used to help fulfill these needs while still offering open water beneath the leaves and between the stems for fish, amphibians, reptiles, and macroinvertebrates to thrive.

May 19, 2025May 26, 2025

Back in the Day: Down on the Farm

Let us travel through time for just a little while to recall those sunny, late-spring days down on the farm—back when the rural landscape was a quiet, semi-secluded realm with little in the way of traffic, housing projects, or industrialized agriculture. Those among us who grew up on one of these family homesteads, or had friends who did, remember the joy of exploring the meadows, thickets, soggy springs, and woodlots they protected.

Low-intensity Farming — During much of the twentieth century, low-intensity agriculture provided a haven for wildlife. Periodic disturbances helped maintain cool-season grassland and early successional habitat for a number of species we currently find in decline.

For many of us, farmland was the first place we encountered and began to understand wildlife. Vast acreage provided an abundance of space to explore. And the discovery of each new creature provided an exciting experience.

Ned Smith's Birds of Field and Garden — Distributed by the Pennsylvania Game Commission, artist Ned Smith’s wildlife posters introduced many residents of the lower Susquehanna region to its birds and mammals. This poster of “Birds of Field and Garden” helped us learn what to expect and search for during our forays to the farm.

Today, high-intensity agriculture, relentless mowing, urban sprawl, and the increasing costs and demand for land have all conspired to seriously deplete habitat quality and quantity for many of the species we used to see on the local farm. Unfortunately for them, farm wildlife has largely been the victim of modern economics.

For old time’s sake, we recently passed a nostalgic afternoon at Middle Creek Wildlife Management Area examining what maintenance of traditional farm habitat has done and can do for breeding birds. Join us for a quick tour to remember how it used to be at the farm next door…

Always found nesting under the forebay of the barn, the Barn Swallow relentlessly pursued flying insects over the pond and meadow.

Eastern Meadowlarks arrived during March and April to begin nesting in their namesake. Their song, “spring-of-the-year”, heralded the new season.

Arriving in meadows and pastures during early May, the Eastern Kingbird provided for its nestlings by ambushing a variety of flying insects. By August, congregations of these birds could be found gathering along ponds and streams ahead of their fall migration.

In the cherry grove down by the creek, the Orchard Oriole would be singing incessantly to defend its territory.

Normally seed eaters through the colder months, American Goldfinches would regularly find a source of protein in the occupants of Eastern Tent Caterpillar nests.

Along the wet margins of the creek, Yellow Warblers would nest in the shrubs and small trees.

The “Traill’s Flycatcher” was a familiar find in low-lying areas of successional shrubs and small trees. Today, “Traill’s Flycatcher” is recognized as two distinct species, the Alder Flycatcher (Empidonax alnorum) and the Willow Flycatcher. In the lower Susquehanna valley, the latter (seen here) is by far the most common of the two.

During the nineteenth century, Eastern Bluebirds became a rarity on lower Susquehanna farms due to a combination of factors: pesticide (DDT) use, habitat loss, and competition with other birds for nest sites. The species saw a resurgence beginning in the 1970s with discontinuation of DDT applications and widespread provision of nesting boxes. Around human habitations, competition with invasive House Sparrows continues to be detrimental to their success.

Purple Martins and Tree Swallows — Purple Martins suffered a similar fate to the bluebirds. The potential for their recovery remains dubious and they continue to be very local breeders, fussy about selection of suitable man-made provisions for nesting. After considerable effort, Purple Martins have at last been attracted to nest in the condos placed for their use at Middle Creek Wildlife Management Area. In the artificial gourds, there are nesting Tree Swallows, a species which also benefits from the placement of boxes intended for bluebirds.

Abandoned fields and other successional habitats were and continue to be favored homes for Field Sparrows.

At almost any time of year, roving bands of Cedar Waxwings would suddenly visit old field habitat looking for berries among the shrubs and other pioneering woody growth. In early summer, after most species have already hatched their young, nesting would commence and these fruit eaters would transform into accomplished fly catchers.

During the twentieth century prior to the 1980s, Ring-necked Pheasant populations in the Lower Susquehanna River Watershed were comprised of breeding descendants of introduced birds supplemented by additional releases to maintain numbers sufficient for hunting. Year-round populations did and can reside in mosaic landscapes of early successional and grassland habitats, the latter including hay fields left unmowed through the nesting season.

Red-winged Blackbirds have always been a fixture of hay fields and meadows on farms. While the increase in mowing frequency has reduced their nesting success, they have persevered as a species by nesting earlier than other birds and by utilizing other landscape features such as densely vegetated stormwater basins for breeding sites.

Do you recall the last time you saw a Bobolink nesting in a hay field near you? Arriving in early May as a Neotropical migrant, the Bobolink requires a cool-season grassland such as hay field through at least July to complete its nesting cycle. Even earlier this century, we remember nesting Bobolinks being more widespread on farms throughout the region. Now, you almost have to go to Middle Creek if you want to see them.

Formerly more widespread in hay fields throughout the lower Susquehanna valley, the native Grasshopper Sparrow is yet another species falling victim to early mowing and intensive farming.

Cool-season Grasses — **Beautiful cool-season grasses waving in the spring breeze.** Meadows and hay fields can be managed to function as cool-season grasslands to provide nesting opportunities for many of the species we used to find down on the farm.

February 26, 2025April 28, 2025

Winter Survival: Generalists and Specialists

We’ve seen worse, but this winter has been particularly tough for birds and mammals in the Lower Susquehanna River Watershed. Due to the dry conditions of late summer and fall in 2024, the wild food crop of seeds, nuts, berries, and other fare has been less than average. The cold temperatures make insects hard to come by. Let’s have a look at how some of our local generalist and specialist species are faring this winter.

House Sparrows and a House Finch — House Sparrows (bottom) and House Finches (top) are generalists. To survive and thrive, they are adapted to a variety of habitats and types of food. House Sparrows live almost anywhere man-made structures are found. They are true omnivores and will eat almost anything, especially if they see something else try to eat it first. The House Sparrow’s close association with humans has allowed it to become the most widespread and successful living avian dinosaur. On a cold night, they’ll take shelter either within dense vegetation alongside a building or within the structure itself. Though not nearly as cosmopolitan, the House Finch has successfully colonized much of the eastern United States after escaping from captivity as a cage bird in New York during the middle of the twentieth century. Upon being trans-located here from the arid southwest, they adapted to suburbs and farmlands consuming primarily a granivore diet of seeds supplemented with seasonally available berries. They quickly became accustomed to offerings at bird-feeding stations as well. To survive the harsh winters in the northern sections of their range, eastern populations of House Finches are developing a pattern of migration. These movements are most evident in late fall when dozens or sometimes hundreds can be seen heading south over regional hawk-counting stations.

Northern Flicker feeding on Poison Ivy — Though they require dead trees for nesting and as places to find the grubs and adult insects upon which they primarily feed, woodpeckers including the Northern Flicker are generalists, seldom passing by a supply of fruits like these Poison Ivy berries as a source of winter food. Flickers regularly visit suburban areas where they’ll drop by at bird-feeding stations for suet. During the warmer months, they are the woodpecker most frequently seen on the ground where swarms of ants garner their full attention.

The Pileated Woodpecker is seldom found outside of mature forests where it digs relentlessly to remove grubs and other infestations from dead wood. But it is not a true specialist…

Pileated Woodpecker eating Poison Ivy — …it too finds a supply of Poison Ivy berries to be indispensable during a cold winter day.

The Ruby-crowned Kinglet is a generalist, feeding mostly on insects, but also consuming small seeds and some berries, including those of Poison Ivy. It nests well to our north in tall spruces and other evergreens. During migration and in winter, the Ruby-crowned Kinglet may be found in deciduous trees, brush, and tall grass in habitats ranging from forests to parks and suburbia. This male is displaying its seldom-seen red crown.

The Golden-crowned Kinglet, seen here on a Poison Ivy vine, is more of a specialist than the Ruby-crowned species, though the two will often occur in mixed groups during the winter. The Golden-crowned Kinglet nests in Spruce-Fir forests and in conifers within mixed woodlands. Even during migration, and particularly in winter, these birds are seldom found far from a stand of large evergreens within which they find shelter for the night.

The Hermit Thrush’s generalist lifestyle allows it to survive cold season weather in the Lower Susquehanna River Watershed. During summer, it breeds in coniferous and mixed woods from the northern parts of our valley north into Canada and feeds primarily on worms, insects, and other arthropods. During migration and in winter, the Hermit Thrush becomes a regular visitor to deciduous forests, woody parks and suburbs, particularly where a supply of wild berries is available to supplement its diet.

Iconic as it pulls earthworms from lawns during the warmer months, the generalist American Robin is fully dependent upon a crop of berries to survive winter conditions in the lower Susquehanna valley. The drought afflicted wild food crop of 2024 has led to fewer robins spending the season here and has delayed the northward push of migrating birds until the ground thaws and the earthworms make the ground rumble once again.

American Crows are an excellent example of a generalist species. They’ll go anywhere to find food and they’ll eat almost anything. Like the House Sparrow and several other generalists, they adapt very well to human activity and actually thrive on it. Garbage anyone?

Another career generalist is the widely worshiped White-tailed Deity, a species adapted to nearly all man-made landscapes with adequate vegetation upon which to browse. Pushed to the limit during severe weather, some individuals will consume carrion and even resort to cannibalism.

You might think the Great Blue Heron is a specialist. Nope, it’s an accomplished generalist. Great blues will live, feed, and breed on almost any body of fresh or brackish water. And their diet includes almost anything that swims. In winter, you’ll even see them in fields hunting mice and voles.

The Red-shouldered Hawk is a generalist with a diet ranging from amphibians and reptiles to small rodents and large insects. Mostly regarded as a species of bottomlands, they’ll frequent woodland edges, roadsides, and suburbia during the winter months.

During its periodic winter visits to the region, the American Tree Sparrow feeds on seeds among the grasses and forbs of semi-open country with scattered short shrubs and trees. A generalist species, it will show up at backyard bird-feeding stations, particularly during periods of inclement weather. In summer, the American Tree Sparrow nests in tundra with growths of stunted willows and spruce and their diet includes insects as a source of protein for themselves and their young.

The White-crowned Sparrow has similar winter habitat preferences to the tree sparrows…

White-crowned Sparrows — …it becomes adaptable and something of a generalist when searching for food during bad winter storms.

The Savannah Sparrow is an omnivore favoring insects in summer and seeds in winter. Though very closely tied to its grassland habitat year-round, snow cover can push these birds to enter woodier environs to consume fruits like these rose hips.

Dusk and dawn during the short days of winter are the prime hunting times of a mammal specialist, the Short-eared Owl. Its presence in the lower Susquehanna valley is dependent on two dominant factors: extensive grassland habitat and an adequate population of the owl’s favored food, the Meadow Vole (Microtus pennsylvanicus). The Short-eared Owl’s requirements as a specialist species make finding a suitable place to live difficult. Unlike the generalist birds and mammals that often adapt to the widespread man-made disturbances in the region, populations of specialists frequently become fragmented, reduced in abundance, and subject to extirpation.

The Meadow Vole is a generalist rodent that can be abundant in grasslands, early successional growth, fallow fields, marshlands, and, of course, meadows. They are primarily herbivores, but will occasionally consume insects and other arthropods. Usually nocturnal, some individuals venture out along their surface runways during daylight hours becoming vulnerable to diurnal raptors including kestrels, harriers, and buteos.

Short-eared Owl patrolling a grassland for Meadow Voles — A Short-eared Owl in near darkness patrolling a grassland for Meadow Voles.

The eastward expansion of the Coyote (Canis latrans), a species of western North America’s grasslands and scrublands, and its progressive mixing with the Wolf (Canis lupus) in these eastern extensions of its range, has produced an expanding population of very adaptable generalists we call Eastern Coyotes (Canis latrans var.). These omnivorous canines colonized the Lower Susquehanna River Watershed during the last four decades to replace extirpated wolves as the top-tier natural predator in the region. Their primary diet includes Meadow Voles in grasslands and other small mammals along woodland edges and in successional habitats. Seasonally, they consume the berries of numerous wild plants. Slightly larger than their western ancestors, Eastern Coyotes with the admixture of Wolf genetics can subdue small ungulates. Like other apex predators, they are attracted to vulnerable prey and thus play a crucial role in culling the weak and diseased among their potential quarry species to assure the health and potential of populations of these species as long-term sources of food energy. The benefit to the prey species is however largely diluted in populous areas of the northeast; most venison consumed by Eastern Coyotes here is in the form of road kill. During periods of extended snow cover when small rodents and other foods become inaccessible, Eastern Coyotes, particularly young individuals, will wander into new areas seeking sustenance. Sometimes they venture into cities and suburbs where they explore the neighborhoods in search of garbage and pet foods placed outside the home. (Video clip courtesy of Tyler and Grace Good. Click image to view.)

Wildlife certainly has a tough time making it through the winter in the lower Susquehanna valley. Establishing and/or protecting habitat that includes plenty of year-round cover and sources of food and water can really give generalist species a better chance of survival. But remember, the goal isn’t to create unnatural concentrations of wildlife, it is instead to return the landscape surrounding us into more of a natural state. That’s why we try to use native plants as much as possible. And that’s why we try to attract not only a certain bird, mammal, or other creature, but we try to promote the development of a naturally functioning ecosystem with a food web, a diversity of pollinating plants, pollinating insects, and so on. Through this experience, we stand a better chance of understanding what it takes to graduate to the bigger job at hand—protecting, enhancing, and restoring habitats needed by specialist species. These are efforts worthy of the great resources that are sometimes needed to make them a success. It takes a mindset that goes beyond a focus upon the welfare of each individual animal to instead achieve the discipline to concentrate long-term on the projects and processes necessary to promote the health of the ecosystems within which specialist species live and breed. It sounds easier than it is—the majority of us frequently become distracted.

Being an individual from a population of a very successful generalist species is no guarantee of survival. This Eastern Gray Squirrel fell from a tall tree when the limbs became ice covered during a storm earlier this month. Just a freak accident? Maybe, but mistakes like this are often fatal in the natural world. This squirrel’s passing may seem brutal, but it provides a better opportunity for other squirrels and animals that share its food and cover requirements to make it through the winter. And those survivors that didn’t suffer such a fatal mistake or, more importantly, don’t possess a vulnerability that may have contributed to such a mishap will have a chance to pass those traits on to a new generation. This squirrel as an individual is gone, its species lives on, and may be stronger for its passing.

Prescribed Burn of a Grassland — Pennsylvania Game Commission crews maintain a grassland ecosystem for Short-eared Owls and other specialists using prescribed fire to prevent succession beyond its earliest stages. Among the additional specialist species benefiting from this management tool are Monarchs and other butterflies whose host plants survive early-season fire, but not competition with woody vines, shrubs, trees, and invasive herbaceous growth.

On the wider scale, it’s of great importance to identify and protect the existing and potential future habitats necessary for the survival of specialist species. And we’re not saying that solely for their benefit. These protection measures should probably include setting aside areas on higher ground that may become the beach intertidal zone or tidal marsh when the existing ones become inundated. And it may mean finally getting out of the wetlands, floodplains, and gullies to let them be the rain-absorbing, storm-buffering, water purifiers they spent millennia becoming. And it may mean it’s time to give up on building stick structures on tinderbox lands, especially hillsides and rocky outcrops with shallow, eroding soils that dry to dust every few years. We need to think ahead and stop living for the view. If you want to enjoy the view from these places, go visit and take plenty of pictures, or a video, that’s always nice—then live somewhere else. Each of these areas includes ecosystems that meet the narrow habitat requirements of many of our specialist species, and we’re building like fools in them. Then we feign victimhood and solicit pity when the calamity strikes: fires, floods, landslides, and washouts—again and again. Wouldn’t it be a whole lot smarter to build somewhere else? It may seem like a lot to do for some specialist animals, but it’s not. Because, you see, we should and can live somewhere else—they can’t.

