


LIFE IN THE LOWER SUSQUEHANNA RIVER WATERSHED
A Natural History of Conewago Falls—The Waters of Three Mile Island
If you’ve visited Middle Creek Wildlife Management Area anytime during the past month, you may have noticed quite a bit of activity around the large pole-mounted nest boxes placed out in the open fields.








LEARN HOW LAND MANAGERS UTILIZE PRESCRIBED FIRE
This coming Saturday, April 18, 2026, beginning at 10 A.M. (rain date April 25), the Pennsylvania Game Commission is hosting a “Prescribed Fire Festival” at Middle Creek Wildlife Management Area. Be certain to come out to the visitor’s center at 100 Museum Road, Stevens, PA, for this event. Land managers will be there to answer questions and to explain the planning and preparations involved in overseeing a prescribed burn. There will be guided walks of habitats preserved using fire of varying intensities. You’ll see the equipment and protective clothing used by certified personnel to administer a live prescribed fire burn right before your eyes. Then you can have lunch—food trucks will be available on site.

After the burn demonstration, why not go for a walk or drive around the refuge. You can take note of how grassland and early successional plant communities are responding to previous doses of prescribed fire…


And how the grassland animals respond as well…








The fate of an avian predator such as a kestrel lies at the mercy of the fate of its quarry. Because, you see, the sun’s energy, after being converted to chemical energy by photosynthesizing plants, flows upward through the trophic levels of the food chain—herbivores (primary consumers) such as rodents and insects to carnivores (secondary and tertiary consumers) including kestrels. Grasslands, when abundant and diverse, are correspondingly abundant and diverse with small mammals and insects and will therefore support thriving populations of American Kestrels and other predators. These secondary and sometimes tertiary consumers fulfill a role in cultivating healthier populations of their prey—the primary and secondary consumers in the food web—as a balanced component of a flourishing grassland ecosystem. Sparse and fragmented grasslands, on the other hand, beget negligible small mammal and insect populations, are stricken with broken food webs, and champion few if any American Kestrels or other predators. If the land it occupies is neat, tidy, manicured, exploited, or sprayed sterile and dead, the energy flow cycle of the ecosystem is dead as well. There’s nothing animal introductions, reintroductions, rescues, culling, stocking, or harvesting can do about it, because in the end, it’s all about the habitat.
With another round of single-digit and possibly sub-zero temperatures on the way, birds and other wildlife are taking advantage of a break in the extreme conditions to re-energize. During the past day, these species were among those attracted to the food and cover provided by the habitat plantings in the susquehannawildlife.net headquarters garden…




Here are a few more late-season migrants you might currently see passing through the lower Susquehanna valley. Where adequate food and cover are available, some may remain into part or all of the winter…






Having experienced our first frost throughout much of the lower Susquehanna valley last night, we can look forward to seeing some changes in animal behavior and distribution in the days and weeks to come. Here are a few examples…







Here’s a short preview of some of the finds you can expect during an outing in the Lower Susquehanna River Watershed’s forests this week…





















Neotropical birds are fairly well acquainted with repetitive periods of thundershowers. With that in mind, we decided not to waste this stormy Tuesday by remaining indoors.












We hope you enjoyed our walk in the rain as much as we did. If you venture out on a similar excursion, please remember this. The majority of the wild animals around us have busy lives, particularly at this time of year. Most don’t take a day off just because it rains—that includes ticks.

As the Flowering Dogwoods remind us, it’s time for adult sea-run shad and other fishes to ascend the Susquehanna to spawn. So yep, we’re off to Fisherman’s Park on the river’s west shoreline below Conowingo Dam in Harford County, Maryland, to check it out.



























Here’s a look at six native shrubs and trees you can find blooming along forest edges in the lower Susquehanna valley right now.







