The Stewards of Inland Shorebird Habitat

As they travel between coastal wintering ranges and breeding territories in Canada and Alaska, vast numbers of shorebirds pass through the lower Susquehanna region each spring and fall—though few stop here to rest, feed, and provide us with an opportunity to observe them. Prior to the construction of man-made dams and other alterations on our lotic (flowing) waterways, shorebirds took advantage of lateral bars, stream deltas, and other alluvial deposits as places to loaf and re-energize. Before they were drained and filled, some wetlands probably included sparsely vegetated flats where shorebirds could drop in for a brief visit. Prior to their extirpation from our region, North American Beavers were the primary providers of quality habitat for shorebirds and other migrating waders. Their widespread network of dams, pools, and marshes maximized floodplain function by keeping streams thoroughly connected to their wetlands, nurturing plant communities that not only provided food and shelter for the beavers and other wildlife, but provided superb buffering against erosion while protecting against sediment and nutrient imbalances in lower Susquehanna waterways.

Beaver Dam and Lodge — Beaver dams need not be large, particularly on low-gradient streams where a structure like this is sufficient to create a pool with depth adequate for building and maintaining a lodge and transporting leafy branches and other food items by water.

A beaver lodge assembled in a pool with less than three feet of water, a depth adequate to provide the family with a measure of protection against terrestrial predators.

Beaver Pool — Beaver pool ecosystems provide homes for hundreds of species of plants and animals, including migrating shorebirds and other waders.

Mud flats in the margins between emergent shrubs and herbaceous plant growth attract migrating shorebirds like this Solitary Sandpiper to the abundance of invertebrate life. Seasonal movements of migrating shorebirds regularly coincide with the reductions of water levels in beaver pools which typically occur between May and September each year.

During its southbound migration in September, a Least Sandpiper searches for arthropods and annelids as it visits a food-rich puddle along the periphery of a beaver pool.

A Pectoral Sandpiper feeds in shallow water near a stand of Common Cattails in a beaver pool.

During migration, rails including the Sora are attracted to dense emergent vegetation in beaver pools. Some will nest in these rodent-managed refugia.

Green Herons visit beaver pools during migration and, due to the reliability of the food supply, will often nest in their vicinity.

So how did this happen? How did the North American Beaver become a keystone species—an animal upon which the majority of other life forms within its ecosystem are so reliant? Well, it’s largely due to the fact that our beavers aren’t particularly fond of a constant stream of noise. More specifically, they don’t like the sound of running water in places where they intend to build and maintain a lodge. And so, as they begin to place sticks, mud, limbs, stones, and other materials within a noisy riffle on a stream, they create a dam, and behind it a pool—a pool that is particularly advantageous for protecting their home and providing a means of conveyance for their construction materials and food supplies.

Cascading Series of Beaver Pools — On a high-gradient segment of stream, beavers will create a cascading series of pools. Because water filters through a beaver dam instead of spilling over it, the work of these meticulous rodents soon silences the sounds of water changing altitude. No more sonorous riffle.

Beaver Dam — **Quiet please.** High seasonal stream flow and damage from storms may create areas where water begins to erode the structure of the dam. Where this condition persists, an adult beaver will soon mend the breach, just to quiet things down. Why would beavers demand such a hush upon their domain? Well, they have poor eyesight, but their hearing is excellent, and they rely upon it to detect danger.

The Louisiana Waterthrush (Parkesia motacilla), one of our earliest-arriving warblers, nests in forests along clear, high-gradient streams. In mid-April, we found this individual and three others squabbling over a breeding territory adjacent to the series of cascading beaver pools shown in the previous images.

Brook Trout and Blacknose Dace — Native denizens of coldwater streams, neither the Brook Trout nor the Eastern Blacknose Dace has any difficulty finding its way through the voids in beaver dams to ascend and descend the sequence of pools.

Beaver Dam and Pool on Low-gradient Stream — On a low-gradient segment of stream, a dam just over a foot in height may be sufficient to create a beaver pool of considerable size. Resembling the water-logged muskeg of the far north, well-established beaver pools form boggy habitats familiar to migrating shorebirds.

Water levels in the pools usually drop along with the stream’s base flow as soon as the effects of the spring thaw and rain showers subside. Feeding areas in shallow water and on muddy ground are often revealed just in time for northbound shorebirds and other waders to stop by in late April and May. These conditions often persist through the growing season and the fall migration of these birds which begins as early as late June and sometimes extends into October.

Tree Swallow at Natural Cavity — Where the pool inundates standing timber such as Red Maple and other species not used by beavers, the dead snags provide vital feeding areas for birds and other wildlife. Cavity nesters like this Tree Swallow seldom find suitable natural housing elsewhere.

Eastern Newts Among Alderfly Exuvia — The mosaic of marshlands and braided stream channels within the beaver pool complex supports an abundance of aquatic life including these breeding Eastern Newts, seen here surrounded by the exuvia left behind following a massive hatch of alderflies (Sialinae). Alderflies are a stream inhabitant during their larval stage and are indicator of clean water conditions.

**A modern-day example of the way fully functional stream floodplains used to look in the Lower Susquehanna River Watershed.** Though the pools may appear pond-like during the cooler and wetter months of the year, by summer the water levels behind the beaver dams recede as the base flow of the stream wanes. Now the true nature of the marsh, shallow pool, and braided stream complex is revealed. Unlike most man-made dams that set a fixed pool elevation regardless of flow by discharging water over the top of the structure or through a spillway or gate, beaver dams merely throttle the flow through their porous construction. Unless the beaver begins plugging small leaks as fast as the stream flow ebbs, the water level in the pools will drop. So they’re not left high and dry, lodges are located in the deepest pools, usually in close proximity to the dam and/or one of the stream courses. Though not inundated during the dry season, the soil in the pool complex is almost always damp and plants grow vigorously there, sequestering nutrients and retaining sediments in beneficial deposition patterns that actually inhibit erosion of the riparian landscape. These streams and floodplains retain their hyporheic zone and freely exchange water with the underlying water table and aquifer. It’s the ultimate floodplain management system, and the beavers don’t even know their doing all the work. (Google Maps base image)

Where another high-gradient segment of stream enters the main pool complex, beavers have created an additional series of cascading pools. The impoundments created by these dams help diffuse stormwater energy and process more nutrients and sediments within the floodplain’s wetland vegetation system. (Google Maps base image)

Sandhill Cranes in Beaver Pool — Beaver pools often refill as stream flows increase following autumn rains. The stockpiles of vegetative foods that grew within the beaver’s domain through the summer then become flooded and are a prime source of nutrition for not only the beavers, but waterfowl during both their autumn and spring migrations. Gleaning and probing Sandhill Cranes often find these habitats to their liking as well.

While North American Beavers have returned to the region, most live as “bank beavers”, residing in the river and larger creeks of the valley where they excavate shelters among the roots of Black Willows and other shoreline trees and shrubs. Floodplain encroachment, legacy sediment deposits, and just plain human intolerance have all conspired to prohibit North American Beavers from performing their magic on smaller local streams. For migrating shorebirds, this continued absence of beaver dam ecosystems has turned much of the lower Susquehanna valley into “fly-over country”. Those travelers that do stop to rest and feed concentrate at the few favorable locations such as the lateral bars and the hydroelectric dam-created delta at the Conejohela Flats on the river in Lancaster County. But centralization has its drawbacks. Migrants spending time at concentration points may have a greater chance of contracting and spreading disease. Protracted heavy foraging can degrade these habitats. And over time, features such the lateral bars and delta deposits, including those on the Conejohela Flats, transition into other habitats—riparian forests. A more widespread selection of favorable stopover points for shorebirds, waders, waterfowl, and other migrants is certainly desirable.

IMITATION IS THE SINCEREST FORM OF FLATTERY

Until public sentiment sways in favor of the North American Beaver, wildlife managers are mimicking some of the attributes of their sound-inspired installations.

New Impoundment at Middle Creek Wildlife Management Area — Created by excavating a depression in heavy clay soils, this new impoundment at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area collects water directly from rainfall and from surface runoff. Its depth at no point is greater than about two or three feet. During the driest times of the year, this space will be a mudflat, and a haven for migrating shorebirds.

Conservation impoundments designed, constructed, and managed for migratory waterfowl and waders including shorebirds are not what we typically refer to as ponds—though they are lentic (still) waters. They more closely resemble the prairie potholes created by glacial scour in the north-central United States and adjacent portions of Canada. Many populations of migratory birds are familiar with these ecosystems and, like the beaver pools and marshes, have relied upon them as waypoints along their journeys for centuries.

Pond and Lake Zonation — Impoundments most beneficial to migrating waterfowl and waders including shorebirds are shallow in depth. They lack the deeper waters of a pond or lake and thus have no limnetic (open water) or aphotic zones. Managed throughout as a littoral zone, impoundments grow plants in shallow water or on damp soil through the summer months to chew up nutrients accumulated in waste from visiting birds. This vegetation is then flooded from late fall through early spring as forage habitat for visiting waterfowl. The timing of the fluctuations in water levels approximates those of the beaver pools and marshes on lotic (flowing) waters.

Impoundment Gate — A gate assembly is a water control structure installed to provide seasonal management of the water levels in a conservation impoundment. More than once, beavers have heard the sounds of tumbling waters inside these types of devices and tried to dam them up!

You’ve heard the line, “If you build it, they will come.” Well, it’s true. Here is a sample of the activity witnessed during the past two weeks at the new impoundment completed just several months ago at Middle Creek Wildlife Management Area along the tour route just beyond the “Stop 3” overlook.

The ubiquitous Snow Geese were among the first migratory occupants of the new impoundment. A few are currently lingering in the vicinity.

Ducks soon followed. These Blue-winged Teal were among the last to pass through earlier this month.

Wilson's Snipe — Wilson’s Snipe were among the first species of migratory shorebirds to visit the new impoundment at Middle Creek…

…as were Killdeer, a species which nests nearby.

Then, earlier this month, flocks of shorebirds including these Least Sandpipers were arriving to feed and rest.

Least Sandpiper and a Semipalmated Sandpiper — A Least Sandpiper (left) and a Semipalmated Sandpiper (right).

White-rumped, Least, and Solitary Sandpipers — Here’s a Solitary Sandpiper (right) feeding alongside a Least Sandpiper (center) and a White-rumped Sandpiper (left).

Least Sandpiper and a White-rumped Sandpiper — A Least Sandpiper (left) and a White-rumped Sandpiper (right).

Rare in our area during spring, the White-rumped Sandpiper (Calidris fuscicollis) flies north using the central flyway, then heads south along the Atlantic flyway in the fall, when it tends to be more regular here.

Rocking and teetering along as it looks for food, the Spotted Sandpiper may be one of the easiest of shorebirds to identify.

Chunky little Dunlin (Calidris alpina) with their conspicuously down-curved bills are another easy-to-identify species, particularly in the spring when their breeding (alternate) plumage includes a black belly.

Black-bellied Plovers acquire a handsome set of plumage in the spring as well.

The Semipalmated Plover doesn’t have nearly the flair for ornament its close relatives the Killdeer do; these little shorebirds wear only one ring around their neck instead of two. Think this plover is cute?,…

Speaking of legs, here’s one of dozens of Lesser Yellowlegs that visited the new impoundment during their recent northbound travels.

Though less numerous than the smaller Lesser Yellowlegs, the Greater Yellowlegs seldom goes unnoticed when dropping by the impoundment.

Wilson's Phalarope — Everybody likes to see something unusual every now and then, and this impoundment delivers. Just yesterday, we photographed this migrating Wilson’s Phalarope (top center) among two Least Sandpipers (top left and right), a Lesser Yellowlegs (left and slightly forward of the phalarope), and a White-rumped Sandpiper (foreground).

Species of wildlife in addition to shorebirds and waterfowl have already found the new impoundment favorable…

Pairs of breeding Spot-winged Gliders (Pantala hymenaea), seen here in tandem while ovipositing, were swarming the impoundment after arriving earlier this week. These ofttimes unpredictable nomads are similar in appearance to the usually more numerous Wandering Gliders. To recolonize seasonal portions of their range, both are famous for hitching a ride en masse on storm systems. They share the behavior of finding ephemeral and new bodies of water favorable for egg laying due to their low density of aquatic predators.

