Do you recall our “Photo of the Day” from seven months ago…
Here’s something to look forward to in the new year. The good citizens of East Donegal Township in Lancaster County have partnered with Alliance for the Chesapeake Bay to establish an extensive wildflower meadow on what had been a mowed field of turf grass at Riverside Park in the Susquehanna floodplain near Marietta. As the photo shows, the lawn plants have been eliminated in preparation for seeding with a diverse assortment of native grasses and wildflowers to provide habitat for birds and pollinators including butterflies, bees, and other insects. Once established, the meadow’s extensive vegetative growth will help reduce stormwater runoff by better infiltrating rainfall to recharge the aquifer. During flood events, the plantings will provide soil stabilization and increase the ability of the acreage to uptake nutrients, thus reducing the negative impact of major storms on the quality of water in the river and in Chesapeake Bay. Check the project’s progress by stopping by from time to time in 2024!
Well, here’s what that site looks like today…
The wildflowers, thousands of them, are now in bloom!Black-eyed Susan and Blue Vervain (Verbena hastata) dominate the assortment currently in flower.
And there are pollinating insects galore, most notably butterflies…
A Cabbage White collecting nectar on Blue Vervain.A Clouded or Orange Sulphur among the grasses in the meadow.A Silver-spotted Skipper.The Least Skipper is our tiniest butterfly.The Little Glassywing (Vernia verna), this one feeding on vervain nectar, deposits its eggs on Purpletop grass, which then functions as the host plant for this butterfly’s larvae.A Summer Azure (Celastrina neglecta) feeding on the nectar of a Black-eyed Susan.A Common Buckeye on a “Gloriosa Daisy”, a showy, large-flowered cultivar of Black-eyed Susan.A Red-winged Blackbird with a caterpillar found among the meadow’s lush growth.An Eastern Tiger Swallowtail on Blue Vervain. Nearby Yellow (Tulip) Poplars and other trees serve as host plants for this butterfly’s larvae.The black morph of the Eastern Tiger Swallowtail shows subdued shading in the wings that closely resembles the brilliant color patterns of the more familiar yellow form.The Sachem, this one visiting a Black-eyed Susan, is a variable species with a range that normally lies south of the 40th parallel, the line of latitude that intersects the Susquehanna in the area of the Conejohela Flats at Washington Boro.A Sachem visiting the blooms of Oxeye sunflower. During recent weeks, scorching winds from the south and southwest have transported an abundance of these vagrant skippers into the lower Susquehanna valley and beyond.A male Sachem approaching the bloom of a “Gloriosa Daisy”. Miles north of the 40th parallel, wandering Sachems are currently the most numerous of the butterflies at the Riverfront Park wildflower meadow.Here and there among the meadow’s plantings we noticed one of our favorites starting to flower, the Partridge Pea.Partridge Pea happens to be a host plant for another vagrant from the south, the big, lime-yellow Cloudless Sulphur. We saw at least half a dozen patrolling the meadow.The stars of the show are the Zebra Swallowtails, gorgeous butterflies that rely on stands of native Common Paw-paw trees in the river floodplain to host their eggs and larvae.The red-white-and-blue underside of a Zebra Swallowtail.WOW!
Why on earth would anyone waste their time, energy, and money mowing grass when they could have this? Won’t you please consider committing graminicide this fall? That’s right, kill that lawn—at least the majority of it. Then visit the Ernst Seed website, buy some “Native Northeast Wildlflower Mix” and/or other blends, and get your meadow planted in time for the 2025 growing season. Just think of all the new kinds of native plants and animals you’ll be seeing. It could change your life as well as theirs.
A Snowberry Clearwing (Hemaris diffinis), more commonly known as a hummingbird moth, visits the flowers of Blue Vervain in the Riverfront Park wildflower meadow.
The bright-red flower buds of the Ninebark (Physocarpus opulifolius) precede clusters of white blooms that will, in coming weeks, attract a variety of butterflies and other pollinators to this indigenous shrub. Its peeling bark and colorful deciduous leaves attract interest throughout the year. In the lower Susquehanna watershed, Ninebark is most frequently found growing along stream banks. It will often thrive on steep slopes with moist soils, so is useful as an erosion control species as well. To add it to your refuge’s landscape, look for it at nurseries that stock native plants. Once there, you’ll find a variety of cultivars that are sure to satisfy even the fussiest of gardeners.
During the past week, we’ve been exploring wooded slopes around the lower Susquehanna region in search of recently arrived Neotropical birds—particularly those migrants that are singing on breeding territories and will stay to nest. Coincidentally, we noticed a good diversity of species in areas where tent caterpillar nests were apparent.
The conspicuous nest of the Eastern Tent Caterpillar (Malacosoma americanum), a native species of moth. The first instar of the larval caterpillars hatch in early spring from egg masses laid on the limbs of the host tree by an adult female moth during the previous spring. Soon after they begin feeding on the host tree’s first tender shoots, these tiny, seldom-noticed larvae start communal construction of a silk tent to act as a shelter and greenhouse-like solar collector that will both provide protection from the elements and expedite their growth.The familiar last instar of the Eastern Tent Caterpillar is the most consumptive stage of the animal’s life. After feeding in the treetops, they will descend to the ground and seek a sheltered location to pupate. Adult moths emerge in several weeks to take to the air, mate, and produce eggs to be deposited on a host tree for hatching next year. The favorite host tree in forests of the Lower Susquehanna River Watershed: native Black Cherry.
Here’s a sample of the variety of Neotropical migrants we found in areas impacted by Eastern Tent Caterpillars. All are arboreal insectivores, birds that feed among the foliage of trees and shrubs searching mostly for insects, their larvae, and their eggs.
The Yellow-throated Vireo nests, feeds, and spends the majority of its time feeding among canopy foliage.The Eastern Wood-Pewee is a flycatcher found in mature woodlands. It feeds not only among the limbs and leaves, but is an aerial predator as well.The Northern Parula nests in mature forests along rivers and on mountainsides, particularly where mature trees are draped with thick vines.The Hooded Warbler (Setophaga citrina) is found among thick understory growth on forested slopes.The Ovenbird builds a domed, oven-like nest on the ground and forages in the canopy.The Kentucky Warbler (Geothlypis formosa) nests in woodland undergrowth, often near steep, forested slopes.The Worm-eating Warbler (Helmitheros vermivorum) nests among woody understory growth on forested hillsides.The Scarlet Tanager is often difficult to observe because of its affinity for the canopy of mature forest trees.
In the locations where these photographs were taken, ground-feeding birds, including those species that would normally be common in these habitats, were absent. There were no Gray Catbirds, Carolina Wrens, American Robins or other thrushes seen or heard. One might infer that the arboreal insectivorous birds chose to establish nesting territories where they did largely due to the presence of an abundance of tent caterpillars as a potential food source for their young. That could very well be true—but consider timing.
Already Gone- By the time Neotropical migrants arrive in our area, the larval stages of the Eastern Tent Caterpillar’s life cycle are already coming to an end. The nests that these native insects constructed to capture the energy of the springtime sun have allowed the larvae to exit and browse foliage when conditions were suitable, then return for shelter when they were not. While inside, the larvae could move among the chambers of their structure to find locations with a temperature that best suited their needs. Therein the solar heating and communal warmth sped up digestion and growth.Eastern Tent Caterpillar larvae are now in their bristly final-instar stage and the majority have already moved to the ground to each seek a place to pupate and metamorphose into an adult moth. Arboreal Neotropical birds have scarcely had a chance to feed upon them, and ground-feeding species seem to lack any temptation. As for the adult moths, they fly only at night and live for just one day, offering little in the way of food for aerial, arboreal, or ground-feeding birds.Having left arboreal environs, Eastern Tent Caterpillar larvae are now food for ground-feeding birds like our resident Wild Turkeys. They need only get past the bristly hairs on the caterpillar’s back and the foul taste that may result from its limited diet of cyanogenic Black Cherry leaves.The arboreal Yellow-billed Cuckoo (seen here) and its close relative the Black-billed Cuckoo (Coccyzus erythropthalmus) are the two species of birds in our area known to regularly feed on bristly tent caterpillars. But having just arrived from the tropics to nest, they’ll need to rely on other insects and their larvae as sources of food for their young.Final-instar Eastern Tent Caterpillars often defoliate Black Cherry trees before moving to the ground to pupate. Their timing allows them to feed on the fresh foliage while it is still young and tender, and to largely avoid becoming food for the waves of Neotropical birds that arrive in the lower Susquehanna basin in May.
So why do we find this admirable variety of Neotropical bird species nesting in locations with tent caterpillars? Perhaps it’s a matter of suitable topography, an appropriate variety of native trees and shrubs, and an attractive opening in the forest.
An American Redstart singing in a Black Cherry. Unlike others in the vicinity, this tree nestled among several very large Eastern White Pines showed no signs of tent caterpillar activity. It may be that for one reason or another, no adult female moth deposited her eggs on this particular tree. During our visits, Black Cherry was but one of the diverse variety of native trees and shrubs found growing on the sloping topography that created attractive habitat for the nesting birds we found. We happened to notice that a majority, but not all, of those Black Cherry trees were impacted by Eastern Tent Caterpillars.The end of the Eastern Tent Caterpillar’s larval surge may spell the end of their nests for the year, but it’s not the end for the Black Cherry and other host trees in the Prunus (cherry) and Malus (apple) genera. Because it’s still early in the season, they have plenty of time to re-leaf and many will still flower and produce fruit. Those flowers and foliage will attract numerous other insects (including pollinators) that benefit breeding birds.The Blue-winged Warbler inhabits shrubby breaks in the forest such as this utility right-of-way where Black Cherry trees have sprouted after their seeds arrived in waste deposited by fruit-eating (frugivorous) birds. Already attractive to a variety of insectivores, these openings soon lure egg-laying Eastern Tent Caterpillar moths to the cherry trees growing therein. Even in dense forest, a small clearing created by a cluster of dead trees makes good bird habitat and will sooner or later be visited by fruit-eating species that will inadvertently sow seeds of Black Cherry, starting yet another stand of host trees for Eastern Tent Caterpillars. It’s the gap in the forest that often attracts the birds, some of which plant the host trees, which sometimes entice Eastern Tent Caterpillar moths to lay their eggs.Adapt and Reuse- A Red-eyed Vireo visits an Eastern Tent Caterpillar nest……and ignores the few remaining occupants that could easily be seized to instead collect silk to reinforce its own nest.
Here in the Lower Susquehanna River Watershed, the presence of Eastern Tent Caterpillar nests can often be an indicator of a woodland opening, natural or man-made, that is being reforested by Black Cherry and other plants which improve the botanical richness of the site. For numerous migratory Neotropical species seeking favorable places to nest and raise young, these regenerative areas and the forests surrounding them can be ideal habitat. For us, they can be great places to see and hear colorful birds.
Our Lucky Break- This Scarlet Tanager descended from the treetops to feed on spiders in a small forest clearing.
The rain and clouds have at last departed. With blue skies and sunshine to remind us just how wonderful a spring afternoon can be, we took a stroll at Memorial Lake State Park in Lebanon County, Pennsylvania, to look for some migratory birds.