Local old timers might remember hearing folklore that equates the northward advance of the blooming of the Flowering Dogwoods with the progress of the American Shad’s spring spawning run up the river. While this is hardly a scientific proclamation, it is likely predicated on what had been some rather consistent observation prior to the construction of the lower Susquehanna’s hydroelectric dams. In fact, we’ve found it to be a useful way to remind us that it’s time for a trip to the river shoreline below Conowingo Dam to witness signs of the spring fish migration each year. We’re headed that way now and will summarize our sightings for you in days to come.

Here are five common forest flowers that the average visitor to these environs may easily overlook during an early April visit.





Be certain to get out and enjoy this year’s blooming seasons of our hundreds of varieties of flowering plants. But, particularly when it comes to native species,…
Our wildlife has been having a tough winter. The local species not only contend with cold and stormy weather, but they also need to find food and shelter in a landscape that we’ve rendered sterile of these essentials throughout much of the lower Susquehanna valley’s farmlands, suburbs, and cities.
Planting trees, shrubs, flowers, and grasses that benefit our animals can go a long way, often turning a ho-hum parcel of property into a privately owned oasis. Providing places for wildlife to feed, rest, and raise their young can help assure the survival of many of our indigenous species. With a little dedication, you can be liberated from the chore of manicuring a lawn and instead spend your time enjoying the birds, mammals, insects, and other creatures that will visit your custom-made habitat.

Fortunately for us, our local county conservation districts are again conducting springtime tree sales offering a variety of native and beneficial cultivated plants at discount prices. Listed here are links to information on how to pre-order your plants for pickup in April. Click away to check out the species each county is offering in 2025!
Cumberland County Conservation District Annual Tree Seedling Sale—
Orders due by: Friday, March 21, 2025
Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025

Dauphin County Conservation District Seedling Sale—
Orders due by: Monday, March 17, 2025
Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025

Franklin County Conservation District Tree Seedling Sale—
Orders due by: Wednesday, March 19, 2025
Pickup on: Thursday, April 24, 2025

Lancaster County Annual Tree Seedling Sale—
Orders due by: Friday, March 7, 2025
Pickup on: Friday, April 11, 2025

Lebanon County Conservation District Tree and Plant Sale—
Orders due by: Monday, March 3, 2025
Pickup on: Friday, April 18, 2025




York County Conservation District Seedling Sale—
Orders due by: March 15, 2025
Pickup on: Thursday, April 10, 2025


If you live in Adams County, Pennsylvania, you may be eligible to receive free trees and shrubs for your property from the Adams County Planting Partnership (Adams County Conservation District and the Watershed Alliance of Adams County). These trees are provided by the Chesapeake Bay Foundation’s Keystone 10-Million Trees Partnership which aims to close a seven-year project in 2025 by realizing the goal of planting 10 million trees to protect streams by stabilizing soils, taking up nutrients, reducing stormwater runoff, and providing shade. If you own property located outside of Adams County, but still within the Chesapeake Bay Watershed (which includes all of the Susquehanna, Juniata, and Potomac River drainages), you still may have an opportunity to get involved. Contact your local county conservation district office or watershed organization for information.