The Allegheny Woodrat (Neotoma magister), a threatened species in Pennsylvania and a critically imperiled species in Maryland, is a habitat specialist requiring the forested rocky slopes, talus-flanked ridgetops, and caves of the Ridge and Valley Province for its nest sites and survival. Isolated populations survived within similar environs in the lower Susquehanna River valley’s Piedmont Province and on South Mountain through at least the first half of the twentieth century, but have since been extirpated. Human encroachment that fragments their habitat and promotes exposure to parasite-hosting mammals including the Raccoon (*Procyon lotor)*, carrier of the Raccoon Roundworm (Baylisascarius procyonis), could prove fatal to remaining populations of this native mammal. (National Park Service image by Rick Olsen)

The American Oystercatcher (Haematopus palliatus) is a specialist species that uses its highly adapted bill to feed on marine invertebrates including mollusks, few of which are actually oysters. Reliant upon tidal ecosystems for its survival, many of the seashore animals that make up this wader’s diet are themselves specialist species. Oystercatchers spend nearly their entire lives in tidal marshes or within the intertidal zone on beaches. They also frequent rocky jetties, particularly during high tide. This individual was photographed near the mouth of Chesapeake Bay in Northhampton County, Virginia, a location that, when the waters of the Atlantic started rising over 10,000 years ago, was the lower Susquehanna valley about 60 miles from the river’s mouth at present-day Norfolk Canyon along the edge of the continental shelf. Get the drift?

February 17, 2025February 24, 2025

Think Spring: Order Your Seedlings Now!

Our wildlife has been having a tough winter. The local species not only contend with cold and stormy weather, but they also need to find food and shelter in a landscape that we’ve rendered sterile of these essentials throughout much of the lower Susquehanna valley’s farmlands, suburbs, and cities.

Planting trees, shrubs, flowers, and grasses that benefit our animals can go a long way, often turning a ho-hum parcel of property into a privately owned oasis. Providing places for wildlife to feed, rest, and raise their young can help assure the survival of many of our indigenous species. With a little dedication, you can be liberated from the chore of manicuring a lawn and instead spend your time enjoying the birds, mammals, insects, and other creatures that will visit your custom-made habitat.

Verdant Neighborhood — What makes some neighborhoods so appealing? It’s the foresight property owners had a half a century or more ago when they planted their lawns and gardens with a variety of sturdy, long-lived trees and shrubs. They’ve not only minimized the need for mowing grass, they’ve provided the present-day residents of their home with added thermal stability during both the blazing heat of summer and the chilling cold of winter.

Fortunately for us, our local county conservation districts are again conducting springtime tree sales offering a variety of native and beneficial cultivated plants at discount prices. Listed here are links to information on how to pre-order your plants for pickup in April. Click away to check out the species each county is offering in 2025!

Cumberland County Conservation District Annual Tree Seedling Sale—

Orders due by: Friday, March 21, 2025

Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025

Zebra Swallowtail on Wildflowers — During its 2025 tree seedling sale, the Cumberland County Conservation District is offering Northeast Native Wildflower seed mix for four dollars per ounce. One ounce plants approximately 200 square feet of bare soil. This is a Zebra Swallowtail visiting nectar-rich flowers during July of the first year after sowing this mix at a site along the Susquehanna.

Dauphin County Conservation District Seedling Sale—

Orders due by: Monday, March 17, 2025

Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025

"Red-twig Dogwood" — Able to thrive in wet soils, Red-osier and Silky Dogwood shrubs are ideal plants for intercepting and polishing stormwater in swales, detention/retention basins, and rain gardens. With their crimson twigs in winter, they look great along borders among clusters of cedars, pines, spruces, and other evergreens. They make an excellent choice for soil stabilization along the shorelines of streams, ponds, and other bodies of water too. Buy a dozen or more to create a showy mass planting in your soggy spot.

Franklin County Conservation District Tree Seedling Sale—

Orders due by: Wednesday, March 19, 2025

Pickup on: Thursday, April 24, 2025

The Franklin County Conservation District is offering American Elm seedlings in bundles of 25 for 36 dollars. Start them in pots for a couple of years to really get ’em going, then find places with damp soil and plenty of room to give ’em a try. During autumn, they look great in the company of spruces, white pines, and other large evergreens.

Lancaster County Annual Tree Seedling Sale—

Orders due by: Friday, March 7, 2025

Pickup on: Friday, April 11, 2025

Norway Spruce and White Pine — We purchased these Eastern White Pine, Norway Spruce, and Common Winterberry plants from the Lancaster County Conservation District Tree Sale about four years ago. They’re filling in as understory growth in the margins beneath some thirty-year-old Eastern Hemlocks to create dense cover for resident and visiting fauna at susquehannawildlife.net headquarters.

Lebanon County Conservation District Tree and Plant Sale—

Orders due by: Monday, March 3, 2025

Pickup on: Friday, April 18, 2025

For 2025, the Lebanon County Conservation District is offering Dawn Redwood (Metasequoia glyptostroboides) trees in packs of ten for twelve dollars. Though native to Asia, these adaptable trees present little threat of naturalizing and have many positive attributes in a conservation planting. Given ample space, the Dawn Redwood, a relative of the sequoias, will rapidly grow to a towering giant. They create a particularly dramatic landscape feature when planted in clumps of three to five trees or more. With age, the trunks become stout and very sturdy. Don’t like raking? The finely divided deciduous foliage can be left where it falls in autumn. It usually disintegrates by spring to enrich the soil and promote more growth.

Three Sequoias — The genus Metasequoia was first described in 1941 based upon fossils collected in Jurassic and earlier strata from widespread locations in the northern hemisphere. Metasequoia were believed extinct until just a few years later when a small number of living Dawn Redwoods were first discovered in southern China. Now distributed around the world for cultivation, direct descendants of this wild population of Metasequoia glyptostroboides are available for nearly anyone in a temperate climate to plant and grow to exceptional size. (National Park Service image)

The Lebanon County Conservation District is also selling Bald Cypress trees. They’re offered in bundles of ten for ten dollars. These long-lived trees resemble the Dawn Redwood. Both are tolerant of damp ground, but the native Bald Cypress is the species to choose for placement along streams, in wetlands, and on other sites with standing water or saturated soil.

Bald Cypress Swamp — Wildlife rich Bald Cypress swamps currently occur on the Atlantic Coastal Plain as far north as Sussex County, Delaware. Just to the south, they’re also found along Chesapeake Bay in areas that, during the last glacial maximum when sea level was 300 to 400 feet below today’s tide lines, were the Lower Susquehanna River Watershed in portions of present-day Maryland and Virginia. The northward post-glacial range extension of Bald Cypresses is now blocked by centuries of human intervention that has eliminated, isolated, or fragmented the wetland habitats where they could potentially become established. Why not lend them a hand? Plant a cypress swamp in your flood-prone bottomland. (National Park Service image by Andrew Bennett)

York County Conservation District Seedling Sale—

Orders due by: March 15, 2025

Pickup on: Thursday, April 10, 2025

A privacy planting of sturdy, native Eastern White Pines and Northern Red Oaks thriving around the border of a parking area where they also provide shade from the sun and help infiltrate a share of the stormwater that would otherwise become runoff.

Native Northeast Wildflower and Grass mix during its first summer — This year, the York County Conservation District is offering a Showy Northeast Native Wildflower and Grass seed mix for $19.99 per quarter pound. Sure beats mowing!

If you live in Adams County, Pennsylvania, you may be eligible to receive free trees and shrubs for your property from the Adams County Planting Partnership (Adams County Conservation District and the Watershed Alliance of Adams County). These trees are provided by the Chesapeake Bay Foundation’s Keystone 10-Million Trees Partnership which aims to close a seven-year project in 2025 by realizing the goal of planting 10 million trees to protect streams by stabilizing soils, taking up nutrients, reducing stormwater runoff, and providing shade. If you own property located outside of Adams County, but still within the Chesapeake Bay Watershed (which includes all of the Susquehanna, Juniata, and Potomac River drainages), you still may have an opportunity to get involved. Contact your local county conservation district office or watershed organization for information.

Planted Riparian Buffer — As they mature, tree and shrub plantings along streams return pollution-controlling functions to floodplains and provide critical habitat for wildlife. These riparian buffers not only improve water quality for fisheries, they also create travel corridors that prevent terrestrial animal populations from becoming isolated.

A CREP Project Area — Do you own a parcel of streamside or wetland acreage that you’d like to set aside and plant for the benefit of wildlife and water quality? Contact your local county conservation district office and ask them to tell you about CREP (Conservation Reserve Enhancement Program) and other programs that may offer incentives including payment of all or a portion of the costs of plantings and other habitat improvements.

We hope you’re already shopping. Need help making your selections? Click on the “Trees, Shrubs, and Woody Vines” tab at the top of this page to check out Uncle Tyler Dyer’s leaf collection. He has most of the species labelled with their National Wetland Plant List Indicator Rating. You can consult these ratings to help find species suited to the soil moisture on your planting site(s). For example: if your site has sloped upland ground and/or the soils sometimes dry out in summer, select plants with a rating such as UPL or FACU. If your planting in soils that remain moist or wet, select plants with the OBL or FACW rating. Plants rated FAC are generally adaptable and can usually go either way, but may not thrive or survive under stressful conditions in extremely wet or dry soils.

NATIONAL WETLAND PLANT LIST INDICATOR RATING DEFINITIONS

- - OBL (Obligate Wetland Plants)—Almost always occur in wetlands.
  - FACW (Facultative Wetland Plants)—Usually occur in wetlands, but may occur in non-wetlands.
  - FAC (Facultative Wetland Plants)—Occur in wetlands and non-wetlands.
  - FACU (Facultative Upland Plants)—Usually occur in non-wetlands, but may occur in wetlands.
  - UPL (Upland Plants)—Almost never occur in wetlands.

Using these ratings, you might choose to plant Pin Oaks (FACW) and Swamp White Oaks (FACW) in your riparian buffer along a stream; Northern Red Oaks (FACU) and White Oaks (FACU) in the lawn or along the street, driveway, or parking area; and Chestnut Oaks (UPL) on your really dry hillside with shallow soil. Give it a try.

December 24, 2024December 24, 2024

Photo of the Day

You may be on the verge of satisfying your appetite with chestnuts from an open fire, but this Red-bellied Woodpecker is just as pleased to retrieve a Shagbark Hickory nut from a hiding place among the tree’s cracks and crevices where it was stashed away earlier in the fall.

December 11, 2024December 14, 2024

Purple Finch Irruption?

Each spring and fall, Purple Finches are regular migrants through the Lower Susquehanna River Watershed. Northbound movements usually peak in April and early May. During the summer, these birds nest primarily in cool coniferous forests to our north. Then, in October and November each year, they make another local appearance on their way to wintering grounds in the southeastern United States. A significant population of Purple Finches remains to our north through the colder months, inhabiting spruce-pine and mixed forests from the Great Lakes east through New England and southeastern Canada. This population can be irruptive, moving south in conspicuous numbers to escape inclement weather, food shortages, and other environmental conditions. Every few years, these irruptive birds can be found visiting suburbs, parks, and feeding stations—sometimes lingering in areas not often visited by Purple Finches. Right now, Purple Finches in flocks larger than those that moved through earlier in the fall are being seen throughout the lower Susquehanna valley. Snow and blustery weather to our north may be prompting these birds to shift south for a visit. Here are some looks at members of a gathering of more than four dozen Purple Finches we’ve been watching in Lebanon County this month…

Male Purple Finch — A male Purple Finch in successional forest edge habitat consumes seeds from a dried stand of goldenrod.

November 19, 2024November 22, 2024

Five Favorite Visitors from the Northern Forests

As the autumn bird migration draws to a close for 2024, we’re delighted to be finding five of our favorite visitors from the coniferous and mixed forests of Canada and the northernmost continental United States.

Male Yellow-bellied Sapsucker — Yellow-bellied Sapsuckers drill evenly spaced holes which they repeatedly visit to harvest “sap” and the insects that become trapped therein. These surgically small excavations penetrate only the bark and the phloem, a shallow, spongy layer of the trunk that carries sugars from the leaves to other parts of the tree. Seldom do they puncture the thin, underlying cambium layer which protects the true sapwood. After healing, sapsucker holes leave linear rows of small scars which are sometimes still discernible decades after a bird’s final visit. This adult male will probably linger in the Lower Susquehanna River Watershed until colder weather renders the tree totally dormant, the flow of “sap” ebbs, and small insects become harder to find. Then he’ll make his way off toward southern forests of oak and pine for the remainder of the winter. Sapsuckers work their way north as sugary “sap” begins flowing again in the spring.

Almost the size of a thrush, the largest of our native sparrows, the Fox Sparrow, is currently at the peak of its southbound migration. While taking a break from their nocturnal flights, some may stray from forest edges to visit suburban feeding stations. Most leave our area by mid-December, then begin returning north as early as mid-February.

Not present every year, flocks of Pine Siskins are beginning to arrive in the lower Susquehanna region. They are currently most common along the mountain tops of the Ridge and Valley Province where they are feeding on the seeds of the Sweet Birch, a tree also known as Black Birch. Look for them around niger seed feeders and on the prickly seed balls of cultivated American Sweetgum, but don’t be surprised if they quickly pass us by to spend the winter on the Atlantic Coastal Plain to our south where this native tree grows in abundance.

Female/Juvenile Male Purple Finch — Since late October, Purple Finches have been widespread throughout the region. These female/juvenile male types are frequently being seen at feeders, particularly near woodlands and other areas with stands of conifers.

While right now is the best time to get out and look for these species from the northern forests, any or all of them could linger into the winter months, particularly where the food supply is sufficient and conifers and other evergreens provide cover from the blustery weather.

October 30, 2024October 31, 2024

A Towering American Elm

During the past week, Uncle Tyler Dyer has been out searching for autumn leaves to add to his collection. One of the species he had not encountered in previous outings was the American Elm (Ulmus americana), so he made a special trip to see a rare mammoth specimen in a small neighborhood park (Park Place) along Chestnut Street between 5th and Quince Streets in Lebanon, Pennsylvania.

American Elm at Park Place in Lebanon, Pennsylvania — Possibly the largest and oldest remaining American Elm in the lower Susquehanna watershed, the Park Place tree in Lebanon exceeds 60 feet in height and may be more than 300 years of age. In the nearly one hundred years since Dutch elm disease (DED) first began killing elms in eastern North America, this tree has been spared the fatal effects of the infection. DED is caused by several species of microfungi (Ophiostoma ulmi, O. himal-ulmi, and O. novo-ulmi) spread by numerous bark beetle (Curculionidae) species. As early as 1928, infected beetles arrived in the United States from the Netherlands among shipments of logs.

There’s still time to get out and see autumn foliage. With warmer weather upon us—at least temporarily—it’s a good time to go for a stroll. Who knows, you might find some spectacular leaves like these collected by Uncle Ty earlier this week. All were found adorning native plants!

Winged Sumac (Rhus copallina), a small native tree. In case you’re wondering, it’s not poisonous.

Black Chokeberry (Photinia melanocarpa), a native shrub also known as Aronia. It produces black-colored fruits in summer.