We watched this Snapping Turtle arrive to apparently find the newly created waters to its liking. Snapping Turtles are important consumers in a wetland ecosystem. Larger specimens may fill the role of an upper trophic level or apex predator, eliminating vulnerable mid-level consumers including other Snapping Turtles.

Because it bounces its tail up and down like a Spotted Sandpiper, you might mistake an American Pipit for a shorebird. Long known as the Water Pipit, these birds have been visiting the impoundment while migrating north and stopping to feed in nearby croplands.

Seen here with Least Sandpipers is a visiting Glossy Ibis.

Western Cattle Egrets visiting Middle Creek this spring have been frequenting the new impoundment.

Western Cattle Egret — With still little in the way of insects such as grasshoppers available in the surrounding landscape,…

…cattle egrets are looking to find prey like this Green Frog.

Managing saturation levels in shallow-water impoundments to resemble the seasonal variations in beaver pool and marsh systems can create lush growth and wildlife-rich environments. Take a look at some images from a project in a headwaters area of a tributary to Conewago Creek (west)…

Well-established Shallow-water Impoundment — Seen in late July, this impoundment is dense with emergent and aquatic plants, the latter kept hydrated in deeper pools of the project by the influx of several captured springs that supplement direct rainfall and sheet runoff to supply its water. Through the heat of mid-summer, southbound shorebirds were already using these mudflats to feed and rest.

Water Lily Growth — In the deeper pools of the impoundment, water lilies and other aquatic plants grow in dense mats to provide cover and feeding areas for resident populations of reptiles and amphibians.

Immature Little Blue Heron — …and this immature Little Blue Heron, a wanderer typical of more southern latitudes.

While walking the road among tall grasses in the supporting landscape surrounding this impoundment, we were at first startled when these Sandhill Cranes strode by going the other direction. We quietly kept moving,…

Sandhill Cranes with Colt — The hatching of this colt is testimony to the vital role wetland ecosystems play in the lives of hundreds of species. Whether they be beaver pools and marshes on lotic waters or man-made shallow lentic waters, each of these habitats is filling a void that left floodplains and other critical lowland biomes faltering. While they can’t replace the full-function floodplain management provided by an active beaver colony, shallow-water impoundments can provide relief for habitat-starved populations of the animals and plants that rely upon them. A constellation of these projects on lands public or private across the lower Susquehanna watershed could help provide refuge for many of our flora and fauna with the most desperately fragmented of ranges.

So that you can relax while observing the comings and goings at a pair of the lower Susquehanna valley’s man-made impoundments, the Pennsylvania Game Commission has erected two viewing pavilions for public use on its lands…

The Haldeman Island pavilion is located on State Game Lands 290 just upstream from the Juniata River’s mouth on the Susquehanna at Clark’s Ferry in Dauphin County.

Haldeman Island Pavilion — It overlooks not only the island’s man-made shallow-water impoundments and neighboring grasslands, but the tower used in the 1970s to reintroduce Bald Eagles from Saskatchewan to the lower Susquehanna. Interpretive signs explain the conservation stories of habitats and the eagle reintroduction program.

Middle Creek W.M.A. Observation Pavilion — The observation pavilion at Middle Creek Wildlife Management Area is of similar construction to the one at Haldeman Island. It is accessed from the parking pull-off along the tour road at its intersection with Chapel Road, just before the right turn and incline that leads to the “Stop 3” grassland overlook.

It too includes numerous interpretive signs to help visitors understand impoundment management.

During the next two weeks, the exodus of migrating shorebirds now staged and feeding upon Atlantic Horseshoe Crab eggs on Delaware Bay will commence. During the evening of their departure from the bay, many of these birds cross portions of the eastern Lower Susquehanna River Watershed. Stormy weather and other climatic conditions may force some of them to seek a place to put down temporarily, so keeping a close eye on the new impoundment at Middle Creek may be a prudent move. After that, waders known as “post-breeding wanderers” can show up at any time. Then, beginning as early as late June, shorebirds begin moving south on a migration that can provide us with viewing opportunities into September and beyond. See you out there!

May 10, 2026May 13, 2026

Scenes from a Managed Grassland Ecosystem

Pretty pictures…

Veery — Photographing Neotropical migrants as they arrive to nest in our local forests can be frustrating under closed canopies and diffused light conditions. We instead decided to focus our efforts and lenses on the grassland species at Middle Creek Wildlife Management Area where, earlier this week, several prescribed fire burns were wrapping up the spring treatments for 2026.

Warm-season grassland — Managed warm-season grassland habitat at Middle Creek Wildlife Management Area.

Already tending nests containing eggs and young, Tree Swallows are scattered around the refuge’s grasslands to welcome newly arriving migrants, and human visitors!

Middle Creek is probably the best place in the lower Susquehanna valley to see ground-nesting Grasshopper Sparrows.

Grasshopper Sparrow and Tree Swallow — It’s a behavior that occasionally puts the newcomer Grasshopper Sparrow in conflict with a well-ensconced Tree Swallow guarding its nest box.

Eastern Bluebirds squeeze in among the Tree Swallows to find a vacant nest box within which they can raise their young.

Song Sparrows may be present year-round in grasslands with interspersed growth of early successional shrubs and briars, however the populations at a given location changes as birds migrate in and out of the area seasonally. Those individuals observed nesting here in the spring and summer are not usually the same birds seen during the late fall and winter.

As a species, the Savannah Sparrow, like the Song Sparrow, may be represented in the grasslands throughout the year, though the individuals vary with the changing seasons. Unlike the Song Sparrow which nests in shrubs, the Savannah Sparrow nests on the ground among thick grasses.

Still munching on the seeds of last year’s wildflowers, American Goldfinches are pairing up around the grasslands waiting for thistle down and other fibers to be available for nest construction. They will then build a small cup within the upper limbs of a small to medium-sized tree to shelter their eggs and nestlings.

The Common Yellowthroat, one of our Neotropical warblers, will reproduce in grasslands with scattered early successional thorny shrubs to afford a safe place to construct a nest. The males sing almost incessantly from the time they arrive in late April and early May until sometime in July, or even later.

Eastern Kingbirds are arriving from tropical wintering grounds to nest in grasslands and pastures throughout the lower Susquehanna region. They are a tyrant flycatcher that will ambush insects from a perch atop a tree, shrub, or tuft of tall grass.

Northern Mockingbirds are considered a resident species, though there may be some southward movement during severe winters when foods such as berries become scarce. At Middle Creek, some of these evacuees have returned to nest along the interface zone between the grassland and early successional shrub habitats.

Nesting Horned Larks apparently take advantage of the interface between the managed grasslands and nearly bare soils in spring-planted croplands at Middle Creek. They may be one of the few, possibly the only, species of bird life to take advantage of no-till farming for nesting on the ground.

Red-winged Blackbirds not only construct nests in cattails and other marsh growth, but in dense grasses and shrubs in and near grasslands and pastures. The males are easily seen displaying their plumage as they sing from a small tree, shrub, or a cattail seed spike.

Female Red-winged Blackbird — Less frequently seen is the female Red-winged Blackbird. She spends the majority of her time in the nest, but will periodically come out to join the ruckus when an intruder is being scolded into leaving the premises.

Another member of the blackbird family is the Eastern Meadowlark. It too is an energetic singer during the breeding season. Meadowlarks build their nests on the ground, often in pastures of cool-season grasses where harvests prior to August are fatal to eggs and young. Middle Creek’s managers delay harvests to allow the birds enough time to complete their reproduction cycle.

Another blackbird is the Bobolink, an obligate grassland specialist and a Neotropical migrant. One of its few uses for trees and shrubs is as a place to burst into display and song, though it will also perform these rituals on the wing and among the grass.

Bobolinks — As the males arrive, competition for suitable nesting sites among the grasses becomes intense, even before a female is anywhere in sight.

Did you know the Baltimore Oriole is a blackbird? Well, it is.

Baltimore Orioles — Competition for these ideal sites can get quite animated.

Western Cattle Egrets and Geese — It’s an unusual experience, watching Western Cattle Egrets feeding in a field of cultivated cool-season grasses alongside Snow Geese, though they certainly aren’t looking for the same thing.

We’ll have more on the Western Cattle Egrets and other interesting migrants at Middle Creek in an upcoming post. Check back soon!

May 8, 2026May 8, 2026

New Floodplain and Stream Restoration Projects 4U2C

The majority of floodplains in the Lower Susquehanna River Watershed are, to put it mildly, dysfunctional. They are frequently disconnected from the lotic (flowing) streams and rivers that created them and are no longer capable of absorbing, purifying, and infiltrating high water during rain and snow-melt events. They are an ongoing source of nutrient and sediment pollution that impairs both our local waterways and Chesapeake Bay. For the plant and wildlife species that rely upon them for survival, their loss has been catastrophic.

Mill dams once impounded nearly every mile of low-gradient streams in the lower Susquehanna region. During the century or more of their existence prior to the implementation of soil conservation practices during the 1930s, sediments consisting mostly of nutrient-loaded clay, silt, and sand eroded from sloped terrain in storm runoff and accumulated behind these dams, filling many to the brim with mud—six, eight, ten, sometimes twelve feet deep! As the dams fail or are removed, the stream is left channelized, incising a path through a floodplain choked with these “legacy sediments”.

Legacy Sediments — **That’s it, a fresh layer of topsoil oughta do it-** Landowners have become so accustomed to legacy sediment pollution and channelized creeks, they think it’s how a natural stream and floodplain ecosystem looks and operates. They’ll even replace washed away legacy sediment with dirt, rocks, cinder blocks, old sidewalks, and other construction debris to “shore things up”. It’s a delusional approach to managing a space that functions as a place to store, treat, and percolate precious water.

Disconnected Stream — Legacy sediments not only displace flood waters into historically unimpacted lands, they clog the hyporheic zone, the area below and alongside the flowing stream where water is exchanged with the aquifer. (United States Geological Survey image)

Legacy Sediment Layer — Restoration of creeks and floodplains impacted by legacy sediment pollution includes the removal of the layer of nutrient-loaded sand, silt, and clay deposited atop the historic stratum of wetland and stream substrates. These lower layers are frequently gray in color and contain gravel from the former stream beds. Within the organic matter contained in this historic layer, seeds of native wetland plants have rested dormant for more than one hundred years. Some are still viable and will germinate upon liberation from the overlying legacy layer.

Legacy Sediments Removed from Swarr Run Floodplain — Stockpiles of legacy sediments and larger fill items await transport to upland sites for reuse following removal from the impaired floodplain during a stream restoration project completed during the past year on Swarr Run, a tributary of Little Conestoga Creek in Lancaster County, Pennsylvania. This excavation, part of the project’s phase-two plan, was preceded in 2020 by similar construction completed as phase one on a segment located just upstream.

Swarr Run has been returned to a meandering stream course in a floodplain cleared of nutrient-rich legacy sediments. How are legacy sediments repurposed? Those from previous projects in the lower Susquehanna valley have been used to create a community garden, as a top coating on athletic fields, as a safety backstop at a shooting range, and as topsoil on some of the croplands from whence they came all those years ago.

Functional Hyporheic Zones — Restoring pool and riffle or meandering stream flow regimes also restores water exchange in the adjoining hyporheic zones. (United States Geological Survey image)

Before and After Legacy Sediment Removal — Removing legacy sediment accumulations from the floodplains of impaired streams allows re-establishment of historic flow regimes and reconnection of the waterway to the nutrient-sequestering functions of fluvial wetlands. In addition, flow into and from the water table is restored.

Swarr Run Stream and Floodplain Restoration — Question: Do you know why nearby businesses have ATMs? Answer: Because there are no longer any banks on this segment of Swarr Run.