We hope you’re already shopping. Need help making your selections? Click on the “Trees, Shrubs, and Woody Vines” tab at the top of this page to check out Uncle Tyler Dyer’s leaf collection. He has most of the species labelled with their National Wetland Plant List Indicator Rating. You can consult these ratings to help find species suited to the soil moisture on your planting site(s). For example: if your site has sloped upland ground and/or the soils sometimes dry out in summer, select plants with a rating such as UPL or FACU. If your planting in soils that remain moist or wet, select plants with the OBL or FACW rating. Plants rated FAC are generally adaptable and can usually go either way, but may not thrive or survive under stressful conditions in extremely wet or dry soils.
NATIONAL WETLAND PLANT LIST INDICATOR RATING DEFINITIONS
Using these ratings, you might choose to plant Pin Oaks (FACW) and Swamp White Oaks (FACW) in your riparian buffer along a stream; Northern Red Oaks (FACU) and White Oaks (FACU) in the lawn or along the street, driveway, or parking area; and Chestnut Oaks (UPL) on your really dry hillside with shallow soil. Give it a try.
Each spring and fall, Purple Finches are regular migrants through the Lower Susquehanna River Watershed. Northbound movements usually peak in April and early May. During the summer, these birds nest primarily in cool coniferous forests to our north. Then, in October and November each year, they make another local appearance on their way to wintering grounds in the southeastern United States. A significant population of Purple Finches remains to our north through the colder months, inhabiting spruce-pine and mixed forests from the Great Lakes east through New England and southeastern Canada. This population can be irruptive, moving south in conspicuous numbers to escape inclement weather, food shortages, and other environmental conditions. Every few years, these irruptive birds can be found visiting suburbs, parks, and feeding stations—sometimes lingering in areas not often visited by Purple Finches. Right now, Purple Finches in flocks larger than those that moved through earlier in the fall are being seen throughout the lower Susquehanna valley. Snow and blustery weather to our north may be prompting these birds to shift south for a visit. Here are some looks at members of a gathering of more than four dozen Purple Finches we’ve been watching in Lebanon County this month…







During the past week, Uncle Tyler Dyer has been out searching for autumn leaves to add to his collection. One of the species he had not encountered in previous outings was the American Elm (Ulmus americana), so he made a special trip to see a rare mammoth specimen in a small neighborhood park (Park Place) along Chestnut Street between 5th and Quince Streets in Lebanon, Pennsylvania.


There’s still time to get out and see autumn foliage. With warmer weather upon us—at least temporarily—it’s a good time to go for a stroll. Who knows, you might find some spectacular leaves like these collected by Uncle Ty earlier this week. All were found adorning native plants!






Back on March 24th, we took a detailed look at the process involved in administering prescribed fire as a tool for managing grassland and early successional habitat. Today we’re going turn back the hands of time to give you a glimpse of how the treated site fared during the five months since the controlled burn. Let’s go back to Middle Creek Wildlife Management Area for a photo tour to see how things have come along…















Elsewhere around the refuge at Middle Creek, prescribed fire and other management techniques are providing high-quality grassland habitat for numerous species of nesting birds…



We hope you enjoyed this short photo tour of grassland management practices. Now, we’d like to leave you with one last set of pictures—a set you may find as interesting as we found them. Each is of a different Eastern Cottontail, a species we found to be particularly common on prescribed fire sites when we took these images in late May. The first two are of the individuals we happened to be able to photograph in areas subjected to fire two months earlier in March. The latter two are of cottontails we happened to photograph elsewhere on the refuge in areas not in proximity to ground treated with a prescribed burn or exposed to accidental fire in recent years.


These first two rabbits are living the good life in a warm-season grass wonderland.


Oh Deer! Oh Deer! These last two rabbits have no clock to track the time; they have only ticks. Better not go for a stroll with them Alice—that’s no wonderland! I know, I know, it’s time to go. See ya later.
The Aphrodite Fritillary (Speyeria aphrodite), also known simply as the Aphrodite, is a brush-footed butterfly of deciduous, coniferous, and mixed forests. We found this female in a grassland margin between woodlots where prescribed fire was administered during the autumn of 2022 to reduce accumulations of natural fuels and an overabundance of invasive vegetation. A goal of the burn was to promote the growth of native species including the violets (Viola species) favored as larval host plants by this and other fritillaries.

By the time these adult butterflies make their reproductive flights in late summer, the violets that serve as larval host plants have gone dormant. To find patches of ground where the violets will come to life in spring, the female Aphrodite Fritillary has an ability to sense the presence of dormant roots, probably by smell. Upon finding an area where suitable violets will begin greening up next year, she’ll deposit her eggs. The eggs overwinter, then hatch to feed on the tender new violet leaves of spring.