Red-osier Dogwood (Cornus sericea), a shrub that prefers wet or damp soils. It yields white fruits.

Maple-leafed Viburnum (Viburnum acerifolium), a native shrub of upland forest understories.

Highbush Blueberry (Vaccinium corymbosum), a native shrub and the source of many cultivated forms of the popular fruit.

To identify your finds, be certain to click the “Trees, Shrubs, and Woody Vines” tab at the top of this page to check out Uncle Ty’s extensive collection.

October 6, 2024

Photo of the Day

Here today, gone tomorrow. This Ruby-throated Hummingbird lingered around the numerous tubular flowers in the headquarters garden until near darkness last evening. It was last seen entering the cover provided by our Eastern Hemlocks where it presumably spent the night. Bright and early this morning, it made repeated visits to Mexican Cigar and this Bat-faced Cuphea to re-energize before resuming its long voyage to the tropics for winter. This marks the third consecutive year we’ve been visited by late-season hummingbirds during the first week of October. All were attracted to our nectar-rich plantings.

August 20, 2024November 22, 2024

Heat Flux Processes in Streams and Their Impact on Coldwater and Coolwater Fishes

The deluge of rain that soaked the lower Susquehanna watershed during last week is now just a memory. Streams to the west of the river, where the flooding courtesy of the remnants of Hurricane Debby was most severe, have reached their crest and receded. Sliding away toward the Chesapeake and Atlantic is all that runoff, laden with a brew of pollutants including but not limited to: agricultural nutrients, sediment, petroleum products, sewage, lawn chemicals, tires, dog poop, and all that litter—paper, plastics, glass, Styrofoam, and more. For aquatic organisms including our freshwater fish, these floods, particularly when they occur in summer, can compound the effects of the numerous stressors that already limit their ability to live, thrive, and reproduce.

One of those preexisting stressors, high water temperature, can be either intensified or relieved by summertime precipitation. Runoff from forested or other densely vegetated ground normally has little impact on stream temperature. But segments of waterways receiving significant volumes of runoff from areas of sun-exposed impervious ground will usually see increases during at least the early stages of a rain event. Fortunately, projects implemented to address the negative impacts of stormwater flow and stream impairment can often have the additional benefit of helping to attenuate sudden rises in stream temperature.

Stream Subjected to Agricultural Runoff — While a row of trees along a creek can help provide protection from the thermal impact of the sun, a vegetative riparian buffer must be much wider to be effective for absorbing, cooling, and treating runoff from fields, lawns, and paved surfaces. This buffer is too narrow to prevent surface runoff from polluting the water.

Of the fishes inhabiting the Lower Susquehanna River Watershed’s temperate streams, the least tolerant of summer warming are the trouts and sculpins—species often described as “coldwater fishes”. Coldwater fishes require water temperatures below 70° Fahrenheit to thrive and reproduce. The optimal temperature range is 50° to 65° F. In the lower Susquehanna valley, few streams are able to sustain trouts and sculpins through the summer months—largely due to the effects of warm stormwater runoff and other forms of impairment.

Sculpins, including the Blue Ridge Sculpin (Cottus caeruleomentum) seen here, are native coldwater fishes which, during the 11,000 years since the last glacial maximum, have had the availability of their favored habitat sharply reduced by warming water temperatures and a rising Atlantic. During this interval, seawater has inundated the path of the “Late” Pleistocene lower Susquehanna which passed through the section of flooded river watershed we now call Chesapeake Bay and continued across the continental shelf to what was, during the glacial maximum, the river’s mouth at Norfolk Canyon. Today, cut off from neighboring drainage basins, sculpins survive exclusively in cold headwaters, and only in those where human alterations including pollution, dams, channelization, and reduced base flow haven’t yet eliminated their isolated populations. Formerly believed to be composed of two widespread North American species, the Slimy Sculpin (Cottus cognatus) and the Mottled Sculpin (Cottus bairdii), study in recent decades is discovering that sculpin populations in the present-day lower Susquehanna and neighboring Potomac headwaters consist of at least three newly delineated species: Blue Ridge Sculpin, Potomac Sculpin (Cottus gerardi), and Checkered Sculpin (Cottus sp.), the latter an as yet undescribed species found only in the refugium of limestone springs in the Potomac drainage in West Virginia; Frederick and Washington Counties, Maryland; and Franklin County, Pennsylvania. (United States Geological Survey image)

Ice Age Susquehanna — Stare at this for a little while, you’ll figure it out…………More than 11,000 years ago, during the last glacial maximum, when sea level was about 275 feet lower than it is today, there was no Chesapeake Bay, just a great Susquehanna River that flowed to the edge of the continental shelf and its mouth at Norfolk Canyon. It was a river draining taiga forests of pine, spruce , and fir, and it carried along the waters of all the present-day bay’s tributaries and more. The section of the river’s watershed we presently call the lower Susquehanna was, at the time, the upper Susquehanna watershed. Brook Trout and sculpins had the run of the river and its tributaries back then. And the entire watershed was a coldwater fishery, with limestone and other groundwater springs providing not refuge from summer heat, but a place to escape freezing water. (United States Geological Survey base image)

Norfolk Canyon, the mouth of the Susquehanna River during the most recent glacial maximum, now lies more than 275 feet below the surface of the ocean and plunges to more than a mile in depth along the finger of out wash from the gorge. (United States Geological Survey image)

Rainbow. Brown, and Brook Trout — Tens of thousands of trout are raised in state-operated and cooperative nurseries for stocking throughout the lower Susquehanna valley. These rearing facilities are located on spring-fed headwaters with sufficient flow to assure cold temperatures year round. While the Rainbow Trout and Brown Trout (Salmo trutta) are the most commonly stocked species, the Brook Trout (Salvelinus fontinalis) is the only one native to American waters. It is the least tolerant of stream warming and still reproduces in the wild only in a few pristine headwaters streams in the region. During spring, all three of these species have been observed on rare occasions entering the fish lift facilities at the hydroelectric dams on the river, presumably returning to the Susquehanna as sea-run trout.

Coldwater fishes are generally found in small spring-fed creeks and headwaters runs. Where stream gradient, substrate, dissolved oxygen, and other parameters are favorable, some species may be tolerant of water warmer than the optimal values. In other words, these temperature classifications are not set in stone and nobody ever explained ichthyology to a fish, so there are exceptions. The Brown Trout for example is sometimes listed as a “coldwater transition fish”, able to survive and reproduce in waters where stream quality is exceptionally good but the temperature may periodically reach the mid-seventies.

The Eastern Blacknose Dace is sometimes classified as a “coldwater transition fish”. It can be found in headwaters runs as well as in creeks with good water quality.

The Longnose Dace is another “coldwater transition fish” known only from clear, clean, flowing waters.

More tolerant of summer heat than the trouts, sculpins, and daces are the “coolwater fishes”—species able to feed, grow, and reproduce in streams with a temperature of less than 80° F, but higher than 60° F. Coolwater fishes thrive in creeks and rivers that hover in the 65° to 70° F range during summer.

Creek Chubs — The Creek Chub is a familiar species of “coolwater fish” seldom found remaining in waters exceeding 80 degrees Fahrenheit.

The Yellow Perch (Perca flavescens) was perhaps the most frequently targeted coolwater “gamefish” in the Lower Susquehanna River Watershed prior to the introduction of the Northern Pike (Esox lucius) and Muskellunge (Esox masquinongy). Today’s prevalence of warmwater streams and the dozens of species of non-native predatory fishes now naturalized within them have left the Yellow Perch populations greatly reduced and all but forgotten by anglers. Out of sight, out of mind. (National Park Service image)

What are the causes of modern-day reductions in coldwater and coolwater fish habitats in the lower Susquehanna River and its hundreds of miles of tributaries? To answer that, let’s take a look at the atmospheric, cosmic, and hydrologic processes that impact water temperature. Technically, these processes could be measured as heat flux—the rate of heat energy transfer per unit area per unit time, frequently expressed as watts per meter squared (W/m²). Without getting too technical, we’ll just take a look at the practical impact these processes have on stream temperatures.

HEAT FLUX PROCESSES IN A SEGMENT OF STREAM

Heat Flux Processes on Streams and Rivers — Heat Flux Processes on Stream and River Segments. These processes could be measured as heat flux—the rate of heat energy transfer per unit area per unit time. (Environmental Protection Agency image)

- - INCOMING TEMPERATURE AND FLOW—The baseline temperature of stream water entering a given segment of waterway is obviously the chief factor determining its temperature when exiting that segment. Incoming temperature and flow also determine the water’s susceptibility to heat absorption or loss while transiting the segment. Lower flows may subject the given volume of water to a greater loss or gain of heat energy during the time needed to pass through the segment than the same volume at a higher flow. Lower flows may also reduce stream velocity and extend a given volume of water’s exposure time to the exchange of heat energy while moving through the segment. Generally speaking…
    1. …the higher the stream flow, the less a given volume of that stream’s water may be impacted by the effects of the heat flux processes within the segment.
    2. …the lower the stream flow, the more a given volume of that stream’s water may be impacted by the effects of the heat flux processes within that segment.
    3. …the temperature and flow rate of precipitation entering the segment are factors that determine the impact of its heat energy transfer to or from a given volume of the stream’s waters.
    4. …the temperature and flow rate of runoff and point-source discharges entering the segment are factors that determine the impact of their heat energy transfer to or from a given volume of the stream’s waters.

Stormwater Discharge into Channelized Creek — Stormwater from impervious surfaces including roads, parking lots, roofs, and lawns quickly impacts temperatures in small creeks. Channelized streams are availed few of the positive attributes provided by many of the heat flux processes we’re about to see. They therefore suffer from severe impairment and are exposed to temperature extremes that few aquatic organisms can survive. Runoff from sun-heated pavement during a summer thunderstorm can often exceed 100 degrees Fahrenheit and can, at sufficient flow rate, quickly raise the temperature of a small stream to well over 90 degrees.

Stormwater Runoff — Stormwater runoff not only poses a thermal threat to waterways, its a significant source of a wide variety of pollutants.

- - GROUNDWATER INPUT—In streams connected to the aquifer, the temperature in a flowing segment can be impacted by the influx of cold groundwater. With temperatures ranging from about 52° to 60° Fahrenheit, groundwater will absorb heat from the stream in summer, and warm it in the winter. In warmwater streams, coldwater and coolwater fishes will often seek areas of the substrate where groundwater is entering for use as refugium from the summer heat. Yellow Perch in the lower Susquehanna are known to exhibit this behavior.

Gaining Stream — Creeks and rivers connected to the aquifer and receiving supplemental flow from it are known as “gaining streams”. These streams frequently feed water into the aquifer as well. (United States Geological Survey image)

Losing Stream — When flowing through an area experiencing drought or an excessive removal of groundwater (lots of wells, etc.), a waterway can become a “losing stream”, one that surrenders a portion of its flow to recharge the aquifer. Further downstream, the reduced flow can make such a creek or river more susceptible to the effects of heat flux processes. (United States Geological Survey image)

Disconnected Stream — Seriously depleted aquifers can lead to a “disconnected stream”. Smaller waterways subjected to these conditions will sometimes lose all their flow to the ground, often causing a catastrophic failure of the aquatic ecosystem supported therein. (United States Geological Survey image)

Urban Flooding and Dry Streambed — Urban runoff overwhelms this small stream with polluted water than can reach temperatures of 100 degrees or more (left), then lets it high and dry with no baseflow during periods of dry weather (right) as the waterway becomes disconnected from the much-depleted aquifer.

Stormwater Retention Basin — Well-designed and properly constructed stormwater retention basins not only recharge groundwater supplies for wells and streams, they can also help prevent thermal pollution in waterways. Planted with native wetland species and allowed to thrive, they can become treasured wildlife islands in otherwise inhospitable environs. The benefits don’t stop there; plants also help sequester nutrients contained in the runoff.

- - HYPORHEIC EXCHANGE—Related to groundwater input, hyporheic exchange is the slow movement of water through the rock, sand, gravel, and soils composing the streambed, saturated shoreline, shallow aquifer, and connected floodplain of a creek or river. As a heat flux process, hyporheic exchange helps moderate extremes in seasonal water temperatures by conducting energy between the solid materials in the zone and the flowing water. Hyporheic zones are important habitats for many species of aquatic invertebrates and spawning fish. Natural chemical processes within these zones convert ammonia-producing wastes into nitrite, then nitrate, allowing it to be absorbed as food by plants growing in the stream or in the alluvium within the zone. Vegetation removal, channelization, legacy sediments, silt deposits, and man-made walls and dams can negate the benefits of hyporheic exchange.

Hyporheic Exchange — Exchange of surface and ground water within the hyporheic zone is most directly associated with high-gradient (left) and meandering (right) segments of streams. (United States Geological Survey image)

Legacy Sediments and Fill — Very common on streams in the lower Susquehanna valley are these accumulations of legacy sediments at the sites of former mill ponds. After the dams were removed, the creeks began eroding their way down through the mire as they tried to reestablish their floodplains and find their native substrate. These trapped waterways are not only cut off from their hyporheic zones, they’re now a major source of nutrient and sediment pollution. Misguided landowners like this one frequently dump fill into these sites to “save their land” and “control flooding”. The fill and materials added to “shore up the banks” do nothing to fix what ails the creek, but instead displace more water to make the impact of flooding even more widespread.

Flooplain and Stream Restoration — Rehabilitation projects that remove legacy sediments help restore hyporheic exchange by reconnecting the stream to its underlying geology, its floodplain, and its wetlands. Rising waters remain in the floodplain where they get a good bio-scrubbing and help replenish the creek and groundwater supply. As the experts say, “floodplains are for flooding.”

- - ATMOSPHERIC EXCHANGE (CONVECTION, EVAPORATION)—Primarily a process by which a stream loses heat energy and cools its waters, atmospheric exchange is also a means by which a warm air mass can relinquish heat to cooler waters and thus increase their temperature. This phenomenon can be dramatically enhanced when a stream passes through a so-called urban heat island where air temperatures remain warm through the night. Convection, the movement of heat energy through a fluid (liquid or gas), causes warmer, less-dense water to rise to the surface of a stream, particularly where there is minimal turbulence. When the air above is cooler than the water’s surface layer, the stream will conduct heat energy across the water/atmosphere interface causing the warmed air molecules to rise in a convection column. If the atmospheric relative humidity is less than 100%, some surface water will vaporize—a process that expends more of the stream’s heat energy. The rate of convective and evaporative cooling in a given stream segment is directly related to the degree of difference between the water temperature and air temperature, and to the relative humidity in the air mass above the lake, creek, or river. The mechanical action of stream turbulence including rapids, riffles, and falls increases the contact area between air and water to maximize the atmospheric exchange of heat energy. The convective air current we call surface wind has a turbulent wave-producing effect on water that can also maximize atmospheric exchange; think of a cold autumn wind robbing heat energy from a warm lake or river or a hot summer wind imparting its heat to a cooler creek. These exchanges are both conductive in nature (air-to-water/water-to-air) and evaporative, the latter being expedited by the movement of dry air over warm water.

Usually classified as one of the coolwater fishes, the bottom-dwelling Tessellated Darter can thrive in the warmer creeks and in the main stem of the Susquehanna by inhabiting riffles where atmospheric exchange in the form of increased evaporation helps reduce temperatures and convective currents carry the cooler, well-oxygenated water to the streambed.

Three mile Island Unit 1 Cooling Towers — Humans utilize the concept of atmospheric exchange, adopting the phenomena of evaporation and convection to cool the hot waters produced during electric generation and other industrial processes before discharge into a lake or river.