For a closer look at the Swarr Run renewal, including early map and aerial photograph views of the site, take a peek at this synopsis illustrating phase one of the project as presented to the North American Association of Wetland Managers by Justin Spangler of LandStudies, designers of the restoration. While perusing the material, be sure to review the significant reductions in nutrient and sediment loads discharged from these sites following the restoration work.

Stormwater Discharge — Floodplain restorations that include legacy sediment removal are often designed to reduce the impact of stormwater from sheet runoff, piped discharges, or conveyances such as eroded and excavated ditches like this one. Projects that restore floodplain functions including storing, filtering, and percolating runoff effectively can sometimes help municipalities and property owners meet regulatory stormwater requirements, including those under the Environmental Protection Agency’s MS4 (Municipal Separate Storm Sewer System) program.

Little Conestoga Stream and Floodplain Restoration — This floodplain restoration was completed during the past year along a segment of Little Conestoga Creek in Lancaster, Pennsylvania. Here the stream receives a heavy dose of stormwater runoff from shopping malls and other commercial properties with acres upon acres of paved parking space. Because they lack connection to any retention or detention systems, storm drains pipe runoff directly from the pavement into the creek from many of these properties.

Little Conestoga Reconnected to Floodplain — Legacy sediment removal eliminates the steep banks along the channelized creek. Braided stream courses were added to again connect the Little Conestoga to its historic floodplain and a complex of wetlands to help store, treat, and infiltrate stormwater.

Removing legacy sediments brings an end to their practice of forcing detrimental high waters out to roam adjacent neighborhoods, putting floods back in the floodplain where they belong.

Conoy Creek Stream and Floodplain Restoration — Completed last year in conjunction with replacement of an aging sanitary sewer interceptor (buried to the right), this floodplain restoration removed legacy sediments and returned Conoy Creek in Lancaster County, Pennsylvania, to a natural, braided stream course.

Cocalico Creek Stream and Floodplain Restoration — Following restoration, a braided stream design reconnects Cocalico Creek in Lancaster County, Pennsylvania, to its floodplain wetlands along this 3,867-foot segment near its headwaters. As part of the construction process, twenty-five acres of invasive growth was treated prior to the planting of 12.7 acres of native riparian buffer species. According to the Pennsylvania Department of Environmental Protection’s (DEP) project summary, annual pollutant load reductions resulting from this restoration are calculated to be 862,980 pounds of Total Suspended Sediments/year, 1,641 pounds of Total Nitrogen/year, and 426 pounds of Total Phosphorus/year.

Hammer Creek Stream and Floodplain Restoration — **Hammer time-** Now under construction: after years of planning and preparations, a floodplain and stream restoration on Hammer Creek near Buffalo Springs in Lebanon County, Pennsylvania, returns this low-gradient segment to a meandering flow regime. Instead of a ditch between fields, it’ll once again be a creek connected to a nutrient-reducing wetland buffer.

Have an impaired stream and a floodplain full of legacy sediments on your land or in your neighborhood? Consider encouraging your municipality, volunteer watershed group, or county conservation district to investigate the possibilities of planning and installing one of these projects. They’re a great enhancement for recreational fisheries, a private refugium, or a community-owned passive park system. Funding assistance may be available, particularly when regulatory or conservation goals can be met by completing restoration.

April 25, 2026May 3, 2026

Preventing Roof Water from Becoming Storm Runoff

Clean rainwater flowing from the roofs of our homes and other buildings is a precious resource that is more often than not directed as quickly as possible into streets and storm drains to become contaminated runoff capable of polluting streams and aggravating flooding conditions. The prudent property steward will manage rainfall as the treasured commodity that it is—retaining on site as much of it as possible.

Whether you happen to be obtaining your domestic water from a community distribution system or from a private well, infiltrating the rain that falls on your roof directly into the soil is a good way to make sure it helps recharge the groundwater stored in the aquifer—banking it for future use.

Better yet, rain barrels provide a means of intercepting, storing, and dispensing a share of the precipitation from your downspouts before it is discharged into the ground. Water from rain barrels can be drained to hydrate garden plants, rinse the patio, wash the dog, or, in the event of an emergency, fill the need for non-potable uses that include bathing and flushing toilets. For potable applications such as cooking, drinking, and brushing teeth, roof water must first be filtered and disinfected.

Rain barrel layout for a 1,200-square-foot house. — Rain barrels collect and save clean roof water for later use. For every one inch of precipitation, the roof on a house or other building with a footprint of just 1,200 square feet sheds enough to fill about fourteen 55-gallon drums. Live in a home supplied by a well? That’s over 700 gallons of soft water available as you need it, such as during a power outage when the pump won’t operate.

Here at susquehannawildlife.net headquarters, our goal is “no net runoff” from our little urban parcel. We use three rain barrels to catch discharge from the spouting system, then direct the overflow into three different management configurations.

RAIN GARDEN

Commercial Rain Barrel — Precipitation from the downspout is captured in this commercially produced rain barrel. The overflow is directed into the black corrugated pipe…

Gravel Beds in Rain Garden — The gravel beds diffuse energy produced not only by water rushing from the barrel’s overflow, but from sheet runoff entering the garden from the driveway and neighboring street.

STOCK TANK PLANTER

Planter Receiving Overflow from Rain Barrel — This home-brew rain barrel was fabricated from a 55-gallon drum originally used to ship and store car wash soap. We scrubbed it out, then plumbed it up with the overflow feeding a 100-gallon stock tank fashioned into a planter. The drain from the planter sometimes discharges a trickle of water into a porous area of the pavement where it soon percolates into the ground. In this “container rain garden”, we grow wildflowers, sedges, rushes, and warm-season grasses.

THE GARDEN POND/POOLS

Pond/Pool Receiving Overflow from Rain Barrel — Here’s another rain barrel made from a soap drum. This one discharges its overflow into the upper pool of our garden pond.

Overflow from Upper Pool Fills Lower Pool (Sump) — …into the lower pool (bottom right). The lower pool consists of a 100-gallon stock tank sunk below grade level and surrounded by rock. Throughout the summer, a pump located in this “sump” circulates water to a spring-like feature in the upper pool. During a downpour, the lower pool will overflow into the encircling boulders, the level often rising half a foot or more above the top rim of the stock tank flooding the cap stones before receding as the excess water slowly infiltrates down through the gravelly soils. It’s a self-automated, hassle-free water change that helps keep our pools and their inhabitants healthy.

Rain Barrel Overflow Management — Our garden pond stormwater management system keeps the pools topped off and the aquatic ecosystem thriving.

Why not give roof water management a try on your property? It’s a great project for exercising your ingenuity and creativity while doing something truly beneficial for the watershed within which you reside. Best of all, it’s really not that expensive to get started.

Container Rain Garden — A virtuous goal in a wasteful world.

April 22, 2026April 22, 2026

The Water Cycle

The dynamics of water’s storage and movement on earth, as well as its transitions through the primary states of matter—gas, liquid, and solid—are known as the water cycle. Understanding how life-supporting water progresses through the global biosphere can provide each of us with a greater appreciation of how fragile a resource clean water is in the presence of 8.3 billion wasteful and polluting humans.

This graphic created by the United States Geological Survey illustrates many of the components of the water cycle’s complex processes. Have a look…

For a PDF version which includes a more detailed description along the bottom margin, click here. We like it so much, we’ve added it to this webpage’s sidebar for future reference.

April 14, 2026May 13, 2026

Managing a Grassland Ecosystem

If you’ve visited Middle Creek Wildlife Management Area anytime during the past month, you may have noticed quite a bit of activity around the large pole-mounted nest boxes placed out in the open fields.

American Kestrels at nest box — American Kestrels, a male and female, at a nest box at Middle Creek Wildlife Management Area.

Kestrels competing for nesting sites and territory. — Within the past week, we noticed that kestrels are still competing for nesting sites and territory, driving away unwanted trespassers. A pair of kestrels seemed to be occupying each of the four box sites we observed. That’s good news, but the presence of these man-made nest cavities is in no way wholly responsible for this positive response from these declining birds. It’s a matter of habitat, much needed grassland habitat.

Cool Season Grasses — Cool-season grasses including fescue (Lolium species) and non-native Orchard Grass (Dactylis glomerata) mature by late spring. If left standing, they continue to provide indispensable habitat for grassland wildlife through the summer. Mowing early and often for hay harvest has rendered most cool-season meadows death traps for nesting birds. By delaying cuttings until at least early August, ground-nesting birds are given adequate time to fledge their young and get the juvenile birds strong enough to fly away from a set of spinning blades.

Grasshopper Sparrow in Timothy — A male Grasshopper Sparrow sings to demarcate its nesting territory in a stand of Timothy, an introduced species cultivated for harvest as hay. An insectivore during the breeding season, he and his mate will attempt to raise their brood exclusively within the cover of these cool-season grasses.

Orchard Grass with Black-eyed Susans — Prior to being mowed, Orchard Grass provides the short, dense cover meadowlarks and other ground-nesting grassland birds need to successfully reproduce. Growing a cool season grassland can be as easy as delaying the mowing of a pasture, field, or oversize lawn until August, then have a farmer friend come and take off a cutting or two of hay or straw to prevent woody plants from becoming established.

Warm-season Grasses — More durable stands of native warm-season grasses including Big Bluestem, Indiangrass, Switchgrass, and Little Bluestem thrive in the summer heat and provide wildlife habitat throughout the year. These perennial “prairie grasses”, fed by root systems five to eight feet deep, are especially drought tolerant . From these entrenched anchors, the plants would quickly bounce back after the once great herds of hungry bison had grazed the landscape bare of surface vegetation before moving on. This adaptation also assured “prairie grass” regeneration following naturally occurring seasonal fires, events mimicked in eastern North America by its pre-Columbian human residents. They used recurrent fire to perpetuate early successional habitat for wildlife propagation, foraging, and agriculture. Today in the lower Susquehanna watershed, establishing warm-season grassland meadows requires soil prep and seeding to get things going again.

Precribed Burn — To prevent succession, warm-season grassland parcels are most commonly maintained using applications of prescribed fire every 3 to 5 years. Among their benefits, these burns invigorate native vegetation while inhibiting the invasive tendencies of many non-native plants. Well-planned periodic fire can significantly reduce fuel accumulations, particularly in tinderbox woodland tracts managed as fire-free zones for the past century or more. Many forest trees including oaks rely on sporadic fire events for regeneration.

LEARN HOW LAND MANAGERS UTILIZE PRESCRIBED FIRE

This coming Saturday, April 18, 2026, beginning at 10 A.M. (rain date April 25), the Pennsylvania Game Commission is hosting a “Prescribed Fire Festival” at Middle Creek Wildlife Management Area. Be certain to come out to the visitor’s center at 100 Museum Road, Stevens, PA, for this event. Land managers will be there to answer questions and to explain the planning and preparations involved in overseeing a prescribed burn. There will be guided walks of habitats preserved using fire of varying intensities. You’ll see the equipment and protective clothing used by certified personnel to administer a live prescribed fire burn right before your eyes. Then you can have lunch—food trucks will be available on site.

Prescribed Fire Demonstration — Visitors witness a prescribed fire demonstration at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area in 2025.

After the burn demonstration, why not go for a walk or drive around the refuge. You can take note of how grassland and early successional plant communities are responding to previous doses of prescribed fire…

Greening up a week or two after the burn, Wingstem is blooming by June in areas treated using prescribed fire earlier in the spring.

Another scene from June: Four-foot-tall Joe-Pye-Weed amidst lush growth of goldenrod in early successional habitat where a prescribed burn eliminated accumulating wildfire fuel and turned back the growth of invasive plant species in March.

And how the grassland animals respond as well…

Ring-necked Pheasants in early successional habitat maintained by the periodic application of prescribed fire.

Eastern Cottontails prosper in the mosaic of warm-season grasses and early successional thickets on lands sustained using prescribed fire. Rabbits are herbivores, primary consumers eating mostly legumes and other plants, the producers that through the process of photosynthesis convert the energy of the the sun into food energy.

