Time to Eat

A glimpse of the rowdy guests crowding the Thanksgiving Day dinner table at susquehannawildlife.net headquarters…

White-breasted Nuthatch
A male White-breasted Nuthatch visits a peanut feeder…
White-breasted Nuthatch
…soon to be joined by a female White-breasted Nuthatch.
Downy Woodpecker
A male Downy Woodpecker gets a bill full of suet.
Carolina Wren
A Carolina Wren nibbles at a peanut.
Eastern Gray Squirrel
An Eastern Gray Squirrel stuffs itself on peanuts dropped by the birds.
Northern Mockingbird
A territorial Northern Mockingbird stands guard over its supply of Common Winterberry fruit.
Eastern Bluebird
To avoid the mockingbird’s aggression, the Eastern Bluebirds opted out of fresh fruit in favor of raisins offered at the feeders.
American Robin
This persistent American Robin has made an art of repeatedly sneaking in to quickly devour a few berries before being chased away by the vigilant mockingbird.
Dark-eyed Junco
After everyone has had their fill, Dark-eyed Juncos clean up the leftovers.

Getting a Head Start on Spring

Recently, we found these Chestnut Oak acorns setting roots into the leaf litter to secure their place among the plants that will turn the forest understory green in the spring.  Individual acorns that germinate soon after falling to the ground in autumn may avoid becoming food for squirrels, turkeys, deer, and other wildlife, thus increasing their chances of surviving to later become adult trees able to produce acorns that pass this quick-development trait to yet another generation of oaks.

Photo of the Day

Blue Wood Aster (Symphyotrichum cordifolium)
Among the hardy wildflowers still in bloom in the forests of the lower Susquehanna valley is the Blue Wood Aster (Symphyotrichum cordifolium), a variable species also known as the Heart-leaved Aster.

Photo of the Day

It looks like Uncle Tyler Dyer had a fun-filled Halloween.  Earlier this afternoon, we caught him drifting off into a nap during his work to make updates to his leaf scans.  As he snoozed away, we found these lying on the floor next to his easy chair and decided to lend a hand by adding them to his collection.  We’ll see how long it takes him to discover our dirty little deed.  You can have a look at our mischief by clicking on the “Trees, Shrubs, and Woody Vines” tab at the top of this page.

If You’re Out Collecting Sweets, It Pays to Look Scary

Only fools mess around with bees, wasps, and hornets as they collect nectar and go about their business while visiting flowering plants.  Relentlessly curious predators and other trouble makers quickly learn that patterns of white, yellow, or orange contrasting with black are a warning that the pain and anguish of being zapped with a venomous sting awaits those who throw caution to the wind.  Through the process of natural selection, many venomous and poisonous animals have developed conspicuously bright or contrasting color schemes to deter would-be predators and molesters from making such a big mistake.

"Red Eft"
The brilliant colors of the “Red Eft”, the terrestrial sub-adult stage of the aquatic Eastern Newt (Notophthalmus viridescens), provide protection not as a form of camouflage, but as a warning to potential predators that “I am inedible” due to the presence of tetrodotoxin, a strong neurotoxin.  Over the generations, natural selection has better enabled the brightest of the individual “Red Efts” to survive to adulthood and reproduce.  Meanwhile, those efts that provided a less obvious visual clue to their toxicity frequently allowed their pursuer to learn of their defense mechanism by the taste-test method.  As one might expect, far fewer of these latter individuals survived to breed and pass along their more cryptic color variation.

Visual warnings enhance the effectiveness of the defensive measures possessed by venomous, poisonous, and distasteful creatures.  Aggressors learn to associate the presence of these color patterns with the experience of pain and discomfort.  Thereafter, they keep their distance to avoid any trouble.  In return, the potential victims of this unsolicited aggression escape injury and retain their defenses for use against yet-to-be-enlightened pursuers.  Thanks to their threatening appearance, the chances of survival are increased for these would-be victims without the need to risk death or injury while deploying their venomous stingers, poisonous compounds, or other defensive measures.

European Paper Wasp
Armed and Dangerous  The yellow-and-black color pattern on this European Paper Wasp signals a potential aggressor that they have come upon a social insect and could be struck with a venomous sting.  The warning colors alone may be all the defenses necessary for this wasp to survive an otherwise fatal encounter.

One shouldn’t be surprised to learn that over time, as these aforementioned venomous, poisonous, and foul-tasting critters developed their patterns of warning colors, there were numerous harmless animals living within close association with these species that, through the process of natural selection, acquired nearly identical color patterns for their own protection from predators.  This form of defensive impersonation is known as Batesian mimicry.

Let’s take a look at some examples of Batesian mimicry right here in the Lower Susquehanna River Watershed.

Suppose for a moment that you were a fly.  As you might expect, you would have plenty to fear while you spend your day visiting flowers in search of energy-rich nectar—hundreds of hungry birds and other animals want to eat you.

Greenbottle Fly
You might not hurt a fly, but plenty of other creatures will.  This Greenbottle Fly relies upon speed and maneuverability to quickly flee predators.
Common Flesh Fly (Sarcophaga species)
Like the Greenbottle Fly, the Common Flesh Fly (Sarcophaga species) needs to be constantly vigilant and survives by being quick to the wing.

If you were a fly and you were headed out and about to call upon numerous nectar-producing flowers so you could round up some sweet treats, wouldn’t you feel a whole lot safer if you looked like those venomous bees, wasps, and hornets in your neighborhood?  Wouldn’t it be a whole lot more fun to look scary—so scary that would-be aggressors fear that you might sting them if they gave you any trouble?

Suppose Mother Nature and Father Time dressed you up to look like a bee or a wasp instead of a helpless fly?  Then maybe you could go out and collect sweets without always worrying about the bullies and the brutes, just like these flies of the lower Susquehanna  do…

FLOWER FLIES/HOVER FLIES

The Common Drone Fly (Eristalis Tenax) is a Honey Bee mimic
The Common Drone Fly (Eristalis tenax) is an unarmed Honey Bee mimic.  This one is gathering nectar on goldenrod flowers.
Transverse Flower Fly (Eristalis transversa)
The Transverse Flower Fly (Eristalis transversa) is another bee mimic.  Members of the genus Eristalis scavenge carcasses in aquatic habitats.  Their larvae are known as rat-tailed maggots, a name that references their long siphons used for breathing surface air while submerged in ponds, streams, and wetlands.
Spilomyia species Flower Fly
Flower flies of the genus Spilomyia are convincing mimics of temperamental yellowjacket wasps.
Yellowjacket Hover Fly
The Yellowjacket Hover Fly (Milesia virginiensis) is usually heard long before it is seen.  It will often approach people and persist with a loud buzzing, sounding more like a bee than a bee does.  Scary, isn’t it?
 Maize Calligrapher
The Maize Calligrapher (Toxomerus politus) is a hover fly mimic of wasps.  Seen here on Indiangrass, it is believed to associate primarily with Corn (Zea mays).
The Narrow-headed Marsh Fly (Helophilus fasciatus) is a wasp mimic.
The Narrow-headed Marsh Fly (Helophilus fasciatus) is a wasp mimic.  Like other mimics of hymenopterans, they are important pollinators of flowering plants.
Syrphus species Hoverfly
This hover fly of the genus Syrphus is another wasp mimic.

TACHINID FLIES

The Feather-legged Fly (Trichopoda species) is a wasp mimic.
The Feather-legged Fly (Trichopoda species) is a wasp mimic.   Its larvae are parasitoids of stink bugs and leaf-footed bugs.

BEE FLIES

Bee Fly (Exoprosopa species)
Bee flies of the genus Exoprosopa convincingly resemble bumble bees.
Bee Fly (Exoprosopa species)
The larvae of Exoprosopa bee flies are believed to be parasitic on the larvae of the parasites of bee and wasp larvae that mature in the soil.  Confused yet?

So let’s review.  If you’re a poor defenseless fly and you want to get your fair share of sweets without being gobbled up by the beasts, then you’ve got to masquerade like a strongly armed member of a social colony—like a bee, wasp, or hornet.  Now look scary and go get your treats.  HAPPY HALLOWEEN!

Visit South Mountain for the Brightest Foliage Display You’ve Ever Seen

Where should you go this weekend to see vibrantly colored foliage in our region?  Where are there eye-popping displays of reds, oranges, yellows, and greens without so much brown and gray?  The answer is Michaux State Forest on South Mountain in Adams, Cumberland, and Franklin Counties.

South Mountain is the northern extension of the Blue Ridge Section of the Ridge and Valley Province in Pennsylvania.  Michaux State Forest includes much of the wooded land on South Mountain.  Within or adjacent to its borders are located four state parks: King’s Gap Environmental Education Center, Pine Grove Furnace State Park, Caledonia State Park, and Mont Alto State Park.  The vast network of trails on these state lands includes the Appalachian Trail, which remains in the mountainous Blue Ridge Section all the way to its southern terminus in Georgia.

Pennsylvania State Parks on South Mounatin
In Pennsylvania, the forested highlands of the Blue Ridge Section of the Ridge and Valley Province are known as South Mountain.  Much of South Mountain lies within the boundaries of Michaux State Forest.  Stars indicate the locations of 1) King’s Gap Environmental Education Center, 2) Pine Grove Furnace State Park, 3) Caledonia State Park, and 4) Mont Alto State Park.  A drive on US 30 between Gettysburg and Chambersburg will take you right through Michaux State Forest along an east to west axis while a scenic northbound or southbound trip along PA 233 will bring you in proximity to each of the state parks located therein.  (Base image from NASA Earth Observatory Collection)

If you want a closeup look at the many species of trees found in Michaux State Forest, and you want them to be labeled so you know what they are, a stop at the Pennsylvania State University’s Mont Alto arboretum is a must.  Located next door to Mont Alto State Park along PA 233, the Arboretum at Penn State Mont Alto covers the entire campus.  Planting began on Arbor Day in 1905 shortly after establishment of the Pennsylvania State Forest Academy at the site in 1903.  Back then, the state’s “forests” were in the process of regeneration after nineteenth-century clear cutting.  These harvests balded the landscape and left behind the combustible waste which fueled the frequent wildfires that plagued reforestation efforts for more than half a century.  The academy educated future foresters on the skills needed to regrow and manage the state’s woodlands.