Do you recall our “Photo of the Day” from seven months ago…

Well, here’s what that site looks like today…


And there are pollinating insects galore, most notably butterflies…


















Why on earth would anyone waste their time, energy, and money mowing grass when they could have this? Won’t you please consider committing graminicide this fall? That’s right, kill that lawn—at least the majority of it. Then visit the Ernst Seed website, buy some “Native Northeast Wildlflower Mix” and/or other blends, and get your meadow planted in time for the 2025 growing season. Just think of all the new kinds of native plants and animals you’ll be seeing. It could change your life as well as theirs.

Resilient to the pressures of flooding, ice scour, drought, and oft times really poor water quality, Water Willow (Dianthera americana, formerly Justicia americana) is the most common herbaceous plant on the Susquehanna’s non-forested alluvial islands. Yet, few know this native wildflower by name or reputation.



The spring of 2024 has been very kind to our beds of Water Willow. Rainfall in the Susquehanna watershed has been frequent enough to maintain river levels just high enough to keep the roots of the plants wet. During the interludes in storm activity, dry spells have rolled back any threat of flooding on the river’s main stem, thus eliminating chances of submerging the plants in muddy water and preventing the sun from keeping them warm, happy, and flowering early. Thundershowers throughout the basin earlier this week have now raised the river a few inches to inundate the base of the plants and make mats of Water Willow favorable places for newly hatched fry and other young fish to take refuge while they grow. Here’s a look…









By now you’ve come to appreciate the importance of Water Willow to the sustainability of our populations of fish and other aquatic life. Like similar habitat features that reduce sediment runoff and nutrient pollution, undisturbed stands of terrestrial, emergent, and submerged native plant species are essential to the viability of our freshwater food webs.

Homo sapiens owes much of its success as a species to an acquired knowledge of how to make, control, and utilize fire. Using fire to convert the energy stored in combustible materials into light and heat has enabled humankind to expand its range throughout the globe. Indeed, humans in their furless incomplete mammalian state may have never been able to expand their populations outside of tropical latitudes without mastery of fire. It is fire that has enabled man to exploit more of the earth’s resources than any other species. From cooking otherwise unpalatable foods to powering the modern industrial society, fire has set man apart from the rest of the natural world.
In our modern civilizations, we generally look at the unplanned outbreak of fire as a catastrophe requiring our immediate intercession. A building fire, for example, is extinguished as quickly as possible to save lives and property. And fires detected in fields, brush, and woodlands are promptly controlled to prevent their exponential growth. But has fire gone to our heads? Do we have an anthropocentric view of fire? Aren’t there naturally occurring fires that are essential to the health of some of the world’s ecosystems? And to our own safety? Indeed there are. And many species and the ecosystems they inhabit rely on the periodic occurrence of fire to maintain their health and vigor.

Man has been availed of the direct benefits of fire for possibly 40,000 years or more. Here in the Lower Susquehanna River Watershed, the earliest humans arrived as early as 12,000 years ago—already possessing skills for using fire. Native plants and animals on the other hand, have been part of the ever-changing mix of ecosystems found here for a much longer period of time—millions to tens of millions of years. Many terrestrial native species are adapted to the periodic occurrence of fire. Some, in fact, require it. Most upland ecosystems need an occasional dose of fire, usually ignited by lightning (though volcanism and incoming cosmic projectiles are rare possibilities), to regenerate vegetation, release nutrients, and maintain certain non-climax habitat types.
But much of our region has been deprived of natural-type fires since the time of the clearcutting of the virgin forests during the eighteenth and nineteenth centuries. This absence of a natural fire cycle has contributed to degradation and/or elimination of many forest and non-forest habitats. Without fire, a dangerous stockpile of combustible debris has been collecting, season after season, in some areas for a hundred years or more. Lacking periodic fires or sufficient moisture to sustain prompt decomposition of dead material, wildlands can accumulate enough leaf litter, thatch, dry brush, tinder, and fallen wood to fuel monumentally large forest fires—fires similar to those recently engulfing some areas of the American west. So elimination of natural fire isn’t just a problem for native plants and animals, its a potential problem for humans as well.