Though running just a few days later than usual, Indigo Buntings have arrived to begin nesting.This Common Loon dropped by Memorial Lake during a storm several days ago and decided to stay awhile. It’s a species that winters in oceanic waters along the Atlantic seaboard and nests on glacial lakes to our north.Because of the low level of turbidity in Memorial Lake, visibility is good enough to allow this benthic feeder an opportunity to see food before expending energy to dive down and retrieve it. Favorable foraging conditions might be part of the reason this bird is hanging around.Clear Water- Memorial Lake is one of the few man-made lakes in the Lower Susquehanna River Watershed to be appropriately vegetated with an abundance of submerged, floating, and emergent plants. As a result, the water from Indiantown Run that passes through the impoundment is minimally impacted by nutrient loads and the algal blooms they can cause. Buffers of woody and herbaceous growth along the lake’s shorelines provide additional nutrient sequestering and help prevent soil erosion and siltation.The breeding season has begun for Neotropical migrants including this Baltimore Oriole, which we found defending a nesting territory in a stand of Black Walnut trees.Along the edge of the lake, this Orchard Oriole and its mate were in yet another stand of tall walnut trees.Early in the season and early in the day, we started seeing Common Nighthawks flying above wooded areas north of the lake at 4 o’clock this afternoon. After all the raw and inclement weather they’ve experienced in recent days, the warm afternoon was probably their first opportunity to feed on flying insects in quite a while.Early birds, Common Nighthawks feeding at 4 P.M.
What? You thought we were gonna drop in on Maryland’s largest city for a couple of ball games and some oysters, clams, and crab cakes—not likely.
In recent days, the peak northbound push of migratory birds that includes the majority of our colorful Neotropical species has been slowed to a trickle by the presence of rain, fog, and low overcast throughout the Mid-Atlantic States. Following sunset last evening, the nocturnal flight resumed—only to be grounded this morning during the pre-dawn hours by the west-to-east passage of a fast-moving line of strong thundershowers. The NOAA/National Weather Service images that follow show the thunderstorms as well as returns created by thousands of migrating birds as they pass through the Doppler Radar coverage areas that surround the lower Susquehanna valley.
Sterling, Virginia, Doppler Radar west of Washington, D.C., at 4:00 A.M. E.D.T. indicates a dense flight of northbound migrating birds located just to the south of the approaching line of rain and thunderstorms over the State College, Pennsylvania, radar coverage area. (NOAA/National Weather Service image)More northbound birds are indicated at 4 A.M. by the radar station located at Dover, Delaware… (NOAA/National Weather Service image)…and by the Mount Holly, New Jersey, radar site. (NOAA/National Weather Service image)Many of the migrating birds shown here over the Binghamton, New York, radar station at 4 A.M. probably overflew the lower Susquehanna region earlier in the night. (NOAA/National Weather Service image)And these birds over Albany, New York’s, radar station at 4 A.M. are mostly migrants that passed north over New Jersey and easternmost Pennsylvania last evening and during the wee hours of this morning. (NOAA/National Weather Service image)
Just after 4 A.M., flashes of lightning in rapid succession repeatedly illuminated the sky over susquehannawildlife.net headquarters. Despite the rumbles of thunder and the din of noises typical for our urban setting, the call notes of nocturnal migrants could be heard as these birds descended in search of a suitable place to make landfall and seek shelter from the storm. At least one Wood Thrush and a Swainson’s Thrush (Catharus ustulatus) were in the mix of species passing overhead. A short time later at daybreak, a Great Crested Flycatcher was heard calling from a stand of nearby trees and a White-crowned Sparrow was seen in the garden searching for food. None of these aforementioned birds is regular here at our little oasis, so it appears that a significant and abrupt fallout has occurred.
A White-crowned Sparrow in the headquarters garden at daybreak. It’s the first visit by this species in a decade or more.
Looks like a good day to take the camera for a walk. Away we go!
Along woodland edges, in thickets, and in gardens, Gray Catbirds were everywhere today. We heard and/or saw hundreds of them.During our travels, American Redstarts were the most frequently encountered warbler. Look for them in low-lying forested habitats.Many early-arriving Baltimore Orioles have already begun building nests. But widespread territorial fighting today may be an indication that some latecomer orioles became trespassers after dropping in on existing territories during the morning fallout.Red-eyed Vireos are difficult to see but easily heard in forested areas throughout the Lower Susquehanna River Watershed.If the oriole isn’t the showiest of the Neotropical migrants, then the Scarlet Tanager is certainly a contender…Listen for their burry, robin-like song in the treetops of mature upland forests.No woodland chorus is complete without the flute-like harmony of the Wood Thrush. Look and listen for them in rich forests with dense understory vegetation.The Eastern Wood-Pewee, another forest denizen, has an easy song to learn…a series of ascending “pee-a-wee” phrases interspersed with an occasional descending “pee-urr”. It was one of the few flycatchers we found today, but more are certainly on the way. Their numbers should peak in coming days.Yellow-rumped Warblers can be especially numerous during migration but tend to peak prior to the arrival of the bulk of the Neotropical species. This was the only “yellow-rump” we encountered today. The majority have already passed through on their way to breeding grounds to our north.If today you were to visit a streamside thicket or any type of early successional habitat, you would probably find this perky little warbler there, the Common Yellowthroat.The Yellow Warbler likes streamside thickets too. You can also find them along lakes, ponds, and wetlands, especially among shrubby willows and alders.While nowhere near the headquarters garden, we ran into another White-crowned Sparrow in less-than-ideal habitat. This one was in a row of trees in a paved parking lot.Not all songbirds migrate at night. The Bobolink is an example of a diurnal (day-flying) migrant. They’re currently arriving in hay fields that are spared the mower until after nesting season.While looking for Neotropical species and other late-season migrants, we also found numerous early arrivals that had already begun their breeding cycles. We discovered this Blue-gray Gnatcatcher on its nest in a Black Walnut tree……then, later in the day, we found this one in its nest, again in a Black Walnut tree. Note the freshly emerging set of leaves and flower clusters. With many tree species already adorned in a full set of foliage, open canopies in stands of walnuts we found growing in reforested areas seemed to be good places to see lots of migrants and other birds today. It’s hard to say whether birds were more numerous in these sections of woods or were just easier to observe among the sparse leaf cover. In either case, the nut-burying squirrels that planted these groves did us and the birds a favor.
There’s obviously more spring migration to come, so do make an effort to visit an array of habitats during the coming weeks to see and hear the wide variety of birds, including the spectacular Neotropical species, that visit the Lower Susquehanna River Watershed each May. You won’t regret it!
Wood Ducks arrived in February and March to breed in the lower Susquehanna valley. Soon after hatching in April or May, the young leave the nest cavity to travel under the watchful gaze of their ever-vigilant mother as they search for food along our local waterways. If you’re fortunate, you might catch a glimpse of a brood and hen while you’re out looking at the more than one hundred species of birds that occur in our region during the first half of May. Good luck!
Homo sapiens owes much of its success as a species to an acquired knowledge of how to make, control, and utilize fire. Using fire to convert the energy stored in combustible materials into light and heat has enabled humankind to expand its range throughout the globe. Indeed, humans in their furless incomplete mammalian state may have never been able to expand their populations outside of tropical latitudes without mastery of fire. It is fire that has enabled man to exploit more of the earth’s resources than any other species. From cooking otherwise unpalatable foods to powering the modern industrial society, fire has set man apart from the rest of the natural world.
In our modern civilizations, we generally look at the unplanned outbreak of fire as a catastrophe requiring our immediate intercession. A building fire, for example, is extinguished as quickly as possible to save lives and property. And fires detected in fields, brush, and woodlands are promptly controlled to prevent their exponential growth. But has fire gone to our heads? Do we have an anthropocentric view of fire? Aren’t there naturally occurring fires that are essential to the health of some of the world’s ecosystems? And to our own safety? Indeed there are. And many species and the ecosystems they inhabit rely on the periodic occurrence of fire to maintain their health and vigor.
For the war effort- The campaign to reduce the frequency of forest fires got its start during World War II with distribution of this poster in 1942. The goal was to protect the nation’s timber resources from accidental or malicious loss due to fire caused by man-made ignition sources. The release of the Walt Disney film “Bambi” during the same year and the adoption of the Smokey the Bear mascot in 1944 softened the message’s delivery, but the public relations outreach continued to be a key element of a no-fire policy to save trees for lumber. Protection and management of healthy forest ecosystems in their entirety has only recently become a priority. (National Archives image)
Man has been availed of the direct benefits of fire for possibly 40,000 years or more. Here in the Lower Susquehanna River Watershed, the earliest humans arrived as early as 12,000 years ago—already possessing skills for using fire. Native plants and animals on the other hand, have been part of the ever-changing mix of ecosystems found here for a much longer period of time—millions to tens of millions of years. Many terrestrial native species are adapted to the periodic occurrence of fire. Some, in fact, require it. Most upland ecosystems need an occasional dose of fire, usually ignited by lightning (though volcanism and incoming cosmic projectiles are rare possibilities), to regenerate vegetation, release nutrients, and maintain certain non-climax habitat types.
But much of our region has been deprived of natural-type fires since the time of the clearcutting of the virgin forests during the eighteenth and nineteenth centuries. This absence of a natural fire cycle has contributed to degradation and/or elimination of many forest and non-forest habitats. Without fire, a dangerous stockpile of combustible debris has been collecting, season after season, in some areas for a hundred years or more. Lacking periodic fires or sufficient moisture to sustain prompt decomposition of dead material, wildlands can accumulate enough leaf litter, thatch, dry brush, tinder, and fallen wood to fuel monumentally large forest fires—fires similar to those recently engulfing some areas of the American west. So elimination of natural fire isn’t just a problem for native plants and animals, its a potential problem for humans as well.
Indiangrass (seen here), Switchgrass, Big Bluestem, and Little Bluestem are native species requiring periodic forms of disturbance to eliminate competition by woody plants. These warm-season grasses develop roots that penetrate deep into the soil, sometimes to depths of six feet or more, allowing them to survive severe drought and flash fire events. In the tall grass prairies, these extensive root systems allow these grasses to return following heavy grazing by roaming herds of American Bison (Bison bison). Without these habitat disturbances, warm season grasslands succumb to succession in about seven years. With their periodic occurrence, the plants thrive and provide excellent wildlife habitat, erosion control, and grazing forage.
To address the habitat ailments caused by a lack of natural fires, federal, state, and local conservation agencies are adopting the practice of “prescribed fire” as a treatment to restore ecosystem health. A prescribed fire is a controlled burn specifically planned to correct one or more vegetative management problems on a given parcel of land. In the Lower Susquehanna River Watershed, prescribed fire is used to…
Eliminate dangerous accumulations of combustible fuels in woodlands.
Reduce accumulations of dead plant material that may harbor disease.
Provide top kill to promote oak regeneration.
Regenerate other targeted species of trees, wildflowers, grasses, and vegetation.
Kill non-native plants and promote growth of native plants.
Prevent succession.
Remove woody growth and thatch from grasslands.
Promote fire tolerant species of plants and animals.
Improve habitat for rare species (Regal Fritillary, etc.)
Recycle nutrients and minerals contained in dead plant material.
Let’s look at some examples of prescribed fire being implemented right here in our own neighborhood…
Prescribed fires are typically planned for the dormant season extending from late fall into early spring with burns best conducted on days when the relative humidity is low.Prescribed fire is used regularly at Fort Indiantown Gap Military Reservation in Lebanon County, Pennsylvania, to keep accumulations of woody and herbaceous fuels from accumulating on and around the training range areas where live ordinance and other sources of ignition could otherwise spark large, hard-to-control wildfires.Prescribed fires replace the periodic natural burns that would normally reduce the fuel load in forested areas. Where these fuels are allowed to accumulate, south-facing slopes are particularly susceptible to extreme fires due to their exposure to the drying effects of intense sunlight for much of the year. The majority of small oaks subjected to treatment by the prescribed fire shown here will have the chance to regenerate without immediate competition from other species including invasive plants. The larger trees are mostly unaffected by the quick exposure to the flames. Note too that these fires don’t completely burn everything on the forest floor, they burn that which is most combustible. There are still plenty of fallen logs for salamanders, skinks, and other animals to live beneath and within.
