Grasshoppers are perhaps best known for the occasions throughout history when an enormous congregation of these insects—a “plague of locusts”—would assemble and rove a region to feed. These swarms, which sometimes covered tens of thousands of square miles or more, often decimated crops, darkened the sky, and, on occasion, resulted in catastrophic famine among human settlements in various parts of the world.
The largest “plague of locusts” in the United States occurred during the mid-1870s in the Great Plains. The Rocky Mountain Locust (Melanoplus spretus), a grasshopper of prairies in the American west, had a range that extended east into New England, possibly settling there on lands cleared for farming. Rocky Mountain Locusts, aside from their native habitat on grasslands, apparently thrived on fields planted with warm-season crops. Like most grasshoppers, they fed and developed most vigorously during periods of dry, hot weather. With plenty of vegetative matter to consume during periods of scorching temperatures, the stage was set for populations of these insects to explode in agricultural areas, then take wing in search of more forage. Plagues struck parts of northern New England as early as the mid-1700s and were numerous in various states in the Great Plains through the middle of the 1800s. The big ones hit between 1873 and 1877 when swarms numbering as many as trillions of grasshoppers did $200 million in crop damage and caused a famine so severe that many farmers abandoned the westward migration. To prevent recurrent outbreaks of locust plagues and famine, experts suggested planting more cool-season grains like winter wheat, a crop which could mature and be harvested before the grasshoppers had a chance to cause any significant damage. In the years that followed, and as prairies gave way to the expansive agricultural lands that presently cover most of the Rocky Mountain Locust’s former range, the grasshopper began to disappear. By the early years of the twentieth century, the species was extinct. No one was quite certain why, and the precise cause is still a topic of debate to this day. Conversion of nearly all of its native habitat to cropland and grazing acreage seems to be the most likely culprit.
The critically endangered Eskimo Curlew (Numenius borealis), a species not photographed since 1962 and not confirmed since 1963, fed on Rocky Mountain Locusts during its spring migration through the Great Plains. Excessive hunting and conversion of grasslands to agriculture are believed responsible for the bird’s demise. (United States Fish and Wildlife Service image by Christina Nelson)
In the Mid-Atlantic States, the mosaic of the landscape—farmland interspersed with a mix of forest and disturbed urban/suburban lots—prevents grasshoppers from reaching the densities from which swarms arise. In the years since the implementation of “Green Revolution” farming practices, numbers of grasshoppers in our region have declined. Systemic insecticides including neonicotinoids keep grasshoppers and other insects from munching on warm-season crops like corn and soybeans. And herbicides including 2,4-D (2,4-Dichlorophenoxyacetic acid) have, in effect, become the equivalent of insecticides, eliminating broadleaf food plants from the pasturelands and hayfields where grasshoppers once fed and reproduced in abundance. As a result, few of the approximately three dozen species of grasshoppers with ranges that include the Lower Susquehanna River Watershed are common here. Those that still thrive are largely adapted to roadsides, waste ground, and small clearings where native and some non-native plants make up their diet.
Here’s a look at four species of grasshoppers you’re likely to find in disturbed habitats throughout our region. Each remains common in relatively pesticide-free spaces with stands of dense grasses and broadleaf plants nearby.
CAROLINA GRASSHOPPER
Dissosteira carolina
The Carolina Grasshopper, also known as the Carolina Locust or Quaker, is one of the band-winged grasshoppers. It is commonly found along roadsides and on other bare ground near stands of tall grass and broadleaf plants.The Carolina Grasshopper is variable in color, ranging from very dark brown……to a rich tan or khaki shade. These earth-tone colors provide the insect with effective camouflage while spending time on the ground.The Carolina Grasshopper is most readily detected and identified when it flies. The colors of the wings resemble those of the Mourning Cloak butterfly.Carolina Grasshoppers are among the preferred victims of Great Black Wasps (Sphex pensylvanicus). A female wasp stings the grasshopper to paralyze it, then drags it away to one of numerous cells in an underground burrow where she lays an egg on it. The body of the disabled grasshopper then provides nourishment for the larval wasp.
DIFFERENTIAL GRASSHOPPER
Melanoplus differentialis
Differential Grasshopper nymph with small “fairy wings”.An adult Differential Grasshopper with fully developed wings.
TWO-STRIPED GRASSHOPPER
Melanoplus bivittatus
An early-stage Two-striped Grasshopper nymph.A Two-striped Grasshopper nymph in a later stage.An adult Two-striped Grasshopper.An adult Two-striped Grasshopper. Note the pale stripe originating at each eye and joining near the posterior end of the wings to form a V-shaped pattern.A female Two-striped Grasshopper.
RED-LEGGED GRASSHOPPER
Melanoplus femurrubrum
The Red-legged Grasshopper may currently be our most abundant and widespread species.A Red-legged Grasshopper hiding in dense urban vegetation.A male Red-legged Grasshopper.A female Red-legged Grasshopper.
Protein-rich grasshoppers are an important late-summer, early-fall food source for birds. The absence of these insects has forced many species of breeding birds to abandon farmland or, in some cases, disappear altogether.
During the mid-twentieth century, the Cattle Egret (Bubulcus ibis), a notoriously nomadic species, transited the Atlantic from Africa to colonize the Americas…and they did it without any direct assistance from humans. During the 1970s and early 1980s, a nesting population of Cattle Egrets on river islands adjacent to the Susquehanna’s Conejohela Flats off Washington Boro was the largest inland rookery in the northeastern United States. The Lancaster County Bird Club censused the birds each August and found peak numbers in 1981 (7,580). During their years of abundance, V-shaped flocks of Cattle Egrets from the rookery islands ventured into grazing lands throughout portions of Lancaster, York, Dauphin, and Lebanon Counties to hunt grasshoppers. These daily flights were a familiar summertime sight for nearly two decades. Then, in the early 1980s, reductions in pastureland acreage and plummeting grasshopper numbers quickly took their toll. By 1988, the rookery was abandoned. The Cattle Egrets had moved on. (Vintage 33 mm image)During the summer and early fall, juvenile and adult Ring-necked Pheasants feed heavily on grasshoppers. Earlier and more frequent mowing along with declining numbers of grasshoppers on farmlands due to an increase in pesticide use were factors contributing to the crash of the pheasant population in the early 1980s.To the delight of Wild Turkeys, each of the four species of grasshoppers shown above frequents clearings and roadsides adjacent to forest areas. While changes in grasshopper distribution have been detrimental to populations of birds like pheasants, they’ve created a feeding bonanza for turkeys.Wild Turkeys feeding on an abundance of grasshoppers along a forest road.An American Kestrel feeds on a grasshopper while ignoring the abundance of Spotted Lanternflies swarming the adjacent utility pole. In Susquehanna valley farmlands, grasshopper and kestrel numbers are down. Lanternflies, on the other hand, have got it made.Maintaining areas bordering roads, forests, wetlands, farmlands, and human development in a state of early succession can provide and ideal mix of mature grasses and broadleaf plants for grasshoppers, pollinators, birds, and other wildlife.
The calls of Fall Field Crickets, a series of chirps one might attempt to replicate by quickly strumming three to five teeth on a hair comb, are a familiar chorus during the late summer and autumn in the Lower Susquehanna River Watershed. The song’s speed changes with temperature, becoming noticeably slower as cold weather sets in. These crickets sing throughout the majority of the day and night, becoming quietest during the chilly hours of dawn. Adults hide in burrows and beneath logs, leaf litter, and other organic debris. Around human habitation, they seek shelter under hundreds of different objects, frequently finding their way into buildings where they do no real harm. Crickets are eaten by almost every predator that exceeds them in size and are especially important as a protein-rich food source for birds feeding their young and preparing for autumn migration. The Fall Field Cricket (Gryllus pennsylvanicus) is nearly identical in appearance to the less common Spring Field Cricket (Gryllus veletis), which calls during the spring and early summer in the Susquehanna valley. The primary difference between the two species? The former overwinter as eggs while the latter pass the colder months as nymphs.
