Ray-finned Fishes (Actinopterygii) – LIFE IN THE LOWER SUSQUEHANNA RIVER WATERSHED

August 20, 2024August 24, 2024

Heat Flux Processes in Streams and Their Impact on Coldwater and Coolwater Fishes

The deluge of rain that soaked the lower Susquehanna watershed during last week is now just a memory. Streams to the west of the river, where the flooding courtesy of the remnants of Hurricane Debby was most severe, have reached their crest and receded. Sliding away toward the Chesapeake and Atlantic is all that runoff, laden with a brew of pollutants including but not limited to: agricultural nutrients, sediment, petroleum products, sewage, lawn chemicals, tires, dog poop, and all that litter—paper, plastics, glass, Styrofoam, and more. For aquatic organisms including our freshwater fish, these floods, particularly when they occur in summer, can compound the effects of the numerous stressors that already limit their ability to live, thrive, and reproduce.

One of those preexisting stressors, high water temperature, can be either intensified or relieved by summertime precipitation. Runoff from forested or other densely vegetated ground normally has little impact on stream temperature. But segments of waterways receiving significant volumes of runoff from areas of sun-exposed impervious ground will usually see increases during at least the early stages of a rain event. Fortunately, projects implemented to address the negative impacts of stormwater flow and stream impairment can often have the additional benefit of helping to attenuate sudden rises in stream temperature.

Stream Subjected to Agricultural Runoff — While a row of trees along a creek can help provide protection from the thermal impact of the sun, a vegetative riparian buffer must be much wider to be effective for absorbing, cooling, and treating runoff from fields, lawns, and paved surfaces. This buffer is too narrow to prevent surface runoff from polluting the water.

Of the fishes inhabiting the Lower Susquehanna River Watershed’s temperate streams, the least tolerant of summer warming are the trouts and sculpins—species often described as “coldwater fishes”. Coldwater fishes require water temperatures below 70° Fahrenheit to thrive and reproduce. The optimal temperature range is 50° to 65° F. In the lower Susquehanna valley, few streams are able to sustain trouts and sculpins through the summer months—largely due to the effects of warm stormwater runoff and other forms of impairment.

Sculpins, including the Blue Ridge Sculpin (Cottus caeruleomentum) seen here, are native coldwater fishes which, during the 11,000 years since the last glacial maximum, have had the availability of their favored habitat sharply reduced by warming water temperatures and a rising Atlantic. During this interval, seawater has inundated the path of the “Late” Pleistocene lower Susquehanna which passed through the section of flooded river watershed we now call Chesapeake Bay and continued across the continental shelf to what was, during the glacial maximum, the river’s mouth at Norfolk Canyon. Today, cut off from neighboring drainage basins, sculpins survive exclusively in cold headwaters, and only in those where human alterations including pollution, dams, channelization, and reduced base flow haven’t yet eliminated their isolated populations. Formerly believed to be composed of two widespread North American species, the Slimy Sculpin (Cottus cognatus) and the Mottled Sculpin (Cottus bairdii), study in recent decades is discovering that sculpin populations in the present-day lower Susquehanna and neighboring Potomac headwaters consist of at least three newly delineated species: Blue Ridge Sculpin, Potomac Sculpin (Cottus gerardi), and Checkered Sculpin (Cottus sp.), the latter an as yet undescribed species found only in the refugium of limestone springs in the Potomac drainage in West Virginia; Frederick and Washington Counties, Maryland; and Franklin County, Pennsylvania. (United States Geological Survey image)

Ice Age Susquehanna — Stare at this for a little while, you’ll figure it out…………More than 11,000 years ago, during the last glacial maximum, when sea level was about 275 feet lower than it is today, there was no Chesapeake Bay, just a great Susquehanna River that flowed to the edge of the continental shelf and its mouth at Norfolk Canyon. It was a river draining taiga forests of pine, spruce , and fir, and it carried along the waters of all the present-day bay’s tributaries and more. The section of the river’s watershed we presently call the lower Susquehanna was, at the time, the upper Susquehanna watershed. Brook Trout and sculpins had the run of the river and its tributaries back then. And the entire watershed was a coldwater fishery, with limestone and other groundwater springs providing not refuge from summer heat, but a place to escape freezing water. (United States Geological Survey base image)

Norfolk Canyon, the mouth of the Susquehanna River during the most recent glacial maximum, now lies more than 275 feet below the surface of the ocean and plunges to more than a mile in depth along the finger of out wash from the gorge. (United States Geological Survey image)

Rainbow. Brown, and Brook Trout — Tens of thousands of trout are raised in state-operated and cooperative nurseries for stocking throughout the lower Susquehanna valley. These rearing facilities are located on spring-fed headwaters with sufficient flow to assure cold temperatures year round. While the Rainbow Trout and Brown Trout (Salmo trutta) are the most commonly stocked species, the Brook Trout (Salvelinus fontinalis) is the only one native to American waters. It is the least tolerant of stream warming and still reproduces in the wild only in a few pristine headwaters streams in the region. During spring, all three of these species have been observed on rare occasions entering the fish lift facilities at the hydroelectric dams on the river, presumably returning to the Susquehanna as sea-run trout.

Coldwater fishes are generally found in small spring-fed creeks and headwaters runs. Where stream gradient, substrate, dissolved oxygen, and other parameters are favorable, some species may be tolerant of water warmer than the optimal values. In other words, these temperature classifications are not set in stone and nobody ever explained ichthyology to a fish, so there are exceptions. The Brown Trout for example is sometimes listed as a “coldwater transition fish”, able to survive and reproduce in waters where stream quality is exceptionally good but the temperature may periodically reach the mid-seventies.

The Eastern Blacknose Dace is sometimes classified as a “coldwater transition fish”. It can be found in headwaters runs as well as in creeks with good water quality.

The Longnose Dace is another “coldwater transition fish” known only from clear, clean, flowing waters.

More tolerant of summer heat than the trouts, sculpins, and daces are the “coolwater fishes”—species able to feed, grow, and reproduce in streams with a temperature of less than 80° F, but higher than 60° F. Coolwater fishes thrive in creeks and rivers that hover in the 65° to 70° F range during summer.

Creek Chubs — The Creek Chub is a familiar species of “coolwater fish” seldom found remaining in waters exceeding 80 degrees Fahrenheit.

The Yellow Perch (Perca flavescens) was perhaps the most frequently targeted coolwater “gamefish” in the Lower Susquehanna River Watershed prior to the introduction of the Northern Pike (Esox lucius) and Muskellunge (Esox masquinongy). Today’s prevalence of warmwater streams and the dozens of species of non-native predatory fishes now naturalized within them have left the Yellow Perch populations greatly reduced and all but forgotten by anglers. Out of sight, out of mind. (National Park Service image)

What are the causes of modern-day reductions in coldwater and coolwater fish habitats in the lower Susquehanna River and its hundreds of miles of tributaries? To answer that, let’s take a look at the atmospheric, cosmic, and hydrologic processes that impact water temperature. Technically, these processes could be measured as heat flux—the rate of heat energy transfer per unit area per unit time, frequently expressed as watts per meter squared (W/m²). Without getting too technical, we’ll just take a look at the practical impact these processes have on stream temperatures.

HEAT FLUX PROCESSES IN A SEGMENT OF STREAM

Heat Flux Processes on Streams and Rivers — Heat Flux Processes on Stream and River Segments. These processes could be measured as heat flux—the rate of heat energy transfer per unit area per unit time. (Environmental Protection Agency image)

- - INCOMING TEMPERATURE AND FLOW—The baseline temperature of stream water entering a given segment of waterway is obviously the chief factor determining its temperature when exiting that segment. Incoming temperature and flow also determine the water’s susceptibility to heat absorption or loss while transiting the segment. Lower flows may subject the given volume of water to a greater loss or gain of heat energy during the time needed to pass through the segment than the same volume at a higher flow. Lower flows may also reduce stream velocity and extend a given volume of water’s exposure time to the exchange of heat energy while moving through the segment. Generally speaking…
    1. …the higher the stream flow, the less a given volume of that stream’s water may be impacted by the effects of the heat flux processes within the segment.
    2. …the lower the stream flow, the more a given volume of that stream’s water may be impacted by the effects of the heat flux processes within that segment.
    3. …the temperature and flow rate of precipitation entering the segment are factors that determine the impact of its heat energy transfer to or from a given volume of the stream’s waters.
    4. …the temperature and flow rate of runoff and point-source discharges entering the segment are factors that determine the impact of their heat energy transfer to or from a given volume of the stream’s waters.

Stormwater Discharge into Channelized Creek — Stormwater from impervious surfaces including roads, parking lots, roofs, and lawns quickly impacts temperatures in small creeks. Channelized streams are availed few of the positive attributes provided by many of the heat flux processes we’re about to see. They therefore suffer from severe impairment and are exposed to temperature extremes that few aquatic organisms can survive. Runoff from sun-heated pavement during a summer thunderstorm can often exceed 100 degrees Fahrenheit and can, at sufficient flow rate, quickly raise the temperature of a small stream to well over 90 degrees.

Stormwater Runoff — Stormwater runoff not only poses a thermal threat to waterways, its a significant source of a wide variety of pollutants.

- - GROUNDWATER INPUT—In streams connected to the aquifer, the temperature in a flowing segment can be impacted by the influx of cold groundwater. With temperatures ranging from about 52° to 60° Fahrenheit, groundwater will absorb heat from the stream in summer, and warm it in the winter. In warmwater streams, coldwater and coolwater fishes will often seek areas of the substrate where groundwater is entering for use as refugium from the summer heat. Yellow Perch in the lower Susquehanna are known to exhibit this behavior.

Gaining Stream — Creeks and rivers connected to the aquifer and receiving supplemental flow from it are known as “gaining streams”. These streams frequently feed water into the aquifer as well. (United States Geological Survey image)

Losing Stream — When flowing through an area experiencing drought or an excessive removal of groundwater (lots of wells, etc.), a waterway can become a “losing stream”, one that surrenders a portion of its flow to recharge the aquifer. Further downstream, the reduced flow can make such a creek or river more susceptible to the effects of heat flux processes. (United States Geological Survey image)

Disconnected Stream — Seriously depleted aquifers can lead to a “disconnected stream”. Smaller waterways subjected to these conditions will sometimes lose all their flow to the ground, often causing a catastrophic failure of the aquatic ecosystem supported therein. (United States Geological Survey image)

Urban Flooding and Dry Streambed — Urban runoff overwhelms this small stream with polluted water than can reach temperatures of 100 degrees or more (left), then lets it high and dry with no baseflow during periods of dry weather (right) as the waterway becomes disconnected from the much-depleted aquifer.

Stormwater Retention Basin — Well-designed and properly constructed stormwater retention basins not only recharge groundwater supplies for wells and streams, they can also help prevent thermal pollution in waterways. Planted with native wetland species and allowed to thrive, they can become treasured wildlife islands in otherwise inhospitable environs. The benefits don’t stop there; plants also help sequester nutrients contained in the runoff.

- - HYPORHEIC EXCHANGE—Related to groundwater input, hyporheic exchange is the slow movement of water through the rock, sand, gravel, and soils composing the streambed, saturated shoreline, shallow aquifer, and connected floodplain of a creek or river. As a heat flux process, hyporheic exchange helps moderate extremes in seasonal water temperatures by conducting energy between the solid materials in the zone and the flowing water. Hyporheic zones are important habitats for many species of aquatic invertebrates and spawning fish. Natural chemical processes within these zones convert ammonia-producing wastes into nitrite, then nitrate, allowing it to be absorbed as food by plants growing in the stream or in the alluvium within the zone. Vegetation removal, channelization, legacy sediments, silt deposits, and man-made walls and dams can negate the benefits of hyporheic exchange.

Hyporheic Exchange — Exchange of surface and ground water within the hyporheic zone is most directly associated with high-gradient (left) and meandering (right) segments of streams. (United States Geological Survey image)

Legacy Sediments and Fill — Very common on streams in the lower Susquehanna valley are these accumulations of legacy sediments at the sites of former mill ponds. After the dams were removed, the creeks began eroding their way down through the mire as they tried to reestablish their floodplains and find their native substrate. These trapped waterways are not only cut off from their hyporheic zones, they’re now a major source of nutrient and sediment pollution. Misguided landowners like this one frequently dump fill into these sites to “save their land” and “control flooding”. The fill and materials added to “shore up the banks” do nothing to fix what ails the creek, but instead displace more water to make the impact of flooding even more widespread.