To address the habitat ailments caused by a lack of natural fires, federal, state, and local conservation agencies are adopting the practice of “prescribed fire” as a treatment to restore ecosystem health. A prescribed fire is a controlled burn specifically planned to correct one or more vegetative management problems on a given parcel of land. In the Lower Susquehanna River Watershed, prescribed fire is used to…
Let’s look at some examples of prescribed fire being implemented right here in our own neighborhood…

























In Pennsylvania, state law provides landowners and crews conducting prescribed fire burns with reduced legal liability when the latter meet certain educational, planning, and operational requirements. This law may help encourage more widespread application of prescribed fire in the state’s forests and other ecosystems where essential periodic fire has been absent for so very long. Currently in the Lower Susquehanna River Watershed, prescribed fire is most frequently being employed by state agencies on state lands—in particular, the Department of Conservation and Natural Resources on State Forests and the Pennsylvania Game Commission on State Game Lands. Prescribed fire is also part of the vegetation management plan at Fort Indiantown Gap Military Reservation and on the land holdings of the Hershey Trust. Visitors to the nearby Gettysburg National Military Park will also notice prescribed fire being used to maintain the grassland restorations there.
For crews administering prescribed fire burns, late March and early April are a busy time. The relative humidity is often at its lowest level of the year, so the probability of ignition of previous years’ growth is generally at its best. We visited with a crew administering a prescribed fire at Middle Creek Wildlife Management Area last week. Have a look…

















Prescribed burns aren’t a cure-all for what ails a troubled forest or other ecosystem, but they can be an effective remedy for deficiencies caused by a lack of periodic episodes of naturally occurring fire. They are an important option for modern foresters, wildlife managers, and other conservationists.
It’s that time of year. Your local county conservation district is taking orders for their annual tree sale and it’s a deal that can’t be beat. Order now for pickup in April.
The prices are a bargain and the selection includes the varieties you need to improve wildlife habitat and water quality on your property. For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue). And don’t forget to order packs of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local native birds. They’re only $12.00 for a bundle of 10.

Cumberland County Conservation District Annual Tree Seedling Sale—
Orders due by: Friday, March 22, 2024
Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024



Dauphin County Conservation District Seedling Sale—
Orders due by: Monday, March 18, 2024
Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024

Lancaster County Annual Tree Seedling Sale—
Orders due by: Friday, March 8, 2024
Pickup on: Friday, April 12, 2024






Lebanon County Conservation District Tree and Plant Sale—
Orders due by: Friday, March 8, 2024
Pickup on: Friday, April 19, 2024




Perry County Conservation District Tree Sale—
Orders due by: Sunday, March 24, 2024
Pickup on: Thursday, April 11, 2024

Again this year, Perry County is offering bluebird nest boxes for sale. The price?—just $12.00.

York County Conservation District Seedling Sale—
Orders due by: Friday, March 15, 2024
Pickup on: Thursday, April 11, 2024

To get your deciduous trees like gums, maples, oaks, birches, and poplars off to a safe start, conservation district tree sales in Cumberland, Dauphin, Lancaster, and Perry Counties are offering protective tree shelters. Consider purchasing these plastic tubes and supporting stakes for each of your hardwoods, especially if you have hungry deer in your neighborhood.

There you have it. Be sure to check out each tree sale’s web page to find the selections you like, then get your order placed. The deadlines will be here before you know it and you wouldn’t want to miss values like these!
Since Tuesday’s snow storm, the susquehannawiildlife.net headquarters garden continues to bustle with bird activity.




Today, there arrived three species of birds we haven’t seen here since autumn. These birds are, at the very least, beginning to wander in search of food. Then too, these may be individuals creeping slowly north to secure an advantage over later migrants by being the first to establish territories on the most favorable nesting grounds.