The Gray Comma is a strikingly colorful late-summer butterfly of rich forests in the Lower Susquehanna River Watershed. It’s sure to catch your attention,……or is it?
Check out this and other local species by clicking the “Butterflies” tab at the top of this page.
During the past two weeks, over 150 American Flamingos (Phoenicopterus ruber) have been reported in the eastern half of the United States, well north of their usual range—northern South America; Cuba, the Bahamas, and other Caribbean Islands; and the Yucatan Peninsula of Mexico. They are also found on the Galapagos Islands. Prior to being extirpated by hunting during the early years of the twentieth century, American Flamingos nested in the Everglades of Florida as well.
The recent influx of these tropical vagrants into the eastern states is attributed to Hurricane Idalia, a storm that formed in the midst of flamingo territory during the final week of August. Idalia got its start over tropical waters off Yucatan before progressing north through the Gulf of Mexico off Cuba to make landfall in the “Big Bend” area of Florida.
Track of Hurricane Idalia. (NOAA/National Weather Service/National Hurricane Center image)
Birds of warm water seas regularly become entangled within tropical storms and hurricanes. In most cases, it’s oceanic seabirds including petrels, shearwaters, terns, and gulls that seek refuge from turbulent conditions by remaining inside the storm’s eye or within the margins of the rotating bands. The duration of the experience and the fitness of each individual bird may determine just how many survive such an ordeal. As a storm makes landfall, the survivors put down on inland bodies of water, often in unexpected places hundreds of miles from the nearest coastline. Some may linger in these far-flung areas to feed and gain strength before finding their way down rivers back to sea.
For birds like American Flamingos, which don’t spend a large portion of their lives on the wing over vast oceanic waters, becoming entangled in a tropical weather system can easily be a life-ending event. American Flamingos, despite their size and lanky appearance, are quite agile fliers. When necessary, some populations travel significant distances in search of new food sources. Some closely related migratory species of flamingos have been recorded at altitudes exceeding 10,000 feet. But becoming trapped in a hurricane for an extended period of time could easily erode the endurance of even the strongest of fliers. And once out over open water, there’s nowhere for a flamingo to come down for a rest.
It appears probable that, during the final week of August, hundreds of American Flamingos did find themselves drawn away from their familiar tidal flats and salty lagoons to be swept into the spinning clouds and gusty downpours of Hurricane Idalia as it churned off the Yucatan coast. Despite the dangers, more than 150 of them completed a journey within its clutches to cross the Gulf of Mexico and land in the Eastern United States.
Amy Davis in a report for the American Birding Association compiled a list of American Flamingo sightings from the days following the landfall of Hurricane Idalia. We’ve plotted those sightings on a satellite image of the storm.
Locations of American Flamingo sightings in the days following landfall of Hurricane Idalia, August 30 through September 7, 2023. Numbers of birds in the plotted flock are indicated. Those flocks marked with an asterisk contained a bird ringed with a leg band marking it as a member of the Ria Largartos population in Yucatan. Click the image to enlarge. (NOAA/GOES-16 base image of Hurricane Idalia landfall)
The majority of the flamingos that survived this harrowing trip found a place to put down along the coast. But as the reader will notice, a significant number of the sightings were outside the path of the storm in areas not reached by even the outermost bands of clouds. Flamingos in Alabama, Tennessee, Kentucky/Indiana, Ohio, and Pennsylvania were found at inland bodies of water, most on the days immediately following the storm’s landfall. The lone exception, the Pennsylvania birds, were discovered a week after landfall (but may have been there days longer). How and why did these birds find their way so far inland at places so distant from the storm…and home? And how did they get there so quickly? The counterclockwise rotation of the hurricane wouldn’t slingshot them in that direction, would it?
Let’s have a look at the surface map showing us the weather in the states outside the direct influence of Hurricane Idalia during the storm’s landfall on August 30, 2023…
Note the pair of cold fronts along the Appalachians by 7 A.M. E.S.T. (8 A.M. E.D.T.). In the hours following Idalia’s landfall, these fronts had begun slowly creeping through the areas where the inland flamingos were subsequently found. (NOAA/National Centers for Environmental Prediction-Weather Prediction Center image)
Now check out this satellite view of the storm during the hours following landfall…
The blue lines mark the approximate locations of the two cold fronts, the “spoilers” that helped steer Idalia out to sea. The pink arrows mark the “flamingo highway”, strong southwest to northeast winds blowing between the cold fronts along the inland edge of Idalia. (GOES-16 imagery from Cooperative Institute for Research in the Atmosphere at Colorado State and the National Oceanic and Atmospheric Administration)
Click the image above to play a GIF animation of GOES-16 satellite photos showing Idalia after landfall and the strong “flamingo highway” winds between the fronts. (It’s a big file, so give it a little time to load.) You’ll also note the ferocious lightning associated with both Huricane Idalia and Hurricane Franklin.
Did the inland flamingos catch a ride on these stiff “flamingo highway” winds? If not, they may have had a little help from a high pressure ridge that became established over the southern Appalachians as Idalia crept out to sea. The clockwise rotation of this ridge kept a similar flow over the “flamingo highway” for much of the four days that followed.
An American Flamingo circling to land in a farm pond near St. Thomas, Franklin County, Pennsylvania.Its a long way from Yucatan to central Pennsylvania. The survivors of the storm were largely at the mercy of the winds that carried them. The good news for this bird…not a single flight is feather missing.An American Flamingo at the foot of the Appalachians in Franklin County, Pennsylvania.An American Flamingo on a farm pond near St. Thomas, Franklin County, Pennsylvania. This bird is just one of at least 150 that arrived in the Eastern United States courtesy of Hurricane Idalia.In deep water, American Flamingos feed by shuffling their feet to stir up benthic invertebrates, plankton, algae, and plant matter. The flamingo that was accompanying this bird apparently shuffled its feet into a Snapping Turtle, which promptly latched onto its leg and injured tendons and other tissue. Following surgery earlier today, the bird remains in hospital.To retrieve their fare from the bottom, flamingos roll their bill upside down, then swish it back-and-forth.Finally, they raise their heads and upright their bills to expel water and debris while simultaneously filtering out the morsels of edibles for consumption.To conserve body heat while under the shade of clouds, this American Flamingo in Franklin County, Pennsylvania, would stop feeding,……then it would tuck its head between its wings. Flamingos will also retain heat by standing on one leg while raising the other into the insulating cover of their belly feathers.
For vagrant birds including those carried away to distant lands by hurricanes and other storms, the challenges of finding food and avoiding hazards including disease, unfamiliar predators, and extreme weather can be a matter of life and death. For American Flamingos now loafing and feeding in the Eastern United States outside Florida, the hardest part of their journey is yet to come. They need to find their way back south to suitable tropical habitat before cold weather sets in. The perils are many. Theirs is not an enviable predicament.
Sorry Pinky, but you can’t stay on the farm forever.
To pass the afternoon, we sat quietly along the edge of a pond created recently by North American Beavers (Castor canadensis). They first constructed their dam on this small stream about five years ago. Since then, a flourishing wetland has become established. Have a look.