A prescribed fire in late winter prevents this grassland consisting of Big Bluestem and native wildflowers from being overtaken by woody growth and invasive species. Fires such as this that are intended to interrupt the process of succession are repeated at least every three to five years.In its wildlife food plots, prescribed fire is used by the Pennsylvania Game Commission to prevent succession and control invasive species such as Multiflora Rose, instead promoting the growth of native plants.An example of a woodlot understory choked with combustible fuels and dense tangles of invasive Multiflora Rose. A forester has the option of prescribing a dose of dormant-season fire for a site like this to reduce the fuel load, top kill non-native vegetation, and regenerate native plants.A dose of prescribed fire was administered on this grassland to kill the woody growth of small trees beginning to overtake the habitat by succession.The Pennsylvania Game Commission employs prescribed fire at Middle Creek Wildlife Management Area and on many of their other holdings to maintain grasslands.Prescribed fire is used to eliminate invasive species including Multiflora Rose from grasslands at Middle Creek W.M.A. Annual burns on the property are conducted in a mosaic pattern so that each individual area of the grassland is exposed to the effects of fire only once every two to five years. Without fire or some type of mechanical or chemical intervention, succession by woody trees and shrubs would take hold after about seven years.Prescribed fire is planned for a fraction of total grassland acreage at Middle Creek W.M.A. each year. Another section of the mosaic is targeted in the following year and yet another in the year that follows that. Because burns are conducted in the spring, grassland cover is available for wildlife throughout the winter. And because each year’s fire burns only a portion of the total grassland acreage, wildlife still has plenty of standing grass in which to take shelter during and after the prescribed fire.Prescribed fire at Middle Creek W.M.A. provides grassland habitat for dozens of species of birds and mammals including the not-so-common Grasshopper Sparrow……and stocked Ring-necked Pheasants that do nest and raise young there.On a few sites in the Lower Susquehanna River Watershed , prescribed fire is being used to establish and maintain savanna-like grasslands. This one, located on a dry, south-facing slope near numerous man-made sources of ignition, can easily be dosed with periodic prescribed burns to both prevent succession and reduce fuel accumulations that may lead to a devastating extreme fire.One year following a prescribed burn, this is the autumn appearance of a savanna-like habitat with fire-tolerant Pitch Pine (Pinus rigida), Bear Oak, warm-season grasses, and a variety of nectar-producing wildflowers for pollinators. These ecosystems are magnets for wildlife and may prove to be a manageable fit on sun-drenched sites adjacent to man-made land disturbances and their sources of ignition.Savanna-like grasslands with oaks and other scattered large trees, some of them dead, make attractive nesting habitat for the uncommon Red-headed Woodpecker.Prescribed fire can benefit hungry Wild Turkeys by maintaining savanna-like grasslands for an abundance of grasshoppers and other insects in summer and improving the success of mast-producing oaks for winter.In the Lower Susquehanna River Watershed, the caterpillar of the rare Eastern Buck Moth feeds on the foliage of the Bear Oak, also known as the Scrub Oak, a shrubby species that relies upon periodic fire to eliminate competition from larger trees in its early successional habitat.Leaves of the Bear Oak in fall. The Bear Oak regenerates readily from top kill caused by fire.Reed Canary Grass (Phalaris arundinacea) is a native cool-season grass with a colorful inflorescence in spring. But given the right situation, it can aggressively overtake other species to create a pure stand lacking biodiversity. It is one of the few native species which is sometimes labelled “invasive”.Prescribed fire can be used to reduce an overabundance of Reed Canary Grass and its thatch in wetlands. Periodic burning can help restore species diversity in these habitats for plants and animals including rare species such as the endangered Bog Turtle (Glyptemys muhlenbergii).On the range areas at Fort Indiantown Gap in Lebanon County, Pennsylvania, disturbances by armored vehicles mimic the effects of large mammals such as the American Bison which periodically trampled grasses to prevent succession and the establishment of woody plants on its prairie habitat. To supplement the activity of the heavy vehicles and to provide suitable habitat for the very rare Regal Fritillary (Speyeria idalia) butterflies found there, prescribed fire is periodically employed to maintain the grasslands on the range. These burns are planned to encourage the growth of “Fort Indiantown Gap Little Bluestem” grass as well as the violets used as host plants by the Regal Fritillary caterpillars. These fires also promote growth of a variety of native summer-blooming wildflowers to provide nectar for the adults butterflies.A last record of a wild American Bison killed in Pennsylvania was an animal taken in the Susquehanna watershed in Union County in 1801. The species is thereafter considered extirpated from the state. Since that time, natural disturbances needed to regenerate warm-season grasses have been limited primarily to fires and riverine ice scour. The waning occurrence of both has reduced the range of these grasses and their prairie-like ecosystems in the commonwealth. (Exhibit: State Museum of Pennsylvania, Harrisburg)A male Regal Fritillary on the range at Fort Indiantown Gap, where armored vehicles and prescribed fire provide suitable prairie-like habitat for this vulnerable species.Prescribed fires return the nutrients and minerals contained in dead plant material to the soil. Following these controlled burns, insects like this Honey Bee can often be seen collecting minerals from the ashes.A Greenbottle Fly gathering minerals from the ash following a prescribed burn.
In Pennsylvania, state law provides landowners and crews conducting prescribed fire burns with reduced legal liability when the latter meet certain educational, planning, and operational requirements. This law may help encourage more widespread application of prescribed fire in the state’s forests and other ecosystems where essential periodic fire has been absent for so very long. Currently in the Lower Susquehanna River Watershed, prescribed fire is most frequently being employed by state agencies on state lands—in particular, the Department of Conservation and Natural Resources on State Forests and the Pennsylvania Game Commission on State Game Lands. Prescribed fire is also part of the vegetation management plan at Fort Indiantown Gap Military Reservation and on the land holdings of the Hershey Trust. Visitors to the nearby Gettysburg National Military Park will also notice prescribed fire being used to maintain the grassland restorations there.
For crews administering prescribed fire burns, late March and early April are a busy time. The relative humidity is often at its lowest level of the year, so the probability of ignition of previous years’ growth is generally at its best. We visited with a crew administering a prescribed fire at Middle Creek Wildlife Management Area last week. Have a look…
Members of a Pennsylvania Game Commission burn crew provide visitors to Middle Creek Wildlife Management Area with an overview of prescribed fire and the equipment and techniques they use to conduct a burn.Pennsylvania Game Commission Southeast Region Forester Andy Weaver will fulfill the role of Burn Boss for administering this day’s dose of fire. His responsibilities include assessing the weather before the burn and calculating a probability of ignition.The Burn Boss briefs personnel with information on site layout, water supply location(s), places of refuge, emergency procedures, the event’s goals and plan of action, crew assignments, and the results of the weather check: wind from the northwest at 5 miles per hour, temperature 48 degrees, and the relative humidity 63%. Today’s patient is a parcel of warm-season grasses receiving a dose of fire to eliminate invasive non-native plants, woody growth, and thatch. The probability of ignition is 20%, but improving by the minute.To begin the burn, a test fire is started in the downwind corner of the parcel, which also happens to be the bottom of the slope. Fuel ignition is good. The burn can proceed.Crews proceed uphill from the location of the test fire while igniting combustibles along both flanks of the area being treated.A drip torch is used to ignite the dried stems and leaves of warm-season grasses and wildflowers. Each member of the burn crew wears Nomex fire-resistant clothing and carries safety equipment including a two-way radio, a hydration pack, and a cocoon-like emergency fire shelter.An all-terrain vehicle equipped with various tools, a fire pump, hose, and a small water tank accompanies the crew on each flank of the fire.A mowed strip of cool-season grasses along the perimeter of the burn area is already green and functions as an ideal fire break. While the drip torch is perfect for lighting combustibles along the fire’s perimeter, the paintball gun-looking device is an effective tool used to lob incendiaries into the center areas of the burn zone for ignition.With green cool-season grasses already growing on the trails surrounding the burn zone, very little water was used to contain this prescribed fire. Where such convenient fire breaks don’t already exist, crews carry tools including chain saws, shovels, and leaf blowers to create their own. They also carry flame swatters, backpack water pumps, shovels, and other tools to extinguish fires if necessary. None of these items were needed to control this particular fire.This fast-burning fire provides enough heat to damage the cambium layer of the woody tree and shrub saplings in this parcel being maintained as a grassland/wildflower plot, thus the process of succession is forestalled. Burns conducted during previous years on this and adjacent fields have also controlled aggressive growth of invasive Multiflora Rose and Olives (Elaeagnus species).Crews proceed up the slope while maintaining the perimeter by igniting dry plant material along the flanks of the burn zone.Ignition complete, the crews monitor the fire.The Burn Boss surveys the final stages of a safe and successful prescribed fire. The fire has left behind a mosaic of burned and unburned areas, just as a naturally occurring event may have done. Wildlife dodging the flames may be taking refuge in the standing grasses, so there is no remedial attempt to go back and ignite these areas. They’ll be burned during prescribed fires in coming years.By June, this grassland will again be lush and green with warm-season grasses and blooming wildflowers like this Common Milkweed being visited by a Great Spangled Fritillary.And later in the summer, Eastern Tiger Swallowtails on Joe-pye Weed.Indiangrass in flower in mid-summer.Bobolinks glow in the late August sun while taking flight from a stand of warm-season grasses maintained using springtime prescribed fire. The small dots on the dark background at the top of the image are multitudes of flying insects, many of them pollinators. The vegetation is predominately Indiangrass, excellent winter cover for birds, mammals, and other wildlife.
Prescribed burns aren’t a cure-all for what ails a troubled forest or other ecosystem, but they can be an effective remedy for deficiencies caused by a lack of periodic episodes of naturally occurring fire. They are an important option for modern foresters, wildlife managers, and other conservationists.
It’s that time of year. Your local county conservation district is taking orders for their annual tree sale and it’s a deal that can’t be beat. Order now for pickup in April.
The prices are a bargain and the selection includes the varieties you need to improve wildlife habitat and water quality on your property. For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue). And don’t forget to order packs of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local native birds. They’re only $12.00 for a bundle of 10.
It’s the most desirable block in town, not because the houses are any different from others built during the post-war years of the mid-twentieth century, but because the first owners of these domiciles had the good taste and foresight to plant long-lived trees on their lots, the majority of them native species. Pin Oak, Northern Red Oak, Yellow Poplar, Flowering Dogwood (Cornus florida), Eastern Red Cedar, Eastern White Pine, Eastern Hemlock, Norway Spruce, and American Holly dominate the landscape and create excellent habitat for birds and other wildlife. These 75-year-old plantings provide an abundance of shade in summer and thermal stability in winter, making it a “cool” place to live or take a stroll at any time of the year.
Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024
Cumberland County Conservation District is taking orders for Common Winterberry, the ideal small shrub for wet soil anywhere on your property. To get berries, you’ll need both males and females, so buy a bunch and plant them in a clump or scattered group.To live for a century or more like this towering giant, a Pin Oak needs to grow in well-drained soils with adequate moisture. These sturdy shade providers do well along streams and on low ground receiving clean runoff from hillsides, roofs, streets, and parking areas. As they age, Pin Oaks can fail to thrive and may become vulnerable to disease in locations where rainfall is not adequately infiltrated into the soil. Therefore, in drier areas such as raised ground or slopes, avoid the Pin Oak and select the more durable Northern Red Oak for planting. This year, Pin Oaks are available from the Cumberland and Lancaster County Conservation Districts, while Dauphin, Lancaster, Lebanon, and York Counties are taking orders for Northern Red Oaks.The Cumberland County Conservation District is again offering a “Showy Northeast Native Wildflower and Grass Mix” for seeding your own pollinator meadow or garden. It consists of more than twenty species including this perennial favorite, Purple Coneflower.
Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024
The Eastern Redbud is small tree native to our forest edges, particularly in areas of the Piedmont Province with Triassic geology (Furnace Hills, Conewago Hills, Gettysburg/Hammer Creek Formations, etc.) Also known as the Judas Tree, the redbud’s brilliant flowers are followed by heart-shaped leaves. As seen here, it is suitable for planting near houses and other buildings. Eastern Redbud seedlings are being offered through tree sales in Dauphin, Cumberland, and Lancaster Counties.
The Yellow Poplar, often called Tuliptree or Tulip Poplar for its showy flowers, is a sturdy, fast-growing deciduous tree native to forests throughout the Lower Susquehanna River Watershed. Its pole-straight growth habit in shady woodlands becomes more spreading and picturesque when the plant is grown as a specimen or shade tree in an urban or suburban setting. The Yellow Poplar can live for hundreds of years and is a host plant for the Eastern Tiger Swallowtail butterfly. It is available this year from the Lancaster County Conservation District.The American Sweetgum, also known as Sweet Gum, is a large, long-lived tree adorned with a mix of vibrant colors in autumn.Ever wonder where all the American Goldfinches and particularly the Pine Siskins go after passing through our region in fall? Well, many are headed to the lowland forests of the Atlantic Coastal Plain where they feed on an abundance of seeds contained in spiky American Sweetgum fruits. In the Piedmont and Ridge and Valley Provinces of the Lower Susquehanna River Watershed, American Sweetgum transplants can provide enough sustenance to sometimes lure our friendly finches into lingering through the winter.The American Sweetgum is a versatile tree. It can be planted on upland sites as well as in wet ground along streams, lakes, and rivers. In the beaver pond seen here it is the dominate tree species. This year, you can buy the American Sweetgum from the Lancaster County Conservation District.“Red-twig Dogwood” is a group of similar native shrubs that, in our region, includes Silky Dogwood and the more northerly Red-Osier Dogwood (Cornus sericea). Both have clusters of white flowers in spring and showy red twigs in winter. They are an excellent choice for wet soils. Landscapers often ruin these plants by shearing them off horizontally a foot or two from the ground each year. To produce flowers and fruit, and to preserve winter attractiveness, trim them during dormancy by removing three-year-old and older canes at ground level, letting younger growth untouched.“Red-twig Dogwoods” make ideal mass plantings for streamside buffers and remain showy through winter, even on a gloomy day. They not only mitigate nutrient and sediment pollution, they provide excellent food and cover for birds and other wildlife. Both Silky and Red-osier Dogwoods are available for sale through the Lancaster County Conservation District as part of their special multi-species offers, the former is included in its “Beauty Pack” and the latter in its “Wildlife Pack”. The similar Gray Dogwood (Cornus racemosa) is being offered for sale by the York County Conservation District.
The unique maroon flowers of the Common Pawpaw produce banana-like fruits in summer. These small native trees grow best in damp, well-drained soils on slopes along waterways, where they often form clonal understory patches. To get fruit, plant a small grove to increase the probability of pollination. The Common Pawpaw is a host plant for the Zebra Swallowtail butterfly. It is available through both the Lebanon and Lancaster County sales.The Eastern Red Cedar provides excellent food, cover, and nesting sites for numerous songbirds. Planted in clumps of dozens or groves of hundreds of trees, they can provide winter shelter for larger animals including deer and owls. The Eastern Red Cedar is being offered for purchase through both the Lebanon and Lancaster County Conservation Districts.Care to try your hand at raising some chestnuts? Lebanon County Conservation District has hybrid American Chestnut seedlings for sale.Lebanon County Conservation District is offering Common Winterberry and Eastern White Pine during their 2024 Tree and Plant Sale. Plant them both for striking color during the colder months. Eastern White Pine is also available from the Cumberland, Dauphin, Lancaster, and York County sales.
In addition to a selection of trees and shrubs, the Perry County Conservation District is again selling wildflower seed mixes for starting your own pollinator meadow or garden. For 2024, they have both a “Northeast Perennials and Annuals Mix” and a “Butterfly and Hummingbird Seed Mix” available. Give them a try so you can give up the mower!
Again this year, Perry County is offering bluebird nest boxes for sale. The price?—just $12.00.
Wait, what?,…twelve bucks,…that’s cheaper than renting!
The Buttonbush, a shrub of wet soils, produces a cosmic-looking flower. It grows well in wetlands, along streams, and in rain gardens. Buttonbush seedlings are for sale from both the York and Lancaster County Conservation Districts.
To get your deciduous trees like gums, maples, oaks, birches, and poplars off to a safe start, conservation district tree sales in Cumberland, Dauphin, Lancaster, and Perry Counties are offering protective tree shelters. Consider purchasing these plastic tubes and supporting stakes for each of your hardwoods, especially if you have hungry deer in your neighborhood.
Tree shelters protect newly transplanted seedlings from browsing deer, klutzy hikers, visually impaired mower operators, and other hazards.
There you have it. Be sure to check out each tree sale’s web page to find the selections you like, then get your order placed. The deadlines will be here before you know it and you wouldn’t want to miss values like these!
As week-old snow and ice slowly disappears from the Lower Susquehanna River Watershed landscape, we ventured out to see what might be lurking in the dense clouds of fog that for more than two days now have accompanied a mid-winter warm spell.
After freezing to a slushy consistency earlier this week, the Susquehanna is already beginning to thaw. Below the York Haven Dam at Conewago Falls, the water is open and ice-free.On frozen man-made lakes and ponds, geese and ducks like these Mallards and American Wigeon are presently concentrated around small pockets of open water.During the past ten days, American Robin numbers have exploded throughout the lower Susquehanna valley. The majority of these birds may be a mix of both those coming south to escape the late onset of wintry conditions to our north and those inching north into our region as early spring migrants.The January thaw has melted the snow from lawns and fields to provide thousands of visiting robins with a chance to forage for earthworms.A visit by this young Cooper’s Hawk to the susquehannnawildlife.net headquarters garden sent songbirds scrambling……but did nothing to unnerve our resident Eastern Gray Squirrels,……which promptly went into tail-waving mode to advertise their presence.But earlier in the week, when heavy snow cover in the rural areas surrounding our urbanized neighborhood made it difficult for rodent-eating raptors to find food, we received brief visits from both a Red-tailed Hawk……and this young Red-shouldered Hawk, an uncommon bird of prey most often found in wet woods and other lowlands.To escape notice during visits by these larger raptors, our squirrels remained motionless and commenced performance of their best bump-on-a-log impressions.Unimpressed, each of our visiting buteos remained for just a few minutes before moving on in search of more favorable hunting grounds and prey.As snow melted and exposed bare ground in fields of early successional growth, we encountered……a flock of White-crowned Sparrows, most in first-winter plumage……and at least a dozen American Tree Sparrows. During the twentieth century, these handsome songbirds were regular winter visitors to the lower Susquehanna region. During recent decades, they’ve become increasingly more difficult to find. Currently, moderate numbers appear to be arriving to escape harsher weather to our north.What could be more appropriate on a foggy, gray evening than finding a “gray ghost” (adult male Northern Harrier) patrolling the fields in search of mice and voles.
If scenes of a January thaw begin to awaken your hopes and aspirations for all things spring, then you’ll appreciate this pair of closing photographs…
The maroon-red flower buds of Silver Maples are beginning to swell. And woodpeckers including Pileated Woodpeckers are beginning to drum, a timber-pounding behavior they use to establish breeding territories in habitats with suitable sites for cavity nesting.In wet soil surrounding spring seeps and streams, Skunk Cabbage is rising through the leaf litter to herald the coming of a new season. Spring must surely be just around the corner.
Just as bare ground along a plowed road attracts birds in an otherwise snow-covered landscape, a receding river or large stream can provide the same benefit to hungry avians looking for food following a winter storm.
Here is a small sample of some of the species seen during a brief stop along the Susquehanna earlier this week.
Along vegetated edges of the Susquehanna and its tributaries, the Song Sparrow is ubiquitous in its search for small seeds and other foods. As the river recedes from the effects of this month’s rains, the shoreline is left bare of more recently deposited snow cover. Song Sparrows and other birds are attracted to streamside corridors of frost-free ground to find sufficient consumables for supplying enough energy to survive the long cold nights of winter.Thousands of American Robins have been widespread throughout the lower Susquehanna valley during the past week. Due to the mild weather during this late fall and early winter, some may still be in the process of working their way south. Currently, many robins are concentrated along the river shoreline where receding water has exposed unfrozen soils to provide these birds with opportunities for finding earthworms (Lumbricidae) and other annelids.This Golden-crowned Kinglet was observed searching the trees and shrubs along the Susquehanna shoreline for tiny insects and spiders. Temperatures above the bare ground along the receding river can be a few degrees higher than in surrounding snow-covered areas, thus improving the chances of finding active prey among the trunks and limbs of the riparian forest.Not far from the kinglet, a Brown Creeper is seen searching the bark of a Silver Maple (Acer saccharinum) for wintering insects, as well as their eggs and larvae. Spiders in all their life stages are a favorite too.American Pipits not only inhabit farm fields during the winter months, they are quite fond of bare ground along the Susquehanna. Seen quite easily along a strip of pebbly shoreline exposed by receding water, these birds will often escape notice when spending time on mid-river gravel and sand bars during periods of low flow.An American Pipit on a bitterly cold afternoon along the Susquehanna.
With the earth at perihelion (its closest approach to the sun) and with our home star just 27 degrees above the horizon at midday, bright low-angle light offered the perfect opportunity for doing some wildlife photography today. We visited a couple of grasslands managed by the Pennsylvania Game Commission to see what we could find…
On this State Game Lands parcel, prescribed fire is used to maintain a mix of grasslands and brushy early successional growth. In nearby areas, both controlled fire and mechanical cutting are used to remove invasive species from hedgerows and the understory of woodlots. Fire tolerant native species then have an opportunity to recolonize the forest and improve wildlife habitat. This management method also reduces the fuel load in areas with the potential for uncontrolled wildfires.The sun-dried fruits of a native Common Persimmon tree found growing in a hedgerow.Just one year ago, mechanical removal of invasive trees and shrubs (including Multiflora Rose) on this State Game Land was followed by a prescribed fire to create this savanna-like grassland.Hundreds of Song Sparrows were found in the grasses and thickets at both locations.White-throated Sparrows were also abundant, but prefer the tangles and shrubs of the thickets.Northern Mockingbirds were vigilantly guarding winter supplies of berries in the woodlots and hedgerows.In grasses and tangles on wetter ground, about a dozen Swamp Sparrows were discovered.The adult White-crowned Sparrow is always a welcome find.And seeing plenty of juvenile White-crowned Sparrows provides some assurance that there will be a steady stream of handsome adult birds arriving to spend the winter during the years to come.Dark-eyed Juncos were encountered only in the vicinity of trees and large shrubs.Several Savannah Sparrows were observed. Though they’re mostly found in treeless country, this particular one happened to pose atop a clump of shrubs located within, you guessed it, the new savanna-like grasslands.A tiny bird, even when compared to a sparrow, the Winter Wren often provides the observer with just a brief glimpse before darting away into the cover of a thicket.Within grasslands, scattered stands of live and dead timber can provide valuable habitat for many species of animals.Woodpeckers and other cavity-nesting birds rely upon an abundance of “snags” (standing dead trees) for breeding sites.This Red-bellied Woodpecker and about a dozen others were found in trees left standing in the project areas.A Yellow-bellied Sapsucker soaks up some sun.This very cooperative Pileated Woodpecker seemed to be preoccupied by insect activity on the sun-drenched bark of the trees. This denizen of mature forests will oft times wander into open country where larger lumber is left intact.