Online resources can help you plan your visit to the Arboretum at Penn State Mont Alto.  More than 800 trees on the campus are numbered with small blue tags.  The “List of arboretum trees by Tag Number” can be downloaded to tell you the species or variety of each.  The interactive map provides the locations of individual trees plotted by tag number while the Grove Map displays the locations of groups of trees on the campus categorized by region of origin.  A Founder’s Tree Map will help you find some of the oldest specimens in the collection and a Commemorative Tree Map will help you find dedicated trees.  There is also a species list of the common and scientific tree names.

Yellow Buckeye
The Yellow Buckeye (Aesculus flava) is a tree found in the forests of the Blue Ridge Section of the Ridge and Valley Province in West Virginia, Virginia, North Carolina and Georgia.  You can see it in Pennsylvania by visiting the collection of trees in the Arboretum at Penn State Mont Alto.
American Chestnut
The American Chestnut can be difficult to find due to the impact of chestnut blight, but you can see it in the Arboretum at Penn State Mont Alto.
Shagbark Hickory
Shagbark Hickory is a common tree in the forests of South Mountain.
Sweet Cherry
The Sweet Cherry (Prunus avium), a native of Europe, is naturalized throughout eastern and south-central Pennsylvania and is one of the more than 150 species of trees in the arboretum’s collection.
Sweet Birch
Sweet Birch (Betula lenta) foliage is particularly bright yellow on South Mountain this autumn.  It really “pops” against the backdrop of the evergreen Eastern White Pines and Eastern Hemlocks.  During less-than-ideal years, Sweet Birch leaves can subtly transition from green to drab brown without much fanfare before falling.
Common Persimmon
You might have a difficult time finding a Common Persimmon (Diospyros virginiana) growing wild in Pennsylvania, but you can find it in the Arboretum at Penn State Mont Alto.

The autumn leaves will be falling fast, so make it a point this weekend to check out the show on South Mountain.

Late Season Ruby-throated Hummingbirds…Again

Last October 3rd, a late-season Ruby-throated Hummingbird stopped by the garden at susquehannawildlife.net headquarters to take shelter from a rainy autumn storm.  It was so raw and chilly that we felt compelled to do something we don’t normally do—put out the sugar water feeder to supplement the nectar produced by our fall-flowering plants.  After several days of constant visits to the feeder and the flowers, our lingering hummer resumed its southbound journey on October 7th.

Fast forward to this afternoon and what do you know, at least two migrating hummingbirds have stopped by to visit the flowers in our garden.  This year, we have an exceptional abundance of blooms on some of their favorite plants.  In the ponds, aquatic Pickerelweed is topped with purple spikes and we still have bright orange tubular flowers on one of our Trumpet Vines—a full two to three months later than usual.

Late-season Ruby-throated Hummingbird
We checked each of our late-season visitors carefully to be reasonably certain that none was a stray western species of hummingbird.  All appear to be female or juvenile “Ruby-throats”.  If you have an abundance of flowering plants and/or you’re going to maintain your hummingbird feeders through the coming weeks, be on the lookout for western species.  Most are more hardy than our Ruby-throats and some have remained in the lower Susquehanna valley through the winter.
Ruby-throated Hummingbird at Mexican Cigar
As is typically the case, Ruby-throated Hummingbirds quickly gravitate toward the tubular flowers of our Cuphea ignea, the Mexican Cigar.  They find these showy plants to be absolutely irresistible.
Ruby-throated Hummingbird at Mexican Cigar
Mexican Cigar grows wild in parts of the Ruby-throated Hummingbird’s winter range.  To them, it’s comfort food.
Ruby-throated Hummingbird at Bat-faced Cuphea
We’re trying some new cultivars of Cuphea to see how they do.  As this composite image shows, the hummingbirds won’t let our Bat-faced Cuphea (Cuphea llavea) alone.  It’s another plant native to Mexico and Central America, right where some of our hummingbirds spend the winter.

Remember, keep those feeders clean and the provisions fresh!  You’ll be glad you did.

Hymanoptera: A Look at Some Bees, Wasps, Hornets, and Ants

What’s all this buzz about bees?  And what’s a hymanopteran?  Well, let’s see.

Hymanoptera—our bees, wasps, hornets and ants—are generally considered to be our most evolved insects.  Some form complex social colonies.  Others lead solitary lives.  Many are essential pollinators of flowering plants, including cultivars that provide food for people around the world.  There are those with stingers for disabling prey and defending themselves and their nests.  And then there are those without stingers.  The predatory species are frequently regarded to be the most significant biological controls of the insects that might otherwise become destructive pests.  The vast majority of the Hymanoptera show no aggression toward humans, a demeanor that is seldom reciprocated.

Late summer and early autumn is a critical time for the Hymanoptera.  Most species are at their peak of abundance during this time of year, but many of the adult insects face certain death with the coming of freezing weather.  Those that will perish are busy, either individually or as members of a colony, creating shelter and gathering food to nourish the larvae that will repopulate the environs with a new generation of adults next year.  Without abundant sources of protein and carbohydrates, these efforts can quickly fail.  Protein is stored for use by the larval insects upon hatching from their eggs.  Because the eggs are typically deposited in a cell directly upon the cache of protein, the larvae can begin feeding and growing immediately.  To provide energy for collecting protein and nesting materials, and in some cases excavating nest chambers, Hymanoptera seek out sources of carbohydrates.  Species that remain active during cold weather must store up enough of a carbohydrate reserve to make it through the winter.  Honey Bees make honey for this purpose.  As you are about to see, members of this suborder rely predominately upon pollen or insect prey for protein, and upon nectar and/or honeydew for carbohydrates.

We’ve assembled here a collection of images and some short commentary describing nearly two dozen kinds of Hymanoptera found in the Lower Susquehanna River Watershed, the majority photographed as they busily collected provisions during recent weeks.  Let’s see what some of these fascinating hymanopterans are up to…

SOLITARY WASPS

Great Black Wasp on goldenrod (Solidago species)
A Great Black Wasp on goldenrod (Solidago species).  Like other solitary wasps, a female  Great Black Wasp will sting and paralyze a host insect upon which she’ll deposit her eggs.  After hatching, the larvae will begin consuming the host’s body as a source of protein.  The parasitized insects are often katydids or grasshoppers.
A Great Black Wasp.
A Great Black Wasp feeding on nectar, a source of carbohydrates.  Unlike social bees and wasps, solitary wasps are equipped with a stinger solely used for immobilizing prey, not defending a nest.  They are therefore quite docile and pose little threat to humans.
A Great Black Wasp powdered with pollen.
A Great Black Wasp powdered with pollen.  Hymanopterans that gather nectar and/or pollen are tremendously important pollinators of hundreds of species of plants.
Thread-waisted Wasp
A female Thread-waisted wasp (Ammophilia species, probably A. nigricans) drags a paralyzed caterpillar to her excavated nest where she’ll deposit an egg on the body.  After hatching, the larval wasp will feed on the disabled caterpillar.  The protein will enable the larvae to grow, pupate, and later emerge as an adult wasp.
The female Eastern Cicada Killer (Sphecius speciosus) excavates an underground nest with branch tunnels connecting a dozen chambers or more.  As the common name suggests, the female wasp paralyzes a cicada, then makes a strenuous effort to fly and drag it back to the nest for placement in a cell.  Each male wasp egg is deposited upon just one immobilized cicada, but a female egg is provided with a cache of several cicadas to provide adequate protein for growth to a larger size.  Nest cells are sealed with soil, then the larvae hatch in just a couple of days.  Within about two weeks, they have consumed the cicada protein and are fully grown.  Wrapped in a cocoon, they spend the winter in the nest, then pupate in the spring before emerging as a new generation of adults.
The Black-and-yellow Mud Dauber
The Black-and-yellow Mud Dauber (Sceliphron caementarium) builds a mud-ball nest within which it packs paralyzed spiders to function as a source of protein for its larvae.
Black-and-yellow Mud Dauber at nest.
A Black-and-yellow Mud Dauber at nest.
Pipe Organ Mud Dauber Nest
The Pipe Organ Mud Dauber builds this elaborate nest in which their eggs and paralyzed spiders are deposited in cells sealed with mud partitions.  After consuming the spiders, the larvae pupate, overwinter, then emerge from their cells as adults during the following spring.  To escape the protection of the nest, the new generation of adults bore through the mud walls.  Adult Pipe Organ Mud Daubers resemble the Great Black Wasp, but have a white or yellow distal segment on their rear legs resembling a pair of light-colored socks.
A closeup of the previous image with the lengths of the nest tubes compressed to show four scavenger flies (Miltogramminae), possibly two species, that have invaded this Pipe Organ Mud Dauber nest.  Scavenger flies are kleptoparasites that victimize various solitary bees and wasps, depositing larvae directly into the host species’ nest cells to consume the protein cache stored therein.

CUCKOO WASPS

Cuckoo Wasp
Cuckoo Wasps (Chrysididae), also known as Emerald Wasps, parasitize the nests of other species of wasps.  Females lay their eggs inside the host’s nest, then flee the scene.  Upon hatching, larval Cuckoo Wasps feed on stockpiles of prey intended for the host species’ offspring.  Like the adult mud daubers that have already matured and departed this nest by digging a hole through the wall of the cell within which they were hatched, the metallic green Cuckoo Wasp in the upper left has just emerged in much the same way.