Vegetation surrounding the inundated floodplain helps sequester nutrients and sediments to purify the water while also providing excellent wildlife habitat.The beaver lodge was built among shrubs growing in shallow water in the middle of the pond.Woolgrass (Scirpus cyperinus) is a bulrush that thrives as an emergent and as a terrestrial plant in moist soils bordering the pond.A male Common Whitetail dragonfly keeping watch over his territory.A Twelve-spotted Skimmer perched on Soft Rush.A Blue Dasher dragonfly seizing a Fall Field Cricket (Gryllus pennsylvanicus).A Spicebush Swallowtail visiting a Cardinal Flower.A Green Heron looking for small fish, crayfish, frogs, and tadpoles.The Green Heron stalking potential prey.A Wood Duck feeding on the tiny floating plant known as Lesser Duckweed (Lemna minor).A Least Sandpiper poking at small invertebrates along the muddy edge of the beaver pond.A Solitary Sandpiper.A Solitary Sandpiper testing the waters for proper feeding depth.A Pectoral Sandpiper searches for its next morsel of sustenance.The Sora (Porzana carolina) is a seldom seen rail of marshlands including those created by North American Beavers. Common Cattails, sedges, and rushes provide these chicken-shaped wetland birds with nesting and loafing cover.
Isn’t that amazing? North American Beavers build and maintain what human engineers struggle to master—dams and ponds that reduce pollution, allow fish passage, and support self-sustaining ecosystems. Want to clean up the streams and floodplains of your local watershed? Let the beavers do the job!
At first glance, this insect might be mistaken for a Brown Marmorated Stink Bug. It is, however, a native species, a Magnolia Leaf-footed Bug (Leptoglossus fulvicornis). Because it feeds almost exclusively on trees in the genus Magnolia, and because the three members of the genus native to the lower Susquehanna valley, Umbrella Magnolia (M. tripetala), Cucumber Tree (M. acuminata), and Sweetbay (M. virginiana), each have a very limited distribution here, this insect’s presence in the watershed is largely dependent on widespread introductions of ornamental magnolias and native transplants including Southern Magnolia (Magnolia grandiflora). Adult Magnolia Leaf-footed Bugs overwinter in leaf litter beneath the host trees. In spring, females lay their eggs in rows along the midrib of the leaves where they are often victims of parasitoid wasps including Gyron pennsylvanicum. The surviving nymphs and the adults feed on the tree’s leaves and fruits, causing little harm to its overall health.
The Broad-necked Root Borer (Prionus laticollis) is one of the Lower Susquehanna River Watershed’s largest insects. Its larvae feed on woody roots for more than two years before pupating and thereafter emerging as adult beetles. The distribution of root borers is not totally random. They are often attracted to distressed trees and shrubs. This particular specimen was found along a forest road where herbicide application, soil dehydration, and a recent wildfire has weakened and killed numerous oaks, poplars, and maples. As they feed, root borer larvae will help begin the process of reducing the remains of these trees into soil nutrients that will benefit regeneration of forest plants at this site.
“Waves” of warblers and other Neotropical songbirds continue to roll along the ridgetops of southern Pennsylvania. The majority of these migrants are headed to wintering habitat in the tropics after departing breeding grounds in the forests of southern Canada. At Second Mountain Hawk Watch, today’s early morning flight kicked off at sunrise, then slowed considerably by 8:30 A.M. E.D.T. Once again, in excess of 400 warblers were found moving through the trees and working their way southwest along the spine of the ridge. Each of the 12 species seen yesterday were observed today as well. In addition, there was a Northern Parula and a Canada Warbler. Today’s flight was dominated by Bay-breasted, Blackburnian, Black-throated Green, and Tennessee Warblers.
A Blackburnian Warbler at sunrise on Second Mountain.A hungry Blackburnian Warbler feeding on insects.Black-throated Green Warblers were a plentiful species among both yesterday’s and today’s waves of Neotropical migrants.A juvenile Black-throated Green Warbler.One of the scores of Tennessee Warblers seen on Second Mountain early this morning.Cape May Warblers were still common today, but not moving through in the numbers seen yesterday.A male Black-throated Blue Warbler.Compared to yesterday’s flight, lesser numbers of Magnolia Warblers were seen today.An adult male Wilson’s Warbler was a good find among the hundreds of birds swarming the ridgetop.This Nashville Warbler spent much of the day in the tangles of Mile-a-minute Weed surrounding the lookout.
Other interesting Neotropical migrants joined the “waves” of warblers…
Red-eyed Vireo numbers were higher than yesterday.This Warbling Vireo was found peering from the cover of the shady forest.A minimum of six Rose-breasted Grosbeaks were identified including the juvenile male seen here in first-fall plumage. Other good sightings were Scarlet Tanagers, an adult male Baltimore Oriole, and a dozen or more Ruby-throated Hummingbirds.Three Least Flycatchers were heard calling and seen chasing one another through a stand of dead timber on the south slope below the lookout.After the warbler flight settled, the task of counting migrating raptors commenced. Five Broad-winged Hawks including this one were tallied as they glided away to the southwest for a winter vacation in the tropics of Central and South America.
During the recent couple of mornings, a tide of Neotropical migrants has been rolling along the crests of the Appalachian ridges and Piedmont highlands of southern Pennsylvania. In the first hours of daylight, “waves” of warblers, vireos, flycatchers, tanagers, and other birds are being observed flitting among the sun-drenched foliage as they feed in trees along the edges of ridgetop clearings. Big fallouts have been reported along Kittattiny Ridge/Blue Mountain at Hawk Mountain Sanctuary and at Waggoner’s Gap Hawk Watch. Birds are also being seen in the Furnace Hills of the Piedmont.
Here are some of the 300 to 400 warblers (a very conservative estimate) seen in a “wave” found working its way southwest through the forest clearing at the Second Mountain Hawk Watch in Lebanon County this morning. The feeding frenzy endured for two hours between 7 and 9 A.M. E.D.T.
Tennessee Warblers (Leiothlypis peregrina) were very common among the migrants seen this morning on Second Mountain.An adult male Tennessee Warbler.A Nashville Warbler.A Chestnut-sided Warbler.Cape May Warblers were another common species during the morning flight.Magnolia Warblers were frequently observed as well.A Black-and-white Warbler exhibiting its nuthatch-like feeding behavior.A Blackburnian Warbler.Black-throated Green Warblers were numerous.A Bay-breasted Warbler.This was the only Wilson’s Warbler discerned among the hundreds of warblers seen in a “wave” on Second Mountain this morning.
Not photographed but observed in the mix of species were several Black-throated Blue Warblers and American Redstarts.
In addition to the warblers, other Neotropical migrants were on the move including two Common Nighthawks, a Broad-winged Hawk, a Least Flycatcher (Empidonax minimus), and…
At least half a dozen Scarlet Tanagers were in the treetops.And no less than 23 Ruby-throated Hummingbirds were counted cruising southbound past the lookout at Second Mountain Hawk Watch today.
Then, there was a taste of things to come…
One of 3 Red-breasted Nuthatches filling the air on the mountaintop with calls reminiscent of a toy tin horn. Will this summer’s forest fires in Canada prompt a significant invasion of this and other birds including winter finches in coming months? Time will tell.
Seeing a “wave” flight is a matter of being in the right place at the right time. Visiting known locations for observing warbler fallouts such as hawk watches, ridgetop clearings, and peninsular shorelines can improve your chances of witnessing one of these memorable spectacles by overcoming the first variable. To overcome the second, be sure to visit early and often. See you on the lookout!
Ruby-throated Hummingbirds will be migrating south through the Lower Susquehanna River Watershed during the next few weeks.While on their journey, they refuel by frequently visiting nectar-producing native wildflowers as well as garden specimens like this Mexican Cigar.Hummingbirds supplement their diet by visiting feeders filled with a mixture of sugar and water. If you’re feeding hummingbirds, or thinking about feeding hummingbirds, be sure to review the helpful tips contained in our post from August 5, 2022, “Two Feeders Are Better Than One”. Their health and your peace of mind may depend on it.