- - STREAMBED CONDUCTIVE EXCHANGE—In the lower Susquehanna watershed, there may be no better natural example of streambed conductive exchange than the Triassic-Jurassic diabase pothole bedrocks of Conewago Falls on the river at the south end of Three Mile Island.

During sunny days, the massive diabase pothole rocks at Conewago Falls absorb solar (shortwave) radiation, then conduct that heat energy into the flowing water, often continuing to pass the accumulated warmth into the river during the night. On cloudy days, the riverbed collects longwave atmospheric radiation, a heat flux process that yields significantly less energy for conduction into the rapids, riffles, and pools of the falls. During periods of low river flow, the heating effect of streambed conductive exchange can become magnified. Compared to conditions that prevail when torrents of turbid water are rushing through the falls, partially exposed bedrock surrounded by clear water collects radiated energy much more efficiently, then conducts the heat to a greatly reduced volume of passing water. During summer and autumn, this process can create a mix of temperature zones within the falls with warmer water lingering in slow-moving pools and cooler water flowing in the deeper fast-moving channels. Along the falls’ mile-long course, a haven is created for aquatic organisms including warmwater and some coolwater fishes, oft times attracting anglers and a variety of hungry migrating birds as well.

Classified as one of our coolwater fishes, the Fallfish finds favorable conditions for feeding, growing, and spawning in the well-oxygenated waters of Conewago Falls.

Though the lower Susquehanna River is classified as a warmwater fishery, the Northern Hog Sucker (Hypentelium nigricans), another of our native coolwater fishes, finds the fast-moving waters of Conewago Falls to its liking. Northern Hog Suckers are known to inhabit streams cold enough to host trout. They exhibit remarkable home range fidelity, sometimes spending their entire lives occupying the same several hundred feet of waterway. Northern Hog Suckers are often designated an indicator of good water quality, intolerant of many stream impairment parameters. Their presence in Conewago Falls provides testament to the quality of the warmwater fishery there.

Severely Impaired Channelized Stream — An unnatural example. The reduced base flow in this channelized and severely impaired creek has been rendered vulnerable to the negative impacts of several heat flux processes including streambed conductive exchange. Urban stormwater/surfacewater inflow, solar (shortwave) radiation, and heat conducted into the stream from the masonry walls, curbs, and raceway can all conspire to cook aquatic organisms with life-quenching summer water temperatures exceeding 90 degrees Fahrenheit.

- - SOLAR (SHORTWAVE) RADIATION—The sun provides the energy that fuels the earth’s complex climate. The primary heat flux process that heats our planet is the absorption of solar radiation in the shortwave spectrum, which includes ultraviolet, visible, and infrared frequencies at the upper end of the longwave spectrum. Streams and other bodies of water absorb the greatest amounts of solar (shortwave) radiation during the weeks around summer solstice when the sun at mid-day is closer to zenith than at any other time of the year. However, the heating impact of the radiation may be greatest when the volume of water in the creek, river, or lake is at its minimum for the year—often during early fall.

The Sun — The rate, measured in watts per square meter, at which solar (shortwave) energy is directly radiated to a given area on the earth’s surface (including streams and other waters) is determined by: solar activity, the angle of the sun in the sky, aspect (slope) of the receiving surface, the opacity of the overlying atmosphere, and the distance of the earth from the sun. The former varies with the year’s seasons, the time of day, and the latitude of a given area. The latter is currently at its annual minimum when earth is at perihelion during the early days of January, thus providing the northern hemisphere with a little bump in radiation during the shortest days of the year when the sun is at its lowest angle in the sky. (NASA image)

Solar (Shortwave) Radiation — A varying portion of the solar (shortwave) radiation reaching the earth is reflected back into space by clouds. A smaller share is absorbed by the atmosphere, thus heating it. An even lesser quantity is reflected back into space by water and land. The remainder of the energy is absorbed by the planet’s surfaces, its water and land. (NASA image)

- - INCIDENT SHORTWAVE RADIATION—Also known as insolation (incoming solar radiation), incident shortwave radiation is the sum total energy of both the direct solar radiation that travels to the earth’s surface unaffected by the atmosphere and the diffuse radiation, waves that have been weakened and scattered by constituents of the atmosphere before reaching the planet’s surface. On a cloudy day, the warming of terrestrial surfaces including streams and other bodies of water is the result of diffuse radiation. On days with any amount of sunshine at all, both direct and diffuse radiation heat our waters and lands.

Pumkinseed — Warmwater fishes such as the native Pumpkinseed (Lepomis gibbosus) thrive in sun-drenched 70-to-85-degree waters as long as other heat flux processes prevent sudden temperature increases and oxygen depletion.

Mowed Stream Bank — Mowed stream banks offer a waterway no protection from incoming solar (shortwave) radiation, nor terrestrial forms of impairment including nutrient-rich stormwater runoff and silt.

- - REFLECTED SHORTWAVE RADIATION—known as albedo, reflected solar (shortwave) radiation is energy directed away from the earth’s surface before being absorbed. A surface’s albedo value is basically determined by its color, black having little reflective value, white and silvery surfaces reflecting nearly all solar (shortwave) radiation away. A surface with no reflective properties has an albedo value of 0, while a totally reflective surface has a value of 1. Clean snow with a value of about 0.85 to 0.9 (85% to 90%) is a highly reflective surface; yellow snow isn’t as good. A stream, river, or lake blanketed with ice and snow will absorb very little solar energy and will rely upon other heat flux processes to trigger a melt and thaw. The surface of open water has a varying albedo value determined mostly by the angle of the sun. Solar radiation striking the water’s surface at a low angle is mostly reflected away, while that originating at an angle closer to zenith is more readily absorbed.

Forested Stream — To avoid the heating effects of solar (shortwave) and atmospheric longwave radiation, coldwater and coolwater fishes require streams offering protection from full exposure to direct sunlight and cloud cover. Runs and creeks flowing beneath a closed canopy of forest trees are shielded from 25% or more of incoming radiation and are thus able to better maintain thermal stability during the most vulnerable period of the year for temperature-sensitive fishes, May through October.

- - LONGWAVE RADIATION—Radiation in the longwave spectrum is composed of infrared waves at frequencies lower than those of the shortwave spectrum. Longwave radiation, sometimes just called infrared radiation, is produced by the earth and its atmosphere and is propagated in all directions, day and night. It warms mostly the lower atmosphere which in turn warms the earth’s surface including its waters. Some longwave energy can even be radiated into the waterway from its own streambed—and the stream can return the favor. Other forms of mass surrounding a stream such as a rocky shoreline or a man-made structure such as bridge pier can trade longwave radiation with a waterway. The effect of these latter exchanges is largely trivial and never rivals the heat flux transfer of warm to cold provided by conduction.

Longwave Radiation — Longwave radiation emissions slow as the temperature of the emitting mass decreases, just as they also increase with temperature of the mass. Longwave radiation emissions therefore decrease with altitude along with the temperature of the water vapor, carbon dioxide, methane, and other gases that produce them. As such, the highest reaches of the atmosphere have a greatly reduced capability of shedding longwave radiation into space. At ground level, lakes, creeks, and streams receive their greatest dose of longwave radiation while beneath the cover of low-lying clouds or fog. (NASA image)

- - CANOPY RADIATION—Trees emit longwave radiation that may have a limited heat flux impact on waterway temperature. This radiation is diffuse, of scattered effect, and scarcely detectable, particularly beneath multilayered dense canopies. Some of the infrared energy transmitted by the tree canopy is radiated skyward as well.
  - WATER RADIATION—Water, like all earthly matter composed of vibrating molecules, emits longwave radiation. This heat flux process provides an ongoing cooling effect to streams, rivers, lakes, and oceans—warmer ones shedding infrared energy at a faster rate than those that are cold.

Now that we have a basic understanding of the heat flux processes responsible for determining the water temperatures of our creeks and rivers, let’s venture a look at a few graphics from gauge stations on some of the lower Susquehanna’s tributaries equipped with appropriate United States Geological Survey monitoring devices. While the data from each of these stations is clearly noted to be provisional, it can still be used to generate comparative graphics showing basic trends in easy-to-monitor parameters like temperature and stream flow.

Each image is self-labeled and plots stream temperature in degrees Fahrenheit (bold blue) and stream discharge in cubic feet per second (thin blue).

West Conewago Creek 90 Days — The West Conewago Creek drains much of the Gettysburg Basin’s Triassic redbeds in Adams and northern York Counties in Pennsylvania and includes a small headwaters area in northern Maryland. The gauge station is located just a over a mile upstream from the waterway’s mouth on the Susquehanna just below Conewago Falls. Right through the summer heatwave, this 90-day graph shows a consistent daily pattern of daytime rises in temperature and nighttime cooling. To the right, a rapid cool down can be seen coinciding with two periods of high water, the first from a series of heavy thundershowers, the second from flooding caused by the remnants of Hurricane Debby. Notice that the early August downpours were so heavy that they cooled the hot surface runoff and waterway quickly, without creating a rise in stream temperature at the gauging station. Had this monitoring device been located on a small tributary in an area with an abundance of impervious surfaces, there would probably have been a brief rise in stream temperature prior to the cooldown. (United States Geological Survey image)

The daily oscillations in temperature reflect the influence of several heat flux processes. During the day, solar (shortwave) radiation and convection from summer air, especially those hot south winds, are largely responsible for the daily rises of about 5° F. Longwave radiation has a round-the-clock influence—adding heat to the stream during the day and mostly shedding it at night. Atmospheric exchange including evaporative cooling may help moderate the rise in stream temperatures during the day, and certainly plays a role in bringing them back down after sunset. Along its course this summer, the West Conewago Creek absorbed enough heat to render it a warmwater fishery in the area of the gauging station. The West Conewago is a shallow, low gradient stream over almost its entire course. Its waters move very slowly, thus extending their exposure time to radiated heat flux and reducing the benefit of cooling by atmospheric exchange. Fortunately for bass, catfish, and sunfish, these temperatures are in the ideal range for warmwater fishes to feed, grow, and reproduce—generally over 80° F, and ideally in the 70° to 85° F range. Coolwater fishes though, would not find this stream segment favorable. It was consistently above the 80° F maximum and the 60° to 70° F range preferred by these species. And coldwater fishes, well, they wouldn’t be caught dead in this stream segment. Wait, scratch that—the only way they would be caught in this segment is dead. No trouts or sculpins here.

The Codorus Creek drains primarily the carbonate valleys of York County to the south of the West Conewago watershed. This gauge station is located about a mile upstream from the creek’s mouth on the Susquehanna just below Haldeman Riffles. The graphic pattern is very similar to that of the West Conewago’s: daily heating and cooling cycles and a noticeable drop in stream temperature in early August caused by a day of thundershowers followed by the remnants of Hurricane Debby. (United States Geological Survey image)

Look closely and you’ll notice that although the temperature pattern on this chart closely resembles that of the West Conewago’s, the readings average about 5 degrees cooler. This may seem surprising when one realizes that the Codorus follows a channelized path through the heart of York City and its urbanized suburbs—a heat island of significance to a stream this size. Before that it passes through numerous impoundments where its waters are exposed to the full energy of the sun. The tempering factor for the Codorus is its baseflow. Despite draining a smaller watershed than its neighbor to the north, the Codorus’s baseflow (low flow between periods of rain) was 96 cubic feet per second on August 5th, nearly twice that of the West Conewago (51.1 cubic feet per second on August 5th). Thus, the incoming heat energy was distributed over a greater mass in the Codorus and had a reduced impact on its temperature. Though the Codorus is certainly a warmwater fishery in its lower reaches, coolwater and transitional fishes could probably inhabit its tributaries in segments located closer to groundwater sources without stress. Several streams in its upper reaches are in fact classified as trout-stocked fisheries.

This is a zoomed-in look at the previous graph showing the impact of a rainfall event on the water temperatures in Codorus Creek. Unlike the sharp declines accompanying the deluge of flood waters during the two events in early August, these lesser storms in late June generated just enough runoff to capture heat energy from impervious surfaces and warm the creek, temporarily breaking the daily heating/cooling cycle. Upstream in the immediate area of the runoff, the impact on the stream and/or its tributaries was probably much more dramatic, certainly raising temperatures into the nineties or above. (United States Geological Survey image)

Kreutz Creek drains a carbonate bedrock area of York County and flows parallel to the Lincoln Highway (US 30) to enter the Susquehanna at Wrightsville. The gauging station is about one mile upstream from the creek’s mouth. (United States Geological Survey image)

The Kreutz Creek gauge shows temperature patterns similar to those in the West Conewago and Codorus data sets, but notice the lower overall temperature trend and the flow. Kreutz Creek is a much smaller stream than the other two, with a flow averaging less than one tenth that of the West Conewago and about one twentieth of that in the Codorus. And most of the watershed is cropland or urban/suburban space. Yet, the stream remains below 80° F through most of the summer. The saving graces in Kreutz Creek are reduced exposure time and gradient. The waters of Kreutz Creek tumble their way through a small watershed to enter the Susquehanna within twenty-four hours, barely time to go through a single daily heating and cooling cycle. As a result, their is no chance for water to accumulate radiant and convective heat over multiple summer days. The daily oscillations in temperature are less amplified than we find in the previous streams—a swing of about three degrees compared to five. This indicates a better balance between heat flux processes that raise temperature and those that reduce it. Atmospheric exchange in the stream’s riffles, forest cover, and good hyporheic exchange along its course could all be tempering factors in Kreutz Creek. From a temperature perspective, Kreutz Creek provides suitable waters for coolwater fishes.

Muddy Creek drains portions of southern York County through rolling farmland and woodlots. There are no large impoundments or widespread urban impacts in the watershed, which may help explain its slightly lower temperature trends. (United States Geological Survey image)

Muddy Creek is a trout-stocked fishery, but it cannot sustain coldwater species through the summer heat. Though temperatures in Muddy Creek may be suitable for coolwater fishes, silt, nutrients, low dissolved oxygen, and other factors could easily render it strictly a warmwater fishery, inhabited by species tolerant of significant stream impairment.

Chiques Creek drains mostly limestone farmland in northwestern Lancaster County. The gauging station is located near the stream’s mouth on the Susquehanna at Chiques (Chickies) Rock. Oscillations in temperature again resemble the other waterways, but daily highs remain almost entirely below 80 degrees. (United States Geological Survey image)

A significant number of stream segments in the Chiques watershed have been rehabilitated to eliminate intrusion by grazing livestock, cropland runoff, and other sources of impairment. Through partnerships between a local group of watershed volunteers and landowners, one tributary, Donegal Creek, has seen riparian buffers, exclusion fencing, and other water quality and habitat improvements installed along nearly ever inch of its run from Donegal Springs through high-intensity farmland to its mouth on the main stem of the Chiques just above its confluence with the Susquehanna. The improved water quality parameters in the Donegal support native coldwater sculpins and an introduced population of reproducing Brown Trout. While coldwater habitat is limited to the Donegal, the main stem of the Chiques and its largest tributary, the Little Chiques Creek, both provide suitable temperatures for coolwater fishes.

Limestone Formation on Little Chiques Creek — Streams in the Chiques Creek and similar limestone watersheds often pass through areas with significant bedrock formations. Heat flux processes including groundwater input, hyporheic exchange, and streambed conductive exchange can have a greater influence on water temperature along these segments.

The Common Shiner (Luxilus cornutus), a fish tolerant of warmwater streams, prefers cool, clear waters for spawning. For protection from late-spring and summer heat, breeding males may seek a section of creek with a streambed inflow of limestone groundwater to defend as their nesting territory.