Just as things were really getting fun, some late afternoon clouds arrived to dim the already fading daylight. Just then, this Northern Harrier made a couple of low passes in search of mice and voles hidden in the grasses.It was a fitting end to a very short, but marvelously sunny, early winter day.
Recently, we found these Chestnut Oak acorns setting roots into the leaf litter to secure their place among the plants that will turn the forest understory green in the spring. Individual acorns that germinate soon after falling to the ground in autumn may avoid becoming food for squirrels, turkeys, deer, and other wildlife, thus increasing their chances of surviving to later become adult trees able to produce acorns that pass this quick-development trait to yet another generation of oaks.
It looks like Uncle Tyler Dyer had a fun-filled Halloween. Earlier this afternoon, we caught him drifting off into a nap during his work to make updates to his leaf scans. As he snoozed away, we found these lying on the floor next to his easy chair and decided to lend a hand by adding them to his collection. We’ll see how long it takes him to discover our dirty little deed. You can have a look at our mischief by clicking on the “Trees, Shrubs, and Woody Vines” tab at the top of this page.
Where should you go this weekend to see vibrantly colored foliage in our region? Where are there eye-popping displays of reds, oranges, yellows, and greens without so much brown and gray? The answer is Michaux State Forest on South Mountain in Adams, Cumberland, and Franklin Counties.
South Mountain is the northern extension of the Blue Ridge Section of the Ridge and Valley Province in Pennsylvania. Michaux State Forest includes much of the wooded land on South Mountain. Within or adjacent to its borders are located four state parks: King’s Gap Environmental Education Center, Pine Grove Furnace State Park, Caledonia State Park, and Mont Alto State Park. The vast network of trails on these state lands includes the Appalachian Trail, which remains in the mountainous Blue Ridge Section all the way to its southern terminus in Georgia.
In Pennsylvania, the forested highlands of the Blue Ridge Section of the Ridge and Valley Province are known as South Mountain. Much of South Mountain lies within the boundaries of Michaux State Forest. Stars indicate the locations of 1) King’s Gap Environmental Education Center, 2) Pine Grove Furnace State Park, 3) Caledonia State Park, and 4) Mont Alto State Park. A drive on US 30 between Gettysburg and Chambersburg will take you right through Michaux State Forest along an east to west axis while a scenic northbound or southbound trip along PA 233 will bring you in proximity to each of the state parks located therein. (Base image from NASA Earth Observatory Collection)
If you want a closeup look at the many species of trees found in Michaux State Forest, and you want them to be labeled so you know what they are, a stop at the Pennsylvania State University’s Mont Alto arboretum is a must. Located next door to Mont Alto State Park along PA 233, the Arboretum at Penn State Mont Alto covers the entire campus. Planting began on Arbor Day in 1905 shortly after establishment of the Pennsylvania State Forest Academy at the site in 1903. Back then, the state’s “forests” were in the process of regeneration after nineteenth-century clear cutting. These harvests balded the landscape and left behind the combustible waste which fueled the frequent wildfires that plagued reforestation efforts for more than half a century. The academy educated future foresters on the skills needed to regrow and manage the state’s woodlands.
Online resources can help you plan your visit to the Arboretum at Penn State Mont Alto. More than 800 trees on the campus are numbered with small blue tags. The “List of arboretum trees by Tag Number” can be downloaded to tell you the species or variety of each. The interactive map provides the locations of individual trees plotted by tag number while the Grove Map displays the locations of groups of trees on the campus categorized by region of origin. A Founder’s Tree Map will help you find some of the oldest specimens in the collection and a Commemorative Tree Map will help you find dedicated trees. There is also a species list of the common and scientific tree names.
The Yellow Buckeye (Aesculus flava) is a tree found in the forests of the Blue Ridge Section of the Ridge and Valley Province in West Virginia, Virginia, North Carolina and Georgia. You can see it in Pennsylvania by visiting the collection of trees in the Arboretum at Penn State Mont Alto.The American Chestnut can be difficult to find due to the impact of chestnut blight, but you can see it in the Arboretum at Penn State Mont Alto.Shagbark Hickory is a common tree in the forests of South Mountain.The Sweet Cherry (Prunus avium), a native of Europe, is naturalized throughout eastern and south-central Pennsylvania and is one of the more than 150 species of trees in the arboretum’s collection.Sweet Birch (Betula lenta) foliage is particularly bright yellow on South Mountain this autumn. It really “pops” against the backdrop of the evergreen Eastern White Pines and Eastern Hemlocks. During less-than-ideal years, Sweet Birch leaves can subtly transition from green to drab brown without much fanfare before falling.You might have a difficult time finding a Common Persimmon (Diospyros virginiana) growing wild in Pennsylvania, but you can find it in the Arboretum at Penn State Mont Alto.
The autumn leaves will be falling fast, so make it a point this weekend to check out the show on South Mountain.
Last October 3rd, a late-season Ruby-throated Hummingbird stopped by the garden at susquehannawildlife.net headquarters to take shelter from a rainy autumn storm. It was so raw and chilly that we felt compelled to do something we don’t normally do—put out the sugar water feeder to supplement the nectar produced by our fall-flowering plants. After several days of constant visits to the feeder and the flowers, our lingering hummer resumed its southbound journey on October 7th.
Fast forward to this afternoon and what do you know, at least two migrating hummingbirds have stopped by to visit the flowers in our garden. This year, we have an exceptional abundance of blooms on some of their favorite plants. In the ponds, aquatic Pickerelweed is topped with purple spikes and we still have bright orange tubular flowers on one of our Trumpet Vines—a full two to three months later than usual.
We checked each of our late-season visitors carefully to be reasonably certain that none was a stray western species of hummingbird. All appear to be female or juvenile “Ruby-throats”. If you have an abundance of flowering plants and/or you’re going to maintain your hummingbird feeders through the coming weeks, be on the lookout for western species. Most are more hardy than our Ruby-throats and some have remained in the lower Susquehanna valley through the winter.As is typically the case, Ruby-throated Hummingbirds quickly gravitate toward the tubular flowers of our Cuphea ignea, the Mexican Cigar. They find these showy plants to be absolutely irresistible.Mexican Cigar grows wild in parts of the Ruby-throated Hummingbird’s winter range. To them, it’s comfort food.We’re trying some new cultivars of Cuphea to see how they do. As this composite image shows, the hummingbirds won’t let our Bat-faced Cuphea (Cuphea llavea) alone. It’s another plant native to Mexico and Central America, right where some of our hummingbirds spend the winter.
Remember, keep those feeders clean and the provisions fresh! You’ll be glad you did.
What’s all this buzz about bees? And what’s a hymanopteran? Well, let’s see.
Hymanoptera—our bees, wasps, hornets and ants—are generally considered to be our most evolved insects. Some form complex social colonies. Others lead solitary lives. Many are essential pollinators of flowering plants, including cultivars that provide food for people around the world. There are those with stingers for disabling prey and defending themselves and their nests. And then there are those without stingers. The predatory species are frequently regarded to be the most significant biological controls of the insects that might otherwise become destructive pests. The vast majority of the Hymanoptera show no aggression toward humans, a demeanor that is seldom reciprocated.
Late summer and early autumn is a critical time for the Hymanoptera. Most species are at their peak of abundance during this time of year, but many of the adult insects face certain death with the coming of freezing weather. Those that will perish are busy, either individually or as members of a colony, creating shelter and gathering food to nourish the larvae that will repopulate the environs with a new generation of adults next year. Without abundant sources of protein and carbohydrates, these efforts can quickly fail. Protein is stored for use by the larval insects upon hatching from their eggs. Because the eggs are typically deposited in a cell directly upon the cache of protein, the larvae can begin feeding and growing immediately. To provide energy for collecting protein and nesting materials, and in some cases excavating nest chambers, Hymanoptera seek out sources of carbohydrates. Species that remain active during cold weather must store up enough of a carbohydrate reserve to make it through the winter. Honey Bees make honey for this purpose. As you are about to see, members of this suborder rely predominately upon pollen or insect prey for protein, and upon nectar and/or honeydew for carbohydrates.
We’ve assembled here a collection of images and some short commentary describing nearly two dozen kinds of Hymanoptera found in the Lower Susquehanna River Watershed, the majority photographed as they busily collected provisions during recent weeks. Let’s see what some of these fascinating hymanopterans are up to…
SOLITARY WASPS
A Great Black Wasp on goldenrod (Solidago species). Like other solitary wasps, a female Great Black Wasp will sting and paralyze a host insect upon which she’ll deposit her eggs. After hatching, the larvae will begin consuming the host’s body as a source of protein. The parasitized insects are often katydids or grasshoppers.A Great Black Wasp feeding on nectar, a source of carbohydrates. Unlike social bees and wasps, solitary wasps are equipped with a stinger solely used for immobilizing prey, not defending a nest. They are therefore quite docile and pose little threat to humans.A Great Black Wasp powdered with pollen. Hymanopterans that gather nectar and/or pollen are tremendously important pollinators of hundreds of species of plants.A female Thread-waisted wasp (Ammophilia species, probably A. nigricans) drags a paralyzed caterpillar to her excavated nest where she’ll deposit an egg on the body. After hatching, the larval wasp will feed on the disabled caterpillar. The protein will enable the larvae to grow, pupate, and later emerge as an adult wasp.The female Eastern Cicada Killer (Sphecius speciosus) excavates an underground nest with branch tunnels connecting a dozen chambers or more. As the common name suggests, the female wasp paralyzes a cicada, then makes a strenuous effort to fly and drag it back to the nest for placement in a cell. Each male wasp egg is deposited upon just one immobilized cicada, but a female egg is provided with a cache of several cicadas to provide adequate protein for growth to a larger size. Nest cells are sealed with soil, then the larvae hatch in just a couple of days. Within about two weeks, they have consumed the cicada protein and are fully grown. Wrapped in a cocoon, they spend the winter in the nest, then pupate in the spring before emerging as a new generation of adults.The Black-and-yellow Mud Dauber (Sceliphron caementarium) builds a mud-ball nest within which it packs paralyzed spiders to function as a source of protein for its larvae.A Black-and-yellow Mud Dauber at nest.The Pipe Organ Mud Dauber builds this elaborate nest in which their eggs and paralyzed spiders are deposited in cells sealed with mud partitions. After consuming the spiders, the larvae pupate, overwinter, then emerge from their cells as adults during the following spring. To escape the protection of the nest, the new generation of adults bore through the mud walls. Adult Pipe Organ Mud Daubers resemble the Great Black Wasp, but have a white or yellow distal segment on their rear legs resembling a pair of light-colored socks.A closeup of the previous image with the lengths of the nest tubes compressed to show four scavenger flies (Miltogramminae), possibly two species, that have invaded this Pipe Organ Mud Dauber nest. Scavenger flies are kleptoparasites that victimize various solitary bees and wasps, depositing larvae directly into the host species’ nest cells to consume the protein cache stored therein.
CUCKOO WASPS
Cuckoo Wasps (Chrysididae), also known as Emerald Wasps, parasitize the nests of other species of wasps. Females lay their eggs inside the host’s nest, then flee the scene. Upon hatching, larval Cuckoo Wasps feed on stockpiles of prey intended for the host species’ offspring. Like the adult mud daubers that have already matured and departed this nest by digging a hole through the wall of the cell within which they were hatched, the metallic green Cuckoo Wasp in the upper left has just emerged in much the same way.