As we enter September, autumn bird migration is well underway. Neotropical species including warblers, vireos, flycatchers, and nighthawks are already headed south. Meanwhile, the raptor migration is ramping up and hawk watch sites throughout the Mid-Atlantic States are now staffed and counting birds. In addition to the expected migrants, there have already been sightings of some unusual post-breeding wanderers. Yesterday, a Wood Stork (Mycteria americana) was seen passing Hawk Mountain Sanctuary and a Swallow-tailed Kite (Elanoides forficatus) that spent much of August in Juniata County was seen from Waggoner’s Gap Hawk Watch while it was hunting in a Perry County field six miles to the north of the lookout! Both of these rarities are vagrants from down Florida way.
A Peregrine Falcon speeds past the lookout at Second Mountain Hawk Watch yesterday.Later this month, numbers of high-flying Broad-winged Hawks moving past counting stations will reach their peak. Most sites will experience one or more days with hundreds or perhaps thousands of these Neotropical migrants streaming by.This Black-and-white Warbler was found among a “wave” of migrating songbirds moving through some ridgetop trees.A juvenile Cape May Warbler peers from the cover of an Eastern Hemlock.
To plan a visit to a hawk watch near you, click on the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab at the top of this page to find a list and brief description of suggested sites throughout the Mid-Atlantic region. “Hawkwatcher’s Helper” also includes an extensive photo guide for identifying the raptors you’re likely to see.
And to identify those confusing fall warblers and other migrants, click the “Birds” tab at the top of this page and check out the photo guide contained therein. It includes nearly all of the species you’re likely to see in the lower Susquehanna valley.
At Lake Redman just to the south of York, Pennsylvania, a draw down to provide drinking water to the city while maintenance is being performed on the dam at neighboring Lake Williams, York’s primary water source, has fortuitously coincided with autumn shorebird migration. Here’s a sample of the numerous sandpipers and plovers seen today on the mudflats that have been exposed at the southeast end of the lake…
One of a hundred or more Least Sandpipers seen on mudflats at Lake Redman today.A Semipalmated Plover and a Least SandpiperPectoral Sandpipers.A Pectoral Sandpiper and two Least Sandpipers.A Semipalmated Sandpiper.A Stilt Sandpiper feeding.Stilt Sandpiper consuming an edible.Stilt Sandpiper at rest.A Solitary SandpiperA Lesser Yellowlegs.A Greater Yellowlegs.Stirring up the shorebird crowd every now and then were several Ospreys, but all would soon be back to the business of feeding in the mud.An Osprey hovers above shallow water near the mudflats as it searches for fish.
Not photographed but present at Lake Redman were at least two additional species of shorebirds, Killdeer and Spotted Sandpiper—bringing the day’s tally to ten. Not bad for an inland location! It’s clearly evident that these waders overfly the lower Susquehanna valley in great numbers during migration and are in urgent need of undisturbed habitat for making stopovers to feed and rest so that they might improve their chances of surviving the long journey ahead of them. Mud is indeed a much needed refuge.
It may be one of the most treasured plants among native landscape gardeners. The Cardinal Flower (Lobelia cardinalis) blooms in August each year with a startling blaze of red color that, believe it or not, will sometimes be overlooked in the wild.
Cardinal Flower is most often found in wet soil along forested bodies of water. The blooms of this shade-loving species may go unnoticed until rays of sunshine penetrate the canopy to strike their brilliant red petals.
The Cardinal Flower grows in wetlands as well as in a variety of moist soils along streams, rivers, lakes, and ponds. Shady locations with short periods of bright sun each day seem to be favored for an abundance of color.
Cardinal Flower in bloom in a riparian forest along the Susquehanna. To its right is its close relative, Great Lobelia, a plant sometimes known as Great Blue Lobelia or Blue Cardinal Flower.Cardinal Flower in a wet bottomland woods.The Cardinal Flower can find favorable growing conditions along stream, river, and lake shores. Even though they are perennial plants, their presence along such waters often seems temporary. Changing conditions cause them to suddenly disappear from known locations, then sometimes reappear at the same place or elsewhere nearby. Some of this phenomenon may be due to the fact that stressed plants can fail to bloom, so they easily escape notice. When producing flowers during favorable years, the plants seem to mysteriously return.Cardinal Flowers along a wave-swept shoreline light up the greenery of erosion-controlling riparian vegetation with glowing red color.
The Cardinal Flower can be an ideal plant for attracting hummingbirds, bees, butterflies, and other late-summer pollinators. It grows well in damp ground, especially in rain gardens and along the edges streams, garden ponds, and stormwater retention pools. If you’re looking to add Cardinal Flower to your landscape, you need first to…
REMEMBER the CARDINAL RULE…
Cardinal Flower plants are available at many nurseries that carry native species of garden and/or pond plants. Numerous online suppliers offer seed for growing your own Cardinal Flowers. Some sell potted plants as well. A new option is to grow Cardinal Flowers from tissue cultures. Tissue-cultured plants are raised in laboratory media, so the pitfalls of disease and hitchhikers like invasive insects and snails are eliminated. These plants are available through the aquarium trade from most chain pet stores. Though meant to be planted as submerged aquatics in fish tank substrate, we’ve reared the tissue-cultured stock indoors as emergent plants in sandy soil and shallow water through the winter and early spring. When it warms up, we transplant them into the edges of the outdoor ponds to naturalize. As a habit, we always grow some Cardinal Flower plants in the fish tanks to take up the nitrates in the water and to provide a continuous supply of cuttings for starting more emergent stock for outdoor use.
A tissue-cultured Cardinal Flower rooted in sandy substrate and being grown as a submerged aquatic plant in a fish tank. Cuttings from this plant will be used to grow emergent specimens in shallow water for transplanting outdoors around the garden pond.A Cardinal Flower grown from an aquarium store tissue culture blooms in the pond at susquehannawildlife.net headquarters.Grown as an emergent, Cardinal Flower may bloom very late in the season. This tissue-cultured specimen in the headquarters pond was photographed in early November, 2022.
Looking for a native wildflower that’s tall, showy, and a great choice for attracting wildlife, especially butterflies and bees? Then check out Wild Senna (Senna hebecarpa).
Wild Senna currently blooming in a roadside wildflower garden on Pennsylvania State Game Lands.
Wild Senna, also known as American Senna, is a host plant for the larvae of Cloudless Sulphur and Sleepy Orange (Eurema nicippe) butterflies. It thrives in almost any moist, well-drained soil in habitats including open woodlands, forest edges, meadows, and gardens like yours. Its height at flowering ranges from three to six feet. If you prefer, this perennial wildflower can even be cultivated as a shrub-like form. It is easily grown from seed, which is available from Ernst Conservation Seeds of Meadville, Pennsylvania, as well as numerous other vendors. And don’t forget to give Wild Senna’s two close relatives, Partridge Pea and Maryland Senna, a try as well. They attract the same species of butterflies and are just as easy to grow. You’ll like ’em.
Cloudless Sulphur butterflies from populations in the south colonize the Lower Susquehanna River Watershed each summer in varying numbers.Close-up view of a Wild Senna flower cluster. Typical of members of the pea family, the seeds of all native Senna species develop within pods after blooming and are sought after by wildfowl (Galliformes), particularly Northern Bobwhite.Partridge Pea with flowers and seed pods in the susquehannawildlife.net garden. Smaller than the other Senna species, Partridge Pea reaches a height of just two to three feet.
Your best bet for finding migrating shorebirds in the lower Susquehanna region is certainly a visit to a sandbar or mudflat in the river. The Conejohela Flats off Washington Boro just south of Columbia is a renowned location. Some man-made lakes including the one at Middle Creek Wildlife Management Area are purposely drawn down during the weeks of fall migration to provide exposed mud and silt for feeding and resting sandpipers and plovers. But with the Susquehanna running high due to recent rains and the cost of fuel trending high as well, maybe you want to stay closer to home to do your observing.