Other small vegetation-eating rodents including mice and voles thrive in managed grasslands. Similar in appearance are the shrews. Though seldom noticed, Northern Short-tailed Shrews spend day and night foraging for food, even in the shallow waters of wet meadows and thickets. Unlike the aforementioned herbivorous rodents, shrews are insectivores, secondary consumers feeding mostly on primary consumers including a variety of insects and other arthropods.

Some years ago, we found this tiny Masked Shrew (Sorex cinereus) in a grassland area being preserved using prescribed fire. Like other shrews, Masked Shrews are secretive, but always on the go. They feed constantly to fuel their vast energy requirements, sometimes consuming three times their body weight in a single day. But their voracious appetite can get them into trouble, causing these incessant eaters to encounter numerous potentially infective parasites during their non-stop foraging missions.

Female American Kestrel — Patrolling the grasslands is the female American Kestrel, a secondary consumer. She’s on the lookout for primary consumers including large insects like grasshoppers or crickets. But perhaps more likely is a small rodent whose less-than-ideal vigor may cause it to slip up, creating an easy target. If she selects a careless shrew as her prey, she may be assuming the role of a tertiary consumer, eating a secondary consumer (the shrew) that fed on insects (the primary consumers) that derived their energy from photosynthesized plant matter.

The strictly nocturnal American Barn Owl (Tyto furcata), another secondary and sometimes tertiary consumer, takes the night shift, hunting unwary voles, mice, and shrews, often by sensing the sounds they make among runways in the grass. As a native predator in its favored habitat, the owl’s selection of each victim actually helps to keep the prey species’ population healthy, eliminating the weak and vulnerable to provide a qualitative service to the surviving wildlife of the grasslands. While we may not think of the barn owl as a direct consumer of insects, its positive influence on insectivorous shrew populations makes it an important functionary in maintaining balance in the ecosystems it calls home.

Male American Kestrel — As food becomes increasing plentiful in the grasslands, a female kestrel will remain mostly out of sight, performing the majority of the egg incubation duties while her colorful mate stands guard nearby.

The fate of an avian predator such as a kestrel lies at the mercy of the fate of its quarry. Because, you see, the sun’s energy, after being converted to chemical energy by photosynthesizing plants, flows upward through the trophic levels of the food chain—herbivores (primary consumers) such as rodents and insects to carnivores (secondary and tertiary consumers) including kestrels. Grasslands, when abundant and diverse, are correspondingly abundant and diverse with small mammals and insects and will therefore support thriving populations of American Kestrels and other predators. These secondary and sometimes tertiary consumers fulfill a role in cultivating healthier populations of their prey—the primary and secondary consumers in the food web—as a balanced component of a flourishing grassland ecosystem. Sparse and fragmented grasslands, on the other hand, beget negligible small mammal and insect populations, are stricken with broken food webs, and champion few if any American Kestrels or other predators. If the land it occupies is neat, tidy, manicured, exploited, or sprayed sterile and dead, the energy flow cycle of the ecosystem is dead as well. There’s nothing animal introductions, reintroductions, rescues, culling, stocking, or harvesting can do about it, because in the end, it’s all about the habitat.

April 1, 2026May 16, 2026

Migrating Waterfowl on Lotic Fresh Waters

Lotic vs. Lentic Freshwater Ecosystems — Some comparative attributes of unspoiled lotic vs. lentic freshwater ecosystems. Low-gradient (slow-moving) lotic waters often create, and remain connected to, accompanying riverine wetlands (a lentic freshwater ecosystem). These swamps, marshes, and ditches absorb, purify, and infiltrate flood waters while supporting a diverse number of plant and animal species.

We frequently perceive all waterfowl migration to be synchronized with the conspicuous movements of familiar species like Snow Geese, Tundra Swans, and Canada Geese—big flights coming south in October and November, then a return to the north in late February and March. And we’re all quite aware of the occurrence of large gatherings of some of these migrants while they make stopovers on some of our largest lentic (still) waters—the man-made lakes and reservoirs created by damming local streams. But did you know that there are populations of colorful waterfowl with dynamic migrations that extend throughout the winter and early spring with movements that are often continuous. Under favorable conditions, these birds favor the lotic (flowing) waters of the river and its larger tributaries as they transit the lower Susquehanna valley. That’s because unpolluted lotic freshwater ecosystems support a greater diversity of plants and animals than lentic waters, and therefore offer more opportunities for hungry migrating waterfowl to find food. Let’s have a look at some of the species that visit the river during their seasonal journeys…

Ice on the Susquehanna — While the urge to head south in the autumn is largely stimulated by the shortening of the photoperiod, it is the presence of ice, particularly on glacial lakes throughout the lands to the north of the lower Susquehanna River basin, that pushes many diving ducks to finally make their way south toward the guarantee of open-water feeding areas along the coast. This movement may occur at anytime between November and late February or, as we have seen during some of the mild winters of recent decades, it may scarcely be noticed at all.

Common Mergansers often lead they way when it comes to migratory diving ducks. They regularly move south in conspicuous numbers by late November and December and are regularly pushing north as soon as the ice begins to melt. During a typical year, it is not unusual for some populations of these large diving birds to remain north of us during the winter. Then, in the days after a sudden rush of frigid polar air, an appreciable increase in ice cover will force a mid-winter movement of birds down the Susquehanna.

Heat Flux Sources Impacting Temperature in Lotic Fresh Waters — Temperatures in lotic fresh waters vary over the length of the stream or river. They are largely determined by the collective impact of the numerous sources of heat flux depicted in this graphic. (Environmental Protection Agency image)

Buffleheads begin passing through the lower Susquehanna region in Novemeber on their way to coastal saltwater bays for the winter. Lingering populations feed by diving in the river’s pools and riffles for benthic invertebrates including snails and insect larvae. Lesser quantities of aquatic plant matter supplement their diet.

Many Common Goldeneyes will remain on shallow, ice-free waters of the northern lakes and rivers sculpted by the most recent glacial event, but only until they are forced south into and through the lower Susquehanna valley by the encroachment of freezing conditions. On the river, they are among the dozen or so species of diving ducks we see visiting or passing through during the typical late fall and early river.

Geomorphological Zonation of a Stream or River — During their visits to the lotic (also known as riverine) fresh waters of the Susquehanna and its largest tributaries, benthic-feeding waterfowl make short dives to take advantage of the plants, small fish, invertebrates, and other food sources inhabiting the stream bottom in the riffles and pools of the free-flowing waterway. Substrates, listed here by size (in descending order), along with other parameters influenced by this flow regime zonation determine the variety and abundance of the forage available to migrating waterfowl and other consumers. Ice or high water and poor visibility due to flooding can render the riffles and pools of the channel unusable for feeding. The birds must then choose to either linger and rest without feeding or leave the lotic freshwater habitat to seek sustenance. During a flood, this may require relocation to a nearby lentic (still) body of fresh water such as a lake or reservoir. The presence of ice will almost invariably force the birds to fly on to the Atlantic Coastal Plain and the tidal waters of its bays and estuaries.

Hooded Mergansers are one of the few species of diving ducks likely to utilize flooded shoreline timber and riverine (fluvial) wetlands as refuge from high water on Susquehanna.

Pied-billed Grebe and Canvasbacks — A Pied-billed Grebe and a pair of Canvasbacks on an ice-free stretch of the lower Susquehanna in mid-winter feed and loaf in a riffle-flanked pool where a large mat of American Eelgrass, a submerged aquatic plant also known as Tapegrass or Wild Celery, grows during the summer. The vast mosaic of riffles and pools in a river this size offers tremendous opportunities for a diverse array of aquatic species to find their niche wherein they can survive and flourish.

The presence of ice forces Buffleheads and other diving ducks to gather in turbulent open water, often below a riffle or dam. Another alternative is to continue on toward the salty bays and estuaries of the coast. High water may push these birds into the shallows among the flooded woods to feed, but they seldom utilize heavily forested riparian wetlands as a refuge due to their need for a running start to get airborne.

**Riffle and Pool Characteristics of High Gradient (A) and Low Gradient (B) Streams**. This graphic illustrates the change in deposition characteristics of a stream or river as its gradient decreases. A high-gradient stream (A) has a rapid velocity, often forms falls, and tends to carry away a high volume of all but the largest of particles of potential substrates as they erode from the surrounding landscape. On a low-gradient stream, the loss in water velocity reduces the water column’s ability to transport even the smallest of substrate particles. Deposition of this gravel, sand, silt, and clay forms lateral bars that over time create the familiar meandering path of a naturally flowing lowland stream. (National Park Service image)

Benthic Life in Lotic Freshwaters — Benthic substrates in lotic freshwater pools and riffles support an abundance of life forms ranging from colorful diatoms on rocks and cobble, to invertebrates including snails and insect larvae, to fishes like this young Channel Catfish. Free of accumulations of sediment, this river bottom not only provides habitat for a healthy fishery, it facilitates the bidirectional exchange of water between the Susquehanna and its underlying aquifer.

On the lower Susquehanna, populations of young Quillback suckers are found almost exclusively in clear, high-gradient pools.

The Harlequin Duck winters along the rocky shores and man-made jetties of the Atlantic coast. In summer, they nest on fast-moving, headwater streams well to our north. Very rare on the Susquehanna, this is the first of two individuals found during March and April of 2025. It was observed feeding in the swift waters of the high-gradient riffles and pools where the river cuts through Blue Mountain north of Harrisburg. During previous weeks, Harlequin Ducks were being seen along the coast as far south as the mouth of the Chesapeake at Cape Charles, Virginia. It’s very possible that some of these birds traveled north through the bay area and up the Susquehanna on their way north.

**Fluvial Geomorphology of a Stream**. Many of the Susquehanna’s tributaries pass through each of these three erosional zones. Along the way, they carry out the process of breaking down the mountains formed by the Allegheny orogeny, the collision of North America and Africa that created the supercontinent Pangea about 325 to 260 million years ago (during portions of both the Carboniferous and Permian periods). Today’s main stem of the lower Susquehanna passes through a transfer zone (Zone 2), carrying eroded materials to a depositional zone (Zone 3) located within the ancient Susquehanna canyon stretching from Havre de Grace, Maryland, to Norfolk Canyon on the edge of the continental shelf. Within this zone, more than a 10,000-year accumulation of post-glacial sediments lies submerged by the rising waters of the Atlantic Ocean and Chesapeake Bay. Although the present-day lower Susquehanna is largely a transfer zone, some deposition occurs along low-gradient segments of the river, particularly where its course parallels the watershed’s ridges both above and below the high-gradient rapids where its path has eroded passage through the highlands. (National Park Service image)

Although the present-day lower Susquehanna is largely a transfer zone, some deposition occurs along low-gradient segments of the river, particularly where its course parallels the watershed’s ridges both above and below the high-gradient rapids where its path continues to erode passage through the bedrock. On this 1908 map of the Susquehanna at Conewago Falls, alluvial terraces of gravel, sand, silt, and clay can readily been seen as pale areas nearly lacking brown contour lines along the shorelines and islands of the river. Deposits within most of these terraces date back to the melt period following the most recent glacial event and beyond. The delta shown at the mouth of Conewago Creek (west) includes massive volumes of material deposited by both the creek and the river. This delta is currently known as Brunner Island, much of it developed as the site of a coal/gas-fired electric generating station. Terrace deposits along the Susquehanna’s shorelines created extensive perched marshes and swamps (wooded marshes) fed by rains, high river water, small streams, and springs, the latter often seeping from the base of the rocky escarpments carved by the ancient Susquehanna and defining the present-day inland border of the floodplain. We call these sites “Alluvial Terrace Wetlands”. Few of these critical components of river morphology survive. Those not drained for farmland were obliterated by urbanization and canal, railroad, and highway construction. (United States Geological Survey base image: Middletown, PA, quadrangle, 1908)

Water Willow is a familiar emergent plant that colonizes lateral bars and other alluvial deposits in low-gradient segments of the Susquehanna.

Water Willow Roots — The fine roots of Water Willow collect sediment and absorb nutrients while creating dense cover for young fish and numerous species of invertebrates including the Virginian River Horn Snails seen here.