SWEAT BEES

A Sweat Bee (Lasioglossum species).
A Sweat Bee (Lasioglossum species) collecting nectar and pollen on White Snakeroot (Ageratina altissima).
A Sweat Bee (Lasioglossum species).
Sweat Bees (Lasioglossum species) visit human skin to lick up the electrolytes left behind by evaporating perspiration.
A Sweat Bee (Lasioglossum species).
Sweat Bees  in the genus Lasioglossum demonstrate various social behaviors ranging from species that are solitary nesters to those that create colonies with work forces ranging in size from as few as four to as many as hundreds of bees.  Some Lasioglossum practice kleptoparasitism, while others are quite accomplished foragers.
An Augochlorine Green Sweat Bee (Augochlorini).
A female Augochlorine Green Sweat Bee (Augochlorini) collecting nectar on White Snakeroot.  Notice the pollen “baskets” on the rear leg.
An Augochlorine Green Sweat Bee (Augochlorini).
An Augochlorine Green Sweat Bee (Augochlorini).  Sweat bees nest in subterranean cavities and in hollowed out sections of trees.
An Augochlorine Green Sweat Bee (Augochlorini).
A copper-colored Augochlorine Green Sweat Bee (Augochlorini) collecting nectar and dusted with pollen.

LEAFCUTTER AND MASON BEES

Leafcutter Bee
A Leafcutter Bee (Megachile species).  Like Mason Bees, female Mason Bees deposit each of their eggs on a “pollen loaf” within an individual cell inside a preexisting tunnel-like cavity in wood, stone, or in the ground.  Unlike Mason Bees, female Leafcutter Bees cut a circular piece of leaf to create each of the cells in their nest.  After hatching, the larval bee feeds on the pollen loaf, pupates, then emerges from the shelter of the nest to start a new generation, usually during the following year.
A Leafcutter Bee (Megachile species).
A Leafcutter Bee (Megachile species) visiting Wild Bergamot.  Female Leafcutter and Mason Bees lack pollen “baskets” on their rear legs but instead have pollen “brushes” on the underside of the abdomen to gather the protein they need to create a “pollen loaf” for each nest cell.
Leafcutter Bee
A Leafcutter Bee (Megachile species) collecting nectar from White Snakeroot.
A Mason Bee (Osmia species) emerging from a nest cell in spring.
A Mason Bee (Osmia species) emerging from a nest in spring.  Mason Bees create nesting cells within preexisting cavities in wood, stone, and other other supporting structures.  Within the nest cavity, each egg is deposited atop a cache of pollen and nectar, a pollen loaf, then enclosed behind a partition of mud.  The female Mason Bee will usually repeat this process until an entire cavity is filled with cells.  During the following spring, a new generation of adult Mason Bees digs its way through the cell walls to emerge and repeat the process.  These bees readily use paper straws or holes drilled in blocks of wood for nesting.
A mason bee nest box with holes drilled into blocks of wood.
A mason bee nest box with holes drilled into blocks of wood.
Parasitized Mason Bee Nest
Mason Bees seal each cell and the outer end of their nest cavity with mud.  These outer nest cells can been parasitized by a variety of wasps.  Here, the outer cell of a Mason Bee nest has been victimized by a tiny chalcid wasp (looks like another one to the lower left).  Several species of female chalcid wasps (native Monodontomerus species or non-native Pteromalus venustus) enlarge weak points in the outer partition of a mason bee nest, then sting and paralyze the larval bee inside before depositing their eggs.  Within the cell. the wasp larvae consume the larval Mason Bee and the “pollen loaf” provided for its growth.  These same parasitic wasps prey upon Leafcutter Bees as well.

BUMBLE BEES, CARPENTER BEES, HONEY BEES, AND DIGGER BEES

Common Eastern Bumble Bee
A Common Eastern Bumble Bee (Bombus impatiens) collecting nectar and pollen on goldenrod.  Bumble bees are our sole native group of social bees.  Their wax nests are built in a burrow or other shelter.  The eggs are deposited in cells along with a supply of pollen for nourishing the larvae upon hatching.  Honey is stored in “honey pots” within the nest.  New queens are produced along with male bees during the late-summer and fall.  Only the new generation of fertilized queens survive the winter to lay eggs and produce workers to construct a new nest.
Common Eastern Bumble Bees
A pair of Common Eastern Bumble Bees collecting nectar and becoming dusted with pollen.  Their fuzzy coats and semi-warm-blooded metabolism allows them to be active in cooler weather than is tolerated by other bees.
A Common Eastern Bumble Bee pollinating a Great Rhododendron flower.
Flowering plants including the Great Rhododendron find success attracting pollinators to their reproductive blossoms by offering carbohydrate-rich nectar to insects like this Eastern Bumble Bee.  The yellow spots on the flower’s upper petal help to guide visitors toward their sweet treat.
Eastern Carpenter Bee
An Eastern Carpenter Bee feeding on goldenrod nectar.  Compare the almost hairless abdomen to that of the bumble bees.  Carpenter bees are semi-social insects.  Females lay their eggs in cells within galleries bored into wood.  These nests are completed with great precision, avoiding creation of any second entrance by mistakenly breaching the outer surface of the excavated wood.  Each egg/larvae is provided with a supply of protein-rich pollen.  Males often hover outside their mate’s nest to prevent competing males from entering the area.
A Honey Bee visiting goldenrod alongside Common Eastern Bumble Bees.
A worker Honey Bee, a female member of a sisterhood of foragers from a nearby hive, visits goldenrod alongside Common Eastern Bumble Bees.  Honey Bees were brought to North America during the 1620s, the earliest years of the trans-Atlantic migration of European colonists, to pollinate cultivated plants and to provide a reliable source of honey and beeswax.  Within the Honey Bee’s social structure, the queen of each hive lays the eggs to produce the female worker bees.  Once each year, male drones are produced along with a new generation of queens.
Honey Bee Hive
In nature, Honey Bees build hives in tree cavities.  Recently, this colony constructed a hive in a screech owl nest box at susquehannawildlife.net headquarters.  To provide protein for the hatching larvae, worker bees collect pollen and deposit it within the hexagonal cells of the vertically aligned beeswax combs.  After an egg is deposited upon the pollen cache, each cell is sealed with more beeswax.  Young females tend these nest combs before maturing and becoming foraging worker bees.
Bee Hive Display
In apiculture, Honey Bees are raised in man-made hives.  This Pennsylvania Association of Beekeepers display gives visitors to the Pennsylvania Farm Show in Harrisburg a look at the inner workings of a live bee hive.  Nectar collected by worker bees is turned into honey to provide the supply of carbohydrates needed to fuel the colony through the winter.  Note the honeycombs on the glass.
A possible Small Carpenter Bee Ceratina species).
A possible Small Carpenter Bee (Ceratina species) visiting White Snakeroot.  Small Carpenter Bees nest inside hollow stems and twigs.  Some species are eusocial, with a queen’s daughters and sisters sharing responsibility for finding food and rearing the young.  Females overwinter inside a one of the excavated stems and begin a new nest there in the spring.
A Digger Bee (possibly Melissodes species).
A Digger Bee (possibly Melissodes species) with “pollen baskets” full of pollen collected from nearby flowers.  Digger Bees in the genus Melissodes are often known as the Long-horned Bees.  These social insects excavate underground nests and many species practice communal living.

SCOLIID WASPS

Two-spotted Scoliid Wasp
The Two-spotted Scoliid Wasp (Scolia dubia), also known as the Blue-winged Scoliid Wasp, is most frequently observed feeding on nectar.  Scoliid wasps are solitary nesters, though they may assemble into groups while visiting flowers.  They often ignore the presence of humans and are seldom disturbed by their presence.  Females seek out the burrowing grubs of beetles including the Green June Bug (Cotinis nitida) and possibly the Japanese Beetle.  After stinging a grub to paralyze it, the wasp will deposit her egg on its body, then bury it.  Upon hatching, the larval wasp will feed on the grub for nourishment as it grows.
June Bugs eating watermelon.
Don’t like having your watermelon overrun by Green June Bugs while you’re eating?  Then you ought to go out of your way to be nice to the Two-spotted Scoliid Wasp.
The Double-banded Scoliid
The Double-banded Scoliid (Scolia bicincta) parasitizes beetle larvae as hosts for its larvae.  For carbohydrates it relishes flower nectar.

PAPER WASPS

Northern Paper Wasp
A Northern Paper Wasp (Polistes fuscatus).  Paper wasps prey upon numerous garden pests, particularly caterpillars, to collect protein.  Though they are social insects equipped with stingers to subdue their victims and defend their nests, paper wasps are surprisingly docile.
The Northern Paper Wasp
A Northern Paper Wasp (Polistes fuscatus) feeding on nectar from a goldenrod flower.
A Northern Paper Wasp harvesting wood pulp
A Northern Paper Wasp harvesting wood pulp from the side of a mason bee nest box at susquehannawildlife.net headquarters.  The pulp is chewed in the wasp’s saliva to create the paper used to construct the colony’s open-cell nest.
Guinea Paper Wasps (Polistes exclamans) at their nest.
Common Paper Wasps (Polistes exclamans), also known as Guinea Paper Wasps, at their open-cell nest.  This and the nests of most other paper wasps are suspended on a filament or a pedicle.  Many paper wasps can excrete an ant repellent on this section of the nest in an effort to prevent invasion.  Like many other social hymenopterans, a defending wasp can secrete a pheromone venom during the stinging process to warn the colony of danger at the nest.  In winter, Common Paper Wasps seek shelter in stumps and other locations to hibernate.
European Paper Wasp
The European Paper Wasp (Polistes dominula) is a non-native species which builds nests in man-made structures including bird houses.  To collect protein, they prey on a wide selection of insects and other invertebrates.  As such, European Paper Wasps are widespread and successful here in North America.