A closeup of the Chiques Creek graph showing what appears to be a little bump in temperature caused by surface runoff during a couple of late-May showers. Stream rehabilitation is an ongoing process and the pressures of land disturbances both old and new present challenges to those who make it their passion to fix the wrongs that have been inflicted upon our local waters. Even the exemplary Donegal Creek faces new threats from urbanization in one of its headwater areas several miles to the northwest of the historic springs. (United States Geological Survey image)

Conewago Creek (East) drains primarily Triassic redbed farmlands in Dauphin, Lancaster, and Lebanon Counties. Much of the headwaters area is forested but is experiencing an increasing rate of encroachment by housing and some commercial development. Conewago Creek (East) enters the Susquehanna on the east side of Conewago Falls at Three Mile Island. The watershed is equipped with three U.S.G.S. gauge stations capable of providing temperature data. This first one is located just over a mile upstream of the creek’s mouth. (United States Geological Survey image)

Despite its meander through and receipt of water from high-intensity farmland, the temperature of the lower Conewago (East) maxes out at about 85° F, making it ideal for warmwater fishes and even those species that are often considered coolwater transition fishes like introduced Smallmouth Bass, Rock Bass, Walleye, and native Margined Madtom. This survivable temperature is a testament to the naturally occurring and planted forest buffers along much of the stream’s course, particularly on its main stem. But the Conewago suffers serious baseflow problems compared to other streams we’ve looked at so far. Just prior to the early August storms, flow was well below 10 cubic feet per second for a drainage area of more than fifty square miles. While some of this reduced flow is the result of evaporation, much of it is anthropogenic in origin as the rate of groundwater removal continues to increase and a recent surge in stream withdraws for irrigation reaches its peak during the hottest days of summer.

Juvenile Rock Bass — A juvenile Rock Bass.

A closer look at the Conewago Creek (East) graphic shows the temperature drop associated with a series of thundershowers and the remnants of Hurricane Debby in early August. Despite the baseflow being below five cubic feet per second, the cooling effect of the downpours as measured in the area of the gauge was significant enough to overwhelm any heating of runoff that may have occurred as precipitation drained across hardened soils or man-made impervious surfaces. (United States Geological Survey image)

A little side note—the flow rate on the Conewago at the Falmouth gauge climbed to about 160 cubic feet per second as a result of the remnants of Hurricane Debby while the gauge on the West Conewago at Manchester skyrocketed to about 20,000 cubic feet per second. Although the West Conewago’s watershed (drainage area) is larger than that of the Conewago on the east shore, it’s larger only by a multiple of two or three, not 125. That’s a dramatic difference in rainfall!

The Bellaire monitoring station on Conewago Creek (East) is located on the stream’s main stem just downstream from the mouth of Little Conewago Creek, a tributary with its origins in farmland and woodlots. (United States Geological Survey image)

The temperatures at the Bellaire monitoring station, which is located upstream of the Conewago’s halfway point between its headwaters in Mount Gretna and its mouth, are quite comparable to those at the Falmouth gauge. Although a comparison between these two sets of data indicate a low net increase in heat absorption along the stream’s course between the two points, it also suggests sources of significant warming upstream in the areas between the Bellaire gauge and the headwaters.

Data from the gauge site on the Little Conewago Creek shows a temperature averaging about five degrees cooler than the gauge several miles downstream on the main stem of the Conewago at Bellaire. (United States Geological Survey image)

The waters of the Little Conewago are protected within planted riparian buffers and mature woodland along much of their course to the confluence with the Conewago’s main stem just upstream of Bellaire. This tributary certainly isn’t responsible for raising the temperature of the creek, but is instead probably helping to cool it with what little flow it has.

Juvenile Eastern Blacknose Dace (top) and a juvenile Longnose Dace. — A stream like the Little Conewago Creek with daily temperatures that remain mostly below 80 degrees and retreat to 75 degrees or less during the night can be suitable for coldwater transition fishes like these juvenile Eastern Blacknose Dace (top) and Longnose Dace.

Though mostly passing through natural and planted forest buffers above its confluence with the Little Conewago, the main stem’s critically low baseflow makes it particularly susceptible to heat flux processes that raise stream temperatures in segments within the two or three large agricultural properties where owners have opted not to participate in partnerships to rehabilitate the waterway. The headwaters area, while largely within Pennsylvania State Game Lands, is interspersed with growing residential communities where potable water is sourced from hundreds of private and community wells—every one of them removing groundwater and contributing to the diminishing baseflow of the creek. Some of that water is discharged into the stream after treatment at the two municipal sewer plants in the upper Conewago. This effluent can become quite warm during processing and may have significant thermal impact when the stream is at a reduced rate of flow. A sizeable headwaters lake is seasonally flooded for recreation in Mount Gretna. Such lakes can function as effective mid-day collectors of solar (shortwave) radiation that both warms the water and expedites atmospheric exchange.

Upper Conewago Creek (East) Watershed — The Conewago Creek (East) Watershed from the Bellaire U.S.G.S. Gauging Station (lower left) upstream to the headwaters in Mount Gretna. (United States Geological Survey image)

Though Conewago Creek (East) is classified as a trout-stocked fishery in its upper reaches in Lebanon County, its low baseflow and susceptibility to warming render it inhospitable to these coldwater fishes by late-spring/early summer.

Despite being considered a warmwater fish, the River Chub (Nocomis micropogon) will ascend streams like the Conewago to seek cooler, gravel-bottomed waters for spawning. Reduced baseflow has probably rendered the stream currently too small for this species on Pennsylvania State Game Lands in Colebrook where this specimen was photographed in 2018.

The Golden Shiner, another warmwater fish, often ascends streams to enter cooler water. Juvenile Golden Shiners like this one will move into shallower headwaters not only to seek reduced temperatures, but to escape large predatory fishes as well.

Irrigation using stream water. — Irrigation of agricultural fields using a large portion of the already diminished baseflow in the Conewago Creek (East) just downstream of the Bellaire gauging station. Despite millions of dollars in investment to rehabilitate this Susquehanna valley stream, the riparian buffers and other practices can have little effect when the creek gets sucked down to just a trickle. Low baseflow is a hard nut to crack. It’s best prevented, not corrected.

Hammer Creek, a trout-stocked fishery, originates, in part, within Triassic conglomerate in the Furnace Hills of Lebanon County, then flows north into the limestone Lebanon Valley where it picks up significant flow from other tributaries before working its way south back through the Furnace Hills into the limestone farmlands of Lancaster County. From there the stream merges with the Cocalico Creek, then the Conestoga River, and at last the Susquehanna. Note the tremendous daily temperature oscillations on this headwaters stream as it surges about 15 degrees each day before recovering back close to groundwater temperature by sunrise the next day. (United States Geological Survey image)

The removal of two water supply dams on the headwaters of Hammer Creek at Rexmont eliminated a large source of temperature fluctuation on the waterway, but did little to address the stream’s exposure to radiant and convective heat flux processes as it meanders largely unprotected out of the forest cover of Pennsylvania State Game Lands and through high-intensity farmlands in the Lebanon Valley. Moderating the temperature to a large degree is the influx of karst water from Buffalo Springs, located about two miles upstream from this gauging station, and other limestone springs that feed tributaries which enter the Hammer from the east and north. Despite the cold water, the impact of the stream’s nearly total exposure to radiative and other warming heat flux processes can readily be seen in the graphic. Though still a coldwater fishery by temperature standards, it is rather obvious that rapid heating and other forms of impairment await these waters as they continue flowing through segments with few best management practices in place for mitigating pollutants. By the time Hammer Creek passes back through the Furnace Hills and Pennsylvania State Game Lands, it is leaning toward classification as a coolwater fishery with significant accumulations of sediment and nutrients. But this creek has a lot going for it—mainly, sources of cold water. A core group of enthusiastic landowners could begin implementing the best management practices and undertaking the necessary water quality improvement projects that could turn this stream around and make it a coldwater treasure. An organized effort is currently underway to do just that. Visit Trout Unlimited’s Don Fritchey Chapter and Donegal Chapter to learn more. Better yet, join them as a volunteer or cooperating landowner!

Male Creek Chub — The male Creek Chub, one of our coolwater fishes, develops head tubercles and becomes flushed with color during spawning season. Hammer Creek not only provides a home for the Creek Chub, its cold headwaters provide refuge for a population of native Brook Trout too.

Like no other example we’ve looked at so far, this closeup of the Hammer Creek graphic shows temperature bumps correlating with the stormwater runoff from early August’s rains. Because the stream flow is small and the precipitation rate was not as great at this location, the effect of heat flux from runoff is more readily apparent. (United States Geological Survey image)

Brook Trout adult and juvenile. (United States Fish and Wildlife Service image by Ryan Hagerty)

For coldwater fishes, the thousands of years since the most recent glacial maximum have seen their range slowly contract from nearly the entirety of the once much larger Susquehanna watershed to the headwaters of only our most pristine streams. Through no fault of their own, they had the misfortune of bad timing—humans arrived and found coldwater streams and the groundwater that feeds them to their liking. Some of the later arrivals even built their houses right on top of the best-flowing springs. Today, populations of these fishes in the region we presently call the Lower Susquehanna River Watershed are seriously disconnected and the prospect for survival of these species here is not good. Stream rehabilitation, groundwater management, and better civil planning and land/water stewardship are the only way coldwater fishes, and very possibly coolwater fishes as well, will survive. For some streams like Hammer Creek, it’s not too late to make spectacular things happen. It mostly requires a cadre of citizens, local government, project specialists, and especially stakeholders to step up and be willing to remain focused upon project goals so that the many years of work required to turn a failing stream around can lead to success.

Riparian Buffer — Riparian buffers with fences to exclude livestock can immediately begin improving water quality. With establishment of such vegetative buffers, the effects of stressors that otherwise eliminate coldwater and coolwater fishes from these segments will begin to diminish.

You’re probably glad this look at heat flux processes in streams has at last come to an end. That’s good, because we’ve got a lot of work to do.

You Don't Have Time to Mow, We've Got Work to Do — Add one more benefit to the wildflower meadow, it infiltrates stormwater to recharge the aquifer much better than mowed grass. And another related plus, it reduces runoff and its thermal pollution. Besides, you don’t have time to mow grass, because we have work to do!

Our native coldwater fishes including the Potomac Sculpin will survive only if we protect and expand the scattered few habitats where they have taken refuge. They have no choice but to live in these seriously threatened places, but we do. So let’s give ’em some space. How ’bout it? (United States Fish and Wildlife Service image by Ryan Hagerty)

August 6, 2024August 6, 2024

Photo of the Day

Berries and foliage of the Black Gum, also known as the Black Tupelo. — In the forests and woodlots of the lower Susquehanna watershed, the Black Gum, also known as the Black Tupelo, is always the first of our native trees to pop with autumn color. Its crimson leaves glow in an otherwise green landscape to help lure hungry birds including waxwings, American Robins, and Pileated Woodpeckers to its soon-to-be-ripe berries. The seeds contained in the fruit are thus well-distributed as a by-product of this avian feast. While not among the most massive of our trees, the Black Gum can live for over five hundred years, an age not exceeded by any other non-clonal indigenous flowering plant.

July 27, 2024July 27, 2024

Wildflower Meadow Update

Do you recall our “Photo of the Day” from seven months ago…

Riverfront Park Wildflower Meadow — Here’s something to look forward to in the new year. The good citizens of East Donegal Township in Lancaster County have partnered with Alliance for the Chesapeake Bay to establish an extensive wildflower meadow on what had been a mowed field of turf grass at Riverside Park in the Susquehanna floodplain near Marietta. As the photo shows, the lawn plants have been eliminated in preparation for seeding with a diverse assortment of native grasses and wildflowers to provide habitat for birds and pollinators including butterflies, bees, and other insects. Once established, the meadow’s extensive vegetative growth will help reduce stormwater runoff by better infiltrating rainfall to recharge the aquifer. During flood events, the plantings will provide soil stabilization and increase the ability of the acreage to uptake nutrients, thus reducing the negative impact of major storms on the quality of water in the river and in Chesapeake Bay. Check the project’s progress by stopping by from time to time in 2024!

Well, here’s what that site looks like today…

And there are pollinating insects galore, most notably butterflies…

A Cabbage White collecting nectar on Blue Vervain.

A Sulphur — A Clouded or Orange Sulphur among the grasses in the meadow.

The Least Skipper is our tiniest butterfly.

The Little Glassywing (Vernia verna), this one feeding on vervain nectar, deposits its eggs on Purpletop grass, which then functions as the host plant for this butterfly’s larvae.

A Summer Azure (Celastrina neglecta) feeding on the nectar of a Black-eyed Susan.

A Common Buckeye on a “Gloriosa Daisy”, a showy, large-flowered cultivar of Black-eyed Susan.

An Eastern Tiger Swallowtail on Blue Vervain. Nearby Yellow (Tulip) Poplars and other trees serve as host plants for this butterfly’s larvae.

Black Morph Eastern Tiger Swallowtail — The black morph of the Eastern Tiger Swallowtail shows subdued shading in the wings that closely resembles the brilliant color patterns of the more familiar yellow form.

The Sachem, this one visiting a Black-eyed Susan, is a variable species with a range that normally lies south of the 40th parallel, the line of latitude that intersects the Susquehanna in the area of the Conejohela Flats at Washington Boro.

A Sachem visiting the blooms of Oxeye sunflower. During recent weeks, scorching winds from the south and southwest have transported an abundance of these vagrant skippers into the lower Susquehanna valley and beyond.

Here and there among the meadow’s plantings we noticed one of our favorites starting to flower, the Partridge Pea.

Partridge Pea happens to be a host plant for another vagrant from the south, the big, lime-yellow Cloudless Sulphur. We saw at least half a dozen patrolling the meadow.

The stars of the show are the Zebra Swallowtails, gorgeous butterflies that rely on stands of native Common Paw-paw trees in the river floodplain to host their eggs and larvae.

Why on earth would anyone waste their time, energy, and money mowing grass when they could have this? Won’t you please consider committing graminicide this fall? That’s right, kill that lawn—at least the majority of it. Then visit the Ernst Seed website, buy some “Native Northeast Wildlflower Mix” and/or other blends, and get your meadow planted in time for the 2025 growing season. Just think of all the new kinds of native plants and animals you’ll be seeing. It could change your life as well as theirs.

A Snowberry Clearwing (Hemaris diffinis), more commonly known as a hummingbird moth, visits the flowers of Blue Vervain in the Riverfront Park wildflower meadow.

June 14, 2024

Photo of the Day

A nectar-seeking Honey Bee visits the fragrant flowers of a Common Milkweed.

June 10, 2024

Photo of the Day

The bright-red flower buds of the Ninebark (Physocarpus opulifolius) precede clusters of white blooms that will, in coming weeks, attract a variety of butterflies and other pollinators to this indigenous shrub. Its peeling bark and colorful deciduous leaves attract interest throughout the year. In the lower Susquehanna watershed, Ninebark is most frequently found growing along stream banks. It will often thrive on steep slopes with moist soils, so is useful as an erosion control species as well. To add it to your refuge’s landscape, look for it at nurseries that stock native plants. Once there, you’ll find a variety of cultivars that are sure to satisfy even the fussiest of gardeners.

May 25, 2024May 26, 2024

Photo of the Day

February 22, 2024February 24, 2024

Time to Order Trees and Shrubs for Spring

It’s that time of year. Your local county conservation district is taking orders for their annual tree sale and it’s a deal that can’t be beat. Order now for pickup in April.

The prices are a bargain and the selection includes the varieties you need to improve wildlife habitat and water quality on your property. For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue). And don’t forget to order packs of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local native birds. They’re only $12.00 for a bundle of 10.