Prior to the nineteenth century, the low-gradient flow regime of the river both above and below the riffles at Chiques Rock (lower right on map) created prime wildlife habitat. The natural accumulations of nutrients and substrates carried into and through the lotic waterway’s pools and riffles were cycled into an ideal growing medium for extensive mats of American Eelgrass and other aquatic plants. This underwater forest hosted a seemingly endless abundance of invertebrates and fishes (both resident and migratory)—supporting a variety of consumer species including various populations of humans. But soon after the mass clearing of much of the watershed’s land for farming and lumber, the mill ponds created by dams constructed on streams to power saw and grain mills became brimful with sediments eroded from the unprotected ground. During storm events, torrents of these sediments then flowed full bore toward the Susquehanna, and began accumulating in the low-gradient segments of the river.

A mat of American Eelgrass growing in the flowing waters of the Susquehanna below Conewago Falls. Eelgrass and other submerged aquatic plants provide essential habitat for a wide variety of small fish and invertebrates while also consuming nutrients deposited in the cobble, gravel, and sand substrate of the river’s pools. Excess quantities of smaller particles of silt and clay can clog the substrate and thus inhibit the hyporheic exchange of water between the stream channel and the underlying aquifer, often diminishing the biomass and diversity of organisms inhabiting this benthic habitat. Buried in these life-choking sediments, the river bottom becomes inhospitable to growth of submerged aquatics including eelgrass.

This low-gradient stretch of the Susquehanna at Marietta flows parallel to the Chickies Quartzite “Hellam Hills” before making a sharp right turn to punch through the ridge as a series of rapids at Chiques Rock. Formerly a fully functional lotic ecosystem and a paradise for migrating waterfowl, this river segment is now impaired by accumulations of nutrient-laden sediments from agricultural and urban runoff.

Conewago Falls Flood Waters — Nutrient and sediment-loaded flood waters roar across the diabase boulders at the York Haven Dam and Conewago Falls, a high-gradient segment of the Susquehanna.

Flood Waters at Haldeman Riffles — They continue past Brunner Island (left power plant stacks) and through Haldeman Riffles and the Shocks Mills Railroad Bridge…

Flooding on the Marietta Broads — …into the stretch of river known as the “Marietta broads” along the base of the Chickies Quartzite “Hellam Hills”. The low stream gradient here produces a slower current and increased deposition of sediments.

Susquehanna Flooding at Chiques Rock — As the flood surges through the riffle and pool complex at Chiques Rock, the high stream gradient maintains a velocity in the water column sufficient to keep additional sediments in suspension until they reach the low-gradient river segment just downstream at Washington Boro, site of a naturally occurring lateral bar area known as the Conejohela Flats. These bars now lie within the man-made depositional zone known as “Lake Clarke”. Created nearly a century ago by construction of the Safe Harbor Dam, this impoundment is accumulating astounding volumes of nutrient-loaded sediments that continue to encapsulate the flats within a stream-impairing delta.

Redheads, Canvasbacks, and a Horned Grebe — Anytime from November to April on the lower Susquehanna, a group of Redheads, Canvasbacks (3 birds to right of the middle of the picture), and a Horned Grebe (lower left) is a welcome sight in a riverine pool known to have a summertime growth of American Eelgrass. Noted Dr. Herbert Beck in 1924 when describing the Canvasback, “Like all ducks, …, it stops to feed within the county (Lancaster) less frequently than formerly, principally because the vast beds of wild celery which existed earlier on broads of the Susquehanna, as at Marietta and Washington Borough, have now been almost entirely wiped out by sedimentation of culm. Prior to 1875 the four or five square miles of quiet water off Marietta were often as abundantly spread with wild fowl as the Susquehanna Flats are now. Sometimes there were as many as 500,000 ducks of various kinds on the Marietta broad at one time.”

Aythya genus ducks. — Today, seeing just dozens of Aythya genus ducks (Redheads, Canvasbacks, scaup) on the lower Susquehanna is a notable event. If they happen to be forced down by inclement weather while migrating through, you might get lucky enough to see several hundred.

During the past two centuries, as food supplies in the Susquehanna grew increasingly compromised for benthic feeders like these Lesser Scaup and other diving ducks, a change in distribution was necessary for survival.

Lesser Scaup in flight. — As individual species, Lesser Scaup and other waterfowl that fail to adapt to natural or man-made changes in their habitats and food supplies may see their overall global numbers falter.

Despite being located in the transfer zone, the lower Susquehanna has become a significant depositional zone along much of its length, mostly courtesy of the placement of sediment-trapping man-made dams.

Following construction of the mill dams and ponds on nearly every mile of the lower Susquehanna’s low-gradient tributary streams, enterprising parties moved on to the river. The first significant spans were constructed using wide timber cribs filled with large rock. They were placed to create water deep enough to allow canal boats to cross the Susquehanna at both Clark’s Ferry at the mouth of the Juniata River in Dauphin County and at Columbia/Wrightsville. These dams also diverted water into the newly excavated canals—the Pennsylvania Eastern Division Canal (completed in 1833) which followed the river’s east shore from Clark’s Ferry to Columbia, and the Susquehanna and Tidewater Canal (completed in 1840) along the west shore from Wrightsville to Havre de Grace, Maryland. Placement of these sediment-trapping man-made dams began a process of converting vast mileage of the lower Susquehanna from a transfer zone into a deposition zone. In addition, layout of the canals and locks followed the contours along the base of the riverside ridges, seriously altering most of the alluvial terrace wetlands that the river had created as a feature of its floodplain during the post-glacial period.

Construction of the canal dams was just the beginning. During the twentieth century, more massive dams would be added to the main stem of the river for hydroelectric energy production at York Haven, Safe Harbor, Holtwood, and Conowingo. Upon their completion, the days of unassisted anadromous fish migrations were over. On both the river and its tributaries, smaller dams including dangerous low-head dams maintain water levels for boating and recreation. They too create current-diminishing, pseudo-lentic waters that blanket the lotic riffle and pool substrates with polluted sediments.

MAN-MADE DAMS TURN LOTIC WATERS INTO UNFLUSHED TOILETS

Sediment Deposition Behind Lower Susquehanna Dams — The construction of dams on the lower Susquehanna has converted vast mileage of the river from a lotic freshwater system into a series of man-made lentic freshwater reservoirs. These areas have lost their function as a lotic transfer zone and are now a sort of dysfunctional series of depositional zones collecting vast volumes of sediment containing nutrients and other pollutants. Within each impoundment, the reduced velocity of the river causes it to drop suspended sand first, then the finer particles of silt and clay closer to the dam. The flow regime of riffles and pools is lost and the hyporheic zone that exchanges water between the river and the underlying aquifer is clogged. These impaired segments of river become ripe for eutrophication: algal blooms followed by die offs that can lead to a fatal reduction of dissolved oxygen in the water column.

Deposits of lateral bars of sediment in low-gradient segments of the Susquehanna can create shallow water feeding habitat for puddle/dabbling ducks like these Gadwall. Where sediment pollution is severe, benthic foods in these areas often consist mostly of invertebrates and plant matter deposited by the current, the buried substrates devoid of a functioning ecosystem and the waters subject to eutrophication.

Common Goldeneyes on a patch of open water on an otherwise ice-covered “Lake Clarke”, the impoundment created by Safe Harbor Dam. While they may find this spot advantageous for loafing, the food supply over the sediment-buried substrate will be limited.

By the end of the twentieth century, accumulations of polluted sediments behind lower Susquehanna dams were nearing capacity. There is no working plan to attenuate the massive release of these pollutants that may be triggered by a catastrophic flood. The effort to reduce nutrient and sediment runoff remains the focus so that new loading is kept to a minimum and won’t add to the capacity problems at the dams or continue downriver to the Chesapeake at full strength when the dams are full. Alleviating the sediment aggregation problem within the river’s impoundments is a tall order and a dilemma not easily solved. (United States Geological Survey image)

Fortunately for everything else living in the benthos, Common Mergansers are big enough to devour invasive, non-native Rusty Crayfish when they find them in our lotic waterways.

TIME TO CLEAN UP OUR ACT

WHERE DOES YOUR STORMWATER GO?

Channelized Urban Stream — **Channelized Urban/Suburban Streams Function as Sewers.** They have no attached lowlands or floodplains to absorb, purify, and infiltrate runoff from rain events. Pollutants including litter, pet waste, lawn chemicals, tire-wear particles, hazardous fluids, and sometimes untreated human excrement flush unchecked from the municipal storm drainage system into the waterway. Thermal shock from summer downpours washing across sun-heated pavements can kill temperature sensitive fishes and other aquatic life. Nutrient and sediment loads from these impaired tributaries later accumulate downstream in low-gradient segments of the Susquehanna, turning the river into an open-air cesspool. Aggressively working to implement projects that eliminate these sources of pollution are the only effective way to keep the problem from getting worse. Making things better requires a lot more dedication and effort. (United States Geological Survey image by Frank Ippolito)

RIPARIAN BUFFERS MAKE A DIFFERENCE…WIDER IS BETTER

FLOODPLAIN RESTORATION

Regardless OF HOW LONG YOU’VE BEEN CONDITIONED TO THINK OTHERWISE, THIS IS A DYSFUNCTIONAL, POLLUTED CREEK—AND IT NEEDS HELP

RESTORED TO ITS HISTORIC FUNCTIONS

Floodplain and Stream Restoration — On an adjacent segment of the same creek, this legacy sediment removal project restored a braided meandering channel and connected it to its newly liberated, historic floodplain. In just their second year, the fluvial wetlands are effectively absorbing stormwater and sequestering nutrients as an attached component of the stream’s riffle and pool complex. During our visit earlier this week, we found American Toads, Northern Leopard Frogs (Lithobates pipiens), and Northern Spring Peepers breeding here. It’s just as Castor canadensis would have it!

“STOP HEMMING AND HAWING AROUND ALREADY”

“HEY COWBOY, HOW ‘BOUT GETTIN’ THEM FILTHY LITTLE DOGIES OUTTA DAT CRICK?”

Impaired Stream — Here’s a polluted stream in a pasture with grazing livestock. The site is a former mill pond within which the creek eroded a channel following removal of the dam. The animals defecate and urinate where access to water is gained at a broken down embankment of the nutrient-loaded legacy sediments deposited in the pond more than a century ago. It’s a haphazard form of animal husbandry and a reminder that all it takes is just one stubborn jackass to foul up the whole waterway.

DIRECT SOURCES OF NITROGEN (AMMONIA) POLLUTION IN STREAMS

DID YOU KNOW that a dairy cow produces about 80 pounds of waste (excrement and urine) every day?

DID YOU KNOW that a horse produces about 50 pounds of waste (excrement and urine) every day?

DID YOU KNOW that a human produces about 3 to 4 pounds of waste (excrement and urine) every day? The exceptions, of course, are those who continue to insist that raising farm animals in and alongside a body of water is okey-doke—a harmless practice. These individuals tend to retain the former constituent of human waste and are thus full of it.

“ATTABOY TEX, THAT’S MORE LIKE IT!”

Stream Fencing and Livestock Crossing — Now that’s better. Legacy sediments have been removed to reconnect the stream to its floodplain. A livestock crossing and exclusion fencing has been installed, and a nutrient-consuming riparian buffer has been planted. This creek segment’s pollution woes have been mitigated. Do you have a neighbor needing this type of remedial work on their farm? Encourage them to contact your local county conservation district office or volunteer waterways restoration organization for assistance. Various programs provide financial assistance covering all or most of the costs of stream improvement as well as monetary incentives for helping to clean up the water.