SWEAT BEES
A Sweat Bee (Lasioglossum species) collecting nectar and pollen on White Snakeroot (Ageratina altissima).Sweat Bees (Lasioglossum species) visit human skin to lick up the electrolytes left behind by evaporating perspiration.Sweat Bees in the genus Lasioglossum demonstrate various social behaviors ranging from species that are solitary nesters to those that create colonies with work forces ranging in size from as few as four to as many as hundreds of bees. Some Lasioglossum practice kleptoparasitism, while others are quite accomplished foragers.A female Augochlorine Green Sweat Bee (Augochlorini) collecting nectar on White Snakeroot. Notice the pollen “baskets” on the rear leg.An Augochlorine Green Sweat Bee (Augochlorini). Sweat bees nest in subterranean cavities and in hollowed out sections of trees.A copper-colored Augochlorine Green Sweat Bee (Augochlorini) collecting nectar and dusted with pollen.
LEAFCUTTER AND MASON BEES
A Leafcutter Bee (Megachile species). Like Mason Bees, female Mason Bees deposit each of their eggs on a “pollen loaf” within an individual cell inside a preexisting tunnel-like cavity in wood, stone, or in the ground. Unlike Mason Bees, female Leafcutter Bees cut a circular piece of leaf to create each of the cells in their nest. After hatching, the larval bee feeds on the pollen loaf, pupates, then emerges from the shelter of the nest to start a new generation, usually during the following year.A Leafcutter Bee (Megachile species) visiting Wild Bergamot. Female Leafcutter and Mason Bees lack pollen “baskets” on their rear legs but instead have pollen “brushes” on the underside of the abdomen to gather the protein they need to create a “pollen loaf” for each nest cell.A Leafcutter Bee (Megachile species) collecting nectar from White Snakeroot.A Mason Bee (Osmia species) emerging from a nest in spring. Mason Bees create nesting cells within preexisting cavities in wood, stone, and other other supporting structures. Within the nest cavity, each egg is deposited atop a cache of pollen and nectar, a pollen loaf, then enclosed behind a partition of mud. The female Mason Bee will usually repeat this process until an entire cavity is filled with cells. During the following spring, a new generation of adult Mason Bees digs its way through the cell walls to emerge and repeat the process. These bees readily use paper straws or holes drilled in blocks of wood for nesting.A mason bee nest box with holes drilled into blocks of wood.Mason Bees seal each cell and the outer end of their nest cavity with mud. These outer nest cells can been parasitized by a variety of wasps. Here, the outer cell of a Mason Bee nest has been victimized by a tiny chalcid wasp (looks like another one to the lower left). Several species of female chalcid wasps (native Monodontomerus species or non-native Pteromalus venustus) enlarge weak points in the outer partition of a mason bee nest, then sting and paralyze the larval bee inside before depositing their eggs. Within the cell. the wasp larvae consume the larval Mason Bee and the “pollen loaf” provided for its growth. These same parasitic wasps prey upon Leafcutter Bees as well.
BUMBLE BEES, CARPENTER BEES, HONEY BEES, AND DIGGER BEES
A Common Eastern Bumble Bee (Bombus impatiens) collecting nectar and pollen on goldenrod. Bumble bees are our sole native group of social bees. Their wax nests are built in a burrow or other shelter. The eggs are deposited in cells along with a supply of pollen for nourishing the larvae upon hatching. Honey is stored in “honey pots” within the nest. New queens are produced along with male bees during the late-summer and fall. Only the new generation of fertilized queens survive the winter to lay eggs and produce workers to construct a new nest.A pair of Common Eastern Bumble Bees collecting nectar and becoming dusted with pollen. Their fuzzy coats and semi-warm-blooded metabolism allows them to be active in cooler weather than is tolerated by other bees.Flowering plants including the Great Rhododendron find success attracting pollinators to their reproductive blossoms by offering carbohydrate-rich nectar to insects like this Eastern Bumble Bee. The yellow spots on the flower’s upper petal help to guide visitors toward their sweet treat.An Eastern Carpenter Bee feeding on goldenrod nectar. Compare the almost hairless abdomen to that of the bumble bees. Carpenter bees are semi-social insects. Females lay their eggs in cells within galleries bored into wood. These nests are completed with great precision, avoiding creation of any second entrance by mistakenly breaching the outer surface of the excavated wood. Each egg/larvae is provided with a supply of protein-rich pollen. Males often hover outside their mate’s nest to prevent competing males from entering the area.A worker Honey Bee, a female member of a sisterhood of foragers from a nearby hive, visits goldenrod alongside Common Eastern Bumble Bees. Honey Bees were brought to North America during the 1620s, the earliest years of the trans-Atlantic migration of European colonists, to pollinate cultivated plants and to provide a reliable source of honey and beeswax. Within the Honey Bee’s social structure, the queen of each hive lays the eggs to produce the female worker bees. Once each year, male drones are produced along with a new generation of queens.In nature, Honey Bees build hives in tree cavities. Recently, this colony constructed a hive in a screech owl nest box at susquehannawildlife.net headquarters. To provide protein for the hatching larvae, worker bees collect pollen and deposit it within the hexagonal cells of the vertically aligned beeswax combs. After an egg is deposited upon the pollen cache, each cell is sealed with more beeswax. Young females tend these nest combs before maturing and becoming foraging worker bees.In apiculture, Honey Bees are raised in man-made hives. This Pennsylvania Association of Beekeepers display gives visitors to the Pennsylvania Farm Show in Harrisburg a look at the inner workings of a live bee hive. Nectar collected by worker bees is turned into honey to provide the supply of carbohydrates needed to fuel the colony through the winter. Note the honeycombs on the glass.A possible Small Carpenter Bee (Ceratina species) visiting White Snakeroot. Small Carpenter Bees nest inside hollow stems and twigs. Some species are eusocial, with a queen’s daughters and sisters sharing responsibility for finding food and rearing the young. Females overwinter inside a one of the excavated stems and begin a new nest there in the spring.A Digger Bee (possibly Melissodes species) with “pollen baskets” full of pollen collected from nearby flowers. Digger Bees in the genus Melissodes are often known as the Long-horned Bees. These social insects excavate underground nests and many species practice communal living.
SCOLIID WASPS
The Two-spotted Scoliid Wasp (Scolia dubia), also known as the Blue-winged Scoliid Wasp, is most frequently observed feeding on nectar. Scoliid wasps are solitary nesters, though they may assemble into groups while visiting flowers. They often ignore the presence of humans and are seldom disturbed by their presence. Females seek out the burrowing grubs of beetles including the Green June Bug (Cotinis nitida) and possibly the Japanese Beetle. After stinging a grub to paralyze it, the wasp will deposit her egg on its body, then bury it. Upon hatching, the larval wasp will feed on the grub for nourishment as it grows.Don’t like having your watermelon overrun by Green June Bugs while you’re eating? Then you ought to go out of your way to be nice to the Two-spotted Scoliid Wasp.The Double-banded Scoliid (Scolia bicincta) parasitizes beetle larvae as hosts for its larvae. For carbohydrates it relishes flower nectar.
PAPER WASPS
A Northern Paper Wasp (Polistes fuscatus). Paper wasps prey upon numerous garden pests, particularly caterpillars, to collect protein. Though they are social insects equipped with stingers to subdue their victims and defend their nests, paper wasps are surprisingly docile.A Northern Paper Wasp (Polistes fuscatus) feeding on nectar from a goldenrod flower.A Northern Paper Wasp harvesting wood pulp from the side of a mason bee nest box at susquehannawildlife.net headquarters. The pulp is chewed in the wasp’s saliva to create the paper used to construct the colony’s open-cell nest.Common Paper Wasps (Polistes exclamans), also known as Guinea Paper Wasps, at their open-cell nest. This and the nests of most other paper wasps are suspended on a filament or a pedicle. Many paper wasps can excrete an ant repellent on this section of the nest in an effort to prevent invasion. Like many other social hymenopterans, a defending wasp can secrete a pheromone venom during the stinging process to warn the colony of danger at the nest. In winter, Common Paper Wasps seek shelter in stumps and other locations to hibernate.The European Paper Wasp (Polistes dominula) is a non-native species which builds nests in man-made structures including bird houses. To collect protein, they prey on a wide selection of insects and other invertebrates. As such, European Paper Wasps are widespread and successful here in North America.
YELLOWJACKETS AND HORNETS
An Eastern Yellowjacket feeding on lanternfly honeydew. Eastern Yellowjackets derive much of their success from being generalists, collecting carbohydrates from nearly any sweet source, natural or man made. They are quite fond of ripe fruits, flower nectar, and sugary snacks and drinks, especially soda. Protein for nourishing their larvae is derived from the wide variety invertebrates upon which they prey and from carrion. These foods are chewed into a paste form in preparation for placement into the brood cells.A subterranean colony of Eastern Yellowjackets is started anew each spring by a young queen that has survived winter hibernation in diapause, a state of interrupted development. She constructs the new nest’s first cells using pulp made by chewing rotting wood. The first brood of workers scales up construction while the queen continues producing eggs. At the nest, these social insects will viciously attack anyone or anything perceived to be a threat, so give them their space and leave them alone. Many yellowjacket infestations of homes and other buildings are the work of non-native German Yellowjacket (Vespula germanica) [not shown], an invasive species that constructs paper nests in void spaces including walls and attics.Yellowjackets may be moody and aggressive, but they do fall victim to a number of predators. A Robber Fly (Promachus species) has taken down and is devouring this Eastern Yellowjacket.A Bald-faced Hornet (Dolichovespula maculata) feeding on Spotted Lanternfly honeydew on a Tree-of-heaven (Ailanthus altissima). In the absence of nectar-producing flowers, many bees, yellowjackets, and hornets have turned to the invasive lanternfly and Ailanthus combo to turn the sun’s energy into the carbohydrates they need. For protein, they prey upon spiders, flies, caterpillars, and a variety of other insects.To create paper for nest construction, this Bald-faced Hornet is collecting wood pulp from the surface of a weathered picnic table. Away from the nest, these hornets demonstrate a calm, carefree demeanor and can be closely observed.A Bald-faced Hornet nest in a pine tree. These hives are strictly temporary. Within the nest, a generation of drones (males) and new queens are produced late each year. These wasps leave the colony to mate. With the arrival of freezing weather, all inhabitants within the nest, including the old queen, perish, as do the drones that departed to breed. Only the new queens survive winter hibernation to propagate the next generation of wasps, starting with the workers needed to construct a fresh nest and reestablish the colony.Did you ever get the feeling you’re being watched? Don’t go messing around with Bald-faced Hornet nests. The occupants therein, like other social bees, wasps, and hornets, are equipped with stingers and venom for defending their colony. This is an adaptation that has developed over time to assure the survival of populations of these insects. Think about it this way, a solitary wasp that loses a nest loses only their individual brood of offspring. There is minimal impact on the wider local population of such insects. Conversely, a social wasp or hornet that loses a nest loses an entire colony, possibly negating the benefits of their cooperative behavior and threatening the survival of the species. Insects that cooperate to build societies for survival can be more vulnerable to the catastrophic impacts of certain circumstances like disease, weather, and invasion of their colonies. Therefore, natural selection has provided them with contingencies for these dangers, for example, the instinct to construct protective shelters and the adaptation of stingers and venom for defense against intruders and would-be predators. Oh, and by the way, the Bald-faced Hornet can spray venom, often aiming for the eyes, so keep your distance.European Hornets (Vespa crabro), an introduced species, are predatory on a variety of flying insects for protein. For carbohydrates they are attracted to sweets like this lanternfly honeydew on Tree-of-heaven.European Hornets constructing a paper nest in a tree cavity.