Flooplain and Stream Restoration — Rehabilitation projects that remove legacy sediments help restore hyporheic exchange by reconnecting the stream to its underlying geology, its floodplain, and its wetlands. Rising waters remain in the floodplain where they get a good bio-scrubbing and help replenish the creek and groundwater supply. As the experts say, “floodplains are for flooding.”

- - ATMOSPHERIC EXCHANGE (CONVECTION, EVAPORATION)—Primarily a process by which a stream loses heat energy and cools its waters, atmospheric exchange is also a means by which a warm air mass can relinquish heat to cooler waters and thus increase their temperature. This phenomenon can be dramatically enhanced when a stream passes through a so-called urban heat island where air temperatures remain warm through the night. Convection, the movement of heat energy through a fluid (liquid or gas), causes warmer, less-dense water to rise to the surface of a stream, particularly where there is minimal turbulence. When the air above is cooler than the water’s surface layer, the stream will conduct heat energy across the water/atmosphere interface causing the warmed air molecules to rise in a convection column. If the atmospheric relative humidity is less than 100%, some surface water will vaporize—a process that expends more of the stream’s heat energy. The rate of convective and evaporative cooling in a given stream segment is directly related to the degree of difference between the water temperature and air temperature, and to the relative humidity in the air mass above the lake, creek, or river. The mechanical action of stream turbulence including rapids, riffles, and falls increases the contact area between air and water to maximize the atmospheric exchange of heat energy. The convective air current we call surface wind has a turbulent wave-producing effect on water that can also maximize atmospheric exchange; think of a cold autumn wind robbing heat energy from a warm lake or river or a hot summer wind imparting its heat to a cooler creek. These exchanges are both conductive in nature (air-to-water/water-to-air) and evaporative, the latter being expedited by the movement of dry air over warm water.

Usually classified as one of the coolwater fishes, the bottom-dwelling Tessellated Darter can thrive in the warmer creeks and in the main stem of the Susquehanna by inhabiting riffles where atmospheric exchange in the form of increased evaporation helps reduce temperatures and convective currents carry the cooler, well-oxygenated water to the streambed.

Three mile Island Unit 1 Cooling Towers — Humans utilize the concept of atmospheric exchange, adopting the phenomena of evaporation and convection to cool the hot waters produced during electric generation and other industrial processes before discharge into a lake or river.

- - STREAMBED CONDUCTIVE EXCHANGE—In the lower Susquehanna watershed, there may be no better natural example of streambed conductive exchange than the Triassic-Jurassic diabase pothole bedrocks of Conewago Falls on the river at the south end of Three Mile Island.

During sunny days, the massive diabase pothole rocks at Conewago Falls absorb solar (shortwave) radiation, then conduct that heat energy into the flowing water, often continuing to pass the accumulated warmth into the river during the night. On cloudy days, the riverbed collects longwave atmospheric radiation, a heat flux process that yields significantly less energy for conduction into the rapids, riffles, and pools of the falls. During periods of low river flow, the heating effect of streambed conductive exchange can become magnified. Compared to conditions that prevail when torrents of turbid water are rushing through the falls, partially exposed bedrock surrounded by clear water collects radiated energy much more efficiently, then conducts the heat to a greatly reduced volume of passing water. During summer and autumn, this process can create a mix of temperature zones within the falls with warmer water lingering in slow-moving pools and cooler water flowing in the deeper fast-moving channels. Along the falls’ mile-long course, a haven is created for aquatic organisms including warmwater and some coolwater fishes, oft times attracting anglers and a variety of hungry migrating birds as well.

Classified as one of our coolwater fishes, the Fallfish finds favorable conditions for feeding, growing, and spawning in the well-oxygenated waters of Conewago Falls.

Though the lower Susquehanna River is classified as a warmwater fishery, the Northern Hog Sucker (Hypentelium nigricans), another of our native coolwater fishes, finds the fast-moving waters of Conewago Falls to its liking. Northern Hog Suckers are known to inhabit streams cold enough to host trout. They exhibit remarkable home range fidelity, sometimes spending their entire lives occupying the same several hundred feet of waterway. Northern Hog Suckers are often designated an indicator of good water quality, intolerant of many stream impairment parameters. Their presence in Conewago Falls provides testament to the quality of the warmwater fishery there.

Severely Impaired Channelized Stream — An unnatural example. The reduced base flow in this channelized and severely impaired creek has been rendered vulnerable to the negative impacts of several heat flux processes including streambed conductive exchange. Urban stormwater/surfacewater inflow, solar (shortwave) radiation, and heat conducted into the stream from the masonry walls, curbs, and raceway can all conspire to cook aquatic organisms with life-quenching summer water temperatures exceeding 90 degrees Fahrenheit.

- - SOLAR (SHORTWAVE) RADIATION—The sun provides the energy that fuels the earth’s complex climate. The primary heat flux process that heats our planet is the absorption of solar radiation in the shortwave spectrum, which includes ultraviolet, visible, and infrared frequencies at the upper end of the longwave spectrum. Streams and other bodies of water absorb the greatest amounts of solar (shortwave) radiation during the weeks around summer solstice when the sun at mid-day is closer to zenith than at any other time of the year. However, the heating impact of the radiation may be greatest when the volume of water in the creek, river, or lake is at its minimum for the year—often during early fall.

The Sun — The rate, measured in watts per square meter, at which solar (shortwave) energy is directly radiated to a given area on the earth’s surface (including streams and other waters) is determined by: solar activity, the angle of the sun in the sky, aspect (slope) of the receiving surface, the opacity of the overlying atmosphere, and the distance of the earth from the sun. The former varies with the year’s seasons, the time of day, and the latitude of a given area. The latter is currently at its annual minimum when earth is at perihelion during the early days of January, thus providing the northern hemisphere with a little bump in radiation during the shortest days of the year when the sun is at its lowest angle in the sky. (NASA image)

Solar (Shortwave) Radiation — A varying portion of the solar (shortwave) radiation reaching the earth is reflected back into space by clouds. A smaller share is absorbed by the atmosphere, thus heating it. An even lesser quantity is reflected back into space by water and land. The remainder of the energy is absorbed by the planet’s surfaces, its water and land. (NASA image)

- - INCIDENT SHORTWAVE RADIATION—Also known as insolation (incoming solar radiation), incident shortwave radiation is the sum total energy of both the direct solar radiation that travels to the earth’s surface unaffected by the atmosphere and the diffuse radiation, waves that have been weakened and scattered by constituents of the atmosphere before reaching the planet’s surface. On a cloudy day, the warming of terrestrial surfaces including streams and other bodies of water is the result of diffuse radiation. On days with any amount of sunshine at all, both direct and diffuse radiation heat our waters and lands.

Pumkinseed — Warmwater fishes such as the native Pumpkinseed (Lepomis gibbosus) thrive in sun-drenched 70-to-85-degree waters as long as other heat flux processes prevent sudden temperature increases and oxygen depletion.

Mowed Stream Bank — Mowed stream banks offer a waterway no protection from incoming solar (shortwave) radiation, nor terrestrial forms of impairment including nutrient-rich stormwater runoff and silt.

- - REFLECTED SHORTWAVE RADIATION—known as albedo, reflected solar (shortwave) radiation is energy directed away from the earth’s surface before being absorbed. A surface’s albedo value is basically determined by its color, black having little reflective value, white and silvery surfaces reflecting nearly all solar (shortwave) radiation away. A surface with no reflective properties has an albedo value of 0, while a totally reflective surface has a value of 1. Clean snow with a value of about 0.85 to 0.9 (85% to 90%) is a highly reflective surface; yellow snow isn’t as good. A stream, river, or lake blanketed with ice and snow will absorb very little solar energy and will rely upon other heat flux processes to trigger a melt and thaw. The surface of open water has a varying albedo value determined mostly by the angle of the sun. Solar radiation striking the water’s surface at a low angle is mostly reflected away, while that originating at an angle closer to zenith is more readily absorbed.

Forested Stream — To avoid the heating effects of solar (shortwave) and atmospheric longwave radiation, coldwater and coolwater fishes require streams offering protection from full exposure to direct sunlight and cloud cover. Runs and creeks flowing beneath a closed canopy of forest trees are shielded from 25% or more of incoming radiation and are thus able to better maintain thermal stability during the most vulnerable period of the year for temperature-sensitive fishes, May through October.

- - LONGWAVE RADIATION—Radiation in the longwave spectrum is composed of infrared waves at frequencies lower than those of the shortwave spectrum. Longwave radiation, sometimes just called infrared radiation, is produced by the earth and its atmosphere and is propagated in all directions, day and night. It warms mostly the lower atmosphere which in turn warms the earth’s surface including its waters. Some longwave energy can even be radiated into the waterway from its own streambed—and the stream can return the favor. Other forms of mass surrounding a stream such as a rocky shoreline or a man-made structure such as bridge pier can trade longwave radiation with a waterway. The effect of these latter exchanges is largely trivial and never rivals the heat flux transfer of warm to cold provided by conduction.

Longwave Radiation — Longwave radiation emissions slow as the temperature of the emitting mass decreases, just as they also increase with temperature of the mass. Longwave radiation emissions therefore decrease with altitude along with the temperature of the water vapor, carbon dioxide, methane, and other gases that produce them. As such, the highest reaches of the atmosphere have a greatly reduced capability of shedding longwave radiation into space. At ground level, lakes, creeks, and streams receive their greatest dose of longwave radiation while beneath the cover of low-lying clouds or fog. (NASA image)

- - CANOPY RADIATION—Trees emit longwave radiation that may have a limited heat flux impact on waterway temperature. This radiation is diffuse, of scattered effect, and scarcely detectable, particularly beneath multilayered dense canopies. Some of the infrared energy transmitted by the tree canopy is radiated skyward as well.
  - WATER RADIATION—Water, like all earthly matter composed of vibrating molecules, emits longwave radiation. This heat flux process provides an ongoing cooling effect to streams, rivers, lakes, and oceans—warmer ones shedding infrared energy at a faster rate than those that are cold.

Now that we have a basic understanding of the heat flux processes responsible for determining the water temperatures of our creeks and rivers, let’s venture a look at a few graphics from gauge stations on some of the lower Susquehanna’s tributaries equipped with appropriate United States Geological Survey monitoring devices. While the data from each of these stations is clearly noted to be provisional, it can still be used to generate comparative graphics showing basic trends in easy-to-monitor parameters like temperature and stream flow.

Each image is self-labeled and plots stream temperature in degrees Fahrenheit (bold blue) and stream discharge in cubic feet per second (thin blue).

West Conewago Creek 90 Days — The West Conewago Creek drains much of the Gettysburg Basin’s Triassic redbeds in Adams and northern York Counties in Pennsylvania and includes a small headwaters area in northern Maryland. The gauge station is located just a over a mile upstream from the waterway’s mouth on the Susquehanna just below Conewago Falls. Right through the summer heatwave, this 90-day graph shows a consistent daily pattern of daytime rises in temperature and nighttime cooling. To the right, a rapid cool down can be seen coinciding with two periods of high water, the first from a series of heavy thundershowers, the second from flooding caused by the remnants of Hurricane Debby. Notice that the early August downpours were so heavy that they cooled the hot surface runoff and waterway quickly, without creating a rise in stream temperature at the gauging station. Had this monitoring device been located on a small tributary in an area with an abundance of impervious surfaces, there would probably have been a brief rise in stream temperature prior to the cooldown. (United States Geological Survey image)

The daily oscillations in temperature reflect the influence of several heat flux processes. During the day, solar (shortwave) radiation and convection from summer air, especially those hot south winds, are largely responsible for the daily rises of about 5° F. Longwave radiation has a round-the-clock influence—adding heat to the stream during the day and mostly shedding it at night. Atmospheric exchange including evaporative cooling may help moderate the rise in stream temperatures during the day, and certainly plays a role in bringing them back down after sunset. Along its course this summer, the West Conewago Creek absorbed enough heat to render it a warmwater fishery in the area of the gauging station. The West Conewago is a shallow, low gradient stream over almost its entire course. Its waters move very slowly, thus extending their exposure time to radiated heat flux and reducing the benefit of cooling by atmospheric exchange. Fortunately for bass, catfish, and sunfish, these temperatures are in the ideal range for warmwater fishes to feed, grow, and reproduce—generally over 80° F, and ideally in the 70° to 85° F range. Coolwater fishes though, would not find this stream segment favorable. It was consistently above the 80° F maximum and the 60° to 70° F range preferred by these species. And coldwater fishes, well, they wouldn’t be caught dead in this stream segment. Wait, scratch that—the only way they would be caught in this segment is dead. No trouts or sculpins here.