Mature Trees in a Suburban Neighborhood — It’s the most desirable block in town, not because the houses are any different from others built during the post-war years of the mid-twentieth century, but because the first owners of these domiciles had the good taste and foresight to plant long-lived trees on their lots, the majority of them native species. Pin Oak, Northern Red Oak, Yellow Poplar, Flowering Dogwood (Cornus florida), Eastern Red Cedar, Eastern White Pine, Eastern Hemlock, Norway Spruce, and American Holly dominate the landscape and create excellent habitat for birds and other wildlife. These 75-year-old plantings provide an abundance of shade in summer and thermal stability in winter, making it a “cool” place to live or take a stroll at any time of the year.

Cumberland County Conservation District Annual Tree Seedling Sale—

Orders due by: Friday, March 22, 2024

Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024

Cumberland County Conservation District is taking orders for Common Winterberry, the ideal small shrub for wet soil anywhere on your property. To get berries, you’ll need both males and females, so buy a bunch and plant them in a clump or scattered group.

To live for a century or more like this towering giant, a Pin Oak needs to grow in well-drained soils with adequate moisture. These sturdy shade providers do well along streams and on low ground receiving clean runoff from hillsides, roofs, streets, and parking areas. As they age, Pin Oaks can fail to thrive and may become vulnerable to disease in locations where rainfall is not adequately infiltrated into the soil. Therefore, in drier areas such as raised ground or slopes, avoid the Pin Oak and select the more durable Northern Red Oak for planting. This year, Pin Oaks are available from the Cumberland and Lancaster County Conservation Districts, while Dauphin, Lancaster, Lebanon, and York Counties are taking orders for Northern Red Oaks.

The Cumberland County Conservation District is again offering a “Showy Northeast Native Wildflower and Grass Mix” for seeding your own pollinator meadow or garden. It consists of more than twenty species including this perennial favorite, Purple Coneflower.

Dauphin County Conservation District Seedling Sale—

Orders due by: Monday, March 18, 2024

Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024

The Eastern Redbud is small tree native to our forest edges, particularly in areas of the Piedmont Province with Triassic geology (Furnace Hills, Conewago Hills, Gettysburg/Hammer Creek Formations, etc.) Also known as the Judas Tree, the redbud’s brilliant flowers are followed by heart-shaped leaves. As seen here, it is suitable for planting near houses and other buildings. Eastern Redbud seedlings are being offered through tree sales in Dauphin, Cumberland, and Lancaster Counties.

Lancaster County Annual Tree Seedling Sale—

Orders due by: Friday, March 8, 2024

Pickup on: Friday, April 12, 2024

The Yellow Poplar, often called Tuliptree or Tulip Poplar for its showy flowers, is a sturdy, fast-growing deciduous tree native to forests throughout the Lower Susquehanna River Watershed. Its pole-straight growth habit in shady woodlands becomes more spreading and picturesque when the plant is grown as a specimen or shade tree in an urban or suburban setting. The Yellow Poplar can live for hundreds of years and is a host plant for the Eastern Tiger Swallowtail butterfly. It is available this year from the Lancaster County Conservation District.

The American Sweetgum, also known as Sweet Gum, is a large, long-lived tree adorned with a mix of vibrant colors in autumn.

American Goldfinches and Pine Siskin on Sweet Gum — Ever wonder where all the American Goldfinches and particularly the Pine Siskins go after passing through our region in fall? Well, many are headed to the lowland forests of the Atlantic Coastal Plain where they feed on an abundance of seeds contained in spiky American Sweetgum fruits. In the Piedmont and Ridge and Valley Provinces of the Lower Susquehanna River Watershed, American Sweetgum transplants can provide enough sustenance to sometimes lure our friendly finches into lingering through the winter.

Sweet Gum in a Beaver Pond — The American Sweetgum is a versatile tree. It can be planted on upland sites as well as in wet ground along streams, lakes, and rivers. In the beaver pond seen here it is the dominate tree species. This year, you can buy the American Sweetgum from the Lancaster County Conservation District.

Silky Dogwood Stream Buffer — “Red-twig Dogwoods” make ideal mass plantings for streamside buffers and remain showy through winter, even on a gloomy day. They not only mitigate nutrient and sediment pollution, they provide excellent food and cover for birds and other wildlife. Both Silky and Red-osier Dogwoods are available for sale through the Lancaster County Conservation District as part of their special multi-species offers, the former is included in its “Beauty Pack” and the latter in its “Wildlife Pack”. The similar Gray Dogwood (Cornus racemosa) is being offered for sale by the York County Conservation District.

Lebanon County Conservation District Tree and Plant Sale—

Orders due by: Friday, March 8, 2024

Pickup on: Friday, April 19, 2024

Common Pawpaw flower — The unique maroon flowers of the Common Pawpaw produce banana-like fruits in summer. These small native trees grow best in damp, well-drained soils on slopes along waterways, where they often form clonal understory patches. To get fruit, plant a small grove to increase the probability of pollination. The Common Pawpaw is a host plant for the Zebra Swallowtail butterfly. It is available through both the Lebanon and Lancaster County sales.

The Eastern Red Cedar provides excellent food, cover, and nesting sites for numerous songbirds. Planted in clumps of dozens or groves of hundreds of trees, they can provide winter shelter for larger animals including deer and owls. The Eastern Red Cedar is being offered for purchase through both the Lebanon and Lancaster County Conservation Districts.

Hybrid American Chestnut — Care to try your hand at raising some chestnuts? Lebanon County Conservation District has hybrid American Chestnut seedlings for sale.

Perry County Conservation District Tree Sale—

Orders due by: Sunday, March 24, 2024

Pickup on: Thursday, April 11, 2024

Pollinator Garden — In addition to a selection of trees and shrubs, the Perry County Conservation District is again selling wildflower seed mixes for starting your own pollinator meadow or garden. For 2024, they have both a “Northeast Perennials and Annuals Mix” and a “Butterfly and Hummingbird Seed Mix” available. Give them a try so you can give up the mower!

Again this year, Perry County is offering bluebird nest boxes for sale. The price?—just $12.00.

Eastern Bluebird — Wait, what?,…twelve bucks,…that’s cheaper than renting!

York County Conservation District Seedling Sale—

Orders due by: Friday, March 15, 2024

Pickup on: Thursday, April 11, 2024

Buttonbush flower — The Buttonbush, a shrub of wet soils, produces a cosmic-looking flower. It grows well in wetlands, along streams, and in rain gardens. Buttonbush seedlings are for sale from both the York and Lancaster County Conservation Districts.

To get your deciduous trees like gums, maples, oaks, birches, and poplars off to a safe start, conservation district tree sales in Cumberland, Dauphin, Lancaster, and Perry Counties are offering protective tree shelters. Consider purchasing these plastic tubes and supporting stakes for each of your hardwoods, especially if you have hungry deer in your neighborhood.

Deciduous Tree Planting Protected by Shelters — Tree shelters protect newly transplanted seedlings from browsing deer, klutzy hikers, visually impaired mower operators, and other hazards.

There you have it. Be sure to check out each tree sale’s web page to find the selections you like, then get your order placed. The deadlines will be here before you know it and you wouldn’t want to miss values like these!

February 15, 2024February 15, 2024

Birds Beginning to Wander

Since Tuesday’s snow storm, the susquehannawiildlife.net headquarters garden continues to bustle with bird activity.

Our Northern Mockingbird remains ever vigilant in its attempts to discourage American Robins and Eastern Bluebirds from feeding on the berry crop. Slowly, the latter species are winning the contest.

Carolina Chickadee feeding on sunflower seed. — A Carolina Chickadee carefully dissects a sunflower seed to snack on the nutritious kernel.

American Robin eating Common Winterberry fruits. — Getting energized for a big move north, the robins keep on gulping berries.

Today, there arrived three species of birds we haven’t seen here since autumn. These birds are, at the very least, beginning to wander in search of food. Then too, these may be individuals creeping slowly north to secure an advantage over later migrants by being the first to establish territories on the most favorable nesting grounds.

This Song Sparrow is the first we’ve seen in the garden since sometime last fall. Is it working its way north or did it just come to town in search of food?

Northern Flickers regularly spend the winter in small numbers in the Lower Susquehanna River Watershed. This is the first one we’ve had visit the garden since late last autumn.

Fish Crows feeding on Eastern Red Cedar berries. — Fish Crows, seen here feeding on the fruits adorning an Eastern Red Cedar, have returned after being absent in our neighborhood since November. In coming weeks, both they and the more numerous American Crows that remained through winter will begin constructing nests in nearby trees.

They say the early bird gets the worm. More importantly, it gets the most favorable nesting spot. What does the early birder get? He or she gets out of the house and enjoys the action as winter dissolves into the miracle of spring. Do make time to go afield and marvel a bit, won’t you? See you there!

February 13, 2024

Robins in a Snowstorm

In mid-February each year, large numbers of American Robins descend upon the susquehannawildlife.net headquarters garden to feast on the ripe fruits that adorn several species of our native shrubs and trees. This morning’s wet snowfall provided the needed motivation for these birds and others to make today the big day for the annual feeding frenzy.

American Robin feeding on Common Winterberry — The fruits of Common Winterberry are always a favorite of visiting robins.

A pair of Carolina Chickadees established a family in the garden during the spring of 2023. At least five of the birds still stop by on a daily basis.

A Blue Jay peeks out from the cover of the Eastern Hemlocks.

Our Carolina Wrens sing throughout the winter,…

…but today we noticed that this Mourning Dove has begun softly cooing to charm a mate…

…and the male House Finches are warbling away with the sounds of spring.

Female Eastern Bluebird — With the local mockingbird busily harassing robins, our Eastern Bluebirds went unmolested long enough to stop by…

Bluebird Feeder — …for some raisins from their enclosed feeder.

Male Eastern Bluebird — A showy male Eastern Bluebird on a snowy day in the garden. Spring must be just around the corner!

January 3, 2024March 30, 2024

Birds of the Sunny Grasslands

With the earth at perihelion (its closest approach to the sun) and with our home star just 27 degrees above the horizon at midday, bright low-angle light offered the perfect opportunity for doing some wildlife photography today. We visited a couple of grasslands managed by the Pennsylvania Game Commission to see what we could find…

Grasslands and Hedgerows — On this State Game Lands parcel, prescribed fire is used to maintain a mix of grasslands and brushy early successional growth. In nearby areas, both controlled fire and mechanical cutting are used to remove invasive species from hedgerows and the understory of woodlots. Fire tolerant native species then have an opportunity to recolonize the forest and improve wildlife habitat. This management method also reduces the fuel load in areas with the potential for uncontrolled wildfires.

The sun-dried fruits of a Common Persimmon tree found growing in a hedgerow. — The sun-dried fruits of a native Common Persimmon tree found growing in a hedgerow.

Savanna-like Grasslands — Just one year ago, mechanical removal of invasive trees and shrubs (including Multiflora Rose) on this State Game Land was followed by a prescribed fire to create this savanna-like grassland.

White-throated Sparrows were also abundant, but prefer the tangles and shrubs of the thickets.

In grasses and tangles on wetter ground, about a dozen Swamp Sparrows were discovered.

Dark-eyed Juncos were encountered only in the vicinity of trees and large shrubs.

A tiny bird, even when compared to a sparrow, the Winter Wren often provides the observer with just a brief glimpse before darting away into the cover of a thicket.

Standing Clump of Timber — Within grasslands, scattered stands of live and dead timber can provide valuable habitat for many species of animals.

A "snag" with an excavated nest cavity. — Woodpeckers and other cavity-nesting birds rely upon an abundance of “snags” (standing dead trees) for breeding sites.

A Yellow-bellied Sapsucker soaks up some sun.

Just as things were really getting fun, some late afternoon clouds arrived to dim the already fading daylight. Just then, this Northern Harrier made a couple of low passes in search of mice and voles hidden in the grasses.

December 27, 2023December 27, 2023

Photo of the Day

Wildflower Meadow Project underway at East Donegal Riverfront Park — Here’s something to look forward to in the new year. The good citizens of East Donegal Township in Lancaster County have partnered with Alliance for the Chesapeake Bay to establish an extensive wildflower meadow on what had been a mowed field of turf grass at Riverside Park in the Susquehanna floodplain near Marietta. As the photo shows, the lawn plants have been eliminated in preparation for seeding with a diverse assortment of native grasses and wildflowers to provide habitat for birds and pollinators including butterflies, bees, and other insects. Once established, the meadow’s extensive vegetative growth will help reduce stormwater runoff by better infiltrating rainfall to recharge the aquifer. During flood events, the plantings will provide soil stabilization and increase the ability of the acreage to uptake nutrients, thus reducing the negative impact of major storms on the quality of water in the river and in Chesapeake Bay. Check the project’s progress by stopping by from time to time in 2024!

November 23, 2023

Time to Eat

A glimpse of the rowdy guests crowding the Thanksgiving Day dinner table at susquehannawildlife.net headquarters…

A male White-breasted Nuthatch visits a peanut feeder…

A male Downy Woodpecker gets a bill full of suet.

November 14, 2023November 15, 2023

Sparrows in the Thicket

As the annual autumn songbird migration begins to reach its end, native sparrows can be found concentrating in fallow fields, early successional thickets, and brushy margins along forest edges throughout the Lower Susquehanna River Watershed.

Brushy Thicket — A streamside thicket composed of seed-producing grasses and wildflowers as well as fruit-bearing shrubs and vines can be ideal habitat for migrating and wintering native sparrows.

Visit native sparrow habitat during mid-to-late November and you have a good chance of seeing these species and more…

The Eastern Towhee is a large native sparrow most often found in early successional growth near woodlands. Look for them in utility right-of-ways.

If you’re lucky enough to live where non-native House Sparrows won’t overrun your bird feeders, you can offer white millet as a supplement to the wild foods these beautiful sparrows might find in your garden sanctuary. Give it a try!

October 21, 2023October 25, 2023

Fall Foliage at the Peak of Color

Have you noticed? Foliage throughout the Lower Susquehanna River Watershed has been painted with the brilliant colors of autumn, so now is the time to get out there and have a look. Why not make a collection? You can pick up an inexpensive scrap book or photo album at the craft store to press and label the varieties you find. Uncle Tyler Dyer is already busy adding to the project he assembled last year. You can use his exhibit as a reference for identifying and learning a little bit more about the leaves you find.

Uncle Tyler Dyer's Leaf Collection — To identify the leaves you discover, click this image.

Leaf Collection Mounted in a Photo Album — The editor’s 1995 collection of leaves from the Susquehanna River floodplain at Conewago Falls.

October 11, 2023

Photo of the Day

A Monarch butterfly takes a break from its southbound migration to energize on nectar from a goldenrod flower.

October 10, 2023October 10, 2023

Photo of the Day

October 5, 2023October 5, 2023

Late Season Ruby-throated Hummingbirds…Again

Last October 3rd, a late-season Ruby-throated Hummingbird stopped by the garden at susquehannawildlife.net headquarters to take shelter from a rainy autumn storm. It was so raw and chilly that we felt compelled to do something we don’t normally do—put out the sugar water feeder to supplement the nectar produced by our fall-flowering plants. After several days of constant visits to the feeder and the flowers, our lingering hummer resumed its southbound journey on October 7th.

Fast forward to this afternoon and what do you know, at least two migrating hummingbirds have stopped by to visit the flowers in our garden. This year, we have an exceptional abundance of blooms on some of their favorite plants. In the ponds, aquatic Pickerelweed is topped with purple spikes and we still have bright orange tubular flowers on one of our Trumpet Vines—a full two to three months later than usual.