Fortunately for us, migratory shorebirds will drop in on almost any biologically active pool of shallow water and mud that they happen to find. This includes flooded portions of fields, construction sites, and especially stormwater retention basins. We stopped by a new basin just west of Hershey, Pennsylvania, and found more than two dozen shorebirds feeding and loafing there. We took each of these photographs from the sidewalk paralleling the south shore of the pool, thus never flushing or disturbing a single bird.
Designed to prevent stream flooding and pollution, this recently installed stormwater retention basin along US 322 west of Hershey, Pennsylvania, has already attracted a variety of migrating plovers and sandpipers.Killdeer stick close to exposed mud as they feed.Two of more than a dozen Least Sandpipers found busily feeding in the inch-deep water.A Lesser Yellowlegs searching for small invertebrates.Two Lesser Yellowlegs work out a disagreement.Male Twelve-spotted Skimmers patrol the airspace above a pair of Least Sandpipers. Dragonflies and other aquatic insects are quick to colonize the waters held in well-engineered retention basins. Proper construction and establishment of a functioning food chain/web in these man-made wetlands prevents them from becoming merely temporary cesspools for breeding mosquitos.
So don’t just drive by those big puddles, stop and have a look. You never know what you might find.
A Semipalmated Sandpiper (middle right) joins a flock of Least Sandpipers.Pectoral Sandpipers (two birds in the center) are regular fall migrants on the Susquehanna at this time of year. They are most frequently seen on gravel and sand bars adjacent to the river’s grassy islands, but unusually high water for this time of year prevents them from using this favored habitat. As a result, you might be lucky enough to discover Pectoral Sandpipers on almost any mudflat in the area.Two Pectoral Sandpipers and five smaller, but otherwise very similar, Least Sandpipers.A Killdeer (right), a Semipalmated Plover (upper right), and Least and Pectoral Sandpipers (left).
An ever-vigilant male Blue Dasher (Pachydiplax longipennis) perches on a small twig overlooking the vegetated shallows along the shoreline of Memorial Lake in Lebanon County, Pennsylvania.
We’ve got the summertime blues for you, right here at susquehannawildlife.net…
In warm-season grass meadows, Big Bluestem is now in flower. This and other species of native prairie grasses provide excellent habitat for birds, mammals, and insects including butterflies. To survive drought and fire, their roots run much deeper than cool season grasses, creeping down four to six feet or more. This adaptation allowed warm season grasses to recover from heavy grazing by large Pre-Anthropocene mammals. Today, it makes them ideal plants for soil stabilization.A male Indigo Bunting has already found ripe seeds among the heads of flowering Big Bluestem.Look closely and you’ll see our Indigo Bunting is beginning a pre-migration molt out of its bright-blue breeding (alternate) plumage and into a gray-brown winter (basic) plumage. The berries of the American Pokeweed upon which it is perched will soon ripen into a dark blue, almost black, color. Though toxic to humans, these fruits find favor with many species of birds and mammals.Another great wildlife food is Silky Dogwood (Cornus amomum), a deciduous shrub that sports blue-colored berries in summer and showy, bright-red twigs in winter. It grows well in wet ground along streams and ponds, as well as in rain gardens.A Great Blue Heron searches the shallows for small fish. This species is also a good mouser, at times seen hunting in grassy meadows. Right now is prime time to see it and a variety of other herons and egrets throughout the Lower Susquehanna River Watershed.
…so don’t let the summertime blues get you down. Grab a pair of binoculars and/or a camera and go for a stroll!
For this White-tailed Deity and her fawns, a dry shoreline provides an opportunity to access the moist, tender greens of emergent plants on a hot summer afternoon.
The Eastern Scissor Grinder (Neotibicen winnemanna), a species of “Annual Cicada” or “Dog-day Cicada”, dries its wings after emerging from its exoskeleton (exuvia). After two to five years of subterranean life as a nymph, this adult will soon be ready to fly into the treetops in search of a mate. There is a sense of urgency. Summer’s end will bring its life to an end as well.
If you’re feeling the need to see summertime butterflies and their numbers just don’t seem to be what they used to be in your garden, then plan an afternoon visit to the Boyd Big Tree Preserve along Fishing Creek Valley Road (PA 443) just east of U.S. 22/322 and the Susquehanna River north of Harrisburg. The Pennsylvania Department of Conservation and Natural Resources manages the park’s 1,025 acres mostly as forested land with more than ten miles of trails. While located predominately on the north slope of Blue Mountain, a portion of the preserve straddles the crest of the ridge to include the upper reaches of the southern exposure.
A grove of American Chestnuts (Castanea dentata) planted at Boyd Big Tree Preserve is part of a propagation program working to restore blight-resistant trees to Pennsylvania and other areas of their former range which included the Appalachians and the upper Ohio River watershed.
Fortunately, one need not take a strenuous hike up Blue Mountain to observe butterflies. Open space along the park’s quarter-mile-long entrance road is maintained as a rolling meadow of wildflowers and cool-season grasses that provide nectar for adult butterflies and host plants for their larvae.
A view looking north at the butterfly meadow and entrance road at Boyd Big Tree Preserve Conservation Area. Second Mountain is in the background.Mowed paths follow the entrance road and a portion of the perimeter of the meadow allowing visitors a chance to wander among the waist-high growth to see butterflies, birds, and blooming plants at close range without trampling the vegetation or risking exposure to ticks.A Silver-spotted Skipper feeding on nectar from the flowers of Indian Hemp (Apocynum cannabinum). Like the milkweeds, Indian Hemp is a member of the dogbane family (Apocynaceae).An Eastern Tiger Swallowtail feeding on Common Milkweed.A Great Spangled Fritillary (Speyeria cybele) on Common Milkweed.A Black Swallowtail (Papilio polyxenes) feeding on Common Milkweed nectar.A Pipevine Swallowtail (Battus philenor) on Common Milkweed.Another Pipevine Swallowtail on Common Milkweed. Note the hook-shaped row of red-orange spots on the underside of the hindwing.A Pipevine Swallowtail visiting the brilliant blooms of Butterfly Weed, a favorite of a wide variety of pollinators.A Black Swallowtail with damaged wings alights atop a Butterfly Weed flower cluster. Note the pair of parallel rows of red-orange spot on the underside of the hindwing.A Monarch feeding on nectar from the flowers of Butterfly Weed.A mating pair of Eastern Tailed Blues on a Timothy (Phleum pratense) spike.A male Great Spangled Fritillary (right) pursuing a female.Butterflies aren’t the only colorful insects patrolling the meadows at Boyd Big Tree Preserve. Dragonflies including Common Green Darners are busily pursuing prey, particularly small flying insects like mosquitos, gnats, and flies.Dragonflies themselves can become prey and are much sought after by Broad-winged Hawks. This very vocal juvenile gave us several good looks as it ventured from the forest into the skies above the upper meadow during midday. It wasn’t yet a good enough flier to snag a dragonfly, but it will have plenty of opportunities for practice during its upcoming fall migration which, for these Neotropical raptors, will get underway later this month.
Do yourself a favor and take a trip to the Boyd Big Tree Preserve Conservation Area. Who knows? It might actually inspire you to convert that lawn or other mowed space into much-needed butterfly/pollinator habitat.
While you’re out, you can identify your sightings using our photographic guide—Butterflies of the Lower Susquehanna River Watershed—by clicking the “Butterflies” tab at the top this page. And while you’re at it, you can brush up on your hawk identification skills ahead of the upcoming migration by clicking the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab. Therein you’ll find a listing and descriptions of hawk watch locations in and around the lower Susquehanna region. Plan to visit one or more this autumn!