The Codorus Creek drains primarily the carbonate valleys of York County to the south of the West Conewago watershed. This gauge station is located about a mile upstream from the creek’s mouth on the Susquehanna just below Haldeman Riffles. The graphic pattern is very similar to that of the West Conewago’s: daily heating and cooling cycles and a noticeable drop in stream temperature in early August caused by a day of thundershowers followed by the remnants of Hurricane Debby. (United States Geological Survey image)

Look closely and you’ll notice that although the temperature pattern on this chart closely resembles that of the West Conewago’s, the readings average about 5 degrees cooler. This may seem surprising when one realizes that the Codorus follows a channelized path through the heart of York City and its urbanized suburbs—a heat island of significance to a stream this size. Before that it passes through numerous impoundments where its waters are exposed to the full energy of the sun. The tempering factor for the Codorus is its baseflow. Despite draining a smaller watershed than its neighbor to the north, the Codorus’s baseflow (low flow between periods of rain) was 96 cubic feet per second on August 5th, nearly twice that of the West Conewago (51.1 cubic feet per second on August 5th). Thus, the incoming heat energy was distributed over a greater mass in the Codorus and had a reduced impact on its temperature. Though the Codorus is certainly a warmwater fishery in its lower reaches, coolwater and transitional fishes could probably inhabit its tributaries in segments located closer to groundwater sources without stress. Several streams in its upper reaches are in fact classified as trout-stocked fisheries.

This is a zoomed-in look at the previous graph showing the impact of a rainfall event on the water temperatures in Codorus Creek. Unlike the sharp declines accompanying the deluge of flood waters during the two events in early August, these lesser storms in late June generated just enough runoff to capture heat energy from impervious surfaces and warm the creek, temporarily breaking the daily heating/cooling cycle. Upstream in the immediate area of the runoff, the impact on the stream and/or its tributaries was probably much more dramatic, certainly raising temperatures into the nineties or above. (United States Geological Survey image)

Kreutz Creek drains a carbonate bedrock area of York County and flows parallel to the Lincoln Highway (US 30) to enter the Susquehanna at Wrightsville. The gauging station is about one mile upstream from the creek’s mouth. (United States Geological Survey image)

The Kreutz Creek gauge shows temperature patterns similar to those in the West Conewago and Codorus data sets, but notice the lower overall temperature trend and the flow. Kreutz Creek is a much smaller stream than the other two, with a flow averaging less than one tenth that of the West Conewago and about one twentieth of that in the Codorus. And most of the watershed is cropland or urban/suburban space. Yet, the stream remains below 80° F through most of the summer. The saving graces in Kreutz Creek are reduced exposure time and gradient. The waters of Kreutz Creek tumble their way through a small watershed to enter the Susquehanna within twenty-four hours, barely time to go through a single daily heating and cooling cycle. As a result, their is no chance for water to accumulate radiant and convective heat over multiple summer days. The daily oscillations in temperature are less amplified than we find in the previous streams—a swing of about three degrees compared to five. This indicates a better balance between heat flux processes that raise temperature and those that reduce it. Atmospheric exchange in the stream’s riffles, forest cover, and good hyporheic exchange along its course could all be tempering factors in Kreutz Creek. From a temperature perspective, Kreutz Creek provides suitable waters for coolwater fishes.

Muddy Creek drains portions of southern York County through rolling farmland and woodlots. There are no large impoundments or widespread urban impacts in the watershed, which may help explain its slightly lower temperature trends. (United States Geological Survey image)

Muddy Creek is a trout-stocked fishery, but it cannot sustain coldwater species through the summer heat. Though temperatures in Muddy Creek may be suitable for coolwater fishes, silt, nutrients, low dissolved oxygen, and other factors could easily render it strictly a warmwater fishery, inhabited by species tolerant of significant stream impairment.

Chiques Creek drains mostly limestone farmland in northwestern Lancaster County. The gauging station is located near the stream’s mouth on the Susquehanna at Chiques (Chickies) Rock. Oscillations in temperature again resemble the other waterways, but daily highs remain almost entirely below 80 degrees. (United States Geological Survey image)

A significant number of stream segments in the Chiques watershed have been rehabilitated to eliminate intrusion by grazing livestock, cropland runoff, and other sources of impairment. Through partnerships between a local group of watershed volunteers and landowners, one tributary, Donegal Creek, has seen riparian buffers, exclusion fencing, and other water quality and habitat improvements installed along nearly ever inch of its run from Donegal Springs through high-intensity farmland to its mouth on the main stem of the Chiques just above its confluence with the Susquehanna. The improved water quality parameters in the Donegal support native coldwater sculpins and an introduced population of reproducing Brown Trout. While coldwater habitat is limited to the Donegal, the main stem of the Chiques and its largest tributary, the Little Chiques Creek, both provide suitable temperatures for coolwater fishes.

Limestone Formation on Little Chiques Creek — Streams in the Chiques Creek and similar limestone watersheds often pass through areas with significant bedrock formations. Heat flux processes including groundwater input, hyporheic exchange, and streambed conductive exchange can have a greater influence on water temperature along these segments.

The Common Shiner (Luxilus cornutus), a fish tolerant of warmwater streams, prefers cool, clear waters for spawning. For protection from late-spring and summer heat, breeding males may seek a section of creek with a streambed inflow of limestone groundwater to defend as their nesting territory.

A closeup of the Chiques Creek graph showing what appears to be a little bump in temperature caused by surface runoff during a couple of late-May showers. Stream rehabilitation is an ongoing process and the pressures of land disturbances both old and new present challenges to those who make it their passion to fix the wrongs that have been inflicted upon our local waters. Even the exemplary Donegal Creek faces new threats from urbanization in one of its headwater areas several miles to the northwest of the historic springs. (United States Geological Survey image)

Conewago Creek (East) drains primarily Triassic redbed farmlands in Dauphin, Lancaster, and Lebanon Counties. Much of the headwaters area is forested but is experiencing an increasing rate of encroachment by housing and some commercial development. Conewago Creek (East) enters the Susquehanna on the east side of Conewago Falls at Three Mile Island. The watershed is equipped with three U.S.G.S. gauge stations capable of providing temperature data. This first one is located just over a mile upstream of the creek’s mouth. (United States Geological Survey image)

Despite its meander through and receipt of water from high-intensity farmland, the temperature of the lower Conewago (East) maxes out at about 85° F, making it ideal for warmwater fishes and even those species that are often considered coolwater transition fishes like introduced Smallmouth Bass, Rock Bass, Walleye, and native Margined Madtom. This survivable temperature is a testament to the naturally occurring and planted forest buffers along much of the stream’s course, particularly on its main stem. But the Conewago suffers serious baseflow problems compared to other streams we’ve looked at so far. Just prior to the early August storms, flow was well below 10 cubic feet per second for a drainage area of more than fifty square miles. While some of this reduced flow is the result of evaporation, much of it is anthropogenic in origin as the rate of groundwater removal continues to increase and a recent surge in stream withdraws for irrigation reaches its peak during the hottest days of summer.

Juvenile Rock Bass — A juvenile Rock Bass.

A closer look at the Conewago Creek (East) graphic shows the temperature drop associated with a series of thundershowers and the remnants of Hurricane Debby in early August. Despite the baseflow being below five cubic feet per second, the cooling effect of the downpours as measured in the area of the gauge was significant enough to overwhelm any heating of runoff that may have occurred as precipitation drained across hardened soils or man-made impervious surfaces. (United States Geological Survey image)

A little side note—the flow rate on the Conewago at the Falmouth gauge climbed to about 160 cubic feet per second as a result of the remnants of Hurricane Debby while the gauge on the West Conewago at Manchester skyrocketed to about 20,000 cubic feet per second. Although the West Conewago’s watershed (drainage area) is larger than that of the Conewago on the east shore, it’s larger only by a multiple of two or three, not 125. That’s a dramatic difference in rainfall!

The Bellaire monitoring station on Conewago Creek (East) is located on the stream’s main stem just downstream from the mouth of Little Conewago Creek, a tributary with its origins in farmland and woodlots. (United States Geological Survey image)

The temperatures at the Bellaire monitoring station, which is located upstream of the Conewago’s halfway point between its headwaters in Mount Gretna and its mouth, are quite comparable to those at the Falmouth gauge. Although a comparison between these two sets of data indicate a low net increase in heat absorption along the stream’s course between the two points, it also suggests sources of significant warming upstream in the areas between the Bellaire gauge and the headwaters.

Data from the gauge site on the Little Conewago Creek shows a temperature averaging about five degrees cooler than the gauge several miles downstream on the main stem of the Conewago at Bellaire. (United States Geological Survey image)

The waters of the Little Conewago are protected within planted riparian buffers and mature woodland along much of their course to the confluence with the Conewago’s main stem just upstream of Bellaire. This tributary certainly isn’t responsible for raising the temperature of the creek, but is instead probably helping to cool it with what little flow it has.

Juvenile Eastern Blacknose Dace (top) and a juvenile Longnose Dace. — A stream like the Little Conewago Creek with daily temperatures that remain mostly below 80 degrees and retreat to 75 degrees or less during the night can be suitable for coldwater transition fishes like these juvenile Eastern Blacknose Dace (top) and Longnose Dace.

Though mostly passing through natural and planted forest buffers above its confluence with the Little Conewago, the main stem’s critically low baseflow makes it particularly susceptible to heat flux processes that raise stream temperatures in segments within the two or three large agricultural properties where owners have opted not to participate in partnerships to rehabilitate the waterway. The headwaters area, while largely within Pennsylvania State Game Lands, is interspersed with growing residential communities where potable water is sourced from hundreds of private and community wells—every one of them removing groundwater and contributing to the diminishing baseflow of the creek. Some of that water is discharged into the stream after treatment at the two municipal sewer plants in the upper Conewago. This effluent can become quite warm during processing and may have significant thermal impact when the stream is at a reduced rate of flow. A sizeable headwaters lake is seasonally flooded for recreation in Mount Gretna. Such lakes can function as effective mid-day collectors of solar (shortwave) radiation that both warms the water and expedites atmospheric exchange.

Upper Conewago Creek (East) Watershed — The Conewago Creek (East) Watershed from the Bellaire U.S.G.S. Gauging Station (lower left) upstream to the headwaters in Mount Gretna. (United States Geological Survey image)

Though Conewago Creek (East) is classified as a trout-stocked fishery in its upper reaches in Lebanon County, its low baseflow and susceptibility to warming render it inhospitable to these coldwater fishes by late-spring/early summer.

Despite being considered a warmwater fish, the River Chub (Nocomis micropogon) will ascend streams like the Conewago to seek cooler, gravel-bottomed waters for spawning. Reduced baseflow has probably rendered the stream currently too small for this species on Pennsylvania State Game Lands in Colebrook where this specimen was photographed in 2018.

The Golden Shiner, another warmwater fish, often ascends streams to enter cooler water. Juvenile Golden Shiners like this one will move into shallower headwaters not only to seek reduced temperatures, but to escape large predatory fishes as well.

Irrigation using stream water. — Irrigation of agricultural fields using a large portion of the already diminished baseflow in the Conewago Creek (East) just downstream of the Bellaire gauging station. Despite millions of dollars in investment to rehabilitate this Susquehanna valley stream, the riparian buffers and other practices can have little effect when the creek gets sucked down to just a trickle. Low baseflow is a hard nut to crack. It’s best prevented, not corrected.

Hammer Creek, a trout-stocked fishery, originates, in part, within Triassic conglomerate in the Furnace Hills of Lebanon County, then flows north into the limestone Lebanon Valley where it picks up significant flow from other tributaries before working its way south back through the Furnace Hills into the limestone farmlands of Lancaster County. From there the stream merges with the Cocalico Creek, then the Conestoga River, and at last the Susquehanna. Note the tremendous daily temperature oscillations on this headwaters stream as it surges about 15 degrees each day before recovering back close to groundwater temperature by sunrise the next day. (United States Geological Survey image)

The removal of two water supply dams on the headwaters of Hammer Creek at Rexmont eliminated a large source of temperature fluctuation on the waterway, but did little to address the stream’s exposure to radiant and convective heat flux processes as it meanders largely unprotected out of the forest cover of Pennsylvania State Game Lands and through high-intensity farmlands in the Lebanon Valley. Moderating the temperature to a large degree is the influx of karst water from Buffalo Springs, located about two miles upstream from this gauging station, and other limestone springs that feed tributaries which enter the Hammer from the east and north. Despite the cold water, the impact of the stream’s nearly total exposure to radiative and other warming heat flux processes can readily be seen in the graphic. Though still a coldwater fishery by temperature standards, it is rather obvious that rapid heating and other forms of impairment await these waters as they continue flowing through segments with few best management practices in place for mitigating pollutants. By the time Hammer Creek passes back through the Furnace Hills and Pennsylvania State Game Lands, it is leaning toward classification as a coolwater fishery with significant accumulations of sediment and nutrients. But this creek has a lot going for it—mainly, sources of cold water. A core group of enthusiastic landowners could begin implementing the best management practices and undertaking the necessary water quality improvement projects that could turn this stream around and make it a coldwater treasure. An organized effort is currently underway to do just that. Visit Trout Unlimited’s Don Fritchey Chapter and Donegal Chapter to learn more. Better yet, join them as a volunteer or cooperating landowner!