Ruby-throated Hummingbird at Mexican Cigar — As is typically the case, Ruby-throated Hummingbirds quickly gravitate toward the tubular flowers of our Cuphea ignea, the Mexican Cigar. They find these showy plants to be absolutely irresistible.

Ruby-throated Hummingbird at Bat-faced Cuphea — We’re trying some new cultivars of Cuphea to see how they do. As this composite image shows, the hummingbirds won’t let our Bat-faced Cuphea (Cuphea llavea) alone. It’s another plant native to Mexico and Central America, right where some of our hummingbirds spend the winter.

Remember, keep those feeders clean and the provisions fresh! You’ll be glad you did.

October 5, 2023March 30, 2024

Hymanoptera: A Look at Some Bees, Wasps, Hornets, and Ants

What’s all this buzz about bees? And what’s a hymanopteran? Well, let’s see.

Hymanoptera—our bees, wasps, hornets and ants—are generally considered to be our most evolved insects. Some form complex social colonies. Others lead solitary lives. Many are essential pollinators of flowering plants, including cultivars that provide food for people around the world. There are those with stingers for disabling prey and defending themselves and their nests. And then there are those without stingers. The predatory species are frequently regarded to be the most significant biological controls of the insects that might otherwise become destructive pests. The vast majority of the Hymanoptera show no aggression toward humans, a demeanor that is seldom reciprocated.

Late summer and early autumn is a critical time for the Hymanoptera. Most species are at their peak of abundance during this time of year, but many of the adult insects face certain death with the coming of freezing weather. Those that will perish are busy, either individually or as members of a colony, creating shelter and gathering food to nourish the larvae that will repopulate the environs with a new generation of adults next year. Without abundant sources of protein and carbohydrates, these efforts can quickly fail. Protein is stored for use by the larval insects upon hatching from their eggs. Because the eggs are typically deposited in a cell directly upon the cache of protein, the larvae can begin feeding and growing immediately. To provide energy for collecting protein and nesting materials, and in some cases excavating nest chambers, Hymanoptera seek out sources of carbohydrates. Species that remain active during cold weather must store up enough of a carbohydrate reserve to make it through the winter. Honey Bees make honey for this purpose. As you are about to see, members of this suborder rely predominately upon pollen or insect prey for protein, and upon nectar and/or honeydew for carbohydrates.

We’ve assembled here a collection of images and some short commentary describing nearly two dozen kinds of Hymanoptera found in the Lower Susquehanna River Watershed, the majority photographed as they busily collected provisions during recent weeks. Let’s see what some of these fascinating hymanopterans are up to…

SOLITARY WASPS

A Great Black Wasp on goldenrod (Solidago species). Like other solitary wasps, a female Great Black Wasp will sting and paralyze a host insect upon which she’ll deposit her eggs. After hatching, the larvae will begin consuming the host’s body as a source of protein. The parasitized insects are often katydids or grasshoppers.

A Great Black Wasp. — A Great Black Wasp feeding on nectar, a source of carbohydrates. Unlike social bees and wasps, solitary wasps are equipped with a stinger solely used for immobilizing prey, not defending a nest. They are therefore quite docile and pose little threat to humans.

A Great Black Wasp powdered with pollen. Hymanopterans that gather nectar and/or pollen are tremendously important pollinators of hundreds of species of plants.

Thread-waisted Wasp — A female Thread-waisted wasp (Ammophilia species, probably A. nigricans) drags a paralyzed caterpillar to her excavated nest where she’ll deposit an egg on the body. After hatching, the larval wasp will feed on the disabled caterpillar. The protein will enable the larvae to grow, pupate, and later emerge as an adult wasp.

The female Eastern Cicada Killer (Sphecius speciosus) excavates an underground nest with branch tunnels connecting a dozen chambers or more. As the common name suggests, the female wasp paralyzes a cicada, then makes a strenuous effort to fly and drag it back to the nest for placement in a cell. Each male wasp egg is deposited upon just one immobilized cicada, but a female egg is provided with a cache of several cicadas to provide adequate protein for growth to a larger size. Nest cells are sealed with soil, then the larvae hatch in just a couple of days. Within about two weeks, they have consumed the cicada protein and are fully grown. Wrapped in a cocoon, they spend the winter in the nest, then pupate in the spring before emerging as a new generation of adults.

The Black-and-yellow Mud Dauber (Sceliphron caementarium) builds a mud-ball nest within which it packs paralyzed spiders to function as a source of protein for its larvae.

Pipe Organ Mud Dauber Nest — The Pipe Organ Mud Dauber builds this elaborate nest in which their eggs and paralyzed spiders are deposited in cells sealed with mud partitions. After consuming the spiders, the larvae pupate, overwinter, then emerge from their cells as adults during the following spring. To escape the protection of the nest, the new generation of adults bore through the mud walls. Adult Pipe Organ Mud Daubers resemble the Great Black Wasp, but have a white or yellow distal segment on their rear legs resembling a pair of light-colored socks.

Scavenger Flies in Pipe Organ Mud Dauber Nest — A closeup of the previous image with the lengths of the nest tubes compressed to show four scavenger flies (Miltogramminae), possibly two species, that have invaded this Pipe Organ Mud Dauber nest. Scavenger flies are kleptoparasites that victimize various solitary bees and wasps, depositing larvae directly into the host species’ nest cells to consume the protein cache stored therein.

CUCKOO WASPS

SWEAT BEES

A Sweat Bee (Lasioglossum species). — A Sweat Bee (Lasioglossum species) collecting nectar and pollen on White Snakeroot (Ageratina altissima).

An Augochlorine Green Sweat Bee (Augochlorini). — A female Augochlorine Green Sweat Bee (Augochlorini) collecting nectar on White Snakeroot. Notice the pollen “baskets” on the rear leg.

LEAFCUTTER AND MASON BEES

A Leafcutter Bee (Megachile species). Like Mason Bees, female Mason Bees deposit each of their eggs on a “pollen loaf” within an individual cell inside a preexisting tunnel-like cavity in wood, stone, or in the ground. Unlike Mason Bees, female Leafcutter Bees cut a circular piece of leaf to create each of the cells in their nest. After hatching, the larval bee feeds on the pollen loaf, pupates, then emerges from the shelter of the nest to start a new generation, usually during the following year.

A Leafcutter Bee (Megachile species). — A Leafcutter Bee (Megachile species) visiting Wild Bergamot. Female Leafcutter and Mason Bees lack pollen “baskets” on their rear legs but instead have pollen “brushes” on the underside of the abdomen to gather the protein they need to create a “pollen loaf” for each nest cell.

A Mason Bee (Osmia species) emerging from a nest cell in spring. — A Mason Bee (Osmia species) emerging from a nest in spring. Mason Bees create nesting cells within preexisting cavities in wood, stone, and other other supporting structures. Within the nest cavity, each egg is deposited atop a cache of pollen and nectar, a pollen loaf, then enclosed behind a partition of mud. The female Mason Bee will usually repeat this process until an entire cavity is filled with cells. During the following spring, a new generation of adult Mason Bees digs its way through the cell walls to emerge and repeat the process. These bees readily use paper straws or holes drilled in blocks of wood for nesting.

A mason bee nest box with holes drilled into blocks of wood.

Parasitized Mason Bee Nest — Mason Bees seal each cell and the outer end of their nest cavity with mud. These outer nest cells can been parasitized by a variety of wasps. Here, the outer cell of a Mason Bee nest has been victimized by a tiny chalcid wasp (looks like another one to the lower left). Several species of female chalcid wasps (native Monodontomerus species or non-native Pteromalus venustus) enlarge weak points in the outer partition of a mason bee nest, then sting and paralyze the larval bee inside before depositing their eggs. Within the cell. the wasp larvae consume the larval Mason Bee and the “pollen loaf” provided for its growth. These same parasitic wasps prey upon Leafcutter Bees as well.

BUMBLE BEES, CARPENTER BEES, HONEY BEES, AND DIGGER BEES

A Common Eastern Bumble Bee (Bombus impatiens) collecting nectar and pollen on goldenrod. Bumble bees are our sole native group of social bees. Their wax nests are built in a burrow or other shelter. The eggs are deposited in cells along with a supply of pollen for nourishing the larvae upon hatching. Honey is stored in “honey pots” within the nest. New queens are produced along with male bees during the late-summer and fall. Only the new generation of fertilized queens survive the winter to lay eggs and produce workers to construct a new nest.

A pair of Common Eastern Bumble Bees collecting nectar and becoming dusted with pollen. Their fuzzy coats and semi-warm-blooded metabolism allows them to be active in cooler weather than is tolerated by other bees.

A Common Eastern Bumble Bee pollinating a Great Rhododendron flower. — Flowering plants including the Great Rhododendron find success attracting pollinators to their reproductive blossoms by offering carbohydrate-rich nectar to insects like this Eastern Bumble Bee. The yellow spots on the flower’s upper petal help to guide visitors toward their sweet treat.

An Eastern Carpenter Bee feeding on goldenrod nectar. Compare the almost hairless abdomen to that of the bumble bees. Carpenter bees are semi-social insects. Females lay their eggs in cells within galleries bored into wood. These nests are completed with great precision, avoiding creation of any second entrance by mistakenly breaching the outer surface of the excavated wood. Each egg/larvae is provided with a supply of protein-rich pollen. Males often hover outside their mate’s nest to prevent competing males from entering the area.

A Honey Bee visiting goldenrod alongside Common Eastern Bumble Bees. — A worker Honey Bee, a female member of a sisterhood of foragers from a nearby hive, visits goldenrod alongside Common Eastern Bumble Bees. Honey Bees were brought to North America during the 1620s, the earliest years of the trans-Atlantic migration of European colonists, to pollinate cultivated plants and to provide a reliable source of honey and beeswax. Within the Honey Bee’s social structure, the queen of each hive lays the eggs to produce the female worker bees. Once each year, male drones are produced along with a new generation of queens.

Honey Bee Hive — In nature, Honey Bees build hives in tree cavities. Recently, this colony constructed a hive in a screech owl nest box at susquehannawildlife.net headquarters. To provide protein for the hatching larvae, worker bees collect pollen and deposit it within the hexagonal cells of the vertically aligned beeswax combs. After an egg is deposited upon the pollen cache, each cell is sealed with more beeswax. Young females tend these nest combs before maturing and becoming foraging worker bees.

Bee Hive Display — In apiculture, Honey Bees are raised in man-made hives. This Pennsylvania Association of Beekeepers display gives visitors to the Pennsylvania Farm Show in Harrisburg a look at the inner workings of a live bee hive. Nectar collected by worker bees is turned into honey to provide the supply of carbohydrates needed to fuel the colony through the winter. Note the honeycombs on the glass.

A possible Small Carpenter Bee Ceratina species). — A possible Small Carpenter Bee (Ceratina species) visiting White Snakeroot. Small Carpenter Bees nest inside hollow stems and twigs. Some species are eusocial, with a queen’s daughters and sisters sharing responsibility for finding food and rearing the young. Females overwinter inside a one of the excavated stems and begin a new nest there in the spring.

A Digger Bee (possibly Melissodes species). — A Digger Bee (possibly Melissodes species) with “pollen baskets” full of pollen collected from nearby flowers. Digger Bees in the genus Melissodes are often known as the Long-horned Bees. These social insects excavate underground nests and many species practice communal living.

SCOLIID WASPS

The Two-spotted Scoliid Wasp (Scolia dubia), also known as the Blue-winged Scoliid Wasp, is most frequently observed feeding on nectar. Scoliid wasps are solitary nesters, though they may assemble into groups while visiting flowers. They often ignore the presence of humans and are seldom disturbed by their presence. Females seek out the burrowing grubs of beetles including the Green June Bug (Cotinis nitida) and possibly the Japanese Beetle. After stinging a grub to paralyze it, the wasp will deposit her egg on its body, then bury it. Upon hatching, the larval wasp will feed on the grub for nourishment as it grows.

June Bugs eating watermelon. — Don’t like having your watermelon overrun by Green June Bugs while you’re eating? Then you ought to go out of your way to be nice to the Two-spotted Scoliid Wasp.

The Double-banded Scoliid (Scolia bicincta) parasitizes beetle larvae as hosts for its larvae. For carbohydrates it relishes flower nectar.

PAPER WASPS

A Northern Paper Wasp (Polistes fuscatus). Paper wasps prey upon numerous garden pests, particularly caterpillars, to collect protein. Though they are social insects equipped with stingers to subdue their victims and defend their nests, paper wasps are surprisingly docile.

The Northern Paper Wasp — A Northern Paper Wasp (Polistes fuscatus) feeding on nectar from a goldenrod flower.

A Northern Paper Wasp harvesting wood pulp from the side of a mason bee nest box at susquehannawildlife.net headquarters. The pulp is chewed in the wasp’s saliva to create the paper used to construct the colony’s open-cell nest.

Guinea Paper Wasps (Polistes exclamans) at their nest. — Common Paper Wasps (Polistes exclamans), also known as Guinea Paper Wasps, at their open-cell nest. This and the nests of most other paper wasps are suspended on a filament or a pedicle. Many paper wasps can excrete an ant repellent on this section of the nest in an effort to prevent invasion. Like many other social hymenopterans, a defending wasp can secrete a pheromone venom during the stinging process to warn the colony of danger at the nest. In winter, Common Paper Wasps seek shelter in stumps and other locations to hibernate.

The European Paper Wasp (Polistes dominula) is a non-native species which builds nests in man-made structures including bird houses. To collect protein, they prey on a wide selection of insects and other invertebrates. As such, European Paper Wasps are widespread and successful here in North America.

YELLOWJACKETS AND HORNETS

An Eastern Yellowjacket feeding on lanternfly honeydew. Eastern Yellowjackets derive much of their success from being generalists, collecting carbohydrates from nearly any sweet source, natural or man made. They are quite fond of ripe fruits, flower nectar, and sugary snacks and drinks, especially soda. Protein for nourishing their larvae is derived from the wide variety invertebrates upon which they prey and from carrion. These foods are chewed into a paste form in preparation for placement into the brood cells.

Robber Fly consuming an Eastern Yellowjacket — Yellowjackets may be moody and aggressive, but they do fall victim to a number of predators. A Robber Fly (Promachus species) has taken down and is devouring this Eastern Yellowjacket.

A Bald-faced Hornet (Dolichovespula maculata) feeding on Spotted Lanternfly honeydew deposits — A Bald-faced Hornet (Dolichovespula maculata) feeding on Spotted Lanternfly honeydew on a Tree-of-heaven (Ailanthus altissima). In the absence of nectar-producing flowers, many bees, yellowjackets, and hornets have turned to the invasive lanternfly and Ailanthus combo to turn the sun’s energy into the carbohydrates they need. For protein, they prey upon spiders, flies, caterpillars, and a variety of other insects.

A Bald-faced Hornet collecting wood pulp from the surface of a weathered picnic table. — To create paper for nest construction, this Bald-faced Hornet is collecting wood pulp from the surface of a weathered picnic table. Away from the nest, these hornets demonstrate a calm, carefree demeanor and can be closely observed.

Bald-faced Hornet Nest — A Bald-faced Hornet nest in a pine tree. These hives are strictly temporary. Within the nest, a generation of drones (males) and new queens are produced late each year. These wasps leave the colony to mate. With the arrival of freezing weather, all inhabitants within the nest, including the old queen, perish, as do the drones that departed to breed. Only the new queens survive winter hibernation to propagate the next generation of wasps, starting with the workers needed to construct a fresh nest and reestablish the colony.