Shorebird identification can be notoriously difficult, but it’s a skill best learned with practice. Now is a good time to become familiar with the two most common species of plovers found in the lower Susquehanna valley. Numbers of both are increasing on local mudflats as waves of southbound migrants begin to arrive. The Killdeer in the lower left is a familiar nesting species of barren fields, stone parking lots, and gravel roofs. Its numbers swell during spring and fall migration, with some lingering into winter until either snow cover or a hard freeze prompts them to finally depart for milder climes. The Semipalmated Plovers in the upper right are strictly a migratory species in our area, nesting in northern Canada and Alaska and wintering along the American coastlines from Oregon and Delmarva south to Chile and Argentina. You’ll notice that the Semipalmated Plover has just a single neck ring while the Killdeer has two. That’s how you can tell them apart in a jiffy. It’s that easy.
Those mid-summer post-breeding wanders continue to delight birders throughout the Mid-Atlantic States. One colorful denizen of ponds and wetlands that has yet to put in an appearance in the Lower Susquehanna River Watershed this year is the Black-bellied Whistling Duck. You might remember this species from earlier posts describing the fortieth anniversary of your editor’s journey to the Lower Rio Grande Valley of Texas. Like many other birds, the Black-bellied Whistling Duck has been extending its range north from Texas, Florida, and other states along the Gulf Coastal Plain. Populations of these waterfowl are chiefly resident birds with some short-distance movement to find suitable habitat for feeding and nesting. They are not usually migratory, so summertime wandering may be the mechanism for their discovery of new habitats advantageous for nesting in areas north of their current home.
Presently, at least two dozen Black-bellied Whistling Ducks are being seen regularly at a stormwater retention pond in a housing subdivision along Amalfi Drive west of Smyrna, Delaware. This small population of avian tourists has spent at least two summers in the area. Just yesterday, Black-bellied Whistling Ducks were seen and photographed about ten miles to the east at Bombay Hook National Wildlife Refuge. Nine were counted there while 27 were being watched simultaneously at the Amalfi site. Earlier this week, a single Black-bellied Whistling Duck visited the John Heinz National Wildlife Refuge in Philadelphia, indicating that the influx of these vagrants has transited the entire Delmarva Peninsula and entered Pennsylvania. So while you’re out watching for those first southbound migrants of the year, be on the lookout for wayward wanderers too—wanderers like Black-bellied Whistling Ducks!
Black-bellied Whistling Ducks in a stormwater retention pond west of Smyrna, Delaware.Summertime exploration of new areas outside their resident turf may enable Black-bellied Whistling Ducks to find favorable habitats for extending their breeding range north.Black-bellied Whistling Ducks favor vegetated ponds, pools, and wetlands for feeding and nesting.Did you remember to go to the post office and buy a Federal Duck Stamp? Your purchase helps provide habitat for Black-bellied Whistling Ducks and so many other magnificent birds. And don’t forget, it’s your ticket for admission to our National Wildlife Refuges for an entire year!
At just after 8:30 this evening, an Antares rocket carrying a Cygnus supply capsule launched from the NASA flight facility on Delmarva Peninsula at Wallops Island, Virginia. We took a walk on the Veterans Memorial Bridge over the Susquehanna at Columbia-Wrightsville to have a look at the spacecraft as it powered its way into orbit for a rendezvous with the International Space Station.
Unfortunately, a smoky haze from wildfires still burning in Canada obscured our view of the ascending rocket as it cleared our horizon and made its way downrange across the Atlantic.
This faint glow from the Antares rocket was visible through thick haze in the southeast sky for less than ten seconds.
But on the brighter side, we were spared any significant disappointment. It just so happened that the Antares/Cygnus rocket wasn’t the only launch visible from the Veteran’s Memorial Bridge this evening…
A rocket climbing skyward above Columbia, Pennsylvania.A National Night Out fireworks display in Columbia preceded the Antares rocket launch.This fireworks grand finale on the Susquehanna ended just minutes prior to the liftoff in Virginia.Smoke and rockets aside, it was an otherwise peaceful and picturesque evening to be on the river.
Here in a series of photographs are just a handful of the reasons why the land stewards at Middle Creek Wildlife Management Area and other properties where conservation and propagation practices are employed delay the mowing of fields composed of cool-season grasses until after August 15 each year.
Eastern Meadowlarks, birds of large pastures, hay lots and other meadows of cool-season grasses, build their nests and raise their young on the ground. In the years since the early twentieth century, loss in the volume of acreage maintained in the lower Susquehanna Valley as grassland habitat types has dramatically reduced the prevalence and abundance of this and other birds with similar nesting requirements. During the most recent fifty years, early and frequent mowing and other practices introduced as part of agriculture’s Green Revolution have all but eliminated ground-nesting grassland species from the region.Like the meadowlarks, Grasshopper Sparrows (Ammodramus savannarum) nest on the ground in fields of cool-season grasses. Mowing prior to the time the young leave the nest and are able to fly away can obliterate a generation of grassland birds. Because their life span is short, widespread loss of an entire year of reproduction can quickly impact overall populations of native sparrows and other small birds. Delayed mowing can improve numbers of Grasshopper Sparrows as well as Savannah Sparrows, Vesper Sparrows (Pooecetes gramineus), and the very rare Henslow’s Sparrow (Centronyx henslowii).The Bobolink, like the meadowlark, is a member of the blackbird family (Icteridae). It too requires grasslands free of disturbances like mowing for the duration of the nesting season which, for this particular bird, lasts until mid-August in the lower Susquehanna region. In places lacking their specific habitat requirements, Bobolinks will seldom be detected except as flyovers during migration.Ring-necked Pheasants were introduced to the lower Susquehanna basin, and their populations were maintained thereafter, by stocking for the purpose of hunting. But throughout the middle twentieth century, there was a substantial population of ring-necks breeding in fields of cool-season grasses in farmlands throughout the region. High-intensity agriculture with frequent mowing eliminated not only nesting habitat in grasslands, but winter cover in areas of early successional growth. Populations of Ring-necked Pheasants, as well as native Northern Bobwhite, crumbled during the late 1970s and early 1980s due to these changes. For these resident birds that don’t migrate or routinely travel great distances to find new places to live and breed, widespread habitat loss can be particularly catastrophic. Not surprisingly, the Northern Bobwhite is no longer found in the Lower Susquehanna River Watershed and has been extirpated from all of Pennsylvania.At places like Middle Creek Wildlife Management Area where a mix of grasslands, early successional growth, and even some cropland are maintained, the Blue Grosbeak has extended its range well north of the Mason-Dixon and has become a regular nesting species during recent decades. Good habitat management does pay dividends.
Right now is a good time to visit Middle Creek Wildlife Management Area to see the effectiveness a delayed mowing schedule can have when applied to fields of cool-season grasses. If you slowly drive, walk, or bicycle the auto tour route on the north side of the lake, you’ll pass through vast areas maintained as cool-season and warm-season grasses and early successional growth—and you’ll have a chance to see these and other grassland birds raising their young. It’s like a trip back in time to see farmlands they way they were during the middle years of the twentieth century.
Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge. At Bombay Hook, they don’t waste your money mowing grass. Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools. Birds there can often be observed at close range. The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools. Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.Two more Short-billed Dowitchers on the way in.Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed. Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge. Mallards, American Avocets, and egrets are gathered on the shore.More Short-billed Dowitchers arriving to feed in Raymond Pool.Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool. A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.American Avocets probe the muddy bottom of Raymond Pool.Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early. Many migrating Dunlin linger at Bombay Hook into October and even November.This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp PoolA Greater Yellowlegs at Bear Swamp Pool.A Caspian Tern patrolling Raymond Pool.The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass. High tide licks at the roots of the cordgrass supporting the temporary domicile.By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice). It is our only dragonfly able to breed in saltwater. Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.Look up! A migrating Bobolink passes over the dike at Shearness Pool.Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool. Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river. The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century. An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers. Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.A Great Egret prowling Shearness Pool.A Snowy Egret in Bear Swamp Pool.Wood Ducks in Bear Swamp Pool.A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.