Male Creek Chub — The male Creek Chub, one of our coolwater fishes, develops head tubercles and becomes flushed with color during spawning season. Hammer Creek not only provides a home for the Creek Chub, its cold headwaters provide refuge for a population of native Brook Trout too.

Like no other example we’ve looked at so far, this closeup of the Hammer Creek graphic shows temperature bumps correlating with the stormwater runoff from early August’s rains. Because the stream flow is small and the precipitation rate was not as great at this location, the effect of heat flux from runoff is more readily apparent. (United States Geological Survey image)

Brook Trout adult and juvenile. (United States Fish and Wildlife Service image by Ryan Hagerty)

For coldwater fishes, the thousands of years since the most recent glacial maximum have seen their range slowly contract from nearly the entirety of the once much larger Susquehanna watershed to the headwaters of only our most pristine streams. Through no fault of their own, they had the misfortune of bad timing—humans arrived and found coldwater streams and the groundwater that feeds them to their liking. Some of the later arrivals even built their houses right on top of the best-flowing springs. Today, populations of these fishes in the region we presently call the Lower Susquehanna River Watershed are seriously disconnected and the prospect for survival of these species here is not good. Stream rehabilitation, groundwater management, and better civil planning and land/water stewardship are the only way coldwater fishes, and very possibly coolwater fishes as well, will survive. For some streams like Hammer Creek, it’s not too late to make spectacular things happen. It mostly requires a cadre of citizens, local government, project specialists, and especially stakeholders to step up and be willing to remain focused upon project goals so that the many years of work required to turn a failing stream around can lead to success.

Riparian Buffer — Riparian buffers with fences to exclude livestock can immediately begin improving water quality. With establishment of such vegetative buffers, the effects of stressors that otherwise eliminate coldwater and coolwater fishes from these segments will begin to diminish.

You’re probably glad this look at heat flux processes in streams has at last come to an end. That’s good, because we’ve got a lot of work to do.

You Don't Have Time to Mow, We've Got Work to Do — Add one more benefit to the wildflower meadow, it infiltrates stormwater to recharge the aquifer much better than mowed grass. And another related plus, it reduces runoff and its thermal pollution. Besides, you don’t have time to mow grass, because we have work to do!

Our native coldwater fishes including the Potomac Sculpin will survive only if we protect and expand the scattered few habitats where they have taken refuge. They have no choice but to live in these seriously threatened places, but we do. So let’s give ’em some space. How ’bout it? (United States Fish and Wildlife Service image by Ryan Hagerty)

June 29, 2024August 30, 2024

Fish Nurseries in the Susquehanna

Resilient to the pressures of flooding, ice scour, drought, and oft times really poor water quality, Water Willow (Dianthera americana, formerly Justicia americana) is the most common herbaceous plant on the Susquehanna’s non-forested alluvial islands. Yet, few know this native wildflower by name or reputation.

Water Willow on Alluvial Island — Pure stands of emergent Water Willow endure at times brutal conditions on non-forested islands in the Susquenanna.

Water Willow (foreground) and Black Willow. — Alluvial deposits of sand, clay, gravel, and silt create ideal substrate for mats of Water Willow along shorelines of the Susquehanna and its larger tributaries. Provided the loose substrate remains moist, this emergent thrives even when water levels retreat during periods of dry weather. The woody plant in the background, the native Black Willow (Salix nigra), shares similar soil preferences but is found growing on slightly higher ground as a non-emergent tree or shrub. It is a member of the willow family (Salicaceae).

Water Willow in bloom. — In bloom now, the orchid-like flower of the Water Willow is a quick giveaway that it is not a close relative of the willow trees but is instead a member of the acanthus family (Acanthaceae) and is allied with the genus Ruellia, the wild petunias.

The spring of 2024 has been very kind to our beds of Water Willow. Rainfall in the Susquehanna watershed has been frequent enough to maintain river levels just high enough to keep the roots of the plants wet. During the interludes in storm activity, dry spells have rolled back any threat of flooding on the river’s main stem, thus eliminating chances of submerging the plants in muddy water and preventing the sun from keeping them warm, happy, and flowering early. Thundershowers throughout the basin earlier this week have now raised the river a few inches to inundate the base of the plants and make mats of Water Willow favorable places for newly hatched fry and other young fish to take refuge while they grow. Here’s a look…

The Spottail Shiner (Notropis hudsonius) is a common native minnow of the Susquehanna. This juvenile was found among several dozen small fish taking refuge in the cover of Water Willow below Conewago Falls.

The Mimic Shiner (Notropis volucellus) is generally regarded to be a native transplant from the Mississippi drainage that has become established in the Susquehanna and many of its tributaries, possibly after introduction by way of bait buckets. However, the fish tends to be very fragile and dies quickly upon handling, so its use and transport as a bait species may be impractical. The Mimic Shiner is very common in around Conewago Falls.

Juvenile Mimic Shiner — A juvenile Mimic Shiner less than one inch in length found among flooded Water Willow below Conewago Falls earlier this week.

Juvenile Quillback — One of about a dozen juvenile Quillbacks (Carpiodes cyprinus) found in Water Willow just below Conewago Falls. For spawning, local populations of this compact native species of carpsucker favor the gravel-bottomed pools among the Jurassic-Triassic boulders of the falls’ pothole rocks. Probably hatched within the last eight weeks, this specimen was just one inch long.

Spotfin/Satinfin Shiner among Water Willow Rhizomes — As summer progresses, stands of emergent Water Willow begin to expand their size by sending out rhizomes. Increasing numbers of small fish like this Spotfin/Satinfin Shiner (Cyprinella species) concentrate in the cover of the thickening vegetation.

Spotfin/Satinfin Shiners — The importance of these patches of emergent wildflowers (sounds weird, doesn’t it?) is demonstrated by the numbers of fish gathered within their underwater forest of stems and leaves by summertime.

Spotfin/Satinfin Shiners (Cyprinella species) — To protect them from burial by silt and to prevent them from being swept away by current, spawning Spotfin/Satinfin Shiners deposit their eggs in crevices of submerged rocks and wood, often in or near mats of Water Willow. Males guard the eggs until hatching. The fry must then take shelter among boulders, cobble, and plant cover. Note the breeding-condition male in the upper right.

Panfish like this non-native Green Sunfish (Lepomis cyanellus) will often choose nesting sites in deeper water adjacent to beds of Water Willow, particularly if submerged growth like this Water Stargrass adds to the availability of cover for their young after hatching.

Smallmouth Bass gather in a pool adjacent to a Water Willow-covered island. These non-native predators rely on beds of these indigenous plants to provide habitat for their young, then, after spawning, lurk in the waters surrounding them to ambush less-than-vigilant minnows and other victims.

By now you’ve come to appreciate the importance of Water Willow to the sustainability of our populations of fish and other aquatic life. Like similar habitat features that reduce sediment runoff and nutrient pollution, undisturbed stands of terrestrial, emergent, and submerged native plant species are essential to the viability of our freshwater food webs.

June 13, 2024July 5, 2024

Schools of Juvenile Largemouth Bass Learning to Survive

Yesterday, while photographing damselflies on a rehabilitated segment of a warmwater lower Susquehanna valley stream, we noticed some oddly chunky small fish gathered on the surface of a pool along the shoreline.

Damselflies and Small Fish — Perched damselflies and some sort of robust little fish feeding nearby.

Upon further inspection, they appeared to be fingerlings of some type of sunfish or bass. Time for a closer look.

Juvenile Largemouth Bass — At just one inch in length, these juvenile Largemouth Bass (Micropterus salmoides) are already showing signs of the dark lateral stripe that so easily identifies the adult fish.

Adult Largemouth Bass on Spawning Bed — Adult Largemouth Bass began spawning among nearby beds of Spatterdock and other emergent and submerged aquatic vegetation about one month ago, just as water temperatures stabilized to a minimum of the low sixties for several days and nights. Each female can lay thousands of eggs. Only those that are successfully fertilized by the attending male have a chance to hatch.

In the Lower Susquehanna River Watershed, the Largemouth Bass is an introduced species, a native transplant from the Mississippi watershed and Atlantic Slope drainages south of the Chesapeake.

March 28, 2024March 28, 2024

Chesapeake Bay Maritime Accidents and Their Impact on Susquehanna Wildlife

Tuesday’s collision of the container ship Dali into Baltimore’s Francis Scott Key Bridge and the nearly immediate collapse of the span into the chilly waters below reminds us just how unforgiving and deadly maritime accidents can be. Upon termination of rescue and recovery operations, salvage and cleanup will be prioritized as the next steps in the long-term process of reopening the navigable waters to ship traffic and construction of a new bridge. Part of the effort will include monitoring for leaks of fuels and other hazardous materials from the ship, its damaged cargo containers, and vehicles and equipment that were on the bridge when it failed.

Dali vs. Francis Scott Key Bridge — Damage to the hull of the Dali and to the cargo containers on her deck could lead to leaks of hazardous liquids or other materials into Chesapeake Bay. (United States Army Corps of Engineers Baltimore image)

On the waters and shores of today’s Chesapeake, numerous county, state, and federal agencies, including the United States Coast Guard, monitor and inspect looking for conditions and situations that could lead to point-source or accidental discharges of petroleum products and other hazardous materials into the bay. Many are trained, equipped, and organized for emergency response to contain and mitigate spills upon detection. But this was not always the case.

Through much of the twentieth century, maritime spills of oil and other chemicals magnified the effects of routine discharges of hazardous materials and sanitary sewer effluent into the Chesapeake and its tributaries. The cumulative effect of these pollutants progressively impaired fisheries and bay ecosystems leading to noticeable declines in numbers of many aquatic species. Rather frequently, spills or discharges resulted in conspicuous fish and/or bird kills.

One of the worst spills occurred near the mouth of the Potomac River on February 2, 1976, when a barge carrying 250,000 gallons of number 6 oil sank in a storm and lost its cargo into the bay. During a month-long cleanup, the United States Coast Guard recovered approximately 167,000 gallons of the spilled oil, the remainder dispersed into the environment. A survey counted 8,469 “sea ducks” killed. Of the total number, the great majority were Horned Grebes (4,347 or 51.3%) and Long-tailed Ducks (2,959 or 34.9%). Other species included Surf Scoter (Melanitta perspicillata) (405 or 4.8%), Common Loon (195 or 2.3%), Bufflehead (166 or 2.0%), Ruddy Duck (107 or 1.3%), Common Goldeneye (78 or 0.9%), Tundra Swan (46 or 0.5%), Greater Scaup (19 or 0.2%), American Black Duck (12 or 0.2%), Common Merganser (11 or 0.1%), Canvasback (10 or 0.1%), Double-crested Cormorant (10 or 0.1%), Canada Goose (8 or 0.1%), White-winged Scoter (Melanitta deglandi) (7 or 0.1%), Redhead (5 or 0.1%), gull species (10 or 0.1%), miscellaneous ducks and herons (13 or 0.2%) and unidentified (61 or 0.7%). During the spring migration, a majority of these birds would have made their way north and passed through the lower Susquehanna valley. The accident certainly impacted the occurrence of the listed species during that spring in 1976, and possibly for a number of years after.

Horned Grebe during migration on the Susquehanna near Haldeman Riffles. — Of the 8,469 birds killed by the February 2, 1976, oil spill on the Chesapeake, 51.3% (4,347) were Horned Grebes. Many of them would have migrated north through the Lower Susquehanna River Watershed during the coming spring.

The Federal Water Pollution Control Act Amendments of 1972, commonly known as the Clean Water Act, put teeth into the original FWCPCA of 1948 and began reversing the accumulation of pollutants in the bay and other bodies of water around the nation. Additional amendments in 1977 and 1987 have strengthened protections and changed the culture of “dump-and-run” disposal and “dilution-is-the-solution” treatment of hazardous wastes. During the late nineteen-seventies and early nineteen-eighties, emergency response teams and agencies began organizing to control and mitigate spill events. The result has been a greater awareness and competency for handling accidental discharges of fuels and other chemicals into Chesapeake Bay and other waterways. These improvements can help minimize the environmental impact of the Dali’s collision with the Francis Scott Key Bridge in Baltimore.