POTTER WASPS
A potter wasp (Eumenes species), probably a Fraternal Potter Wasp (E. fraternus), hovering near a European Paper Wasp on Partridge Pea. The female potter wasp builds a small mud nest resembling a tiny clay pot. One of her eggs is inserted and left hanging on a thin thread. Then a paralyzed caterpillar is deposited as a source of protein to nourish the larva upon hatching. Lastly, the pot is sealed with a lid made of wet mud. Upon maturing, the new generation of adult wasps perform a pottery breaking to emerge and take flight.
ANTS
Field Ants (Formica species, possibly Formica pallidefulva) clear the entrance to their underground nest. Field ants are eusocial insects, they work in concert to build, maintain, and defend the nest, rear young, and find food. There is no social caste system. Field Ants are predators and scavengers when collecting protein. For carbohydrates they often rely on the honeydew produced by aphids. As a method of improving and sustaining the production of honeydew, some ant species will tend colonies of aphids by moving the younger individuals from depleted portions of plants to more healthy tissue. Field Ant nests contain chambers used for a variety of functions including raising young and storing food. Some nests include multiple queens and some colonies consist of more than one nest. Ants in the genus Formica are weaponized; they can spray formic acid to repel intruders and defend their colony.
We hope this brief but fascinating look at some of our more common bees, wasps, hornets, and ants has provided the reader with an appreciation for the complexity with which their food webs and ecology have developed over time. It should be no great mystery why bees and other insects, particularly native species, are becoming scarce or absent in areas of the Lower Susquehanna River Watershed where the landscape is paved, hyper-cultivated, sprayed, mowed, and devoid of native vegetation, particularly nectar-producing plants. Late-summer and autumn can be an especially difficult time for hymanopterans seeking the sources of proteins and carbohydrates needed to complete preparations for next year’s generations of these valuable insects. An absence of these staples during this critical time of year quickly diminishes the diversity of species and begins to tear at the fabric of the food web. This degradation of a regional ecosystem can have unforeseen impacts that become increasingly widespread and in many cases permanent.
How can anyone be surprised by the absence of bees and other pollinators in farmland? Manicured and cultivated ground offers little in the way of year-round shelter and food sources for insects and other wildlife.This savanna-like habitat on a south-facing slope provides the abundance of nectar-producing, pollen-rich wildflowers needed to nourish a diverse population of insects including bees, wasps, hornets, and ants. Goldenrods, asters, and White Snakeroot are some of their late-season favorites.
Editor’s Note: No bees, wasp, hornets, or ants were harmed during this production. Neither was the editor swarmed, attacked, or stung. Remember, don’t panic, just observe.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York, NY.
Ruby-throated Hummingbirds will be migrating south through the Lower Susquehanna River Watershed during the next few weeks.While on their journey, they refuel by frequently visiting nectar-producing native wildflowers as well as garden specimens like this Mexican Cigar.Hummingbirds supplement their diet by visiting feeders filled with a mixture of sugar and water. If you’re feeding hummingbirds, or thinking about feeding hummingbirds, be sure to review the helpful tips contained in our post from August 5, 2022, “Two Feeders Are Better Than One”. Their health and your peace of mind may depend on it.
Looking for a native wildflower that’s tall, showy, and a great choice for attracting wildlife, especially butterflies and bees? Then check out Wild Senna (Senna hebecarpa).
Wild Senna currently blooming in a roadside wildflower garden on Pennsylvania State Game Lands.
Wild Senna, also known as American Senna, is a host plant for the larvae of Cloudless Sulphur and Sleepy Orange (Eurema nicippe) butterflies. It thrives in almost any moist, well-drained soil in habitats including open woodlands, forest edges, meadows, and gardens like yours. Its height at flowering ranges from three to six feet. If you prefer, this perennial wildflower can even be cultivated as a shrub-like form. It is easily grown from seed, which is available from Ernst Conservation Seeds of Meadville, Pennsylvania, as well as numerous other vendors. And don’t forget to give Wild Senna’s two close relatives, Partridge Pea and Maryland Senna, a try as well. They attract the same species of butterflies and are just as easy to grow. You’ll like ’em.
Cloudless Sulphur butterflies from populations in the south colonize the Lower Susquehanna River Watershed each summer in varying numbers.Close-up view of a Wild Senna flower cluster. Typical of members of the pea family, the seeds of all native Senna species develop within pods after blooming and are sought after by wildfowl (Galliformes), particularly Northern Bobwhite.Partridge Pea with flowers and seed pods in the susquehannawildlife.net garden. Smaller than the other Senna species, Partridge Pea reaches a height of just two to three feet.
If you’re feeling the need to see summertime butterflies and their numbers just don’t seem to be what they used to be in your garden, then plan an afternoon visit to the Boyd Big Tree Preserve along Fishing Creek Valley Road (PA 443) just east of U.S. 22/322 and the Susquehanna River north of Harrisburg. The Pennsylvania Department of Conservation and Natural Resources manages the park’s 1,025 acres mostly as forested land with more than ten miles of trails. While located predominately on the north slope of Blue Mountain, a portion of the preserve straddles the crest of the ridge to include the upper reaches of the southern exposure.
A grove of American Chestnuts (Castanea dentata) planted at Boyd Big Tree Preserve is part of a propagation program working to restore blight-resistant trees to Pennsylvania and other areas of their former range which included the Appalachians and the upper Ohio River watershed.
Fortunately, one need not take a strenuous hike up Blue Mountain to observe butterflies. Open space along the park’s quarter-mile-long entrance road is maintained as a rolling meadow of wildflowers and cool-season grasses that provide nectar for adult butterflies and host plants for their larvae.
A view looking north at the butterfly meadow and entrance road at Boyd Big Tree Preserve Conservation Area. Second Mountain is in the background.Mowed paths follow the entrance road and a portion of the perimeter of the meadow allowing visitors a chance to wander among the waist-high growth to see butterflies, birds, and blooming plants at close range without trampling the vegetation or risking exposure to ticks.A Silver-spotted Skipper feeding on nectar from the flowers of Indian Hemp (Apocynum cannabinum). Like the milkweeds, Indian Hemp is a member of the dogbane family (Apocynaceae).An Eastern Tiger Swallowtail feeding on Common Milkweed.A Great Spangled Fritillary (Speyeria cybele) on Common Milkweed.A Black Swallowtail (Papilio polyxenes) feeding on Common Milkweed nectar.A Pipevine Swallowtail (Battus philenor) on Common Milkweed.Another Pipevine Swallowtail on Common Milkweed. Note the hook-shaped row of red-orange spots on the underside of the hindwing.A Pipevine Swallowtail visiting the brilliant blooms of Butterfly Weed, a favorite of a wide variety of pollinators.A Black Swallowtail with damaged wings alights atop a Butterfly Weed flower cluster. Note the pair of parallel rows of red-orange spot on the underside of the hindwing.A Monarch feeding on nectar from the flowers of Butterfly Weed.A mating pair of Eastern Tailed Blues on a Timothy (Phleum pratense) spike.A male Great Spangled Fritillary (right) pursuing a female.Butterflies aren’t the only colorful insects patrolling the meadows at Boyd Big Tree Preserve. Dragonflies including Common Green Darners are busily pursuing prey, particularly small flying insects like mosquitos, gnats, and flies.Dragonflies themselves can become prey and are much sought after by Broad-winged Hawks. This very vocal juvenile gave us several good looks as it ventured from the forest into the skies above the upper meadow during midday. It wasn’t yet a good enough flier to snag a dragonfly, but it will have plenty of opportunities for practice during its upcoming fall migration which, for these Neotropical raptors, will get underway later this month.
Do yourself a favor and take a trip to the Boyd Big Tree Preserve Conservation Area. Who knows? It might actually inspire you to convert that lawn or other mowed space into much-needed butterfly/pollinator habitat.
While you’re out, you can identify your sightings using our photographic guide—Butterflies of the Lower Susquehanna River Watershed—by clicking the “Butterflies” tab at the top this page. And while you’re at it, you can brush up on your hawk identification skills ahead of the upcoming migration by clicking the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab. Therein you’ll find a listing and descriptions of hawk watch locations in and around the lower Susquehanna region. Plan to visit one or more this autumn!
Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge. At Bombay Hook, they don’t waste your money mowing grass. Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools. Birds there can often be observed at close range. The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools. Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.Two more Short-billed Dowitchers on the way in.Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed. Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge. Mallards, American Avocets, and egrets are gathered on the shore.More Short-billed Dowitchers arriving to feed in Raymond Pool.Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool. A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.American Avocets probe the muddy bottom of Raymond Pool.Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early. Many migrating Dunlin linger at Bombay Hook into October and even November.This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp PoolA Greater Yellowlegs at Bear Swamp Pool.A Caspian Tern patrolling Raymond Pool.The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass. High tide licks at the roots of the cordgrass supporting the temporary domicile.By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice). It is our only dragonfly able to breed in saltwater. Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.Look up! A migrating Bobolink passes over the dike at Shearness Pool.Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool. Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river. The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century. An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers. Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.A Great Egret prowling Shearness Pool.A Snowy Egret in Bear Swamp Pool.Wood Ducks in Bear Swamp Pool.A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.
As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.
Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay. We found just this one during our visit to Bombay Hook. Man has certainly decimated populations of this ancient crustacean during recent decades.As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.A male Atlantic Marsh Fiddler Crab peers from its den.Herons and egrets including this Great Egret are quite fond of fiddler crabs. As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.An Osprey patrols the vast tidal areas opposite Shearness Pool.
No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook. This one was in shrubby growth along the dike at the north end of Shearness Pool.These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries. The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware. The seed balls are a favorite winter food of goldfinches and siskins.Finis Pool has no frontage on the tidal marsh but is still worth a visit. It lies along a spur road on the tour route and is located within a deciduous coastal plain forest. Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.The White-tailed Deity is common along the road to Finis Pool.Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well. This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.
We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.
Here’s a look at some native plants you can grow in your garden to really help wildlife in late spring and early summer.
The showy bloom of a Larger Blue Flag (Iris versicolor) and the drooping inflorescence of Soft Rush (Juncus effusus). These plants favor moist soils in wetlands and damp meadows where they form essential cover and feeding areas for insects, amphibians, and marsh birds. Each is an excellent choice for helping to absorb nutrients in a rain garden or stream-side planting. They do well in wet soil or shallow water along the edges of garden ponds too.The fruits of Smooth Shadbush (Amelanchier laevis), also known as Allegheny Serviceberry, Smooth Serviceberry, or Smooth Juneberry, ripen in mid-June and are an irresistible treat for catbirds, robins, bluebirds, mockingbirds, and roving flocks of Cedar Waxwings.Also in mid-June, the fragrant blooms of Common Milkweed attract pollinators like Eastern Carpenter Bees,……Honey Bees,……and butterflies including the Banded Hairstreak (Satyrium calanus). In coming weeks, Monarch butterflies will find these Common Milkweed plants and begin laying their eggs on the leaves. You can lend them a hand by planting milkweed species (Asclepias) in your garden. Then watch the show as the eggs hatch and the caterpillars begin devouring the foliage. Soon, they’ll pupate and, if you’re lucky, you’ll be able to watch an adult Monarch emerge from a chrysalis!
Let’s have a look at some of the magnificent native wildflowers blooming on this Mother’s Day in forests and thickets throughout the Lower Susquehanna River Watershed.