YELLOWJACKETS AND HORNETS

An Eastern Yellowjacket
An Eastern Yellowjacket feeding on lanternfly honeydew.  Eastern Yellowjackets derive much of their success from being generalists, collecting carbohydrates from nearly any sweet source, natural or man made.  They are quite fond of ripe fruits, flower nectar, and sugary snacks and drinks, especially soda.  Protein for nourishing their larvae is derived from the wide variety invertebrates upon which they prey and from carrion.  These foods are chewed into a paste form in preparation for placement into the brood cells.
An Eastern Yellowjacket.
A subterranean colony of Eastern Yellowjackets is started anew each spring by a young queen that has survived winter hibernation in diapause, a state of interrupted development.  She constructs the new nest’s first cells using pulp made by chewing rotting wood.  The first brood of workers scales up construction while the queen continues producing eggs.  At the nest, these social insects will viciously attack anyone or anything perceived to be a threat, so give them their space and leave them alone.  Many yellowjacket infestations of homes and other buildings are the work of non-native German Yellowjacket (Vespula germanica) [not shown], an invasive species that constructs paper nests in void spaces including walls and attics.
Robber Fly consuming an Eastern Yellowjacket
Yellowjackets may be moody and aggressive, but they do fall victim to a number of predators.  A Robber Fly (Promachus species) has taken down and is devouring this Eastern Yellowjacket.
A Bald-faced Hornet (Dolichovespula maculata) feeding on Spotted Lanternfly honeydew deposits
A Bald-faced Hornet (Dolichovespula maculata) feeding on Spotted Lanternfly honeydew on a Tree-of-heaven (Ailanthus altissima).  In the absence of nectar-producing flowers, many bees, yellowjackets, and hornets have turned to the invasive lanternfly and Ailanthus combo to turn the sun’s energy into the carbohydrates they need.  For protein, they prey upon spiders, flies, caterpillars, and a variety of other insects.
A Bald-faced Hornet collecting wood pulp from the surface of a weathered picnic table.
To create paper for nest construction, this Bald-faced Hornet is collecting wood pulp from the surface of a weathered picnic table.  Away from the nest, these hornets demonstrate a calm, carefree demeanor and can be closely observed.
Bald-faced Hornet Nest
A Bald-faced Hornet nest in a pine tree.  These hives are strictly temporary.  Within the nest, a generation of drones (males) and new queens are produced late each year.  These wasps leave the colony to mate.  With the arrival of freezing weather, all inhabitants within the nest, including the old queen, perish, as do the drones that departed to breed.  Only the new queens survive winter hibernation to propagate the next generation of wasps,  starting with the workers needed to construct a fresh nest and reestablish the colony.
Bald-faced Hornets Peering from Nest
Did you ever get the feeling you’re being watched?  Don’t go messing around with Bald-faced Hornet nests.  The occupants therein, like other social bees, wasps, and hornets, are equipped with stingers and venom for defending their colony.  This is an adaptation that has developed over time to assure the survival of populations of these insects.  Think about it this way, a solitary wasp that loses a nest loses only their individual brood of offspring.  There is minimal impact on the wider local population of such insects.  Conversely, a social wasp or hornet that loses a nest loses an entire colony, possibly negating the benefits of their cooperative behavior and threatening the survival of the species.  Insects that cooperate to build societies for survival can be more vulnerable to the catastrophic impacts of certain circumstances like disease, weather, and invasion of their colonies.  Therefore, natural selection has provided them with contingencies for these dangers, for example, the instinct to construct protective shelters and the adaptation of stingers and venom for defense against intruders and would-be predators.  Oh, and by the way, the Bald-faced Hornet can spray venom, often aiming for the eyes, so keep your distance.
European Hornets
European Hornets (Vespa crabro), an introduced species, are predatory on a variety of flying insects for protein.  For carbohydrates they are attracted to sweets like this lanternfly honeydew on Tree-of-heaven.
European Hornets constructing a nest in a tree cavity.
European Hornets constructing a paper nest in a tree cavity.

POTTER WASPS

A Potter Wasp (Eumenes species, probably E. fraturnus) hovering near a European Paper Wasp.
A potter wasp (Eumenes species), probably a Fraternal Potter Wasp (E. fraternus), hovering near a European Paper Wasp on Partridge Pea.  The female potter wasp builds a small mud nest resembling a tiny clay pot.  One of her eggs is inserted and left hanging on a thin thread.  Then a paralyzed caterpillar is deposited as a source of protein to nourish the larva upon hatching.  Lastly, the pot is sealed with a lid made of wet mud.  Upon maturing, the new generation of adult wasps perform a pottery breaking to emerge and take flight.

ANTS

Field Ants (Formica species, possibly Formica pallidefulva) clearing the entrance to their underground nest.
Field Ants (Formica species, possibly Formica pallidefulva) clear the entrance to their underground nest.  Field ants are eusocial insects, they work in concert to build, maintain, and defend the nest, rear young, and find food.  There is no social caste system.  Field Ants are predators and scavengers when collecting protein.  For carbohydrates they often rely on the honeydew produced by aphids.  As a method of improving and sustaining the production of honeydew, some ant species will tend colonies of aphids by moving the younger individuals from depleted portions of plants to more healthy tissue.  Field Ant nests contain chambers used for a variety of functions including raising young and storing food.  Some nests include multiple queens and some colonies consist of more than one nest.   Ants in the genus Formica are weaponized; they can spray formic acid to repel intruders and defend their colony.

We hope this brief but fascinating look at some of our more common bees, wasps, hornets, and ants has provided the reader with an appreciation for the complexity with which their food webs and ecology have developed over time.  It should be no great mystery why bees and other insects, particularly native species, are becoming scarce or absent in areas of the Lower Susquehanna River Watershed where the landscape is paved, hyper-cultivated, sprayed, mowed, and devoid of native vegetation, particularly nectar-producing plants.  Late-summer and autumn can be an especially difficult time for hymanopterans seeking the sources of proteins and carbohydrates needed to complete preparations for next year’s generations of these valuable insects.  An absence of these staples during this critical time of year quickly diminishes the diversity of species and begins to tear at the fabric of the food web.  This degradation of a regional ecosystem can have unforeseen impacts that become increasingly widespread and in many cases permanent.

A farmland desert.

A farmland desert.
How can anyone be surprised by the absence of bees and other pollinators in farmland? Manicured and cultivated ground offers little in the way of year-round shelter and food sources for insects and other wildlife.
A savanna-like habitat.
This savanna-like habitat on a south-facing slope provides the abundance of nectar-producing, pollen-rich wildflowers needed to nourish a diverse population of insects including bees, wasps, hornets, and ants.  Goldenrods, asters, and White Snakeroot are some of their late-season favorites.

Editor’s Note: No bees, wasp, hornets, or ants were harmed during this production.  Neither was the editor swarmed, attacked, or stung.  Remember, don’t panic, just observe.

SOURCES

Eaton, Eric R., and Kenn Kaufman.  2007.  Kaufman Field Guide to Insects of North America.  Houghton Mifflin Company.  New York, NY.

(If you’re interested in insects, get this book!)

Four Common Grasshoppers

Grasshoppers are perhaps best known for the occasions throughout history when an enormous congregation of these insects—a “plague of locusts”—would assemble and rove a region to feed.  These swarms, which sometimes covered tens of thousands of square miles or more, often decimated crops, darkened the sky, and, on occasion, resulted in catastrophic famine among human settlements in various parts of the world.

The largest “plague of locusts” in the United States occurred during the mid-1870s in the Great Plains.  The Rocky Mountain Locust (Melanoplus spretus), a grasshopper of prairies in the American west, had a range that extended east into New England, possibly settling there on lands cleared for farming.  Rocky Mountain Locusts, aside from their native habitat on grasslands, apparently thrived on fields planted with warm-season crops.  Like most grasshoppers, they fed and developed most vigorously during periods of dry, hot weather.  With plenty of vegetative matter to consume during periods of scorching temperatures, the stage was set for populations of these insects to explode in agricultural areas, then take wing in search of more forage.  Plagues struck parts of northern New England as early as the mid-1700s and were numerous in various states in the Great Plains through the middle of the 1800s.  The big ones hit between 1873 and 1877 when swarms numbering as many as trillions of grasshoppers did $200 million in crop damage and caused a famine so severe that many farmers abandoned the westward migration.  To prevent recurrent outbreaks of locust plagues and famine, experts suggested planting more cool-season grains like winter wheat, a crop which could mature and be harvested before the grasshoppers had a chance to cause any significant damage.  In the years that followed, and as prairies gave way to the expansive agricultural lands that presently cover most of the Rocky Mountain Locust’s former range, the grasshopper began to disappear.  By the early years of the twentieth century, the species was extinct.  No one was quite certain why, and the precise cause is still a topic of debate to this day.  Conversion of nearly all of its native habitat to cropland and grazing acreage seems to be the most likely culprit.

The critically endangered Eskimo Curlew (Numenius borealis), a species not photographed since 1962 and not confirmed since 1963, fed on Rocky Mountain Locusts during its spring migration through the Great Plains.  Excessive hunting and conversion of grasslands to agriculture are believed responsible for the bird’s demise.  (United States Fish and Wildlife Service image by Christina Nelson)

In the Mid-Atlantic States, the mosaic of the landscape—farmland interspersed with a mix of forest and disturbed urban/suburban lots—prevents grasshoppers from reaching the densities from which swarms arise.  In the years since the implementation of “Green Revolution” farming practices, numbers of grasshoppers in our region have declined.  Systemic insecticides including neonicotinoids keep grasshoppers and other insects from munching on warm-season crops like corn and soybeans.  And herbicides including 2,4-D (2,4-Dichlorophenoxyacetic acid) have, in effect, become the equivalent of insecticides, eliminating broadleaf food plants from the pasturelands and hayfields where grasshoppers once fed and reproduced in abundance.  As a result, few of the approximately three dozen species of grasshoppers with ranges that include the Lower Susquehanna River Watershed are common here.  Those that still thrive are largely adapted to roadsides, waste ground, and small clearings where native and some non-native plants make up their diet.

Here’s a look at four species of grasshoppers you’re likely to find in disturbed habitats throughout our region.  Each remains common in relatively pesticide-free spaces with stands of dense grasses and broadleaf plants nearby.