Oil spills and other pollution in the Chesapeake can impact populations of migratory fish including the anadromous Hickory Shad which are presently transiting the bay on their way to the waters of the Susquehanna below Conowingo Dam.

SOURCES

Roland, John V., Moore, Glenn E., and Bellanca, Michael A. 1977. “The Chesapeake Bay Oil Spill—February 2, 1976: A Case History”. International Oil Spill Conference Proceedings (1977). 1977 (1): 523-527.

January 10, 2024January 18, 2024

Piscivorous Waterfowl Visiting Lakes and Ponds

Heavy rains and snow melt have turned the main stem of the Susquehanna and its larger tributaries into a muddy torrent. For fish-eating (piscivorous) ducks, the poor visibility in fast-flowing turbid waters forces them to seek better places to dive for food. With man-made lakes and ponds throughout most of the region still ice-free, waterfowl are taking to these sources of open water until the rivers and streams recede and clear.

Common Mergansers — The Common Merganser is a species of diving duck with a primary winter range that, along the Atlantic Coast, reaches its southern extreme in the lower Susquehanna and Potomac watersheds. Recently, many have left the main stem of the muddy rivers to congregate on waters with better visibility at some of the area’s larger man-made lakes.

Common Mergansers Feeding — Common Mergansers dive to locate and capture prey, primarily small fish. During this century, their numbers have declined along the southern edge of their winter range, possibly due to birds remaining to the north on open water, particularly on the Great Lakes. In the lower Susquehanna valley, some of these cavity-nesting ducks can now be found year-round in areas where heavy timber again provides breeding sites in riparian forests. After nesting, females lead their young to wander widely along our many miles of larger rivers and streams to feed.

Several Common Mergansers Intimidating a Male with a Freshly Caught Fish — The behavior of these mergansers demonstrates the stiff competition for food that can result when predators are forced away from ideal habitat and become compressed into less favorable space. On the river, piscivores can feed on the widespread abundance of small fish including different species of minnows, shiners, darters, and more. In man-made lakes stocked for recreational anglers with sunfish, bass, and other predators (many of them non-native), small forage species are usually nonexistent. As a result, fish-eating birds can catch larger fish, but are successful far less often. Seen here are several mergansers resorting to intimidation in an effort to steal a young bass away from the male bird that just surfaced with it. While being charged by the aggressors, he must quickly swallow his oversize catch or risk losing it.

With a hard freeze on the way, the fight for life will get even more desperate in the coming weeks. Lakes will ice over and the struggle for food will intensify. Fortunately for mergansers and other piscivorous waterfowl, high water on the Susquehanna is expected to recede and clarify, allowing them to return to their traditional environs. Those with the most suitable skills and adaptations to survive until spring will have a chance to breed and pass their vigor on to a new generation of these amazing birds.

May 28, 2023June 3, 2023

Forty Years Ago in the Lower Rio Grande Valley: Day Eight

Back in late May of 1983, four members of the Lancaster County Bird Club—Russ Markert, Harold Morrrin, Steve Santner, and your editor—embarked on an energetic trip to find, observe, and photograph birds in the Lower Rio Grande Valley of Texas. What follows is a daily account of that two-week-long expedition. Notes logged by Markert some four decades ago are quoted in italics. The images are scans of 35 mm color slide photographs taken along the way by your editor.

DAY EIGHT—May 28, 1983

“Bentsen State Park, Texas”

“Alarm at 6:00 A.M. After breakfast we traveled to Falcon State Park and toured the whole camp area, stopping many places to observe birds. We ran up a good list.”

And so we left what had been our home for the last several days and headed west. In the forty years since our departure that morning, Bentsen-Rio Grande State Park has experienced a number of operational changes. Today, it is a World Birding Center site. For conducting the seasonal hawk census, a tower has been erected to provide counters and observes with an unrestricted view above the treetops. If you wanted to camp in the park now, you would need reservations and would have to hike your gear in to one of only a few primitive campsites. Trailer and motor home accommodations no longer exist. A tram service is now available for touring the park by motor vehicle.

McAllen, Texas, to Falcon Dam — West of Rio Grande City, we exited the river’s outflow delta and entered the Texas scrubland, an area mostly devoid of large trees except in moist soils immediately adjacent to the Rio Grande where the lush vegetation creates a dense subtropical riparian forest in many places. The reservoir itself is known to attract migrating and vagrant waterfowl, waders, shorebirds, gulls, terns, and seabirds. (United States Fish and Wildlife Service base image)

Falcon State Park is located along the east shore of Falcon Reservoir. There are no shade trees beneath which one can escape the scorching rays of the sun on a hot day. This is the easternmost section of the scrubland’s Tamaulipan Saline Thornscrub, a xeric plant community of head-high brush found only on clay soils with a particularly high salinity. Many of the plants look similar to other varieties of shrubs and small trees with which one may be familiar, except nearly all of them are covered with nasty thorns and prickles. And yes, there are cactus. You can’t make your way bushwhacking cross country without obtaining cuts, gashes, and scars to show for it. The Falcon State Recreation Area bird checklist published in 1977 has a nice description of the plants found there—mesquite, ebano, guaycan, blackbrush and catclaw acacia, granjeno, coyotillo, huisache, tasajillo, prickly pear, allthorn, cenizo, colima, and yucca. In the margins between the thornscrub growth, there is an abundance of grasses and wildflowers. On nearby ridges, Tamaulipan Calcareous Thornscrub, a similar xeric plant community, occupies soils with a higher content of calcium carbonate. Together, these communities comprise much of the Tamaulipan Mezquital ecoregion of scrublands in Starr County and western Hidalgo County in the Rio Grande valley of Texas.

After being greeted by a Greater Roadrunner at the campsite, we took a walk to the nearby shoreline of the reservoir. We spotted Olivaceous Cormorants perched on some dead limbs in the water nearby. Known today as Neotropic Cormorant (Phalacrocorax brasilianus), it is yet another specialty of the Rio Grande Valley. Elsewhere on or near the water—Cattle Egret, Great Egret, Black-bellied Whistling Duck, Osprey, Common Gallinule, Killdeer, Laughing Gull, Forster’s Tern (Sterna forsteri), Least Tern, and Caspian Tern were seen.

In the thornscrub around the campground, which, like Bentsen-Rio Grande State Park, we had pretty much to ourselves, we saw Scissor-tailed Flycatcher, Curve-billed Thrasher, White-winged Dove, Mourning Dove, Ground Dove, Inca Dove, and White-tipped Dove. A single Chihuahuan Raven was a fly by. We saw and smelled several road-killed Nine-banded Armadillos (Dasypus novemcinctus), but never found one alive.

Then, it started to rain. Not just a shower, but a soaker that persisted through much of the day. Rainy days can make for great birding, so we kept at it. Unfortunately, such days aren’t too ideal for photography, so we did only what we could without ruining our equipment.

In the campground at Falcon State Park, a Cactus Wren (*Campylorhynchus brunneicapillus*) takes shelter from the rain beneath a canopy protecting a picnic table.

“Finally we drove to the spillway of the dam and parked.”

Falcon Dam was another of the numerous flood-control projects built on the Rio Grande during the middle of the twentieth century. Behind it, Falcon Reservoir stores water for irrigation and operation of a hydroelectric generating station located within the dam complex. Construction of the dam and power plant was a joint venture shared by Mexico and the United States. The project was dedicated by Presidents Adolfo Ruiz Cortines and Dwight D. Eisenhower in 1953.

Rainy days aside, the route precipitation takes to reach the Falcon Reservoir and the Lower Rio Grande Valley includes hundreds of miles through arid grasslands and scrublands. Along the way, much of that water is lost to natural processes including evaporation and aquifer recharge, but an increasing percentage of the volume is being removed by man for civil, industrial, and agricultural uses. Can the Rio Grande and its tributaries continue to meet demand?

Rio Grande Drainage Basin — The Rio Grande’s headwaters can be found in south-central Colorado where spring snow melt is vital to establishing adequate flow to allow the river to recharge aquifers in the hundreds of miles of arid lands through which it flows. Presently, dams in the upper reaches of the river are operated to hold water during the spring thaw, then release it slowly to compensate for base flow lost to withdrawals for irrigation in extensive areas of the middle reaches of the watershed. With everyone wanting their take, is there enough water to go around? Diminishing ground water levels suggest the answer is no. (National Oceanic and Atmospheric Administration base image)

Ground Water Recharge in the Rio Grande Watershed — Using the streamflow recharge process, the flowing Rio Grande provides water to the aquifers in the arid regions through which it passes. Click the image for a slightly larger version. (United States Geological Survey image)

“On the way in we saw and photographed an apparent sick or injured Swainson’s Hawk. We approached it very close.”

“At the spillway we sat in the camper, except when the rain slackened, then we stood out and watched in vain for the Green or Ringed Kingfisher, which we never did see.”

At the spillway House Sparrows, Rough-winged Swallows, and Cliff Swallows were nesting on the dam, the latter two species grabbing flying insects above the waters of the Rio Grande.

Despite the rain, anglers were fishing along the spillway walls where this young man caught a Longnose Gar (Lepisosteus osseus). The reservoir has been stocked with Alligator Gar (Atractosteus spatula), Spotted Gar (Lepisosteus oculatus), and a variety of bass, catfish, and other species to establish a trophy sport fishery. Many specimens grow to become trophy-size there due to the warm water temperatures. These guys though were fishing for food, not trophies.

“I made dinner here at this spillway and we continued to watch. The rain almost stopped, so we walked down the road about 1 1/2 miles, during which time we saw a lifer for Harold — Hook-billed Kite. We followed Father Tom’s directions to a spot for the Ferruginous Owl — no luck.”

A Red-billed Pigeon in a willow tree in the subtropical riparian forest along the banks of the Rio Grande below Falcon Dam. Prior to this trip, neither Steve nor I had ever seen this tropical species before.

“Back at the spillway we had supper and then repeated the hike — no Ferruginous Owl, but a Barn Owl and Great Horned Owl. Back to our #201 campsite and wrote up the day’s log.”

Trees along the river provided habitat for orioles and other species. Since the rain had subsided, we decided to see what might come out and begin feeding. Soon, we not only saw an Altamira Oriole, but found Hooded Oriole (Icterus cucullatus) and the yellow and black tropical species, Audubon’s Oriole (Icterus graduacauda), formerly known as Black-headed Oriole. Three species of orioles on a backdrop of lush green subtropical foliage, it was magnificent.

Along the dirt road below the dam, the mix of scrubland and subtropical riparian forest made for excellent birding. We not only found a soaring Hook-billed Kite, one of the target birds for the trip, but we had good looks at both a Great Horned Owl, then a Barn Owl (Tyto alba) that we flushed from the bare ground in openings among the vegetation as we walked the through. Both had probably pounced on some sort of small prey species prior to our arrival. Because there are seldom crows or ravens to bother them, owls here are more active during than day than they are elsewhere. The subject of this afternoon’s intensive search, the elusive and diminutive Ferruginous Pygmy Owl (Glaucidium brasilianum), is routinely diurnal. Other sightings on our two walks included Turkey Vulture, Black Vulture, White-tailed Kite, Northern Bobwhite, Yellow-billed Cuckoo, Golden-fronted Woodpecker, Ladder-backed Woodpecker, Couch’s Kingbird, Brown-crested Flycatcher, Green Jay, Black-crested Titmouse, Mockingbird, Long-billed Thrasher, Great-tailed Grackle, Bronzed Cowbird, Northern Cardinal, and Painted Bunting.

The day finished as so many others had earlier during the trip—with insect-hunting Common Nighthawks calling from the skies around our campsite.

December 19, 2022

Photo of the Day

A juvenile Ring-billed Gull with a freshly caught Gizzard Shad is pursued by a hungry adult Great Black-backed Gull on the Susquehanna at Conowingo Dam.

August 10, 2022August 13, 2022

Shakedown Cruise of the S. S. Haldeman

First there was the Nautilus. Then there was the Seaview. And who can forget the Yellow Submarine? Well, now there’s the S. S. Haldeman, and today we celebrated her shakedown cruise and maiden voyage. The Haldeman is powered by spent fuel that first saw light of day near Conewago Falls at a dismantled site that presently amounts to nothing more than an electrical substation. Though antique in appearance, the vessel discharges few emissions, provided there aren’t any burps or hiccups while underway. So, climb aboard as we take a cruise up the Susquehanna at periscope depth to have a quick look around!

Brunner Island as seen from the east channel.

Close-in approach to emergent Water Willow growing on an alluvial Island.

The approach to York Haven Dam and Conewago Falls from the west channel.

A pair of Powdered Dancers on a midriver log.