They say the early bird gets the worm. More importantly, it gets the most favorable nesting spot. What does the early birder get? He or she gets out of the house and enjoys the action as winter dissolves into the miracle of spring. Do make time to go afield and marvel a bit, won’t you? See you there!
In mid-February each year, large numbers of American Robins descend upon the susquehannawildlife.net headquarters garden to feast on the ripe fruits that adorn several species of our native shrubs and trees. This morning’s wet snowfall provided the needed motivation for these birds and others to make today the big day for the annual feeding frenzy.

















With the earth at perihelion (its closest approach to the sun) and with our home star just 27 degrees above the horizon at midday, bright low-angle light offered the perfect opportunity for doing some wildlife photography today. We visited a couple of grasslands managed by the Pennsylvania Game Commission to see what we could find…



















A glimpse of the rowdy guests crowding the Thanksgiving Day dinner table at susquehannawildlife.net headquarters…









Where should you go this weekend to see vibrantly colored foliage in our region? Where are there eye-popping displays of reds, oranges, yellows, and greens without so much brown and gray? The answer is Michaux State Forest on South Mountain in Adams, Cumberland, and Franklin Counties.
South Mountain is the northern extension of the Blue Ridge Section of the Ridge and Valley Province in Pennsylvania. Michaux State Forest includes much of the wooded land on South Mountain. Within or adjacent to its borders are located four state parks: King’s Gap Environmental Education Center, Pine Grove Furnace State Park, Caledonia State Park, and Mont Alto State Park. The vast network of trails on these state lands includes the Appalachian Trail, which remains in the mountainous Blue Ridge Section all the way to its southern terminus in Georgia.

If you want a closeup look at the many species of trees found in Michaux State Forest, and you want them to be labeled so you know what they are, a stop at the Pennsylvania State University’s Mont Alto arboretum is a must. Located next door to Mont Alto State Park along PA 233, the Arboretum at Penn State Mont Alto covers the entire campus. Planting began on Arbor Day in 1905 shortly after establishment of the Pennsylvania State Forest Academy at the site in 1903. Back then, the state’s “forests” were in the process of regeneration after nineteenth-century clear cutting. These harvests balded the landscape and left behind the combustible waste which fueled the frequent wildfires that plagued reforestation efforts for more than half a century. The academy educated future foresters on the skills needed to regrow and manage the state’s woodlands.
Online resources can help you plan your visit to the Arboretum at Penn State Mont Alto. More than 800 trees on the campus are numbered with small blue tags. The “List of arboretum trees by Tag Number” can be downloaded to tell you the species or variety of each. The interactive map provides the locations of individual trees plotted by tag number while the Grove Map displays the locations of groups of trees on the campus categorized by region of origin. A Founder’s Tree Map will help you find some of the oldest specimens in the collection and a Commemorative Tree Map will help you find dedicated trees. There is also a species list of the common and scientific tree names.






The autumn leaves will be falling fast, so make it a point this weekend to check out the show on South Mountain.
Last October 3rd, a late-season Ruby-throated Hummingbird stopped by the garden at susquehannawildlife.net headquarters to take shelter from a rainy autumn storm. It was so raw and chilly that we felt compelled to do something we don’t normally do—put out the sugar water feeder to supplement the nectar produced by our fall-flowering plants. After several days of constant visits to the feeder and the flowers, our lingering hummer resumed its southbound journey on October 7th.
Fast forward to this afternoon and what do you know, at least two migrating hummingbirds have stopped by to visit the flowers in our garden. This year, we have an exceptional abundance of blooms on some of their favorite plants. In the ponds, aquatic Pickerelweed is topped with purple spikes and we still have bright orange tubular flowers on one of our Trumpet Vines—a full two to three months later than usual.




Remember, keep those feeders clean and the provisions fresh! You’ll be glad you did.