Bald-faced Hornets Peering from Nest — Did you ever get the feeling you’re being watched? Don’t go messing around with Bald-faced Hornet nests. The occupants therein, like other social bees, wasps, and hornets, are equipped with stingers and venom for defending their colony. This is an adaptation that has developed over time to assure the survival of populations of these insects. Think about it this way, a solitary wasp that loses a nest loses only their individual brood of offspring. There is minimal impact on the wider local population of such insects. Conversely, a social wasp or hornet that loses a nest loses an entire colony, possibly negating the benefits of their cooperative behavior and threatening the survival of the species. Insects that cooperate to build societies for survival can be more vulnerable to the catastrophic impacts of certain circumstances like disease, weather, and invasion of their colonies. Therefore, natural selection has provided them with contingencies for these dangers, for example, the instinct to construct protective shelters and the adaptation of stingers and venom for defense against intruders and would-be predators. Oh, and by the way, the Bald-faced Hornet can spray venom, often aiming for the eyes, so keep your distance.

European Hornets (Vespa crabro), an introduced species, are predatory on a variety of flying insects for protein. For carbohydrates they are attracted to sweets like this lanternfly honeydew on Tree-of-heaven.

European Hornets constructing a nest in a tree cavity. — European Hornets constructing a paper nest in a tree cavity.

POTTER WASPS

A Potter Wasp (Eumenes species, probably E. fraturnus) hovering near a European Paper Wasp. — A potter wasp (Eumenes species), probably a Fraternal Potter Wasp (E. fraternus), hovering near a European Paper Wasp on Partridge Pea. The female potter wasp builds a small mud nest resembling a tiny clay pot. One of her eggs is inserted and left hanging on a thin thread. Then a paralyzed caterpillar is deposited as a source of protein to nourish the larva upon hatching. Lastly, the pot is sealed with a lid made of wet mud. Upon maturing, the new generation of adult wasps perform a pottery breaking to emerge and take flight.

ANTS

We hope this brief but fascinating look at some of our more common bees, wasps, hornets, and ants has provided the reader with an appreciation for the complexity with which their food webs and ecology have developed over time. It should be no great mystery why bees and other insects, particularly native species, are becoming scarce or absent in areas of the Lower Susquehanna River Watershed where the landscape is paved, hyper-cultivated, sprayed, mowed, and devoid of native vegetation, particularly nectar-producing plants. Late-summer and autumn can be an especially difficult time for hymanopterans seeking the sources of proteins and carbohydrates needed to complete preparations for next year’s generations of these valuable insects. An absence of these staples during this critical time of year quickly diminishes the diversity of species and begins to tear at the fabric of the food web. This degradation of a regional ecosystem can have unforeseen impacts that become increasingly widespread and in many cases permanent.

A farmland desert. — How can anyone be surprised by the absence of bees and other pollinators in farmland? Manicured and cultivated ground offers little in the way of year-round shelter and food sources for insects and other wildlife.

A savanna-like habitat. — This savanna-like habitat on a south-facing slope provides the abundance of nectar-producing, pollen-rich wildflowers needed to nourish a diverse population of insects including bees, wasps, hornets, and ants. Goldenrods, asters, and White Snakeroot are some of their late-season favorites.

Editor’s Note: No bees, wasp, hornets, or ants were harmed during this production. Neither was the editor swarmed, attacked, or stung. Remember, don’t panic, just observe.

SOURCES

Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York, NY.

(If you’re interested in insects, get this book!)

September 24, 2023June 19, 2024

Four Common Grasshoppers

Grasshoppers are perhaps best known for the occasions throughout history when an enormous congregation of these insects—a “plague of locusts”—would assemble and rove a region to feed. These swarms, which sometimes covered tens of thousands of square miles or more, often decimated crops, darkened the sky, and, on occasion, resulted in catastrophic famine among human settlements in various parts of the world.

The largest “plague of locusts” in the United States occurred during the mid-1870s in the Great Plains. The Rocky Mountain Locust (Melanoplus spretus), a grasshopper of prairies in the American west, had a range that extended east into New England, possibly settling there on lands cleared for farming. Rocky Mountain Locusts, aside from their native habitat on grasslands, apparently thrived on fields planted with warm-season crops. Like most grasshoppers, they fed and developed most vigorously during periods of dry, hot weather. With plenty of vegetative matter to consume during periods of scorching temperatures, the stage was set for populations of these insects to explode in agricultural areas, then take wing in search of more forage. Plagues struck parts of northern New England as early as the mid-1700s and were numerous in various states in the Great Plains through the middle of the 1800s. The big ones hit between 1873 and 1877 when swarms numbering as many as trillions of grasshoppers did $200 million in crop damage and caused a famine so severe that many farmers abandoned the westward migration. To prevent recurrent outbreaks of locust plagues and famine, experts suggested planting more cool-season grains like winter wheat, a crop which could mature and be harvested before the grasshoppers had a chance to cause any significant damage. In the years that followed, and as prairies gave way to the expansive agricultural lands that presently cover most of the Rocky Mountain Locust’s former range, the grasshopper began to disappear. By the early years of the twentieth century, the species was extinct. No one was quite certain why, and the precise cause is still a topic of debate to this day. Conversion of nearly all of its native habitat to cropland and grazing acreage seems to be the most likely culprit.

The critically endangered Eskimo Curlew (Numenius borealis), a species not photographed since 1962 and not confirmed since 1963, fed on Rocky Mountain Locusts during its spring migration through the Great Plains. Excessive hunting and conversion of grasslands to agriculture are believed responsible for the bird’s demise. (United States Fish and Wildlife Service image by Christina Nelson)

In the Mid-Atlantic States, the mosaic of the landscape—farmland interspersed with a mix of forest and disturbed urban/suburban lots—prevents grasshoppers from reaching the densities from which swarms arise. In the years since the implementation of “Green Revolution” farming practices, numbers of grasshoppers in our region have declined. Systemic insecticides including neonicotinoids keep grasshoppers and other insects from munching on warm-season crops like corn and soybeans. And herbicides including 2,4-D (2,4-Dichlorophenoxyacetic acid) have, in effect, become the equivalent of insecticides, eliminating broadleaf food plants from the pasturelands and hayfields where grasshoppers once fed and reproduced in abundance. As a result, few of the approximately three dozen species of grasshoppers with ranges that include the Lower Susquehanna River Watershed are common here. Those that still thrive are largely adapted to roadsides, waste ground, and small clearings where native and some non-native plants make up their diet.

Here’s a look at four species of grasshoppers you’re likely to find in disturbed habitats throughout our region. Each remains common in relatively pesticide-free spaces with stands of dense grasses and broadleaf plants nearby.

CAROLINA GRASSHOPPER

Dissosteira carolina

Carolina Grasshopper wing — The Carolina Grasshopper is most readily detected and identified when it flies. The colors of the wings resemble those of the Mourning Cloak butterfly.

Great Black Wasp on goldenrod. — Carolina Grasshoppers are among the preferred victims of Great Black Wasps (Sphex pensylvanicus). A female wasp stings the grasshopper to paralyze it, then drags it away to one of numerous cells in an underground burrow where she lays an egg on it. The body of the disabled grasshopper then provides nourishment for the larval wasp.

DIFFERENTIAL GRASSHOPPER

Melanoplus differentialis

An adult female Differential Grasshopper with fully developed wings.

An adult female Differential Grasshopper

TWO-STRIPED GRASSHOPPER

Melanoplus bivittatus

An adult female Two-striped Grasshopper.

RED-LEGGED GRASSHOPPER

Melanoplus femurrubrum

The Red-legged Grasshopper may currently be our most abundant and widespread species.

Protein-rich grasshoppers are an important late-summer, early-fall food source for birds. The absence of these insects has forced many species of breeding birds to abandon farmland or, in some cases, disappear altogether.

Beginning in the early 1930s, the Western Cattle Egret (Bubulcus ibis), a notoriously nomadic species, transited the Atlantic from Africa to colonize the Americas…and they did it without any direct assistance from humans. During the 1970s and early 1980s, a nesting population of Western Cattle Egrets on river islands adjacent to the Susquehanna’s Conejohela Flats off Washington Boro was the largest inland rookery in the northeastern United States. The Lancaster County Bird Club censused the birds each August and found peak numbers in 1981 (7,580). During their years of abundance, V-shaped flocks of cattle egrets from the rookery islands ventured into grazing lands throughout portions of Lancaster, York, Dauphin, and Lebanon Counties to hunt grasshoppers. These daily flights were a familiar summertime sight for nearly two decades. Then, in the early 1980s, reductions in pastureland acreage and plummeting grasshopper numbers quickly took their toll. By 1988, the rookery was abandoned. The cattle egrets had moved on. (Vintage 33 mm image)

To the delight of Wild Turkeys, each of the four species of grasshoppers shown above frequents clearings and roadsides adjacent to forest areas. While changes in grasshopper distribution have been detrimental to populations of birds like pheasants, they’ve created a feeding bonanza for turkeys.

Wild Turkeys feeding on grasshoppers along a forest road. — Wild Turkeys feeding on an abundance of grasshoppers along a forest road.

American Kestrel Feeding on Grasshopper — An American Kestrel feeds on a grasshopper while ignoring the abundance of Spotted Lanternflies swarming the adjacent utility pole. In Susquehanna valley farmlands, grasshopper and kestrel numbers are down. Lanternflies, on the other hand, have got it made.

Early Successional Growth — Maintaining areas bordering roads, forests, wetlands, farmlands, and human development in a state of early succession can provide and ideal mix of mature grasses and broadleaf plants for grasshoppers, pollinators, birds, and other wildlife.

September 11, 2023September 12, 2023

A Visit to a Beaver Pond

To pass the afternoon, we sat quietly along the edge of a pond created recently by North American Beavers (Castor canadensis). They first constructed their dam on this small stream about five years ago. Since then, a flourishing wetland has become established. Have a look.

A beaver lodge. — The beaver lodge was built among shrubs growing in shallow water in the middle of the pond.

Woolgrass in a beaver pond. — Woolgrass (Scirpus cyperinus) is a bulrush that thrives as an emergent and as a terrestrial plant in moist soils bordering the pond.

A male Common Whitetail dragonfly keeping watch over his territory.

A Twelve-spotted Skimmer perched on Soft Rush.

A Blue Dasher dragonfly seizing a Fall Field Cricket (Gryllus pennsylvanicus).

A Spicebush Swallowtail visiting Cardinal Flower. — A Spicebush Swallowtail visiting a Cardinal Flower.

A Green Heron looking for small fish, crayfish, frogs, and tadpoles.

A Green Heron stalks potential prey. — The Green Heron stalking potential prey.

A Wood Duck feeding on Lesser Duckweed. — A Wood Duck feeding on the tiny floating plant known as Lesser Duckweed (Lemna minor).

A Least Sandpiper feeding along the muddy edge of a beaver pond. — A Least Sandpiper poking at small invertebrates along the muddy edge of the beaver pond.

A Solitary Sandpiper testing the waters for proper feeding depth.

A Pectoral Sandpiper searches for its next morsel of sustenance.

A Sora rail in a beaver pond. — The Sora (Porzana carolina) is a seldom seen rail of marshlands including those created by North American Beavers. Common Cattails, sedges, and rushes provide these chicken-shaped wetland birds with nesting and loafing cover.

Isn’t that amazing? North American Beavers build and maintain what human engineers struggle to master—dams and ponds that reduce pollution, allow fish passage, and support self-sustaining ecosystems. Want to clean up the streams and floodplains of your local watershed? Let the beavers do the job!

September 2, 2023September 2, 2023

Ruby-throated Hummingbirds on the Go

Ruby-throated Hummingbirds will be moving south through the Lower Susquehanna River Watershed during the next few weeks. — Ruby-throated Hummingbirds will be migrating south through the Lower Susquehanna River Watershed during the next few weeks.

Ruby-throated Hummingbird at feeder. — Hummingbirds supplement their diet by visiting feeders filled with a mixture of sugar and water. If you’re feeding hummingbirds, or thinking about feeding hummingbirds, be sure to review the helpful tips contained in our post from August 5, 2022, “Two Feeders Are Better Than One”. Their health and your peace of mind may depend on it.

August 22, 2023September 7, 2023

One of Nature’s Finest: The Cardinal Flower

It may be one of the most treasured plants among native landscape gardeners. The Cardinal Flower (Lobelia cardinalis) blooms in August each year with a startling blaze of red color that, believe it or not, will sometimes be overlooked in the wild.

Cardinal Flower on a Stream — Cardinal Flower is most often found in wet soil along forested bodies of water. The blooms of this shade-loving species may go unnoticed until rays of sunshine penetrate the canopy to strike their brilliant red petals.

The Cardinal Flower grows in wetlands as well as in a variety of moist soils along streams, rivers, lakes, and ponds. Shady locations with short periods of bright sun each day seem to be favored for an abundance of color.

Cardinal Flower and Great Blue Lobelia — Cardinal Flower in bloom in a riparian forest along the Susquehanna. To its right is its close relative, Great Lobelia, a plant sometimes known as Great Blue Lobelia or Blue Cardinal Flower.

Cardinal Flower in a wet bottomland woods.

The Cardinal Flower can find favorable growing conditions along stream, river and lake shores. — The Cardinal Flower can find favorable growing conditions along stream, river, and lake shores. Even though they are perennial plants, their presence along such waters often seems temporary. Changing conditions cause them to suddenly disappear from known locations, then sometimes reappear at the same place or elsewhere nearby. Some of this phenomenon may be due to the fact that stressed plants can fail to bloom, so they easily escape notice. When producing flowers during favorable years, the plants seem to mysteriously return.

Cardinal Flowers along a wave-swept shoreline light up the greenery of erosion-controlling riparian vegetation with glowing red color.

The Cardinal Flower can be an ideal plant for attracting hummingbirds, bees, butterflies, and other late-summer pollinators. It grows well in damp ground, especially in rain gardens and along the edges streams, garden ponds, and stormwater retention pools. If you’re looking to add Cardinal Flower to your landscape, you need first to…

REMEMBER the CARDINAL RULE…

Cardinal Flower plants are available at many nurseries that carry native species of garden and/or pond plants. Numerous online suppliers offer seed for growing your own Cardinal Flowers. Some sell potted plants as well. A new option is to grow Cardinal Flowers from tissue cultures. Tissue-cultured plants are raised in laboratory media, so the pitfalls of disease and hitchhikers like invasive insects and snails are eliminated. These plants are available through the aquarium trade from most chain pet stores. Though meant to be planted as submerged aquatics in fish tank substrate, we’ve reared the tissue-cultured stock indoors as emergent plants in sandy soil and shallow water through the winter and early spring. When it warms up, we transplant them into the edges of the outdoor ponds to naturalize. As a habit, we always grow some Cardinal Flower plants in the fish tanks to take up the nitrates in the water and to provide a continuous supply of cuttings for starting more emergent stock for outdoor use.

Tissue culture Cardinal Flower being grown as a submerged aquatic in a fish aquarium. — A tissue-cultured Cardinal Flower rooted in sandy substrate and being grown as a submerged aquatic plant in a fish tank. Cuttings from this plant will be used to grow emergent specimens in shallow water for transplanting outdoors around the garden pond.

Cardinal Flower from Tissue Culture — A Cardinal Flower grown from an aquarium store tissue culture blooms in the pond at susquehannawildlife.net headquarters.

Cardinal Flower blooming in November. — Grown as an emergent, Cardinal Flower may bloom very late in the season. This tissue-cultured specimen in the headquarters pond was photographed in early November, 2022.