Watertight and working fine. Let’s flood the tanks and have a peek at the benthos. Dive, all dive!

American Eelgrass/Tapegrass — American Eelgrass, also known as Tapegrass, looks to be growing well in the channels. Historically, vast mats of this plant were the primary food source for the thousands of Canvasback ducks that once visited the lower Susquehanna each autumn.

As is Water Stargrass (Heteranthera dubia). When mature, both of these native plants provide excellent cover for young fish. Note the abundance of shells from deceased Asiatic Clams (Corbicula fluminea) covering the substrate.

Mayfly nymph — A three-tailed mayfly (Ephemeroptera) nymph and a several exoskeletons cling to the downstream side of a rock.

Comb-lipped Casemaker Caddisfly larva and case. — This hollowed-out stick may be a portable protective shelter belonging to a Comb-lipped Casemaker Caddisfly larva (Calamoceratidae). The larva itself appears to be extending from the end of the “case” in the upper right of the image. Heteroplectron americanum, a species known for such behavior, is a possibility.

In the Susquehanna and its tributaries, the Rusty Crayfish (Faxonius rusticus) is an introduced invasive species. It has little difficulty displacing native species due to its size and aggressiveness.

Freshwater Snails Susquehanna: Virginian River Horn Snail — Summers with conditions that promote eelgrass and stargrass growth tend to be big years for Virginian River Horn Snails (Elimia virginica). 2022 appears to be one of those years. They’re abundant and they’re everywhere on the rocks and gravel substrate in midriver. Feeding almost incessantly on algae and detritus, these snails are an essential component of the riverine ecosystem, breaking down organic matter for final decomposition by bacteria and fungi.

Susquehanna Snails: Virginian River Horn Snails and Lesser Mystery Snails — Virginian River Horn Snails with Lesser Mystery Snails (Campeloma decisum), another native species commonly encountered at Conewago Falls and in surrounding waters.

Freshwater Snails Susquehanna: River Snail and Virginian River Horn Snail — A River Snail (Leptoxis carinata), also known as a Crested Mudalia, hitching a ride on a Virginian River Horn Snail. The two species are frequently found together.

Mollusks of the Susquehanna: Yellow Lampmussel and River Snail — A River Snail cleaning the shell of a native freshwater Unionidae mussel, Lampsilis cariosa, commonly called the Yellow Lampmussel or Carried Lampmussel. Because of their general decline in abundance and range, all Unionidae mussels are protected in Pennsylvania.

Fishes of the Susquehanna: Banded Darter — The Banded Darter (Etheostoma zonale) is a member of the perch family (Percidae).

Fishes of the Susquehanna: Smallmouth Bass — A Smallmouth Bass in strong current.

Fishes of the Susquehanna: Spotfin or Satinfin Shiners — Along the edge of an alluvial island at midriver, Cyprinella (Spotfin or Satinfin) Shiners gather in the cover of an emergent stand of Water Willow. The closely related Spotfin Shiner (Cyprinella spiloptera) and Satinfin Shiner (Cyprinella analostanus) are nearly impossible to differentiate in the field.

Fishes of the Susquehanna: Spotfin or Satinfin Shiner — A breeding condition male Cyprinella (Spotfin or Satinfin) Shiner.

Fishes of the Susquehanna; Juvenile Channel Catfish — A juvenile Channel Catfish.

We’re finding that a sonar “pinger” isn’t very useful while running in shallow water. Instead, we should consider bringing along a set of Pings—for the more than a dozen golf balls seen on the river bottom. It appears they’ve been here for a while, having rolled in from the links upstream during the floods. Interestingly, several aquatic species were making use of them.

A golf ball used as an anchor point for silk cases woven by Net-spinning Caddisfly (Hydropsychidae) larvae to snare food from the water column.

Freshwater Snails (Gastropods) of the Lower Susquehanna River Watershed: Creeping Ancylid (Ferrissia species) — A Creeping Ancylid (Ferrissia species), a tiny gastropod also known as a Coolie Hat Snail, River Limpet, or Brook Freshwater Limpet, inhabits the dimple on a “Top Flight”.

We now know why your golf balls always end up in the drink, it’s where they go to have their young.

Well, it looks like the skipper’s tired and grumpy, so that’s all for now. Until next time, bon voyage!

July 25, 2022August 15, 2022

If I Had a Pool, Here’s What I Would Do

On these hot and steamy days of summer, I get to thinking about how great it would be to have a swimming pool. Maybe I would take a little dip, you know, just to cool down. But when I really get to thinking about it, I just might do what the professor has done with his pool.

Or maybe I would do what this lady did with her pool.

Or what this lady did.

Yep, I’d do this.

This is what I’d do.

Wouldn’t you?

April 25, 2021April 22, 2022

Conowingo Dam: Cormorants, Eagles, Snakeheads and a Run of Hickory Shad

Meet the Double-crested Cormorant, a strangely handsome bird with a special talent for catching fish. You see, cormorants are superb swimmers when under water—using their webbed feet to propel and maneuver themselves with exceptional speed in pursuit of prey.

Like many species of birds that dive for their food, Double-crested Cormorants run across the surface of the water to gain speed for a takeoff. Smaller wings may make it more difficult to get airborne, but when folded, they provide improved streamlining for submerged swimming.

Double-crested Cormorants, hundreds of them, are presently gathered along with several other species of piscivorous (fish-eating) birds on the lower Susquehanna River below Conowingo Dam near Rising Sun, Maryland. Fish are coming up the river and these birds are taking advantage of their concentrations on the downstream side of the impoundment to provide food to fuel their migration or, in some cases, to feed their young.

Double-crested Cormorants, mostly adult birds migrating toward breeding grounds to the north, are gathered on the rocks on the east side of the river channel below Conowingo Dam. A Great Blue Heron from a nearby rookery can be seen at the center of the image.

Bald Eagles normally gather in large numbers at Conowingo Dam in the late fall and early winter. Presently there are more than 50 there, and the majority of them are breeding age adults. Presumably they are still on their way north to nest. Meanwhile, local pairs are already feeding young, so it seems these transient birds are running a bit late. Many of them can be seen on the rocks along the east side of the river channel,…

…on the powerline trestles on the island below the dam…

…and in the trees surrounding Fisherman’s Park on the west shore.

In addition to the birds, the movements of fish attract larger fish, and even larger fishermen.

Anglers gather to fish the placid waters below the dam’s hydroelectric powerhouse . Only a few of the generating turbines are operating, so the flow through the dam is minimal.

Some water is being released along the west shoreline to attract migratory river herring to the west fish lift for sorting and retention as breeding stock for a propagation program. The east lift, the passage that hoists American Shad (Alosa sapidissima) to a trough that allows them to swim over the top of the dam to waters upriver, will begin operating as soon as these larger migratory fish begin arriving.

The excitement starts when the sirens start to wail and the red lights begin flashing. Yes friends, it’s showtime.

Red lights and sirens are a warning that additional flow is about to be released from the dam. Boaters should anticipate rough water and persons in and along the river need to seek higher ground immediately.

Gates are opened at mid-river to release a surge of water through the dam.

The wake from the release quickly reaches the shoreline, raising the water level in moments.

Experienced anglers know that the flow through the dam gets fish moving and can improve the catch significantly, especially in spring when many species are ascending the river.

Within minutes of the renewed flow, birds are catching fish.

A Double-crested Cormorant with a young Channel Catfish (Ictalurus punctatus).

A Double-crested Cormorant fleeing others trying to steal its Channel Catfish.

Another Double-crested Cormorant eating a Channel Catfish. Did you realize that Channel Catfish were an introduced species in the Susquehanna River system?

An Osprey with a stick, it’s too busy building a nest right now to fish.

Great Blue Herons swallow their prey at the spot of capture, then fly back to the nest to regurgitate a sort of “minced congealed fish product” to their young.

Then the anglers along the wave-washed shoreline began catching fish too.

This young man led off a flurry of catches that would last for the remainder of the afternoon.

Though Gizzard Shad are filter feeders that don’t readily take baits and lures, they are regularly foul-hooked and reeled in from the large schools that ascend the river in spring.

Gizzard Shad are very abundant in the lower Susquehanna, providing year-round forage for many species of predatory animals including Bald Eagles.

A Double-crested Cormorant swallowing a Gizzard Shad.

This angler soon helped another fisherman by landing his large catch, a Northern Snakehead (Channa argus).

The teeth of a Northern Snakehead are razor sharp. It is an aggressive non-native invasive species currently overtaking much of the Lower Susquehanna River Watershed. Anglers are encouraged to fish for them, catch them, keep them, and kill them at the site of capture. **Never transport a live Northern Snakehead anywhere at any time. It is illegal in both Maryland and Pennsylvania to possess a live snakehead.**

Northern Snakehead advisory sign posted at Exelon Energy’s Conowingo Fishermen’s Park.

A stringer of Northern Snakeheads. This species was imported from Asia as a food fish, so it has excellent culinary possibilities. It’s better suited for a broiler or frying pan than a river or stream.

Another stringer of Northern Snakeheads. It’s pretty safe to say that they have quickly become one of the most abundant predatory fish in the river. Their impact on native species won’t be good, so catch and eat as many as you can. Remember, snakeheads swim better in butter and garlic than in waters with native fish.

This foul-hooked Shorthead Redhorse (Moxostoma macrolepidotum), a native species of sucker, was promptly released.

Striped Bass are anadromous fish that leave the sea in spring to spawn in fresh water. They ascend the Susquehanna in small numbers, relying upon the operation of the fish passages at the Conowingo, Holtwood, Safe Harbor, and York Haven Dams to continue their journey upstream. During spring spawning, Striped Bass in the Susquehanna River and on the Susquehanna Flats portion of the upper Chesapeake Bay are not in season and may not be targeted, even for catch-and-release. This accidental catch was immediately turned loose.

After removal from the hook, this hefty Smallmouth Bass was returned to the river. Many anglers are surprised to learn that Smallmouth Bass are not native to the Susquehanna basin.

This angler’s creel contains a Northern Snakehead (left) and a Walleye (right). Did you know that the Walleye (Sander vitreus) is an introduced species in the Susquehanna watershed?

By late afternoon, anglers using shad darts began hooking into migrating Hickory Shad (Alosa mediocris), a catch-and-release species in Maryland.

Hickory Shad are recognized by their lengthy lower jaw. They are anadromous herring that leave the sea to spawn in freshwater streams. Hickory Shad ascend the Susquehanna as far as Conowingo Dam each year, but shy away from the fish lifts. Downriver from the dam, they do ascend Deer Creek along the river’s west shore and Octoraro Creek on the east side. In Pennsylvania, the Hickory Shad is an endangered species.

A Hickory Shad angled on a dual shad dart rig. During the spring spawning run, they feed mostly on small fish, and are the most likely of the Susquehanna’s herring to take the hook.

Simultaneous hook-ups became common after fours hours worth of release water from the dam worked its way toward the mouth of the river and got the schools moving. Water temperatures in the mid-to-upper-fifties trigger the ascent of Hickory Shad. On the Susquehanna, those temperatures were slow to materialize in the spring of 2021, so the Hickory Shad migration is a bit late.

Catch-and-release fishing for Hickory Shad appears to be in full swing not only at the dam, but along the downstream shoreline to at least the mouth of Deer Creek at Susquehanna State Park too.

Many Hickory Shad could be seen feeding on some of the millions of caddisflies (Trichoptera) swarming on the river. These insects, along with earlier hatches of Winter Stoneflies (Taeniopterygidae), not only provide forage for many species of fish, but are a vital source of natural food for birds that migrate up the river in March and April each year. Swallows, Ring-billed Gulls, and Bonaparte’s Gulls are particularly fond of snatching them from the surface of the water.

A Winter Stonefly (Taeniopterygidae) from an early-season hatch on the Susquehanna River at the Veteran’s Memorial Bridge at Columbia/Wrightsville, Pennsylvania. (March 3, 2021)

Just below Conowingo Dam, a lone fly fisherman was doing a good job mimicking the late-April caddisfly hatch, successfully reeling in numerous surface-feeding Hickory Shad.

You may have noticed the extraordinary number of introduced fish species listed in this account of a visit to Conowingo Dam. Sorry to say that there are two more: the Flathead Catfish (Pylodictis olivaris) and the Blue Catfish (Ictalurus furcatus). Like the Northern Snakehead, each has become a plentiful invasive species during recent years. Unlike the Northern Snakehead, these catfish are “native transplants”, species introduced from populations in the Mississippi River and Gulf Slope drainages of the United States. So if you visit the area, consider getting a fishing license and catching a few. Like the snakeheads, they too are quite palatable.