As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.
Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay. We found just this one during our visit to Bombay Hook. Man has certainly decimated populations of this ancient crustacean during recent decades.As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.A male Atlantic Marsh Fiddler Crab peers from its den.Herons and egrets including this Great Egret are quite fond of fiddler crabs. As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.An Osprey patrols the vast tidal areas opposite Shearness Pool.
No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook. This one was in shrubby growth along the dike at the north end of Shearness Pool.These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries. The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware. The seed balls are a favorite winter food of goldfinches and siskins.Finis Pool has no frontage on the tidal marsh but is still worth a visit. It lies along a spur road on the tour route and is located within a deciduous coastal plain forest. Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.The White-tailed Deity is common along the road to Finis Pool.Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well. This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.
We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.
Visitors stopping by Middle Creek Wildlife Management Area this week found yet another post-breeding wanderer feeding in the shallows of the main lake and adjacent pond along Hopeland Road—a juvenile Little Blue Heron.
The juvenile Little Blue Heron is a white bird resembling an egret during its first year. At about one year of age, it begins molt into a deep blue adult plumage. Young birds are notorious for roaming inland and north from breeding areas along the Atlantic coast and throughout the south. They are a post-breeding wanderer nowhere near as rare as the Limpkin seen at Middle Creek a week ago; a few are found each summer in the Lower Susquehanna River Watershed.
A juvenile Little Blue Heron currently at Middle Creek Wildlife Management Area. Note the yellow legs and pale beak, field marks that help separate this species from the Great Egret and Snowy Egret.
As oft times happens, birders attracted to see one unusual bird find another in the vicinity. So with fall shorebird migration ramping up, the discovery of something out of the ordinary isn’t a total surprise, particularly where habitat is good and people are watching.
A Hudsonian Godwit (Limosa haemastica) has arrived on mudflats at Middle Creek Wildlife Management Area.
The arrival of a Hudsonian Godwit is not an unheard of occurrence in the lower Susquehanna region, but locating one that sticks around and provides abundant viewing opportunities is a rarity. This adult presumably left the species’ breeding areas in Alaska or central/western Canada in recent weeks to begin its southbound movement. Hudsonian Godwits pass through the eastern United States only during the autumn migration, and the majority fly by without being noticed along a route that mostly takes them offshore of the Mid-Atlantic States.
The Hudsonian Godwit (right) at Middle Creek W.M.A. feeding with other migrating shorebirds…a Lesser Yellowlegs (top) and a Least Sandpiper (left).To probe the muddy bottom for invertebrates, Hudsonian Godwits will often wade in deeper water than accompanying species.This godwit seemed to be capturing small snails, presumably the young of the thousands of adult Acute Bladder Snails (Physellla acuta) seen here covering the surface of the mud. This species of air-breathing freshwater snail is tolerant of low levels of dissolved oxygen and is frequently the only mollusk found in polluted waters of the lower Susquehanna valley.For feeding shorebirds, the young snails may be more edible than the adults due to their fragile shells. Small birds like the Least Sandpiper may also be consuming the gelatinous egg masses. In North America, the Acute Bladder Snail was believed to be an introduced species from Europe. DNA testing has now determined that it is actually a native species that was instead transported into Europe from North America early in the nineteenth century or before. Locally, the snails were known as Physella heterostropha and were thought to be native. However, the recent genetic tests have shown Physella heterostropha and Physella integra, a snail first described by Lancaster County naturalist Samuel S. Haldeman, to be synonyms of Physella acuta. Click the “Freshwater Snails” tab at the top of this page to learn more about these mollusks.The Hudsonian Godwit at Middle Creek W.M.A., you may never get a better look!
Mid-summer can be a less than exciting time for those who like to observe wild birds. The songs of spring gradually grow silent as young birds leave the nest and preoccupy their parents with the chore of gathering enough food to satisfy their ballooning appetites. To avoid predators, roving families of many species remain hidden and as inconspicuous as possible while the young birds learn how to find food and handle the dangers of the world.
But all is not lost. There are two opportunities for seeing unique birds during the hot and humid days of July.
First, many shorebirds such as sandpipers, plovers, dowitchers, and godwits begin moving south from breeding grounds in Canada. That’s right, fall migration starts during the first days of summer, right where spring migration left off. The earliest arrivals are primarily birds that for one reason on another (age, weather, food availability) did not nest this year. These individuals will be followed by birds that completed their breeding cycles early or experienced nest failures. Finally, adults and juveniles from successful nests are on their way to the wintering grounds, extending the movement into the months we more traditionally start to associate with fall migration—late August into October.
For those of you who find identifying shorebirds more of a labor than a pleasure, I get it. For you, July can bring a special treat—post-breeding wanderers. Post-breeding wanderers are birds we find roaming in directions other than south during the summer months, after the nesting cycle is complete. This behavior is known as “post-breeding dispersal”. Even though we often have no way of telling for sure that a wandering bird did indeed begin its roving journey after either being a parent or a fledgling during the preceding nesting season, the term post-breeding wanderer still applies. It’s a title based more on a bird sighting and it’s time and place than upon the life cycle of the bird(s) being observed. Post-breeding wanderers are often southern species that show up hundreds of miles outside there usual range, sometimes traveling in groups and lingering in an adopted area until the cooler weather of fall finally prompts them to go back home. Many are birds associated with aquatic habitats such as shores, marshes, and rivers, so water levels and their impact on the birds’ food supplies within their home range may be the motivation for some of these movements. What makes post-breeding wanderers a favorite among many birders is their pop. They are often some of our largest, most colorful, or most sought-after species. Birds such as herons, egrets, ibises, spoonbills, stilts, avocets, terns, and raptors are showy and attract a crowd.
While it’s often impossible to predict exactly which species, if any, will disperse from their typical breeding range in a significant way during a given year, some seem to roam with regularity. Perhaps the most consistent and certainly the earliest post-breeding wanderer to visit our region is the “Florida Bald Eagle”. Bald Eagles nest in “The Sunshine State” beginning in the fall, so by early spring, many of their young are on their own. By mid-spring, many of these eagles begin cruising north, some passing into the lower Susquehanna valley and beyond. Gatherings of dozens of adult Bald Eagles at Conowingo Dam during April and May, while our local adults are nesting and after the wintering birds have gone north, probably include numerous post-breeding wanderers from Florida and other Gulf Coast States.
So this week, what exactly was it that prompted hundreds of birders to travel to Middle Creek Wildlife Management Area from all over the Mid-Atlantic States and from as far away as Colorado?
Birders observing something special at Middle Creek Wildlife Management Area on July 10, 2023.
Was it the majestic Great Blue Herons and playful Killdeer?
Great Blue Heron and a Killdeer.
Was it the colorful Green Herons?
Green Heron
Was it the Great Egrets snapping small fish from the shallows?
Great Egret
Was it the small flocks of shorebirds like these Least Sandpipers beginning to trickle south from Canada?
Least Sandpipers
All very nice, but not the inspiration for traveling hundreds or even thousands of miles to see a bird.
It was the appearance of this very rare post-breeding wanderer…
A Limpkin at Middle Creek Wildlife Management Area in Lancaster County. The Limpkin (Aramus guarauna) is the only surviving member of the family Aramidae.
…Pennsylvania’s first record of a Limpkin, a tropical wading bird native to Florida, the Caribbean Islands, and South America. Many observers visiting Middle Creek Wildlife Management Area had never seen one before, so if they happen to be a “lister”, a birder who keeps a tally of the wild bird species they’ve seen, this Limpkin was a “lifer”.
The Limpkin is an inhabitant of vegetated marshlands where it feeds almost exclusively upon large snails of the family Ampullariidae, including the Florida Applesnail (Pomacea paludosa), the largest native freshwater snail in the United States.