CAROLINA GRASSHOPPER

Dissosteira carolina

Carolina Grasshopper
The Carolina Grasshopper, also known as the Carolina Locust or Quaker, is one of the band-winged grasshoppers.  It is commonly found along roadsides and on other bare ground near stands of tall grass and broadleaf plants.
Carolina Grasshopper
The Carolina Grasshopper is variable in color, ranging from very dark brown…
Carolina Grasshopper
…to a rich tan or khaki shade.  These earth-tone colors provide the insect with effective camouflage while spending time on the ground.
Carolina Grasshopper wing
The Carolina Grasshopper is most readily detected and identified when it flies.  The colors of the wings resemble those of the Mourning Cloak butterfly.
Great Black Wasp on goldenrod.
Carolina Grasshoppers are among the preferred victims of Great Black Wasps (Sphex pensylvanicus).  A female wasp stings the grasshopper to paralyze it, then drags it away to one of numerous cells in an underground burrow where she lays an egg on it.  The body of the disabled grasshopper then provides nourishment for the larval wasp.

DIFFERENTIAL GRASSHOPPER

Melanoplus differentialis

Differential Grasshopper nymph.
Differential Grasshopper nymph with small “fairy wings”.
Differential Grasshopper
An adult female Differential Grasshopper with fully developed wings.
An adult female Differential Grasshopper
An adult female Differential Grasshopper

TWO-STRIPED GRASSHOPPER

Melanoplus bivittatus

Two-striped Grasshopper nymph.
An early-stage Two-striped Grasshopper nymph.
Two-striped Grasshopper nymph.
A Two-striped Grasshopper nymph in a later stage.
Two-striped Grasshopper
An adult female Two-striped Grasshopper.
Two-striped Grasshopper
An adult female Two-striped Grasshopper.  Note the pale stripe originating at each eye and joining near the posterior end of the wings to form a V-shaped pattern.
Two-striped Grasshopper
An adult female Two-striped Grasshopper.

RED-LEGGED GRASSHOPPER

Melanoplus femurrubrum

A Red-legged Grasshopper hiding in dense urban vegetation.
An adult male Red-legged Grasshopper hiding in dense urban vegetation.
Red-legged Grasshopper
The Red-legged Grasshopper may currently be our most abundant and widespread species.
Red-legged Grasshopper
An adult male Red-legged Grasshopper.
Red-legged Grasshopper
An adult female Red-legged Grasshopper.

Protein-rich grasshoppers are an important late-summer, early-fall food source for birds.  The absence of these insects has forced many species of breeding birds to abandon farmland or, in some cases, disappear altogether.

During the mid-twentieth century, the Cattle Egret (Bubulcus ibis), a notoriously nomadic species, transited the Atlantic from Africa to colonize the Americas…and they did it without any direct assistance from humans.  During the 1970s and early 1980s, a nesting population of Cattle Egrets on river islands adjacent to the Susquehanna’s Conejohela Flats off Washington Boro was the largest inland rookery in the northeastern United States.  The Lancaster County Bird Club censused the birds each August and found peak numbers in 1981 (7,580).  During their years of abundance, V-shaped flocks of Cattle Egrets from the rookery islands ventured into grazing lands throughout portions of Lancaster, York, Dauphin, and Lebanon Counties to hunt grasshoppers.  These daily flights were a familiar summertime sight for nearly two decades.  Then, in the early 1980s, reductions in pastureland acreage and plummeting grasshopper numbers quickly took their toll.  By 1988, the rookery was abandoned.  The Cattle Egrets had moved on.  (Vintage 33 mm image)
During the summer and early fall, juvenile and adult Ring-necked Pheasants feed heavily on grasshoppers.  Earlier and more frequent mowing along with declining numbers of grasshoppers on farmlands due to an increase in pesticide use were factors contributing to the crash of the pheasant population in the early 1980s.
Wild Turkey
To the delight of Wild Turkeys, each of the four species of grasshoppers shown above frequents clearings and roadsides adjacent to forest areas.  While changes in grasshopper distribution have been detrimental to populations of birds like pheasants, they’ve created a feeding bonanza for turkeys.
Wild Turkeys feeding on grasshoppers along a forest road.
Wild Turkeys feeding on an abundance of grasshoppers along a forest road.
An American Kestrel feeds on a grasshopper while ignoring the abundance of Spotted Lanternflies swarming the adjacent utility pole.  In Susquehanna valley farmlands, grasshopper and kestrel numbers are down.  Lanternflies, on the other hand, have got it made.
Early Successional Growth
Maintaining areas bordering roads, forests, wetlands, farmlands, and human development in a state of early succession can provide and ideal mix of mature grasses and broadleaf plants for grasshoppers, pollinators, birds, and other wildlife.

A Visit to a Beaver Pond

To pass the afternoon, we sat quietly along the edge of a pond created recently by North American Beavers (Castor canadensis).  They first constructed their dam on this small stream about five years ago.  Since then, a flourishing wetland has become established.  Have a look.

A Beaver Pond
Vegetation surrounding the inundated floodplain helps sequester nutrients and sediments to purify the water while also providing excellent wildlife habitat.
A beaver lodge.
The beaver lodge was built among shrubs growing in shallow water in the middle of the pond.
Woolgrass in a beaver pond.
Woolgrass (Scirpus cyperinus) is a bulrush that thrives as an emergent and as a terrestrial plant in moist soils bordering the pond.
A male Common Whitetail dragonfly keeping watch over his territory.
A male Common Whitetail dragonfly keeping watch over his territory.
A Twelve-spotted Skimmer perched on Soft Rush.
A Twelve-spotted Skimmer perched on Soft Rush.
A Blue Dasher dragonfly seizing a Fall Field Cricket (Gryllus pennsylvanicus).
A Blue Dasher dragonfly seizing a Fall Field Cricket (Gryllus pennsylvanicus).
A Spicebush Swallowtail visiting Cardinal Flower.
A Spicebush Swallowtail visiting a Cardinal Flower.
Green Heron
A Green Heron looking for small fish, crayfish, frogs, and tadpoles.
A Green Heron stalks potential prey.
The Green Heron stalking potential prey.
A Wood Duck feeding on Lesser Duckweed.
A Wood Duck feeding on the tiny floating plant known as Lesser Duckweed (Lemna minor).
A Least Sandpiper feeding along the muddy edge of a beaver pond.
A Least Sandpiper poking at small invertebrates along the muddy edge of the beaver pond.
Solitary Sandpiper
A Solitary Sandpiper.
A Solitary Sandpiper testing the waters for proper feeding depth.
A Solitary Sandpiper testing the waters for proper feeding depth.
Pectoral Sandpiper
A Pectoral Sandpiper searches for its next morsel of sustenance.
A Sora rail in a beaver pond.
The Sora (Porzana carolina) is a seldom seen rail of marshlands including those created by North American Beavers.  Common Cattails, sedges, and rushes provide these chicken-shaped wetland birds with nesting and loafing cover.

Isn’t that amazing?  North American Beavers build and maintain what human engineers struggle to master—dams and ponds that reduce pollution, allow fish passage, and support self-sustaining ecosystems.  Want to clean up the streams and floodplains of your local watershed?  Let the beavers do the job!

Photo of the Day

Magnolia Leaf-footed Bug
At first glance, this insect might be mistaken for a Brown Marmorated Stink Bug.  It is, however, a native species, a Magnolia Leaf-footed Bug (Leptoglossus fulvicornis).  Because it feeds almost exclusively on trees in the genus Magnolia, and because the three members of the genus native to the lower Susquehanna valley, Umbrella Magnolia (M. tripetala), Cucumber Tree (M. acuminata), and Sweetbay (M. virginiana), each have a very limited distribution here, this insect’s presence in the watershed is largely dependent on widespread introductions of ornamental magnolias and native transplants including Southern Magnolia (Magnolia grandiflora).  Adult Magnolia Leaf-footed Bugs overwinter in leaf litter beneath the host trees.  In spring, females lay their eggs in rows along the midrib of the leaves where they are often victims of parasitoid wasps including Gyron pennsylvanicum.  The surviving nymphs and the adults feed on the tree’s leaves and fruits, causing little harm to its overall health.

Surf’s Up: The Waves Keep Rolling In

“Waves” of warblers and other Neotropical songbirds continue to roll along the ridgetops of southern Pennsylvania.  The majority of these migrants are headed to wintering habitat in the tropics after departing breeding grounds in the forests of southern Canada.  At Second Mountain Hawk Watch, today’s early morning flight kicked off at sunrise, then slowed considerably by 8:30 A.M. E.D.T.  Once again, in excess of 400 warblers were found moving through the trees and working their way southwest along the spine of the ridge.  Each of the 12 species seen yesterday were observed today as well.  In addition, there was a Northern Parula and a Canada Warbler.  Today’s flight was dominated by Bay-breasted, Blackburnian, Black-throated Green, and Tennessee Warblers.

A Blackburnian Warbler at sunrise on Second Mountain.
A Blackburnian Warbler at sunrise on Second Mountain.
A hungry Blackburnian Warbler feeding on insects.
A hungry Blackburnian Warbler feeding on insects.
Black-throated Green Warbler on Second Mountain.
Black-throated Green Warblers were a plentiful species among both yesterday’s and today’s waves of Neotropical migrants.
A juvenile Black-throated Green Warbler on Second Mountain
A juvenile Black-throated Green Warbler.
Tennessee Warbler on Second Mountain.
One of the scores of Tennessee Warblers seen on Second Mountain early this morning.
Cape May Warbler
Cape May Warblers were still common today, but not moving through in the numbers seen yesterday.
A male Black-throated Blue Warbler on Second Mountain.
A male Black-throated Blue Warbler.
Magnolia Warbler on Second Mountain.
Compared to yesterday’s flight, lesser numbers of Magnolia Warblers were seen today.
An adult male Wilson's Warbler on Second Mounatin.
An adult male Wilson’s Warbler was a good find among the hundreds of birds swarming the ridgetop.
Nashville Warbler
This Nashville Warbler spent much of the day in the tangles of Mile-a-minute Weed surrounding the lookout.