The arrival of migrating Hickory Shad heralds the start of a movement that will soon include White Perch, anadromous American Shad, and dozens of other fish species that swim upstream during the springtime. Do visit Fisherman’s Park at Conowingo Dam to see this spectacle before it’s gone. The fish and birds have no time to waste, they’ll soon be moving on.

To reach Exelon’s Conowingo Fisherman’s Park from Rising Sun, Maryland, follow U.S. Route 1 south across the Conowingo Dam, then turn left onto Shuresville Road, then make a sharp left onto Shureslanding Road. Drive down the hill to the parking area along the river. The park’s address is 2569 Shureslanding Road, Darlington, Maryland.

A water release schedule for the Conowingo Dam can be obtained by calling Exelon Energy’s Conowingo Generation Hotline at 888-457-4076. The recording is updated daily at 5 P.M. to provide information for the following day.

And remember, the park can get crowded during the weekends, so consider a weekday visit.

January 9, 2021January 14, 2021

2020: A Good Year

You say you really don’t want to take a look back at 2020? Okay, we understand. But here’s something you may find interesting, and it has to do with the Susquehanna River in 2020.

As you may know, the National Weather Service has calculated the mean temperature for the year 2020 as monitored just upriver from Conewago Falls at Harrisburg International Airport. The 56.7° Fahrenheit value was the highest in nearly 130 years of monitoring at the various stations used to register official climate statistics for the capital city. The previous high, 56.6°, was set in 1998.

Though not a prerequisite for its occurrence, record-breaking heat was accompanied by a drought in 2020. Most of the Susquehanna River drainage basin experienced drought conditions during the second half of the year, particularly areas of the watershed upstream of Conewago Falls. A lack of significant rainfall resulted in low river flows throughout late summer and much of the autumn. Lacking water from the northern reaches, we see mid-river rocks and experience minimal readings on flow gauges along the lower Susquehanna, even if our local precipitation happens to be about average.

Back in October, when the river was about as low as it was going to get, we took a walk across the Susquehanna at Columbia-Wrightsville atop the Route 462/Veteran’s Memorial Bridge to have a look at the benthos—the life on the river’s bottom.

As we begin our stroll across the river, we quickly notice Mallards and a Double-crested Cormorant (far left) feeding among aquatic plants. You can see the leaves of the vegetation just breaking the water’s surface, particularly behind the feeding waterfowl. Let’s have a closer look.

An underwater meadow of American Eelgrass (Vallisneria americana) as seen from atop the Veteran’s Memorial Bridge at Columbia-Wrightsville. Also known as Freshwater Eelgrass, Tapegrass, and Wild Celery, it is without a doubt the Susquehanna’s most important submerged aquatic plant. It grows in alluvial substrate (gravel, sand, mud, etc.) in river segments with moderate to slow current. Water three to six feet deep in bright sunshine is ideal for its growth, so an absence of flooding and the sun-blocking turbidity of muddy silt-laden water is favorable.

Plants in the genus Vallisneria have ribbon-like leaves up to three feet in length that grow from nodes rooted along the creeping stems called runners. A single plant can, over a period of years, spread by runners to create a sizable clump or intertwine with other individual plants to establish dense meadows and an essential wildlife habitat.

An uprooted segment of eelgrass floats over a thick bed of what may be parts of the same plant. Eelgrass meadows on the lower Susquehanna River were decimated by several events: deposition of anthracite coal sediments (culm) in the late-nineteenth and early-twentieth centuries, dredging of the same anthracite coal sediments during the mid-twentieth century, and the ongoing deposition of sediments from erosion occurring in farm fields, logged forests, abandoned mill ponds, and along denuded streambanks. Not only has each of these events impacted the plants physically by either burying them or ripping them out by the roots, each has also contributed to the increase in water turbidity (cloudiness) that blocks sunlight and impairs their growth and recovery.

A submerged log surrounded by beds of eelgrass forms a haven for fishes in sections of the river lacking the structure found in rock-rich places like Conewago Falls. A period absent of high water and sediment runoff extended through the growing season in 2020 to allow lush clumps of eelgrass like these to thrive and further improve water quality by taking up nutrients, particularly nitrogen and phosphorus. Nutrients used by vascular plants including eelgrass become unavailable for feeding detrimental algal blooms in downstream waters including Chesapeake Bay.

Small fishes and invertebrates attract predatory fishes to eelgrass beds. We watched this Smallmouth Bass leave an ambush site among eelgrass’s lush growth to shadow a Common Carp as it rummaged through the substrate for small bits of food. The bass would snatch up crayfish that darted away from the cover of stones disturbed by the foraging carp.

Sunfishes are among the species taking advantage of eelgrass beds for spawning. They’ll build a nest scrape in the margins between clumps of plants allowing their young quick access to dense cover upon hatching. The abundance of invertebrate life among the leaves of eelgrass nourishes feeding fishes, and in turn provides food for predators including Bald Eagles, this one carrying a freshly-caught Bluegill.

These improvements in water quality and wildlife habitat can have a ripple effect. In 2020, the reduction in nutrient loads entering Chesapeake Bay from the low-flowing Susquehanna may have combined with better-than-average flows from some of the bay’s lesser-polluted smaller tributaries to yield a reduction in the size of the bay’s oxygen-deprived “dead zones”. These dead zones typically occur in late summer when water temperatures are at their warmest, dissolved oxygen levels are at their lowest, and nutrient-fed algal blooms have peaked and died. Algal blooms can self-enhance their severity by clouding water, which blocks sunlight from reaching submerged aquatic plants and stunts their growth—making quantities of unconsumed nutrients available to make more algae. When a huge biomass of algae dies in a susceptible part of the bay, its decay can consume enough of the remaining dissolved oxygen to kill aquatic organisms and create a “dead zone”. The Chesapeake Bay Program reports that the average size of this year’s dead zone was 1.0 cubic miles, just below the 35-year average of 1.2 cubic miles.

Back on a stormy day in mid-November, 2020, we took a look at the tidal freshwater section of Chesapeake Bay, the area known as Susquehanna Flats, located just to the southwest of the river’s mouth at Havre de Grace, Maryland. We wanted to see how the restored American Eelgrass beds there might have fared during a growing season with below average loads of nutrients and life-choking sediments spilling out of the nearby Susquehanna River. Here’s what we saw.

We followed the signs from Havre de Grace to Swan Harbor Farm Park.

Harford County Parks and Recreation’s Swan Harbor Farm Park consists of a recently-acquired farming estate overlooking the tidal freshwater of Susquehanna Flats.

Along the bay shore, a gazebo and a fishing pier have been added. Both provide excellent observation points.

The shoreline looked the way it should look on upper Chesapeake Bay, a vegetated buffer and piles of trees and other organic matter at the high-water line. There was less man-made garbage than we might find following a summer that experienced an outflow from river flooding, but there was still more than we should be seeing.

Judging by the piles of fresh American Eelgrass on the beach, it looks like it’s been a good year. Though considered a freshwater plant, eelgrass will tolerate some brackish water, which typically invades upper Chesapeake Bay each autumn due to a seasonal reduction in freshwater inflow from the Susquehanna and other tributaries. Saltwater can creep still further north when the freshwater input falls below seasonal norms during years of severe drought. The Susquehanna Flats portion of the upper bay very rarely experiences an invasion by brackish water; there was none in 2020.

As we scanned the area with binoculars and a spotting scope, a raft of over one thousand ducks and American Coots (foreground) could be seen bobbing among floating eelgrass leaves and clumps of the plants that had broken away from their mooring in the mud. Waterfowl feed on eelgrass leaves and on the isopods and other invertebrates that make this plant community their home.

While coots and grebes seemed to favor the shallower water near shore, a wide variety of both diving and dabbling ducks were widespread in the eelgrass beds more distant. Discernable were Ring-necked Ducks, scaup, scoters, Long-tailed Ducks, Redheads, American Wigeons, Gadwall, Ruddy Ducks, American Black Ducks, and Buffleheads.

We noticed a few Canvasbacks (Aythya valisineria) on the Susquehanna Flats during our visit. Canvasbacks are renowned as benthic feeders, preferring the tubers and other parts of submerged aquatic plants (a.k.a. submersed aquatic vegetation or S.A.V.) including eelgrass, but also feeding on invertebrates including bivalves. The association between Canvasbacks and eelgrass is reflected in the former’s scientific species name valisineria, a derivitive of the genus name of the latter, Vallisneria.

Canvasbacks on Chesapeake Bay. (United States Fish and Wildlife Service image by Ryan Hagerty)

The plight of the Canvasback and of American Eelgrass on the Susquehanna River was described by Herbert H. Beck in his account of the birds found in Lancaster County, Pennsylvania, published in 1924:

“Like all ducks, however, it stops to feed within the county less frequently than formerly, principally because the vast beds of wild celery which existed earlier on broads of the Susquehanna, as at Marietta and Washington Borough, have now been almost entirely wiped out by sedimentation of culm (anthracite coal waste). Prior to 1875 the four or five square miles of quiet water off Marietta were often as abundantly spread with wild fowl as the Susquehanna Flats are now.”

Beck quotes old Marietta resident and gunner Henry Zink:

“Sometimes there were as many as 500,000 ducks of various kinds on the Marietta broad at one time.”

The abundance of Canvasbacks and other ducks on the Susquehanna Flats would eventually plummet too. In the 1950s, there were an estimated 250, 000 Canvasbacks wintering on Chesapeake Bay, primarily in the area of the American Eelgrass, a.k.a. Wild Celery, beds on the Susquehanna Flats. When those eelgrass beds started disappearing during the second half of the twentieth century, the numbers of Canvasbacks wintering on the bay took a nosedive. As a population, the birds moved elsewhere to feed on different sources of food, often in saltier estuarine waters.

Canvasbacks were able to eat other foods and change their winter range to adapt to the loss of habitat on the Susquehanna River and Chesapeake Bay. But not all species are the omnivores that Canvasbacks happen to be, so they can’t just change their diet and/or fly away to a better place. And every time a habitat like the American Eelgrass plant community is eliminated from a region, it fragments the range for each species that relied upon it for all or part of its life cycle. Wildlife species get compacted into smaller and smaller suitable spaces and eventually their abundance and diversity are impacted. We sometimes marvel at large concentrations of birds and other wildlife without seeing the whole picture—that man has compressed them into ever-shrinking pieces of habitat that are but a fraction of the widespread environs they once utilized for survival. Then we sometimes harass and persecute them on the little pieces of refuge that remain. It’s not very nice, is it?

By the end of 2020, things on the Susquehanna were getting back to normal. Near normal rainfall over much of the watershed during the final three months of the year was supplemented by a mid-December snowstorm, then heavy downpours on Christmas Eve melted it all away. Several days later, the Susquehanna River was bank full and dishing out some minor flooding for the first time since early May. Isn’t it great to get back to normal?

The rain-and-snow-melt-swollen Susquehanna from Chickies Rock looking upriver toward Marietta during the high-water crest on December 27th.

Cresting at Columbia as seen from the Route 462/Veteran’s Memorial Bridge. A Great Black-backed Gull monitors the waters for edibles.

All back to normal on the Susquehanna to end 2020.

Yep, back to normal on the Susquehanna. Maybe 2021 will turn out to be another good year, or maybe it’ll just be a Michelin or Firestone.

SOURCES

Beck, Herbert H. 1924. A Chapter on the Ornithology of Lancaster County, Pennsylvania. The Lewis Historical Publishing Company. New York, NY.

White, Christopher P. 1989. Chesapeake Bay, Nature of the Estuary: A Field Guide. Tidewater Publishers. Centreville, MD.

November 17, 2020November 17, 2020

Bald Eagles Arriving at Conowingo Dam

You need to see this to believe it—dozens, sometimes hundreds, of Bald Eagles doing their thing and you can stand or sit in just one place to take it all in.

Conowingo Dam on the Susquehanna River near Darlington, Maryland, attracts piscivores galore. Young Gizzard Shad (Dorosoma cepedianum) and other small fishes are temporarily stunned as they pass through the turbines and gated discharges at the hydroelectric facility’s power house. Waiting for them in the rapids below are predatory fishes including Striped Bass (Morone saxatilis), White Perch (Morone americana), several species of catfishes, and more. From above, fish-eating birds are on the alert for a disoriented turbine-traveler they can easily seize for a quick meal.

U.S. Route 1 crosses the Susquehanna River atop the Conowingo Dam. Conowingo Fisherman’s Park, the observation site for the dam’s Bald Eagles and other birds, is located downstream of the turbine building along the river’s west (south) shore. As the name implies, the park is a superb location for angling.