AND FINALLY

WHEN IT COMES TO BUILDING DAMS ON LOTIC FRESH WATERS…

…LEAVE IT TO THE BEAVERS

North American Beavers (Castor canadensis) create habitats that connect the riffle and pool regime of a low-gradient stream to a surrounding fluvial wetland that retains sediments, cycles nutrients, and provides essential habitat for hundreds of plant and animal species. Floodplains are for flooding, and if a beaver floods an area, you can be guaranteed that it was already part of a floodplain. You see, beavers don’t encroach upon humans, it’s humans engaging in the encroachment upon beavers. (National Park Service image)

MAD HATTERS

DID YOU KNOW that even before the landscape was cleared for farms and a supply of timber, and before mill dams on local creeks began accumulating soil runoff from the consequently barren hillsides, all the North American Beavers, the keystone species of lower Susquehanna stream ecology, were killed and sold to make hats? It’s no wonder things are fubar!

COMING SOON…

Horned Grebes are regular migrants and sometimes winter residents on ice-free stretches of the lower Susquehanna. They spend their time plying the benthic substrate of the river’s clear riffles and pools for a variety of invertebrates and small fish. Look for them moving north in coming days sporting this beautiful breeding plumage.

April and early May are prime time for observing Common Loons on the Susquehanna as they undertake a journey from the Atlantic surf where they spent the winter to nesting sites on northern lakes. For this migrant in breeding plumage, clear water for sighting plenty of benthic life in the river’s riffles and pools assures a successful dive in search of energy-replenishing forage.

March 23, 2026

Photo of the Day

March 15, 2026March 15, 2026

The Next Wave

Our first big wave of northbound migrants has come and gone, but don’t despair; for spring is still a few days away, and we have yet to reach the end of the beginning …

Migrating Canada Geese — All but the last of the migratory Canada Geese have passed through the lower Susquehanna basin en route to their nesting grounds in the land for which they are named. Those that remain are mostly members of a resident population established here during the latter decades of the twentieth century.

Migrating Tundra Swans — The majority of the northwest-flying Tundra Swans that wintered on the Atlantic Coastal Plain and will nest in Alaska and western Canada have now traversed our skies and are on their way through the Eastern Great Lakes region.

Snow Goose — Snow Geese too have departed. Only stragglers like this individual with a wounded wing linger.

Migrating Ring-billed Gulls — Still moving up the river corridor are thousands of Ring-billed Gulls. Their movements are prolonged, extending through much of the late winter and into early spring with primarily adult birds leading the way.

Displaying Common Grackle — Male Common Grackles are coming north to lay claim to favorable nesting habitat too. In an effort to fend off rivals, a male will thrust back his head and ruffle his feathers to display the colorful sheen adorning his plumage.

Recognized by their nasal two-syllable calls, Fish Crows have been filtering up the valleys of the river’s main stem and its tributaries for almost a month now. While the closely related American Crow is numerous throughout the winter in the lower Susquehanna watershed, most of our Fish Crows retreat to the Atlantic Coastal Plain during the colder months.

The Fish Crows that return early stake claims on the best nest sites, often in lowland areas where they are sometimes in direct competition with a portion of the population of American Crows that commonly breeds here.

With the arrival of the spring equinox, longer periods of daylight will accompany the return of many more species. Be sure to get outside, then have a look around.

March 8, 2026March 8, 2026

Look, Up in the Sky, It’s a Bird, No, Wait, It’s a Bunch of Birds!

Step outside, take a listen, and look up. A substantial northward movement of early migrants is, at this writing, underway across the lower Susquehanna basin—primarily in the skies within five to maybe ten miles of the river corridor. Within the last hour, we witnessed a steady parade of geese, swans, and blackbirds in the skies above the headquarters garden. These are not birds that are staging or are in the midst of a stopover here during their journey north. They are instead flocks taking advantage of favorable conditions to make a big jump from Chesapeake Bay and adjacent areas of the Atlantic Coastal Plain into territory to our north and northwest that was just two weeks ago in the middle of a deep freeze. Compared to the slowpokes whose northbound movements are a bit more cautious, the shared determination of these birds to get to the nesting grounds first may be advantageous for their breeding success.

Canada Geese and Blackbirds — Below them, flocks of hundreds of blackbirds including these Common Grackles and Red-winged Blackbirds filtered through.

Tundra Swans — …because they’re gone in just moments for another year.

Winds from the southwest are forecast for the coming couple of days, so the movements could continue. Head outside, enjoy the weather, and look up.

March 2, 2026March 6, 2026

Do Chipmunks Really Eat All That Food They Stash Away in the Fall?

Eastern Chipmunks are familiar mammals of our deciduous forests and wooded neighborhoods. They are surprisingly solitary and territorial, defending their well-concealed underground excavations vigorously to protect them from would-be intruders—including other chipmunks. Their individual effort to stockpile a cache of seeds, nuts, and other non-perishable foods during the autumn will largely determine whether or not they survive winter’s worst.

Eastern Chipmunk gathering deeds in autumn — Everyone knows that Eastern Chipmunks spend much of the fall storing provisions for consumption as snacks during breaks in their winter slumbers, but did you realize that those chipmunks with the best inventory will have a pronounced advantage over less successful neighbors come late winter and spring?

Male Eastern Chipmunk — Upon exiting their subterranean shelters earlier this week, Eastern Chipmunks commenced the courting ritual which includes males like this one chasing other males away from both their dens…

Female Eastern Chipmunk — …and prospective mates like this female trying to escape notice by hiding in a hollow stump.

Eastern Chipmunks — Being well-nourished throughout the winter has adequately prepared this male (right) for the strenuous activity of securing both his territory and his mate against other males.

Eastern Chipmunks Copulating — …she becomes receptive to his advances. A well-fed pair of chipmunks can complete courtship and mating within hours of emerging from their winter abodes.

February 27, 2026February 27, 2026

It’s Snow Goose Time at Middle Creek

In case you haven’t already heard, the Snow Geese are at last filtering north from the Atlantic Coastal Plain to congregate at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area in Lancaster and Lebanon Counties.

Snow Geese at Middle Creek W.M.A. — As many as 60,000 Snow Geese have been enumerated in recent days at the refuge.

Tundra Swans at Midle Creek WMA — And more than 500 Tundra Swans are making a stopover.

Snow Geese at Willow Point — When not in the fields feeding, Snow Geese are concentrated around the small area of open water on the otherwise frozen lake, making them easily observed from the Willow Point viewing area. If you’re planning to visit and have a look, don’t wait. The urge to begin their 2,000-mile journey to breeding areas on the arctic tundra is strong, and as soon as conditions permit, they’ll be resuming their excursion. This could very well be an abbreviated stay.

February 25, 2026February 25, 2026

Start Your Own Tree Nursery

Just a reminder—there’s still time to order trees and shrubs from your local county conservation district’s annual sale, but you need to act soon…

Cumberland County Conservation District 48th Annual Tree Seedling Sale

Order by Friday, March 20, 2026

Pick Up on Thursday, April 16, 2026, or Friday, April 17, 2026

Dauphin County Conservation District Seedling Sale

Order by Monday, March 16, 2026

Pick Up on Thursday, April 23, 2026, or Friday, April 24, 2026

Franklin County Conservation District Tree Seedling Sale

Order by Tuesday, March 17, 2026

Pick Up on Thursday, April 23, 2026

Huntingdon County Conservation District Tree/Seedling Sale

Order by Friday, April 3, 2026

Pick Up on Thursday, April 16, 2026, or Friday, April 17, 2026

Lancaster County Conservation District Annual Tree Seedling Sale

Order by Friday, March 6, 2026

Pick Up on Friday, April 10, 2026

Lebanon County Conservation District ALL NATIVE! Tree & Plant Sale

Order by Monday, March 2, 2026

Pick Up on Friday, April 17, 2026

Mifflin County Conservation District Tree Sale

Order by Friday, March 6, 2026

Pick Up on Wednesday, April 15, 2026, or Thursday, April 16, 2026

Schuylkill County Conservation District/Sweet Arrow Lake Conservation Association 51st Annual Seedling Sale

Order by Saturday, March 21, 2026

Pick Up on Saturday, May 2, 2026

Snyder County Conservation District Tree Seedling Sale

Order by Monday, March 30, 2026

Pick Up on Wednesday, April 15, 2026

York County Conservation District Seedling Sale

Order by Sunday, March 15, 2026

Pick Up on Thursday, April 16, 2026

If maybe you would like to order trees but you’re not quite ready to put them in the ground, why not pot them up and start your own plant nursery. It’s a great way to build an inventory of hardy stock for planting around your own property or for use in community or civic conservation projects.

Potting Sweetgum Seedlings — Earlier this week we potted up some bare-root seedlings in a mix of compost and sand to give them a head start before planting them in the ground either during the coming fall or in years to come. These happen to be some young American Sweetgum trees we purchased from a nursery in Perkasie, Bucks County, Pennsylvania.

Native Tree Nursery — After watering them in, we added our new trees to the inventory we have available for stream buffers, rain gardens, reforestation, and other rewilding projects. We’ll either stake or trim them to remedy the curved stems.

Yellow Birch Sapling — In just a season or two, we have nice robust saplings ready to install in a project or in the headquarters garden. Starting plants in the nursery gives them the size they need to improve their chances of survival.

This is one of ten Norway Spruce trees we purchased in a pack from a local conservation district sale almost ten years ago. We started each two-year-old seedling in a pot, then transplanted them into the ground a year later. All have been thriving ever since.

Since being installed as a potted sapling reared in the headquarters nursery about fifteen years ago, this sturdy Eastern Sycamore has matured and is producing seeds relished by American Goldfinches and other birds.

Eastern Sycamore and Conifers — Growing in a streamside woodland, a really massive sycamore that got its start in your nursery could continue to provide valuable wildlife habitat over a lifespan extending one hundred, two hundred, maybe even three hundred years or more. These long-lived denizens of the floodplain provide homes for such beloved breeding birds as Baltimore Orioles, Bald Eagles, Warbling Vireos, Yellow-throated Warblers, and several species of owls,…

Great Horned Owl Nest in Eastern Sycamore — …including this Great Horned Owl, seen currently incubating eggs inside a legacy sycamore that has been an active nest site among its neighboring Norway Spruces and Eastern White Pines for decades.

Yellow-crowned Night Herons at Nest — Yellow-crowned Night Herons, an endangered species in Pennsylvania, nesting in an Eastern Sycamore planted during the early twentieth century as a street tree very near the Susquehanna in mid-town Harrisburg. Ordering and planting trees today is essential for their tomorrow.

February 14, 2026February 16, 2026

Adapting to Winter Extremes

During winter’s harshest conditions, one must frequently marvel at the methods various forms of wildlife have to survive. Take a look at some of the animals we found using their life-sustaining adaptations to find food amidst the snow-covered landscape and bitter cold air.

We watched this Blue Jay digging to retrieve an acorn from beneath the tuft of leaves and dried grasses where it had apparently concealed it earlier in the season.

Blue Jay Eating Acorn — It then carried it to a nearby limb and chiseled away the husk to devour the nutritious contents. Blue Jays are known to cache hundreds or even thousands of seeds, nuts, and acorns for winter consumption. Want to see it for yourself? Just put out some unshelled no-salt peanuts and watch the jays haul them away, that is if the hoarding squirrels don’t get them first!

Like other members of the Picidae family, Red-headed Woodpeckers pry and chisel away at decaying and insect-infested trees to find food during the winter.

First-winter Red-headed Woodpecker — Like jays, Red-headed Woodpeckers including this first-winter bird are very fond of acorns and will often collect them from the ground. And like jays, they’ll cache acorns for use as a backup supply when a blanket of snow may prevent them from gathering those that remain beneath the oaks.

Native to eastern Asia, Wintercreeper (Euonymus fortunei) has escaped from mostly urban cultivation to become naturalized and often invasive in some wooded areas of the lower Susquehanna River watershed. It can be quite aggressive, usually found growing as a climbing vine or less frequently as a shrub. In the absence of native and more palatable foods, the berries are an attractive survival fare for members of the Turidae (thrush) family and other birds.

We found this and several hundred more American Robins surviving the current winter weather while feeding on Wintercreeper in a suburban woods with a heavy growth of the invasive plant.