Other interesting Neotropical migrants joined the “waves” of warblers…

Red-eyed Vireo
Red-eyed Vireo numbers were higher than yesterday.
Warbling Vireo
This Warbling Vireo was found peering from the cover of the shady forest.
Rose-breasted Grosbeaks
A minimum of six Rose-breasted Grosbeaks were identified including the juvenile male seen here in first-fall plumage.  Other good sightings were Scarlet Tanagers, an adult male Baltimore Oriole, and a dozen or more Ruby-throated Hummingbirds.
Least Flycatcher on second Mountain.
Three Least Flycatchers were heard calling and seen chasing one another through a stand of dead timber on the south slope below the lookout.
Broad-winged Hawk
After the warbler flight settled, the task of counting migrating raptors commenced.  Five Broad-winged Hawks including this one were tallied as they glided away to the southwest for a winter vacation in the tropics of Central and South America.

Ruby-throated Hummingbirds on the Go

Ruby-throated Hummingbirds will be moving south through the Lower Susquehanna River Watershed during the next few weeks.
Ruby-throated Hummingbirds will be migrating south through the Lower Susquehanna River Watershed during the next few weeks.
Ruby-throated Hummingbird at Mexican Cigar.
While on their journey, they refuel by frequently visiting nectar-producing native wildflowers as well as garden specimens like this Mexican Cigar.
Ruby-throated Hummingbird at feeder.
Hummingbirds supplement their diet by visiting feeders filled with a mixture of sugar and water.  If you’re feeding hummingbirds, or thinking about feeding hummingbirds, be sure to review the helpful tips contained in our post from August 5, 2022, “Two Feeders Are Better Than One”.  Their health and your peace of mind may depend on it.

One of Nature’s Finest: The Cardinal Flower

It may be one of the most treasured plants among native landscape gardeners.  The Cardinal Flower (Lobelia cardinalis) blooms in August each year with a startling blaze of red color that, believe it or not, will sometimes be overlooked in the wild.

Cardinal Flower on a Stream
Cardinal Flower is most often found in wet soil along forested bodies of water.  The blooms of this shade-loving species may go unnoticed until rays of sunshine penetrate the canopy to strike their brilliant red petals.

The Cardinal Flower grows in wetlands as well as in a variety of moist soils along streams, rivers, lakes, and ponds.  Shady locations with short periods of bright sun each day seem to be favored for an abundance of color.

Cardinal Flower and Great Blue Lobelia
Cardinal Flower in bloom in a riparian forest along the Susquehanna.  To its right is its close relative, Great Lobelia, a plant sometimes known as Great Blue Lobelia or Blue Cardinal Flower.
Cardinal Flower in a wet bottomland woods.
Cardinal Flower in a wet bottomland woods.
The Cardinal Flower can find favorable growing conditions along stream, river and lake shores.
The Cardinal Flower can find favorable growing conditions along stream, river, and lake shores.  Even though they are perennial plants, their presence along such waters often seems temporary.  Changing conditions cause them to suddenly disappear from known locations, then sometimes reappear at the same place or elsewhere nearby.  Some of this phenomenon may be due to the fact that stressed plants can fail to bloom, so they easily escape notice.  When producing flowers during favorable years, the plants seem to mysteriously return.
Cardinal Flowers along a wave-swept shoreline light up the greenery of erosion-controlling riparian vegetation with glowing red color.
Cardinal Flowers along a wave-swept shoreline light up the greenery of erosion-controlling riparian vegetation with glowing red color.

The Cardinal Flower can be an ideal plant for attracting hummingbirds, bees, butterflies, and other late-summer pollinators.  It grows well in damp ground, especially in rain gardens and along the edges streams, garden ponds, and stormwater retention pools.  If you’re looking to add Cardinal Flower to your landscape, you need first to…

REMEMBER the CARDINAL RULE…

Cardinal Flower plants are available at many nurseries that carry native species of garden and/or pond plants.  Numerous online suppliers offer seed for growing your own Cardinal Flowers.  Some sell potted plants as well.  A new option is to grow Cardinal Flowers from tissue cultures.  Tissue-cultured plants are raised in laboratory media, so the pitfalls of disease and hitchhikers like invasive insects and snails are eliminated.  These plants are available through the aquarium trade from most chain pet stores.  Though meant to be planted as submerged aquatics in fish tank substrate, we’ve reared the tissue-cultured stock indoors as emergent plants in sandy soil and shallow water through the winter and early spring.  When it warms up, we transplant them into the edges of the outdoor ponds to naturalize.  As a habit, we always grow some Cardinal Flower plants in the fish tanks to take up the nitrates in the water and to provide a continuous supply of cuttings for starting more emergent stock for outdoor use.

Tissue culture Cardinal Flower being grown as a submerged aquatic in a fish aquarium.
A tissue-cultured Cardinal Flower rooted in sandy substrate and being grown as a submerged aquatic plant in a fish tank.  Cuttings from this plant will be used to grow emergent specimens in shallow water for transplanting outdoors around the garden pond.
Cardinal Flower from Tissue Culture
A Cardinal Flower grown from an aquarium store tissue culture blooms in the pond at susquehannawildlife.net headquarters.
Cardinal Flower blooming in November.
Grown as an emergent, Cardinal Flower may bloom very late in the season.  This tissue-cultured specimen in the headquarters pond was photographed in early November, 2022.

Wild Senna, a Showy Background Plant for Your Wildflower Garden

Looking for a native wildflower that’s tall, showy, and a great choice for attracting wildlife, especially butterflies and bees?  Then check out Wild Senna (Senna hebecarpa).

Wild Senna in a roadside wildflower garden on Pennsylvania State Gamelands.
Wild Senna currently blooming in a roadside wildflower garden on Pennsylvania State Game Lands.

Wild Senna, also known as American Senna, is a host plant for the larvae of Cloudless Sulphur and Sleepy Orange (Eurema nicippe) butterflies.  It thrives in almost any moist, well-drained soil in habitats including open woodlands, forest edges, meadows, and gardens like yours.  Its height at flowering ranges from three to six feet.  If you prefer, this perennial wildflower can even be cultivated as a shrub-like form.  It is easily grown from seed, which is available from Ernst Conservation Seeds of Meadville, Pennsylvania, as well as numerous other vendors.  And don’t forget to give Wild Senna’s two close relatives, Partridge Pea and Maryland Senna, a try as well.  They attract the same species of butterflies and are just as easy to grow.  You’ll like ’em.

Cloudless Sulphur
Cloudless Sulphur butterflies from populations in the south colonize the Lower Susquehanna River Watershed each summer in varying numbers.
Close-up view of a Wild Senna flower cluster.
Close-up view of a Wild Senna flower cluster.  Typical of members of the pea family, the  seeds of all native Senna species develop within pods after blooming and are sought after by wildfowl (Galliformes), particularly Northern Bobwhite.
Partridge Pea in flower with seed pods.
Partridge Pea with flowers and seed pods in the susquehannawildlife.net garden.  Smaller than the other Senna species, Partridge Pea reaches a height of just two to three feet.

Blue Tuesday

We’ve got the summertime blues for you, right here at susquehannawildlife.net…

Big Bluestem
In warm-season grass meadows, Big Bluestem is now in flower.  This and other species of native prairie grasses provide excellent habitat for birds, mammals, and insects including butterflies.  To survive drought and fire, their roots run much deeper than cool season grasses, creeping down four to six feet or more.  This adaptation allowed warm season grasses to recover from heavy grazing by large Pre-Anthropocene mammals.  Today, it makes them ideal plants for soil stabilization.
A male Indigo Bunting has already found some ripe seeds among the heads of flowering Big Bluestem.
A male Indigo Bunting has already found ripe seeds among the heads of flowering Big Bluestem.
Molting Indigo Bunting
Look closely and you’ll see our Indigo Bunting is beginning a pre-migration molt out of its bright-blue breeding (alternate) plumage and into a gray-brown winter (basic) plumage.  The berries of the American Pokeweed upon which it is perched will soon ripen into a dark blue, almost black, color.  Though toxic to humans, these fruits find favor with many species of birds and mammals.
Silky Dogwood
Another great wildlife food is Silky Dogwood (Cornus amomum), a deciduous shrub that sports blue-colored berries in summer and showy, bright-red twigs in winter.  It grows well in wet ground along streams and ponds, as well as in rain gardens.
Great Blue Heron
A Great Blue Heron searches the shallows for small fish.  This species is also a good mouser, at times seen hunting in grassy meadows.  Right now is prime time to see it and a variety of other herons and egrets throughout the Lower Susquehanna River Watershed.

…so don’t let the summertime blues get you down.  Grab a pair of binoculars and/or a camera and go for a stroll!

Butterflies and More at Boyd Big Tree Preserve Conservation Area

If you’re feeling the need to see summertime butterflies and their numbers just don’t seem to be what they used to be in your garden, then plan an afternoon visit to the Boyd Big Tree Preserve along Fishing Creek Valley Road (PA 443) just east of U.S. 22/322 and the Susquehanna River north of Harrisburg.  The Pennsylvania Department of Conservation and Natural Resources manages the park’s 1,025 acres mostly as forested land with more than ten miles of trails.  While located predominately on the north slope of Blue Mountain, a portion of the preserve straddles the crest of the ridge to include the upper reaches of the southern exposure.

American Chestnut at Boyd Big Tree Preserve
A grove of American Chestnuts (Castanea dentata) planted at Boyd Big Tree Preserve is part of a propagation program working to restore blight-resistant trees to Pennsylvania and other areas of their former range which included the Appalachians and the upper Ohio River watershed.

Fortunately, one need not take a strenuous hike up Blue Mountain to observe butterflies.  Open space along the park’s quarter-mile-long entrance road is maintained as a rolling meadow of wildflowers and cool-season grasses that provide nectar for adult butterflies and host plants for their larvae.