Heed this warning. Close your windows and sunroof or the vultures will subject your vehicle’s contents to a thorough search for food. Then they’ll deposit a little consolation prize on your paint.

Scavenging Black Vultures congregate by the hundreds at Conowingo Dam to clean up the scraps left behind by people and predators. They’ll greet you right in the parking lot.

Photographers line up downstream of the turbine building for an opportunity to get the perfect shot of a Bald Eagle.

The operator of the Conowingo Hydroelectric Generating Station, Exelon Energy, provides clean comfortable facilities for fishing, sightseeing, and wildlife observation.

There’s almost always a Peregrine Falcon zooming around the dam to keep the pigeons on their toes.

Double-crested Cormorants on the boulders that line the channel below the dam. Hundreds are there right now.

Double-crested Cormorants dive for fish near the power house discharge, which, while just one small generator is operating, seems nearly placid. The feeding frenzy really gets going when Conowingo begins generating with multiple large turbines and these gently flowing waters become torrential rapids filled with disoriented fishes.

Ring-billed Gulls seek to snag a small fish from the water’s surface.

After successfully nabbing shad or perch, these Double-crested Cormorants need to swallow their catch fast or risk losing it. Stealing food is a common means of survival for the gulls, eagles, and other birds found here.

Where do migrating eagles go? There are, right now, at least 50 Bald Eagles at Conowingo Dam, with more arriving daily. Numbers are likely to peak during the coming weeks.

Eagles can be seen perched in the woods along both river shorelines, even in the trees adjacent to the Conowingo Fisherman’s Park car lot. Others take stand-by positions on the boulders below the dam.

To remind visiting eagles that they are merely guests at Conowingo, a resident Bald Eagle maintains a presence at its nest on the wooded slope above Fisherman’s Park. Along the lower Susquehanna, female Bald Eagles lay eggs and begin incubation in January.

When an eagle decides to venture out and attempt a dive at a fish, that’s when the photographers rush to their cameras for a chance at a perfect shot.

The extraordinary concentrations of Bald Eagles at Conowingo make it an excellent place to study the plumage differences between birds of various ages.

Here’s a first-year Bald Eagle, also known as a hatch-year or juvenile bird.

A second-year or Basic I immature Bald Eagle. Note the long juvenile secondaries giving the wings a ragged-looking trailing edge.

A third-year or Basic II immature Bald Eagle.

A second-year/Basic I immature Bald Eagle (top) and a third-year/Basic II immature Bald Eagle (bottom).

A second-year/Basic I immature Bald Eagle (bottom) and a third-year/Basic II immature Bald Eagle (top). Note the white feathers on the backs of eagles in these age classes.

A third-year/Basic II immature Bald Eagle perched in a tree alongside the parking area. Note the Osprey-like head plumage.

A sixth-year or older adult Bald Eagle in definitive plumage (left) and a fourth-year or Basic III immature Bald Eagle (right).

If you want to see the Bald Eagles at Conowingo Dam, don’t wait. While many birds are usually present throughout the winter, the large concentrations may start dispersing as early as December when eagles begin wandering in search of other food sources, particularly if the river freezes.

A pair of Bald Eagles is already working on a nest atop this powerline trestle downstream of Conowingo Dam. By late December, most adult eagles will depart Conowingo to begin spending their days establishing and defending breeding territories elsewhere. Any non-adult eagles still loitering around the dam will certainly begin receiving encouragement from the local nesting pair(s) to move along as well.

As Bald Eagle numbers continue to increase, expect the parking lot to become full during weekends and over the Thanksgiving holiday. To avoid the crowds, plan to visit during a weekday.

You can get the generating schedule for the Conowingo Dam by calling the Conowingo Generation Hotline at 888-457-4076. The recording is updated daily at 5 P.M. to provide information for the following day.

May 28, 2020May 28, 2020

How Much Can a Bald Eagle Lift?

Here it is—just as it happened, recently in the lower Susquehanna valley.

The eagle glides in and grabs an unsuspecting Common Carp, the bird’s momentum and a head wind helping it to raise the fish out of the water. This particular carp appears to be eighteen to twenty-four inches in length. In that range, it could weigh anywhere from four to ten pounds or more.

But as the eagle tries to flap its wings to carry the carp skyward, it loses lift, and the weight of the fish drags the raptor down. A carp exceeding twenty inches in length usually weighs more than five pounds, approximately half the mass of a male (about 9 lbs.) or a female (about 12 lbs.) Bald Eagle. From a dead stop, trying to extricate a submerged fish weighing half as much again as it does is an impossible task for this or any other Bald Eagle.

The eagle fights briefly to pull the carp from the water, then abandons the effort to instead release its talons from the fish to prevent its own demise by drowning.

Free of its hold on the carp, the eagle flaps its wings briskly to get up and out of the potentially deadly situation.

Both the eagle and, presumably, the carp survive the encounter.

The eagle finds a nearby tree limb where it takes a much-needed break from fishing.

Such an event may have been fatal to an inexperienced younger Bald Eagle. Though not an adult, this bird is no spring chicken. Its plumage, particularly its white head with a dark line through the eye (often called an “osprey face”), is indicative of a bird in its fourth year of life. By this time next year, it should be sporting a full set of adult feathers. In the meantime, it should stick to eating suckers; there are plenty of those to go around.

Common Carp are classified as a minnow (Cyprinidae). They are indigenous to Europe and are raised as a food fish in many parts of the world. In the Lower Susquehanna River Watershed, Common Carp are an introduced species with negative impacts on native fisheries. Those exceeding twenty inches in length begin to gain girth and weight, sometimes reaching thirty pounds at thirty inches.

SOURCES

Sedaghat, Safoura, Seyed Abbas Hoseini, Mohammad Larijani, and Khadijeh Shamekhi Ranjbar. 2013. “Age and Growth of Common Carp (Cyprinus carpio Linnaeus, 1758) in Southern Caspian Sea, Iran”. World Journal of Fish and Marine Sciences. 5:1. pp.71-73.

June 28, 2018July 1, 2024

Shocking Fish Photos!

There are two Conewago Creek systems in the Lower Susquehanna River Watershed. One drains the Gettysburg Basin west of the river, mostly in Adams and York Counties, then flows into the Susquehanna at the base of Conewago Falls. The other drains the Gettysburg Basin east of the river, flowing through Triassic redbeds of the Gettysburg Formation and York Haven Diabase before entering Conewago Falls near the south tip of Three Mile Island. Both Conewago Creeks flow through suburbia, farm, and forest. Both have their capacity to support aquatic life impaired and diminished by nutrient and sediment pollution.

This week, some of the many partners engaged in a long-term collaboration to restore the east shore’s Conewago Creek met to have a look at one of the prime indicators of overall stream habitat health—the fishes. Kristen Kyler of the Lower Susquehanna Initiative organized the effort. Portable backpack-mounted electrofishing units and nets were used by crews to capture, identify, and count the native and non-native fishes at sampling locations which have remained constant since prior to the numerous stream improvement projects which began more than ten years ago. Some of the present-day sample sites were first used following Hurricane Agnes in 1972 by Stambaugh and Denoncourt and pre-date any implementation of sediment and nutrient mitigation practices like cover crops, no-till farming, field terracing, stormwater control, nutrient management, wetland restoration, streambank fencing, renewed forested stream buffers, or modernized wastewater treatment plants. By comparing more recent surveys with this baseline data, it may be possible to discern trends in fish populations resulting not only from conservation practices, but from many other variables which may impact the Conewago Creek Warmwater Stream ecosystem in Dauphin, Lancaster, and Lebanon Counties.

So here they are. Enjoy these shocking fish photos.

Electrofishing on the Conewago Creek in Lebanon County, PA — Matt Kofroth, Watershed Specialist with the Lancaster County Conservation District, operates the electrofishing wand in Conewago Creek while his team members prepare to net and collect momentarily-stunned fish. Three other electrofishing units operated by staff from the Susquehanna River Basin Commission and aided by teams of netters were in action at other sample locations along the Conewago on this day.

Fishes of the Lower Susquehanna River Watershed: Common Carp — Really big fish, such as this Common Carp (Cyprinus carpio), were identified, counted, and immediately returned to the water downstream of the advancing electrofishing team.

Fishes of the Lower Susquehanna River Watershed: Swallowtail Shiner, Fallfish, Red-breast Sunfish, and suckers. — Other fish, such as the Swallowtail Shiner, Redbreast Sunfish (Lepomis auritus), Fallfish, and suckers seen here, were placed in a sorting tank.

Fishes of the Lower Susquehanna River Watershed: Fallfish — Fallfish (Semotilus corporalis) are very active and require plenty of dissolved oxygen in the water to survive. Fallfish, Rainbow Trout (Oncorhynchus mykiss), and Smallmouth Bass (Micropterus dolomieu) were quickly identified and removed from the sorting tank for release back into the stream. Other larger, but less active fish, including suckers, quickly followed.

Fishes of the Lower Susquehanna River Watershed: Fathead Minnow — Small fish like minnows were removed from the sorting tank for a closer look in a hand-held viewing tank. This Fathead Minnow (Pimephales promelas) was identified, added to the tally sheet, and released back into the Conewago. The Fathead Minnow is not native to the Susquehanna drainage. It is the minnow most frequently sold as bait by vendors.

Fishes of the Lower Susquehanna River Watershed: a breeding male Bluntnose Minnow — A breeding condition male Bluntnose Minnow (Pimephales notatus).

Fishes of the Lower Susquehanna River Watershed: Cutlips Minnow — The Cutlips Minnow (Exoglossum maxillingua) is a resident of clear rocky streams. Of the more than 30 species collected during the day, two native species which are classified as intolerant of persisting stream impairment were found: Cutlips Minnow and Swallowtail Shiner.

Fishes of the Lower Susquehanna River Watershed: Central Stoneroller — The Central Stoneroller (Campostoma anomalum) is a benthic feeder in creeks over gravel and sand.

Fishes of the Lower Susquehanna River Watershed: Eastern Blacknose Dace — The Eastern Blacknose Dace (Rhinichthys atratulus) is found in clear water over pebble and stone substrate.

Fishes of the Lower Susquehanna River Watershed: Longnose Dace — The Longnose Dace (Rhinichthys cataractae) is another species of pebbly rocky streams.

Fishes of the Lower Susquehanna River Watershed: juvenile Golden Shiner — A juvenile Golden Shiner (Notemigonus crysoleucas). Adults lack the side stripe and grow to the size of a sunfish.

Fishes of the Lower Susquehanna River Watershed: Swallowtail Shiner — A Swallowtail Shiner (Notropis procne) and a very young White Sucker (Catostomus commersonii) in the upper left of the tank.

Fishes of the Lower Susquehanna River Watershed: Spotfin Shiner — A probable Spotfin Shiner (Cyprinella spiloptera).

Fishes of the Lower Susquehanna River Watershed: Margined Madtom — The Margined Madtom (Noturus insignis) is a small native catfish of pebbly streams.

Fishes of the Lower Susquehanna River Watershed: Banded Killifish — The Banded Killifish (Fundulus diaphanus) is adept at feeding upon insects, including mosquitos.

Fishes of the Lower Susquehanna River Watershed: a juvenile Rock Bass — A young Rock Bass (Ambloplites rupestris). This species was introduced to the Susquehanna and its tributaries.

Fishes of the Lower Susquehanna River Watershed: Greenside Darter — The Greenside Darter (Etheostoma blennioides) is not native to the Susquehanna basin. The species colonized the Conewago Creek (east) from introduced local populations within the last five years.

Fishes of the Lower Susquehanna River Watershed: Tessellated Darter — The Tessellated Darter (Etheostoma olmstedi) is a native inhabitant of the Susquehanna and its tributaries.

Fishes of the Lower Susquehanna River Watershed: American Eel — The stars of the day were the American Eels (Anguilla rostrata).

American Eels serve as an intermediate host for the microscopic parasitic glochidia (larvae) of the Eastern Elliptio (Elliptio complanata), a declining native freshwater mussel of the Lower Susquehanna River Watershed. While feeding on their host (usually in its gills), the glochidia cause little injury and soon drop off to continue growth, often having assured distribution of their species by accepting the free ride. Freshwater mussels are filter feeders and improve water quality. They grow slowly and can live for decades.

SOURCES

Normandeau Associates, Inc. and Gomez and Sullivan. 2018. Muddy Run Pumped Storage Project Conowingo Eel Collection Facility FERC Project 2355. Prepared for Exelon.

Stambaugh, Jr., John W., and Robert P. Denoncourt. 1974. A Preliminary Report on the Conewago Creek Faunal Survey, Lancaster County, Pennsylvania. Proceedings of the Pennsylvania Academy of Sciences. 48: 55-60.