The Hermit Thrush, a species that nests in the coniferous and mixed forests of the northern United States and Canada, winters sparingly in the lower Susquehanna valley. We found this one among the robins and bluebirds of the Wintercreeper thicket where it too was probably attracted by the supply of berries.

Frequently escaping notice in the winter woods is the Brown Creeper, another species that nests primarily in coniferous and mixed forests to our north, though it does breed in our area at scattered locations, primarily in mountainous or swampy terrain.

The petite Winter Wren exhibits mouse-like behavior as it hops and crawls among logs, rocks, and brush along the banks of wooded waterways. Like the creeper, it is primarily an insectivore, spending much of its time escaping notice searching beneath streamside structure for its daily nourishment. In summer, Winter Wrens nest in damp coniferous forests.

Seeing a Red Fox repeatedly during daylight hours is typically considered to be a sign that one may be observing a diseased animal, but we soon determined that there was nothing at all wrong with this vulpid we found patrolling a large woodland lying along the outskirts of a lower Susquehanna city.

February 13, 2026February 14, 2026

Today’s Middle Creek Snow Goose Count: One

For those of you who may be wondering if there are Snow Geese at Middle Creek Wildlife Management Area, the answer is yes—just one!

So where are the thousands of Snow Geese we’ve grown accustomed to seeing during recent decades as they gather at the refuge in February while preparing to fly north for the summer?

With much of the river and nearly all of the lower Susquehanna basin’s lakes and ponds frozen, Snow Geese and many other migratory waterfowl remain concentrated on the Atlantic Coastal Plain where salty tidewater provides ice-free conditions for feeding and loafing. In fact, some ducks, particularly diving species, may still be evacuating freshwater locations to our north such as the Eastern Great Lakes where, during the past week, even more surface area has succumbed to freezing. Look for migrants to begin pushing north in bigger numbers as soon as rising temperatures start to melt local ice. (NOAA/U.S. National Ice Center image)

February 11, 2026February 11, 2026

To Stave Off Starvation, Nip It in the Bud

It looks like the worst of the cold may be behind us. With temperatures trending upwards during the coming days and weeks, some species of wildlife will soon find their search for food made a whole lot easier.

White-crowned Sparrows spend the winter seeking shelter in dense tangles of early successional growth, particularly among thorny shrubs and vines. Extended periods with snow cover can make their search for food a real challenge, but we spotted this one hitting the jackpot on a bare patch of sun-drenched ground along the edge of a thicket of briars. It has found the earliest green buds of a clover or other herbaceous plant. Many songbirds in the finch, cardinal, and sparrow families turn to buds in late winter and early spring as a source of energy-rich sugars.

In woodlands, sweet sap will soon swell the buds of maples, much to the delight of Eastern Gray Squirrels whose favorite dining spot will suddenly become the treetops instead of the leaf litter. Often selected by property owners for their colorful autumn foliage and fast growth, the prevalence of both native and non-native maples in the suburban landscape helps more squirrels survive the winter to expand their numbers in the spring.

Nibbling late-winter buds is a daylong pastime for the hungry White-tailed Deity. While high-intensity twig browsing can alter forest ecology and practically eliminate understory growth and regeneration, bud snacking only tends to make some of the trees more stout than others. It’s similar to the trimming and snipping a bonsai artist may do to maintain a prized specimen.

Tree Shelters Protecting Saplings — Those of you installing stream buffers or reforesting all or part of your property may not want your newly planted trees turned into bonsai or lost entirely to hungry ungulates. If you’re seeking some relief from a browsing problem, you’ve got to nip it in the bud. Tree shelters offer protection during the first half a decade or more of your sapling’s new life. You can find these tubes for sale from many online sources including some of the local county conservation districts during their annual spring tree sales (which happen to be underway right now, hint-hint). Installing these plastic protectors may not only relieve you from the frustration of worrying about your new trees, it might also excuse you from risking involvement in the sometimes heated quarrel between the worshipers of the White-tailed Deity who practice sacrifice and those who religiously oppose it.

February 5, 2026

An Intermission Between Deep Freezes

With another round of single-digit and possibly sub-zero temperatures on the way, birds and other wildlife are taking advantage of a break in the extreme conditions to re-energize. During the past day, these species were among those attracted to the food and cover provided by the habitat plantings in the susquehannawildlife.net headquarters garden…

Dark-eyed Juncos, one of our numerous species of native sparrows, are sometimes called “snowbirds”, and for good reason. They spend the majority of their time on the ground searching for seeds and are particularly noticeable when the landscape is blanketed in white.

As they mature, our plantings of junipers, hollies, Eastern Hemlock, Norway Spruce, and Eastern White Pine offer the environs needed by evergreen-loving species like this Red-breasted Nuthatch to remain as winter residents instead of just seasonal transients.

January 29, 2026

In the Frozen Foods Aisle

For overwintering birds and other animals, finding enough food is especially difficult when there’s snow on the ground. And nighttime temperatures in the single digits make critical the need to replenish energy during the daylight hours. Earlier this afternoon, we found these American Robins seizing the berry-like cones from ornamental junipers in a grocery store parking lot. It was an urgent effort in their struggle for survival.

American Robin on Juniper — Though seldom considered ideal wildlife plants, low-growing ornamental junipers often produce an abundance of seed-containing cones that resemble berries.

American Robins Eating Juniper — In a pinch, they’ll attract dozens of robins and other fruit-eating birds as a quick source of nourishment on a windy, bitter-cold day.

American Robins Feeding on Juniper "Berries" — After all, these landscape shrubs are often derived from native species including the Creeping Juniper (Juniperus horizontalis) of Canada and the northernmost United States, a plant resembling a recumbent version of the local Eastern Red Cedar (Juniperus virginiana) with which these birds are quite familiar.

County conservation district offices will soon be taking orders for their spring tree sales. Be sure to load up on plenty of the species that offer food, cover, and nesting sites for birds and other wildlife. These sales are an economical way of adding dense-growing clusters or temperature-moderating groves of evergreens to your landscape. Plus, selecting four or five shrubs for every tree you plant can help establish a shelter-providing understory or hedgerow on your refuge. Nearly all of the varieties included in these sales produce some form of wildlife food, whether it be seeds, nuts, cones, berries, or nectar. Many are host plants for butterflies too. Acquiring plants from your county conservation district is a great opportunity to reduce the amount of ground you’re mowing and thus exposing to runoff and erosion as well!

January 21, 2026January 27, 2026

Thirty Years Ago: The Ice-jam Floods of 1996

It all started rather innocently as a typically cold early January with sheets of ice covering the river and with soils on the lands in the remainder of the lower Susquehanna watershed frozen solid. Then came the “Blizzard of ’96”, blanketing much of the valley with between 20 and 30 inches of wind-driven snow.

"Blizzard of '96" — Snowfall totals for the “Blizzard of ’96”, January 6-8, 1996. (NOAA/National Weather Service image)

In affected areas of Pennsylvania, Governor Tom Ridge closed all state roads for the duration of the snowfall event. Many would remain closed for much of the following week as drifting hampered exhaustive efforts to get impassable routes open.

Just as the recovery entered its second week, a change in the weather pattern took hold as milder air and spring-like rains hastened the melt. Clogged by snow often several feet in depth, storm sewer inlets and other drainage features failed to collect the runoff. Street and urban flooding became widespread. Some buildings, particular those with large flat roofs, experienced structural damage due to any remaining snow soaking up the additional weight of the rain. As local creeks swelled to stain snowy meadows brown, attention shifted to the icy river.

On the Susquehanna, rising waters started moving ice into accumulating piles of car-sized chunks behind dams and at choke points along the river’s course. During warmer weather, stream gauges provide a depth of water reference known as a stage (measured in feet) that corresponds to a rate of flow passing the gauge site (measured in cubic feet per second). On occasions when ice and debris block river channels during winter, these readings can fluctuate wildly and the relationship between stage and flow can become dubious. When the water is running ice-free at Harrisburg for example, a gauge reading of about 11.1 feet is indicated when the river flow rate is approximately 162,000 cubic feet per second. But due to its impact on the capability of the river channels to pass water, the presence of slow-moving and jammed ice can cause rapid and sometimes unpredictable variations in gauge readings, even when the flow rate remains steady. Impaired by an ice jam at the gauging station or just downriver, a flow of 162,000 cubic feet per second could lead to a stage measurement significantly higher than 11.1 feet, and the area of adjacent floodplain inundated by rising waters will increase to a corresponding degree. Conversely, a jam upstream of the gauging station could cause the reading to drop below 11.1 feet—at least temporarily—then look out, a dangerous surge could be forthcoming!

At Harrisburg, the ice jam behind the Dock Street Dam in January, 1996, caused devastating flooding in the city’s Shipoke neighborhood, on City Island, and in low-lying areas along the river’s west shore.

USGS Gauging Station at Harrisburg — Just upriver from the Dock Street Dam at Harrisburg, the Susquehanna at the United States Geological Survey gauging station on the east side of City Island crested at 25.08 feet on January 20, 1996. As the rising waters finally began heaving the jammed ice up and over the dam, the scouring energy of the departing glacier-like mounds would carry away two spans of the truss-construction Walnut Street Bridge on the west side of the island, and eventually collapse a third. After receding during the remainder of the day and through the night, yet another aggregation of chunked ice would cause the river to crest again on January 21st, this time at 24.66 feet. Under free-flowing conditions, these readings would correspond to a river flow of just under 600,000 cubic feet per second; that’s more than 250 million gallons per minute!

Downriver at Conewago Falls, ice jams behind the York Haven Dam and at several choke points within the riverine archipelago that extends from Haldeman Island to Haldeman Riffles in Lancaster County would be relieved as the river crested there during the afternoon of January 21st. The following photograph accurately relates the scene, minus the stench of mammalian feces and petroleum emanating from the polluted water of course.

Our friends (L to R) Rick Morton, Bob Zeager, Dan Seitz and his daughter Sierra witness the ice-choked Susquehanna River cresting below Conewago Falls on January 21, 1996. Bob awoke the previous morning inside his mobile home (left rear) to discover he was rapidly being surrounded by fast-rising water. With the ice-covered road now submerged in over three feet of river, he had no hope of getting out with his van, so he started removing clothing and a few other essentials by canoe. Dan soon arrived to move a few of his things to higher ground and de-energize both properties. We did our best to lend a hand, then we all got out of the floodplain. A short time after we had departed, Rick arrived and became concerned when he saw Bob’s van, now in almost four feet of water alongside his home. Rick parked his conveyance, also a van, on a rise in the lowland terrain, then made his way back the old Pennsylvania Canal towpath to make sure Bob wasn’t stranded. After concluding that he had already evacuated, Rick returned via the towpath to his van. He then discovered that yet another surge of quickly rising water had inundated the ice-covered section of access road by which he had arrived, so he was forced to wade out, leaving his vehicle behind. A short time later, he reached us by telephone proclaiming, “I’m out!”, and filled us in on the details. As the flood receded, Rick was able to retrieve his van on January 22nd. Bob’s van was totally destroyed and both his and Dan’s mobile homes were left uninhabitable. Though by no means their first experience with river flooding, this event proved particularly costly and stressful for Bob and Dan…the effects life-altering. The moral of the story: floodplains are for flooding, not filling, not building, and certainly not dwelling. Floodplains happen to make outstanding wildlife preserves, and that’s what this water-purifying alluvial terrace wetland is today. (Vintage 35 mm image)

January 7, 2026January 8, 2026

The Variety of Medium and Large-size Gulls Visiting for the Winter

You may find this hard to believe, but during the colder months in the lower Susquehanna valley, gulls aren’t as numerous as they used to be. In the years since their heyday in the late twentieth century, many of these birds have chosen to congregate in other areas of the Mid-Atlantic region where the foods they crave are more readily available.