Butterfly Meadow at Boyd Big Tree Preserve
A view looking north at the butterfly meadow and entrance road at Boyd Big Tree Preserve Conservation Area.  Second Mountain is in the background.
Walking a Meadow Path
Mowed paths follow the entrance road and a portion of the perimeter of the meadow allowing visitors a chance to wander among the waist-high growth to see butterflies, birds, and blooming plants at close range without trampling the vegetation or risking exposure to ticks.
A Silver-spotted Skipper feeding on nectar from Dogbane (Apocynum cannabinum) flowers.
A Silver-spotted Skipper feeding on nectar from the flowers of Indian Hemp (Apocynum cannabinum).  Like the milkweeds, Indian Hemp is a member of the dogbane family (Apocynaceae).
An Eastern Tiger Swallowtail feeding on Common Milkweed.
An Eastern Tiger Swallowtail feeding on Common Milkweed.
Great Spangled Fritillary on Common Milkweed.
A Great Spangled Fritillary (Speyeria cybele) on Common Milkweed.
A Black Swallowtail feeding on Common Milkweed nectar.
A Black Swallowtail (Papilio polyxenes) feeding on Common Milkweed nectar.
A Pipevine Swallowtail on Common Milkweed.
A Pipevine Swallowtail (Battus philenor) on Common Milkweed.
A Pipevine Swallowtail on Common Milkweed.
Another Pipevine Swallowtail on Common Milkweed.  Note the hook-shaped row of red-orange spots on the underside of the hindwing.
A Pipevine Swallowtail on visiting Butterfly Weed.
A Pipevine Swallowtail visiting the brilliant blooms of Butterfly Weed, a favorite of a wide variety of pollinators.
A Black Swallowtail on Butterfly Weed
A Black Swallowtail with damaged wings alights atop a Butterfly Weed flower cluster.  Note the pair of parallel rows of red-orange spot on the underside of the hindwing.
A Monarch on Butterfly Weed
A Monarch feeding on nectar from the flowers of Butterfly Weed.
A mating pair of Eastern Tailed Blues.
A mating pair of Eastern Tailed Blues on a Timothy (Phleum pratense) spike.
A female (left) and male Great Spangled Fritillary.
A male Great Spangled Fritillary (right) pursuing a female.
Common Green Darner
Butterflies aren’t the only colorful insects patrolling the meadows at Boyd Big Tree Preserve.  Dragonflies including Common Green Darners are busily pursuing prey, particularly small flying insects like mosquitos, gnats, and flies.
Juvenile Broad-winged Hawk
Dragonflies themselves can become prey and are much sought after by Broad-winged Hawks. This very vocal juvenile gave us several good looks as it ventured from the forest into the skies above the upper meadow during midday.  It wasn’t yet a good enough flier to snag a dragonfly, but it will have plenty of opportunities for practice during its upcoming fall migration which, for these Neotropical raptors, will get underway later this month.

Do yourself a favor and take a trip to the Boyd Big Tree Preserve Conservation Area.  Who knows?  It might actually inspire you to convert that lawn or other mowed space into much-needed butterfly/pollinator habitat.

While you’re out, you can identify your sightings using our photographic guide—Butterflies of the Lower Susquehanna River Watershed—by clicking the “Butterflies” tab at the top this page.  And while you’re at it, you can brush up on your hawk identification skills ahead of the upcoming migration by clicking the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab.  Therein you’ll find a listing and descriptions of hawk watch locations in and around the lower Susquehanna region.  Plan to visit one or more this autumn!

Shorebirds and More at Bombay Hook National Wildlife Refuge

Have you purchased your 2023-2024 Federal Duck Stamp?  Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points.  Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States.  So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.

2023-2024 Federal Duck Stamp. Your Federal Duck Stamp is your free pass to visit the nation's National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.

Still not convinced that a Federal Duck Stamp is worth the money?  Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge.  Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.

Northern Bobwhite
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge.  At Bombay Hook, they don’t waste your money mowing grass.  Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.
Shearness Pool at Bombay Hook N.W.R.
Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools.  Birds there can often be observed at close range.  The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools.  Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.
Great Blue Heron
A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.
Semipalmated Sandpipers
Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.
Semipalmated Sandpipers and Short-billed Dowitcher
More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.
Short-billed Dowitchers
Two more Short-billed Dowitchers on the way in.
Sandpipers, Avocets, Egrets, and Mallards
Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed.  Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge.  Mallards, American Avocets, and egrets are gathered on the shore.
Short-billed Dowitchers
More Short-billed Dowitchers arriving to feed in Raymond Pool.
Semipalmated Sandpipers
Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.
Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.
Semipalmated Sandpipers, Forster's Terns, and a Short-billed Dowitcher
A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).
Short-billed Dowitchers
Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.
Short-billed Dowitchers, American Avocets, and a Snowy Egret
Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool.  A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.
Stilt Sandpiper among Short-billed Dowitchers
Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.
American Avocets
American Avocets probe the muddy bottom of Raymond Pool.
Dunlin and Short-billed Dowitchers
Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early.  Many migrating Dunlin linger at Bombay Hook into October and even November.
Least Sandpiper
This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.
Lesser Yellowlegs
Lesser Yellowlegs at Bear Swamp Pool.
Lesser Yellowlegs
Lesser Yellowlegs at Bear Swamp Pool
Greater Yellowlegs
A Greater Yellowlegs at Bear Swamp Pool.
Caspian Tern
A Caspian Tern patrolling Raymond Pool.
Marsh Wren singing
The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.
Marsh Wren Nest
This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass.  High tide licks at the roots of the cordgrass supporting the temporary domicile.
Seaside Dragonlet
By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice).  It is our only dragonfly able to breed in saltwater.  Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.
Red-winged Blackbird
Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.
Bobolink
Look up!   A migrating Bobolink passes over the dike at Shearness Pool.
Mute Swans and Canada Geese
Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.
Trumpeter Swans
A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool.  Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river.  The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century.  An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers.  Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.
Great Egret
A Great Egret prowling Shearness Pool.
Snowy Egret
A Snowy Egret in Bear Swamp Pool.
A hen Wood Duck (second from right) escorts her young.
Wood Ducks in Bear Swamp Pool.
Black-necked Stilt and young.
A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.

As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes.  Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.

Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.
Atlantic Horseshoe Crab
The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay.  We found just this one during our visit to Bombay Hook.  Man has certainly decimated populations of this ancient crustacean during recent decades.
As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.
Atlantic Marsh Fiddler Crabs
Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).
Atlantic Marsh Fiddler Crabs
Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.
Atlantic Marsh Fiddler Crab
A male Atlantic Marsh Fiddler Crab peers from its den.
Great Egret
Herons and egrets including this Great Egret are quite fond of fiddler crabs.  As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.
Green Heron
A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.
Clapper Rail
A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.
Glossy Ibis
Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.
Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide.
Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.
Ospey
An Osprey patrols the vast tidal areas opposite Shearness Pool.

No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.

Indigo Bunting
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.
Blue Grosbeak
Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook.  This one was in shrubby growth along the dike at the north end of Shearness Pool.
Trumpet Creeper and Poison Ivy
These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.
Pileated Woodpecker in Sweet Gum
The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries.  The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware.  The seed balls are a favorite winter food of goldfinches and siskins.
Red-bellied Slider and Painted Turtle
Finis Pool has no frontage on the tidal marsh but is still worth a visit.  It lies along a spur road on the tour route and is located within a deciduous coastal plain forest.  Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.
White-tailed Deity
The White-tailed Deity is common along the road to Finis Pool.
Fowler's Toad
Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.
Turk's Cap Lily
The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well.  This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.
Wild Turkey
Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.

We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon.  And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.

A Limpkin’s Journey to Pennsylvania: A Waffle House Serving Escargot at Every Exit

Mid-summer can be a less than exciting time for those who like to observe wild birds.  The songs of spring gradually grow silent as young birds leave the nest and preoccupy their parents with the chore of gathering enough food to satisfy their ballooning appetites.  To avoid predators, roving families of many species remain hidden and as inconspicuous as possible while the young birds learn how to find food and handle the dangers of the world.

But all is not lost.  There are two opportunities for seeing unique birds during the hot and humid days of July.

First, many shorebirds such as sandpipers, plovers, dowitchers, and godwits begin moving south from breeding grounds in Canada.  That’s right, fall migration starts during the first days of summer, right where spring migration left off.  The earliest arrivals are primarily birds that for one reason on another (age, weather, food availability) did not nest this year.  These individuals will be followed by birds that completed their breeding cycles early or experienced nest failures.  Finally, adults and juveniles from successful nests are on their way to the wintering grounds, extending the movement into the months we more traditionally start to associate with fall migration—late August into October.

For those of you who find identifying shorebirds more of a labor than a pleasure, I get it.  For you, July can bring a special treat—post-breeding wanderers.  Post-breeding wanderers are birds we find roaming in directions other than south during the summer months, after the nesting cycle is complete.  This behavior is known as “post-breeding dispersal”.  Even though we often have no way of telling for sure that a wandering bird did indeed begin its roving journey after either being a parent or a fledgling during the preceding nesting season, the term post-breeding wanderer still applies.  It’s a title based more on a bird sighting and it’s time and place than upon the life cycle of the bird(s) being observed.  Post-breeding wanderers are often southern species that show up hundreds of miles outside there usual range, sometimes traveling in groups and lingering in an adopted area until the cooler weather of fall finally prompts them to go back home.  Many are birds associated with aquatic habitats such as shores, marshes, and rivers, so water levels and their impact on the birds’ food supplies within their home range may be the motivation for some of these movements.  What makes post-breeding wanderers a favorite among many birders is their pop.  They are often some of our largest, most colorful, or most sought-after species.  Birds such as herons, egrets, ibises, spoonbills, stilts, avocets, terns, and raptors are showy and attract a crowd.