In the United States, the native range of the Limpkin lies within the native range of the Florida Applesnail, shown here in gold. Introduced populations of the snail are shown in brown. (United States Geological Survey Nonindigenous Aquatic Species image)A spectacular nineteenth-century rendition of the Florida Applesnail, including an egg mass, illustrated by Helen E. Lawson in Samuel S. Haldeman’s “Monograph of the Freshwater Univalve Mollusca of the United States”.
Observations of the Limpkin lingering at Middle Creek Wildlife Management Area have revealed a pair of interesting facts. First, in the absence of Florida Applesnails, this particular Limpkin has found a substitute food source, the non-native Chinese Mystery Snail (Cipangopaludina chinensis). And second, Chinese Mystery Snails have recently become established in the lakes, pools, and ponds at the refuge, very likely arriving as stowaways on Spatterdock (Nupharadvena) and/or American Lotus (Nelumbo lutea), native transplants brought in during recent years to improve wetland habitat and process the abundance of nutrients (including waterfowl waste) in the water.
The Chinese Mystery Snail is the largest freshwater snail in the Lower Susquehanna River Watershed. (Vintage 35 mm image)By hitching a ride on aquatic transplants like this Spatterdock, non-native freshwater snails are easily vectored into new areas outside their previous range.Spatterdock, a native species also known as Yellow Pond Lily or Cow Lily, flowering in August.Blooming American Lotus transplants in a pool at Middle Creek Wildlife Management Area during August.The Limpkin at Middle Creek W.M.A. capturing a Chinese Mystery Snail.The Limpkin at Middle Creek carrying a Chinese Mystery Snail.The Limpkin is seen here maneuvering the the snail in its bill, a set of mandibles specially adapted for extracting the bodies of large freshwater snails from their shells.The tweezers-like tip of the bill is used to grasp the shell by the rim of the opening or by the “trapdoor” (operculum) that protects the snail inside.A posed Chinese Mystery Snail showing its “trapdoor”, the operculum protecting the soft body tissue when the animal withdraws inside. The tips of the Limpkin’s bill close tightly like the end of a tweezers to grasp the operculum and remove it and the snail’s body from the shell. (Vintage 35 mm image)The tweezers-tipped bill, which is curved slightly to the right in some Limpkins, is slid into the shell to grasp the snails body and remove it for consumption. The entire extraction process takes 10 to 30 seconds.
The Middle Creek Limpkin’s affinity for Chinese Mystery Snails may help explain how it was able to find its way to Pennsylvania in apparent good health. Look again at the map showing the range of the Limpkin’s primary native food source, the Florida Applesnail. Note that there are established populations (shown in brown) where these snails were introduced along the northern coast of Georgia and southern coast of South Carolina…
Native (gold) and non-native (brown) ranges of the Florida Applesnail. (United States Geological Survey Nonindigenous Aquatic Species image)
…now look at the latest U.S.G.S. Nonindigenous Aquatic Species map showing the ranges (in brown) of established populations of non-native Chinese Mystery Snails…
Range (in brown) of established populations of non-native Chinese Mystery Snails. (United States Geological Survey Nonindigenous Aquatic Species image)
…and now imagine that you’re a happy-go-lucky Limpkin working your way up the Atlantic Coastal Plain toward Pennsylvania and taking advantage of the abundance of food and sunshine that summer brings to the northern latitudes. It’s a new frontier. Introduced populations of Chinese Mystery Snails are like having a Waffle House serving escargot at every exit along the way!
Be sure to click the “Freshwater Snails” tab at the top of this page to learn more about the Chinese Mystery Snail and its arrival in the Lower Susquehanna River Watershed. Once there, you’ll find some additional commentary about the Limpkin and the likelihood of Everglade Snail Kites taking advantage of the presence of Chinese Mystery Snails to wander north. Be certain to check it out.
The endangered Everglade Snail Kite (Rostrhamus sociabilis plumbeus), a Florida Applesnail specialist, has survived in part due to its ability to adapt to eating the non-native Pomacea maculata applesnails which have become widespread in Florida following releases from aquaria. The adaptation?…a larger body and bill for eating larger snails. (National Park Service image)
(Public Domain image by RCA Manufacturing Company)
A brief test procedure has rubbed the epithelial cells in one of your editor’s critical organs the wrong way, thus prompting them to let loose with cytokine proteins all over his body. Shortly put, it’s like experiencing the aftermath of a major heart attack for a second time. We’ll be back when the action-packed reenactment is over.
The gasoline and gunpowder gang’s biggest holiday of the year has arrived yet again. Where does all the time go?
In observance of this festive occasion, we’ve decided to take a look at all the stuff that’s floating around in the atmosphere before all the motor travel, celebratory fires, and exciting explosions get underway.
We’ll start with the smoke from wildfires in Canada…
In the margins between the water vapor clouds, a smoky haze can be seen across the Mid-Atlantic States this morning. To warn residents of the potential health impacts, air quality alerts have been issued by numerous state and local agencies. (NOAA/GOES image)Near the top of this image, ember-red areas denote the locations of some of the hottest forest fires presently burning in remote portions of northern Quebec. (NOAA/GOES Fire Temperature Composite image)Wildfires are now burning in every province in Canada. Note the smoky haze that is visible between the water vapor clouds as it drifts from Alberta, Manitoba, and Saskatchewan through the Great Lakes and into the Mid-Atlantic States. (NOAA/GOES image)
If you think that smoke accounts for all the particulate matter now obscuring skies in the northern half of the western hemisphere, then have a gander at this…
As it often does at this time of year, dust from the Sahara Desert in northern Africa is blowing into the Caribbean Islands and Amazonia. This morning’s full-disk satellite image shows both the smoke from wildfires in Canada and a well-defined earth-tone cloud of desert dust streaming west across the Atlantic from Africa. (NOAA/GOES image)
As you can see, natural processes are currently providing a plentiful load of particulates in our skies. There’s no real need to aggravate yourself and the situation by sitting in traffic or burning your groceries on the barbecue. And you can let those cult-like homeowner chores for later. After all, running the mower, whacker, and blower will only add to the airborne pollutants. While celebrating this Fourth of July, why risk mangling fingers on your throwing hand or catching the neighbor’s house on fire when you could just relax and quietly eat ice cream or watermelon? Yeah, that’s more like it.
Don’t want to feed suet to the birds around your home during the blazing heat of summer? Well, you might be glad to know that peanuts offered in one of these expanded metal tube feeders make a great substitute. They provide a nutritious supplement to naturally occurring foods for nuthatches, chickadees, Tufted Titmice, Carolina Wrens, Blue Jays, finches, and woodpeckers including this Red-bellied Woodpecker. Secured to a vertical length of wire hung from a horizontal tree limb, these feeders have proven so puzzling to the squirrels at susquehannawildlife.net headquarters that they no longer make any effort to raid them. Though marketed primarily to dispense suet nuggets, powder-coated metal mesh feeders can be used for sunflower seeds too. This juvenile House Finch plucks the black oil variety from one of the tubes in our garden. Seeds that fall are quickly scarfed up by ground-feeding species including Northern Cardinals, Eastern Chipmunks (Tamias striatus), and frustrated squirrels. Fewer seeds are lost if the larger varieties of sunflower such as “grey stripe” are used.
When you were young, you may have selected your tennis shoes because they promised to make you run faster and jump higher. Remember those? Then as you got a bit older, you may have really wanted the brand that would get you noticed—those overpriced status-symbol athletic shoes. As the years went by and you put on the extra pounds in all the wrong places, maybe it occurred to you that it might be a really good idea to get some exercise. So you went out and bought some stylish and expensive fitness footwear that promised to help you run faster and jump higher—and wore that pair to go shopping at the grocery store. Then you finally realized…
…that the best sneakers are just something comfortable to loaf around in.Wisdom does come with age.