As you may have guessed, the population boom of the 1980s and 1990s was largely predicated on human activities. These four factors were particularly beneficial for wintering gulls…

- 1. Disposal of food-bearing waste in open landfills
  2. High-intensity agriculture with disc plowing
  3. Gizzard Shad population boom in nutrient-impaired river/Hydroelectric power generation
  4. Fumbled Fast Food

Earthworms lifted to the soil's surface by plowing attracted Ring-billed Gulls by the thousands to Susquehanna Valley farmlands during the late twentieth century. — Earthworms lifted to the soil’s surface by plowing attracted Ring-billed Gulls by the thousands to Susquehanna Valley farmlands during the late twentieth century. (Vintage 35 mm image)

Young Gizzard Shad — Filter-feeding Gizzard Shad populations thrive in nutrient-rich waters like the Susquehanna. Their rambunctious feeding style stirs up benthic sediment deposits to release more nutrients into the water column and promote the algal blooms that often lead to detrimental eutrophic conditions. Decades ago, hundreds and sometimes thousands of gulls gathered below the river’s hydroelectric dams to feed on the seemingly endless supply of small Gizzard Shad disoriented by their passage through the turbines during electric generation. (Vintage 35 mm image)

So what happened? Why are wintering gulls going elsewhere and no longer concentrating on the Susquehanna? Well, let’s look at what has changed with our four man-made factors…

1. 1. A larger percentage of the lower Susquehanna basin’s household and food industry waste is now incinerated/Landfills practice “cover as you go” waste burial.
  2. Implementation of “no-till” farming has practically eliminated availability of earthworms and other sub-surface foods for gulls.
  3. The population explosion of invasive Asiatic Clams has reduced the Gizzard Shad’s relative abundance and biomass among filter feeders.
  4. Hold on tight! Fast food has become too expensive to waste.

The non-native population of Asiatic Clams in the Susquehanna and most of its larger tributaries exploded during the last two decades of the twentieth century. Its present-day biomass in the river is exceeded by no other macroinvertebrate species. The share of plankton and other tiny foods that the Asiatic Clam harvests from the water column is no longer available to native filter-feeders including Gizzard Shad. Hence, Gizzard Shad biomass has been reduced and far fewer are available to attract amazingly enormous flocks of hungry gulls to hydroelectric dams. (Vintage 35 mm image)

GULLS THIS WINTER

Despite larid abundance on the lower Susquehanna not being the spectacle it was during the man-made boom days, an observer can still find a variety of medium and large-sized gulls wintering in the region. We ventured out to catch a glimpse of some of the species being seen both within the watershed and very nearby.

Ring-billed Gulls on Lake Clarke — By far, most of the gulls you’ll encounter in the Lower Susquehanna River Watershed right now are Ring-billed Gulls. Some of them still drop by at the business end of the drive-thru lane looking for a lost order of fries and maybe a cheeseburger in paradise.

Ring-billed Gulls in winter plumage. — First-winter (left), second-winter (center), and adult Ring-billed Gulls on the Susquehanna.

Ring-billed and American Herring Gulls — Nearly every flock of gulls found in our area right now is composed exclusively of Ring-billed Gulls. The trick to finding other species, particularly rarities, is to look for slightly larger birds mixed among them, particularly American Herring Gulls like the first-winter (back row left) and third-winter (back row right) birds seen here. Herring and other species of similar-sized gulls seem to prefer each other’s company while on the wintering grounds.

American Herring Gulls with Ring-billed Gulls — Five American Herring Gulls with smaller Ring-billed Gulls. The bird to the upper left and the bird hunkered down to its right are adult American Herring Gulls, the three brownish birds are in their first winter.

Non-adult American Herring Gulls in flight. Always look for birds with all-pale wing tips when encountering herring gulls in flight.

Four Species of Medium-sized Gulls — Midway in size between the Ring-billed Gulls in the foreground and the first-winter American Herring Gulls in the upper left and middle right of this image is a Lesser Black-backed Gull (dark-mantled bird resting at center). The similar-sized bird in the water behind it is a first-winter Iceland Gull (Larus glaucoides), a rare visitor from the arctic.

First-winter Iceland Gull — Another look at the first-winter Iceland Gull from the previous image. Did you notice the all-white primary feathers and compare them to the dark wing tips on the Ring-billed Gulls seen here in its company?

Iceland Gull Bathing — The conspicuously pale wings of the first-winter Iceland Gull, seen here bathing in the presence of two first-winter American Herring Gulls.

Lesser Black-backed Gulls like this one observed at Blue Marsh Lake are uncommon in the Lower Susquehanna River Watershed. They are progressively more likely as you travel east toward the aforementioned Bucks County landfills where hundreds of these birds make up a core Atlantic seaboard population.

Great-black-backed Gull with Ring-billed Gulls — Great Black-backed Gulls are the most frequently encountered large gull on the Susquehanna. They’re easily identified by their enormous size and, as adults, their dark mantle.

First-winter Glaucous Gull — Rivaling the Great Black-backed Gull in size is the Glaucous Gull (Larus hyperboreus), another rare arctic visitor with pale wing tips. There are numerous reports of these unusual winter visitors from sites on Delaware Bay north to the Tullytown landfills on Delaware River where half a dozen or more have been occurring. Seen near the mouth of the Susquehanna on Chesapeake Bay at North East, Maryland, has been a first-winter bird similar to this one that we photographed at Blue Marsh Lake in Berks County, Pennsylvania.

For us, seeing a Glaucous Gull brought back memories of the last time we saw the species. It was forty-five years ago on New Year’s Day 1981 that we discovered two first-winter birds feeding on Gizzard Shad in open water on an otherwise ice-choked Susquehanna below the York Haven Dam powerhouse at Conewago Falls. Hey Doc Robert, do you remember that day?

December 21, 2025December 21, 2025

Photo of the Day

The first day of winter is the darkest day of the year. In the midst of the cold weather that accompanies it, one hardly thinks of looking for insect activity. But in subterranean environs where the temperature remains relatively constant, life goes on. We recently discovered this handsome Camel Cricket while performing some maintenance in our below-grade utility entrance. Worldwide in distribution, Camel Crickets, also known as Cave Crickets, Sand Treaders, and Spider Crickets, include hundreds of species in the family Rhaphidophoridae. Most are omnivorous residents of caves, mines, cellars, hollow logs, wood piles, wells, and other dark, temperature-stable situations where they feel their way around with their antennae while searching for food. Camel Crickets usually escape notice, keeping only nocturnal hours when venturing out from light-deprived cover and lacking the organs to produce the familiar “chirps” that call attention to other crickets during the breeding season.

December 10, 2025December 11, 2025

Observing Susquehanna Bald Eagles: A Chart for Determining Age

As part of an update to our “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” page, we’ve just added this chart for determining the age of the Bald Eagles you might observe on the lower Susquehanna River and elsewhere in coming weeks.

Bald Eagle age classes in late autumn in the Lower Susquehanna River Watershed. — Bald Eagle age classes as they typically appear during late autumn in the Lower Susquehanna River Watershed. (Click the image for a full-size PDF version of the chart)

Bald Eagles in each age class often retain their fall appearance through much of the winter. However, beginning January 1st, each bird is reclassified into the next in the series of chronological plumage designations. Consequently, during the early part of the new year until a new generation of eagles is hatched in late winter and spring, there are no birds in our area designated “hatch-year/juvenile”. After fledging, these youngest eagles, the new generation of juveniles, often show little change in appearance until after their first birthday, by which time they are already classified as second-year/Basic I immature birds. For birds other than the new generation of hatch-year/juvenile eagles, the majority of the molt that produces their new autumnal appearance each year occurs during spring and especially summer, when food is abundant and the bird’s energy needs for purposes other than growing feathers are at a minimum. Hence, by the time fall migration rolls around, the next in the successive progression of plumage changes is evident. For more details and year-round images of Bald Eagles, check out our “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” page by clicking the tab at the top of this page.

December 8, 2025

Cuisine to Crow About!

Red-shouldered Hawk and thigh of frog (cuisses de grenouille). — Our VIP guest, a discerning Red-shouldered Hawk, dining yet again on cuisses de grenouille (thigh of frog) at susquehannawildlife.net headquarters.

December 6, 2025December 6, 2025

Eagle Time At Conowingo Dam’s Fisherman’s Park

Wintering Bald Eagles are again congregating on the lower Susquehanna River, particularly in the area of Conowingo Dam near Rising Sun, Maryland. To catch a glimpse of the action earlier this week, we took a drive on U.S. Route 1 atop Conowingo’s impounding structure to reach Fisherman’s Park on the river’s west shore below the powerhouse.

Eagle Watching at Conowingo Dam — Scores of dedicated eagle watchers and photographers brave the raw weather to see and document the concentration of eagles that gather to feed and roost in the vicinity of Exelon Energy’s Fisherman’s Park.

Susquehanna at Conowingo Dam — The panoramic view of the Susquehanna from Fisherman’s Park offers excellent opportunities to witness Bald Eagle activity.

Immature Bald Eagles — When you arrive, it’s not unusual to hear the sounds of squabbling eagles immediately upon exiting the shelter of your vehicle. During our visit, we sighted probably 60 to 80 individuals of various age classes among the rocks and trees along the river shorelines below the dam.

Second-year (Basic I) Bald Eagle — Soon enough, we experienced a close fly-by from this second-year Bald Eagle.

Hatch-year (Juvenile) Bald Eagle — We were a little bit concerned to see only one hatch-year (juvenile) Bald Eagle among the birds at Fisherman’s Park. Perhaps the aggressive behavior of the large number of older and more experienced eagles in the area has these first-year individuals shying away. We discerned no third-year birds either, though they may certainly have been present.

Fourth-year (Basic III) Bald Eagle — A probable fourth-year Bald Eagle shows a white head with the remains of a dark line through the eye, a trait often more conspicuous in third-year birds often known as “osprey face” eagles.

Fifth-year (Basic IV) Bald Eagle — This probable fifth-year Bald Eagle has nearly lost the dark markings on the head and tail that differentiate immature birds from adults. A molt during the coming year will yield adult plumage and mark the completion of this bird’s sexual maturity.

Eagle Photographers at Fisherman's Park — Meanwhile, a little action gets the shutters clicking,…

Eagle Dogfight in the Incipient Stage — …a fourth-year Bald Eagle (top) is drawing the ire of an adult bird,…

Eagle Dogfight — …necessitating a reprisal for the taunting behavior.

To the delight of photographers at Conowingo, some of the eagles can be seen grabbing fish, mostly Gizzard Shad, from the tailrace area of the river below the powerhouse. But Bald Eagles are opportunistic feeders, and their feeding habits are similar to those of numerous other birds found in the vicinity of the dam at this time of year—they’re scavengers. Here’s a glimpse of some of the other scavengers found in the midst of this Bald Eagle realm…

Fish Crows are recognized by their nasal call. They’ll eat almost anything they can find including garbage, fish remains, discarded bait, lunch scraps, road kill, and more.

Visitors to Fisherman’s Park are warned to keep out of sight any food they may have stored in their cars. Black Vultures are known to peel rubber away from windows as they search for something to eat, a habit they possibly learned during productive forays to landfills where the edges of rubber coverings sometimes hide a freshly dumped buffet of potential sustenance.

During our stop at Conowingo Dam earlier this week, we saw only one Turkey Vulture, though more are certainly in the vicinity feeding on road kill and other carrion.

Like eagles, Ring-billed Gulls are opportunistic feeders, seen here looking for disoriented Gizzard Shad and other fish,…

Ring-billed Gull — …then quickly changing focus to check the humans along the shoreline for discarded bait or fumbled snack foods.

Juvenile Ring-billed Gull — Even young Ring-billed Gulls learn the value of watching people for activities that provide an opportunity to scavenge food.

While Ring-billed Gulls and other scavengers aren’t particularly fussy about what they eat, Double-crested Cormorants are;…

Double-crested Cormorant — …they’re targeting Gizzard Shad and other fish in the waters below the dam. Thus, we would categorize cormorants as predators, eating mollusks and other aquatic organisms as well.

Bald Eagles at Heron Rookery — But if perhaps winter isn’t your thing, don’t despair. These Bald Eagles came upon last year’s Great Blue Heron rookery on the island below the dam and it seems to be giving them some ideas. If you think like an eagle, spring is just weeks away!

November 28, 2025