Rue Anemone (Thalictrum thalictroides) is a fragile little wildflower of open woods.The Blackhaw (Viburnum prunifolium) is a shrub of thickets and forest borders. The flower clusters mature into an abundance of berries relished by birds and other wildlife.A large Black Cherry tree adorned in white blossoms can be pleasing to the eye and very beneficial for pollinating insects.The Common Pawpaw is a small tree of damp well-drained soils. Where conditions are ideal, it may form clonal colonies in the forest understory. The deep maroon blooms mature into banana-like fruits, hence its alternate common name “Custard Apple”.The Mayapple or Mandrake (Podophyllum peltatum) commonly blankets the forest floor. It, like other ephemeral wildflowers, is vulnerable to invasions by non-native shrubs that leaf out earlier in spring than native species and rob them of sunlight during their already abbreviated growing season.
Aren’t they spectacular?
Why not take a stroll to have a look at these and other plants you can find growing along a nearby woodland trail? You could check out some place new and get a little exercise too!
If you go, please tread lightly, take your camera, mind your manners, and…
Potting small bare root tree and shrub seedlings in a compost mixture allows you to nurture them with some tender loving care in your home nursery at least through the summer. The extra attention they’ll get allows you to plant them as larger specimens this autumn or thereafter. We’ve found that giving native species this quick head start in the nursery significantly improves their survival rate in the field.
Have you noticed a purple haze across the fields right now? If so, you may have wondered, “What kind of flowers are they?”
A purple haze of color stretches across fields not already green with cold-season crops like winter wheat.
Say hello to Purple Dead Nettle (Lamium purpureum), a non-native invasive species that has increased its prevalence in recent years by finding an improved niche in no-till cropland. Purple Dead Nettle, also known as Red Dead Nettle, is native to Asia and Europe. It has been a familiar early spring “weed” in gardens, along roadsides, and in other disturbed ground for decades.
Purple Dead Nettle owes its new-found success to the timing of its compressed growing season. Its tiny seeds germinate during the fall and winter, after crops have been harvested and herbicide application has ended for the season. The plants flower early in the spring and are thus particularly attractive to Honey Bees and other pollinators looking for a source of energy-rich nectar as they ramp up activity after winter lock down. In many cases, Purple Dead Nettle has already completed its flowering cycle and produced seeds before there is any activity in the field to prepare for planting the summer crop. The seeds spend the warmer months in dormancy, avoiding the hazards of modern cultivation that expel most other species of native and non-native plants from the agricultural landscape.
Flowering Purple Dead Nettle as a volunteer cover crop among last year’s corn stubble in a no-till field.Like the flowers of orchids, Purple Hedge Nettle blossoms are described as yoke shaped or bilateral (zygomorphic). Psychedelic experiences are produced only through observation, not by ingestion. A member of the mint family, its edible young leaves and tops have nutritional value, making a unique addition to salads and soups.We call them “weeds”, but what do we know? Purple Dead Nettle, Common Dandelion, and Shepherd’s Purse (Capsella bursa-pastoris), three edible non-native invasive species with similar life cycles seen here flowering along a rural road among fields where intensive farming is practiced. Shepard’s Purse, like many members of the mustard family, is already producing seeds at the bottom of the flower cluster by the time the uppermost buds open for business.In preparation for seeding of a warm-season crop, herbicide is applied on a no-till field to kill Shepherd’s Purse and other cool-season plants. To help prevent sediment and nutrient discharge from lands where high-intensity agriculture is practiced, no-till methods are used to reduce runoff from the areas of bare soil that would otherwise be created by traditional plowing.
While modern farming has eliminated a majority of native plant and animal species from agricultural lands of the lower Susquehanna valley, its crop management practices have simultaneously invited vigorous invasion by a select few non-native species. High-intensity farming devotes its acreage to providing food for a growing population of people—not to providing wildlife habitat. That’s why it’s so important to minimize our impact on non-farm lands throughout the remainder of the watershed. If we continue subdividing, paving, and mowing more and more space, we’ll eventually be living in a polluted semi-arid landscape populated by little else but non-native invasive plants and animals. We can certainly do better than that.
This linear grove of mature trees, many of them nearly one hundred years old, is a planting of native White Oaks (Quercus alba) and Swamp White Oaks (Quercus bicolor).
Imagine the benefit of trees like this along that section of stream you’re mowing or grazing right now. The Swamp White Oak in particular thrives in wet soils and is available now for just a couple of bucks per tree from several of the lower Susquehanna’s County Conservation District Tree Sales. These and other trees and shrubs planted along creeks and rivers to create a riparian buffer help reduce sediment and nutrient pollution. In addition, these vegetated borders protect against soil erosion, they provide shade to otherwise sun-scorched waters, and they provide essential wildlife habitat. What’s not to love?
Autumn leaf of a Swamp White Oak
The following native species make great companions for Swamp White Oaks in a lowland setting and are available at bargain prices from one or more of the County Conservation District Tree Sales now underway…
The Red Maple is an ideal tree for a stream buffer project. They do so well that you should limit them to 10% or less of the plants in your project so that they don’t overwhelm slower-growing species.The River Birch (Betula nigra) is a multi-trunked tree of lowlands. Large specimens with arching trunks help shade waterways and provide a source of falling insects for surface-feeding fish. Its peeling bark is a distinctive feature.The Common Winterberry with its showy red winter-time fruit is a slow-growing shrub of wet soils. Only female specimens of this deciduous holly produce berries, so you need to plant a bunch to make sure you have both genders for successful pollination.An American Robin feeding on Common Winterberry.Common Spicebush is a shrub of moist lowland soils. It is the host plant for the Spicebush Swallowtail butterfly and produces small red berries for birds and other wildlife. Plant it widely among taller trees to provide native vegetation in the understory of your forest.Common Spicebush foliage and berries in the shade beneath a canopy of tall trees.The Common Pawpaw a small shade-loving tree of the forest understory.Common Pawpaw is a colony-forming small tree which produces a fleshy fruit. It is the host plant for the caterpillars of the Zebra Swallowtail.The Buttonbush is a shrub of wet soils. It produces a round flower cluster, followed by this globular seed cluster.And don’t forget the Eastern Sycamore, the giant of the lowlands. At maturity, the white-and-tan-colored bark on massive specimens makes them a spectacular sight along stream courses and river shores. Birds ranging from owls, eagles, and herons to smaller species including the Yellow-throated Warbler rely upon them for nesting sites.Yellow-crowned Night Herons, an endangered species in Pennsylvania, nesting in an Eastern Sycamore.
So don’t mow, do something positive and plant a buffer!
Act now to order your plants because deadlines are approaching fast. For links to the County Conservation District Tree Sales in the Lower Susquehanna River Watershed, see our February 18th post.
County Conservation District Tree Sales are underway throughout the Lower Susquehanna River Watershed. Now is the time to order for pickup in April. The prices are a bargain and the selection is fabulous. For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue). And don’t forget to order bundles of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local birds. They’re only $12.00 for a bundle of 10—can’t beat that deal!
Pickup on: Thursday, April 20, 2023 or Friday, April 21, 2023
Don’t mow it. Plant a meadow or pollinator garden instead.Both Cumberland and Perry Counties are offering a native warm-season grass and wildflower seed mix for planting your own meadow or pollinator garden. Perry County is also taking orders for a seed mix specifically formulated to grow plants for attracting hummingbirds and butterflies.
Pickup on: Thursday, April 20, 2023 or Friday, April 21, 2023
The Eastern Hemlock, Pennsylvania’s official state tree, is an excellent choice for addition to your landscape or reforestation project. It tolerates rocky soils and its cones are an prime source of food for birds ranging from chickadees to finches.
The handsome yet underused Northern Red Oak is a sturdy long-lived native tree that is ideal for street-side, lawn, and reforestation plantings. In spring, it can be a magnet for migrating Neotropical birds when its flowers attract a wide variety of tiny insects to its upper reaches. Unlike many other oaks, this species is a relatively fast grower.
In autumn, even after the bright red foliage is gone, the berries of mature Black Gum (Nyssa sylvatica) trees attract a wide variety of birds like this Pileated Woodpecker. The Lebanon County Conservation District is offering Black Gum, also known as Black Tupelo, during their 2023 tree sale. Why not order and plant a half dozen or more?
The Perry County Conservation District is not only offering plants during this year’s sale, you can also purchase bluebird nest boxes for just $12.00 each!For less than the cost of one year of mowing, this stream corridor in Conewago Township, Dauphin County was reforested by the owner with hundreds of native trees, the majority purchased through County Conservation District Tree Sale events spanning a period of several years. By replacing bare soil and mowed areas, the riparian buffer created by these plantings has significantly reduced the nutrient and sediment loads that were polluting the small stream therein known as Brill’s Run. With determination and not a lot of money, you can do it too.But don’t forget the Eastern White Pines!
From a lookout atop an Eastern Hemlock, a Northern Mockingbird maintains a vigil over a garden full of berry-producing plants. To assure their survival during cold weather, these bold birds will vigorously defend winter food supplies on hollies, viburnums, poison-ivy, bittersweet, and other fruit-producing trees, shrubs, and vines, often swooping in to startle and flush birds and other animals that approach its stores too closely.
By November, as insects become more difficult to come by, migrating Yellow-rumped Warblers are seldom found far from a supply of berries. This one was in the vicinity of the white fruits of Poison Ivy vines.
Many cultivars of ornamental maples for the home garden are derived from this native species, and it’s easy to see why. Despite the variation in the foliage of each, every tree in this view is a Red Maple (Acer rubrum).The Red Maple is the most common and widespread tree in all of eastern North America. It is easily recognized by its three-lobed leaves. Not only do the fall foliage colors vary among a population of trees, but some individual specimens have a well-stocked palette of their own. This collection came from a single Red Maple. Visit a forested area today and check them out!
The harmless, herbivorous Northern Walkingstick (Diapheromera femorata), also known as the Common Walkingstick, is well camouflaged when among the twigs and stems of deciduous trees. As foliage drops in autumn, these wingless insects often descend to ground level and sometimes venture into view. This individual was found in the vicinity of ideal habitat: a portion of forest including Black Cherry and Black Oak (Quercus velutina) trees. Juvenile Northern Walkingsticks feed on the leaves of the former, adults on those of the latter.
Earlier today, these migrants were found feeding on berries along the edge of a forest clearing in northern Lebanon County, Pennsylvania. Can you find the three Hermit Thrushes among the early successional growth seen here? For extra credit, identify the three species of berry-producing pioneer plants that are shown. For additional credit, which one of these plants is a non-native invasive species? Click the image to see how you did.
During autumn, fuzzy caterpillars crossing roads and trails are a familiar sight throughout the Lower Susquehanna River Watershed. In oak-hickory forests you may encounter the stunningly beautiful Hickory Tussock Moth (Lophocampa caryae) caterpillar, but don’t touch it; the spine-like bristles are mildly venomous. Some people react with skin irritation and rashes after contact. This native species rarely defoliates its host trees. When it does, it’s usually just weeks before the deciduous leaves would fall, so the overall health of the plant is little affected. To balance the population, there are numerous avian and terrestrial predators that feed upon both the larval and adult stages of the Hickory Tussock Moth.
A small butterfly with flights extending into October, the Horace’s Duskywing (Erynnis horatius) is an easily overlooked inhabitant of forest clearings in the vicinity of oaks, particularly Chinkapin Oak (Quercus muehlenbergii), a favorite larval foodplant.
Having arrived yesterday during a break in the storm, this Ruby-throated Hummingbird spent a rain soaked fourth of October feeding on nectar among the abundant blooms covering the Mexican Cigar plants at susquehannawildlife.net headquarters. When the remains of Hurricane Ian finally depart the Lower Susquehanna River Watershed, it’ll resume its flight to the tropics for the winter. And yes, October is late for Ruby-throated Hummingbirds in our region. Throughout autumn, each passing day increases the likelihood of a hummingbird found here being one of the hardy species from western North America, so look carefully at any you see this fall.