While it’s often impossible to predict exactly which species, if any, will disperse from their typical breeding range in a significant way during a given year, some seem to roam with regularity.  Perhaps the most consistent and certainly the earliest post-breeding wanderer to visit our region is the “Florida Bald Eagle”.  Bald Eagles nest in “The Sunshine State” beginning in the fall, so by early spring, many of their young are on their own.   By mid-spring, many of these eagles begin cruising north, some passing into the lower Susquehanna valley and beyond.  Gatherings of dozens of adult Bald Eagles at Conowingo Dam during April and May, while our local adults are nesting and after the wintering birds have gone north, probably include numerous post-breeding wanderers from Florida and other Gulf Coast States.

So this week, what exactly was it that prompted hundreds of birders to travel to Middle Creek Wildlife Management Area from all over the Mid-Atlantic States and from as far away as Colorado?

Birders observing something special at Middle Creek Wildlife Management Area on July 10, 2023.

Was it the majestic Great Blue Herons and playful Killdeer?

Great Blue Heron
Great Blue Heron and a Killdeer.

Was it the colorful Green Herons?

Green Heron
Green Heron

Was it the Great Egrets snapping small fish from the shallows?

Great Egret
Great Egret

Was it the small flocks of shorebirds like these Least Sandpipers beginning to trickle south from Canada?

Least Sandpipers
Least Sandpipers

All very nice, but not the inspiration for traveling hundreds or even thousands of miles to see a bird.

It was the appearance of this very rare post-breeding wanderer…

Limpkin at Middle Creek Wildlife Management Area
A Limpkin at Middle Creek Wildlife Management Area in Lancaster County.  The Limpkin (Aramus guarauna) is the only surviving member of the family Aramidae.

…Pennsylvania’s first record of a Limpkin, a tropical wading bird native to Florida, the Caribbean Islands, and South America.  Many observers visiting Middle Creek Wildlife Management Area had never seen one before, so if they happen to be a “lister”, a birder who keeps a tally of the wild bird species they’ve seen, this Limpkin was a “lifer”.

The Limpkin is an inhabitant of vegetated marshlands where it feeds almost exclusively upon large snails of the family Ampullariidae, including the Florida Applesnail (Pomacea paludosa), the largest native freshwater snail in the United States.

Native and Non-native Range of Florida Applesnail
In the United States, the native range of the Limpkin lies within the native range of the Florida Applesnail, shown here in gold.  Introduced populations of the snail are shown in brown.  (United States Geological Survey Nonindigenous Aquatic Species image)
A spectacular nineteenth-century rendition of the Florida Applesnail, including an egg mass, illustrated by Helen E. Lawson in Samuel S. Haldeman’s “Monograph of the Freshwater Univalve Mollusca of the United States”.

Observations of the Limpkin lingering at Middle Creek Wildlife Management Area have revealed a pair of interesting facts.  First, in the absence of Florida Applesnails, this particular Limpkin has found a substitute food source, the non-native Chinese Mystery Snail (Cipangopaludina chinensis).  And second, Chinese Mystery Snails have recently become established in the lakes, pools, and ponds at the refuge, very likely arriving as stowaways on Spatterdock (Nuphar advena) and/or American Lotus (Nelumbo lutea), native transplants brought in during recent years to improve wetland habitat and process the abundance of nutrients (including waterfowl waste) in the water.

A Chinese Mystery Snail.
The Chinese Mystery Snail is the largest freshwater snail in the Lower Susquehanna River Watershed.  (Vintage 35 mm image)
By hitching a ride on aquatic transplants like this Spatterdock, non-native freshwater snails are easily vectored into new areas outside their previous range.
Spatterdock, a native species also known as Yellow Pond Lily or Cow Lily.
Spatterdock, a native species also known as Yellow Pond Lily or Cow Lily, flowering in August.
American Lotus in flower at Middle Creek Wildlife Management Area.
Blooming American Lotus transplants in a pool at Middle Creek Wildlife Management Area during August.
The Limpkin at Middle Creek W.M.A. capturing a Chinese Mystery Snail.
The Limpkin at Middle Creek W.M.A. capturing a Chinese Mystery Snail.
The Limpkin at Middle Creek carrying a Chinese Mystery Snail.
The Limpkin at Middle Creek carrying a Chinese Mystery Snail.
Limpkin holding Chinese Mystery Snail
The Limpkin is seen here maneuvering the the snail in its bill, a set of mandibles specially adapted for extracting the bodies of large freshwater snails from their shells.
Limpkin Grasping Chinese Mystery Snail
The tweezers-like tip of the bill is used to grasp the shell by the rim of the opening or by the “trapdoor” (operculum) that protects the snail inside.
Chinese Mystery Snail
A posed Chinese Mystery Snail showing its “trapdoor”, the operculum protecting the soft body tissue when the animal withdraws inside.  The tips of the Limpkin’s bill close tightly like the end of a tweezers to grasp the operculum and remove it and the snail’s body from the shell.  (Vintage 35 mm image)
Limpkin Removing Chinese Mystery Snail from Shell
The tweezers-tipped bill, which is curved slightly to the right in some Limpkins, is slid into the shell to grasp the snails body and remove it for consumption.  The entire extraction process takes 10 to 30 seconds.

The Middle Creek Limpkin’s affinity for Chinese Mystery Snails may help explain how it was able to find its way to Pennsylvania in apparent good health.  Look again at the map showing the range of the Limpkin’s primary native food source, the Florida Applesnail.  Note that there are established populations (shown in brown) where these snails were introduced along the northern coast of Georgia and southern coast of South Carolina…

Native and Non-native Range of Florida Applesnail
Native (gold) and non-native (brown) ranges of the Florida Applesnail.  (United States Geological Survey Nonindigenous Aquatic Species image)

…now look at the latest U.S.G.S. Nonindigenous Aquatic Species map showing the ranges (in brown) of established populations of non-native Chinese Mystery Snails…

Range (in brown) of established populations of non-native Chinese Mystery Snails.  (United States Geological Survey Nonindigenous Aquatic Species image)

…and now imagine that you’re a happy-go-lucky Limpkin working your way up the Atlantic Coastal Plain toward Pennsylvania and taking advantage of the abundance of food and sunshine that summer brings to the northern latitudes.  It’s a new frontier.  Introduced populations of Chinese Mystery Snails are like having a Waffle House serving escargot at every exit along the way!

Be sure to click the “Freshwater Snails” tab at the top of this page to learn more about the Chinese Mystery Snail and its arrival in the Lower Susquehanna River Watershed.  Once there, you’ll find some additional commentary about the Limpkin and the likelihood of Everglade Snail Kites taking advantage of the presence of Chinese Mystery Snails to wander north.  Be certain to check it out.

Everglade Snail Kite
The endangered Everglade Snail Kite (Rostrhamus sociabilis plumbeus), a Florida Applesnail specialist, has survived in part due to its ability to adapt to eating the non-native Pomacea maculata applesnails which have become widespread in Florida following releases from aquaria.  The adaptation?…a larger body and bill for eating larger snails.  (National Park Service image)

A Few Plants with Wildlife Impact in June

Here’s a look at some native plants you can grow in your garden to really help wildlife in late spring and early summer.

The Larger Blue Flag (Iris versicolor) and Soft Rush (Juncus effusus) in flower in mid-June.
The showy bloom of a Larger Blue Flag (Iris versicolor) and the drooping inflorescence of Soft Rush (Juncus effusus).  These plants favor moist soils in wetlands and damp meadows where they form essential cover and feeding areas for insects, amphibians, and marsh birds.  Each is an excellent choice for helping to absorb nutrients in a rain garden or stream-side planting.  They do well in wet soil or shallow water along the edges of garden ponds too.
Smooth Shadbush
The fruits of Smooth Shadbush (Amelanchier laevis), also known as Allegheny Serviceberry, Smooth Serviceberry, or Smooth Juneberry, ripen in mid-June and are an irresistible treat for catbirds, robins, bluebirds, mockingbirds, and roving flocks of Cedar Waxwings.
Common Milkweed and Eastern Carpenter Bee
Also in mid-June, the fragrant blooms of Common Milkweed attract pollinators like Eastern Carpenter Bees,…
Common Milkweed and Honey Bee
…Honey Bees,…
Common Milkweed and Banded Hairstreak
…and butterflies including the Banded Hairstreak (Satyrium calanus).  In coming weeks, Monarch butterflies will find these Common Milkweed plants and begin laying their eggs on the leaves.  You can lend them a hand by planting milkweed species (Asclepias) in your garden.  Then watch the show as the eggs hatch and the caterpillars begin devouring the foliage.  Soon, they’ll pupate and, if you’re lucky, you’ll be able to watch an adult Monarch emerge from a chrysalis!

Mother’s Day Flowers

Let’s have a look at some of the magnificent native wildflowers blooming on this Mother’s Day in forests and thickets throughout the Lower Susquehanna River Watershed.

Rue Anemone
Rue Anemone (Thalictrum thalictroides) is a fragile little wildflower of open woods.
Blackhaw
The Blackhaw (Viburnum prunifolium) is a shrub of thickets and forest borders. The flower clusters mature into an abundance of berries relished by birds and other wildlife.
Black Cherry
A large Black Cherry tree adorned in white blossoms can be pleasing to the eye and very beneficial for pollinating insects.
Common Pawpaw
The Common Pawpaw is a small tree of damp well-drained soils.  Where conditions are ideal, it may form clonal colonies in the forest understory.  The deep maroon blooms mature into banana-like fruits, hence its alternate common name “Custard Apple”.
Mayapple
The Mayapple or Mandrake (Podophyllum peltatum) commonly blankets the forest floor.  It, like other ephemeral wildflowers, is vulnerable to invasions by non-native shrubs that leaf out earlier in spring than native species and rob them of sunlight during their already abbreviated growing season.

Aren’t they spectacular?

Why not take a stroll to have a look at these and other plants you can find growing along a nearby woodland trail?  You could check out some place new and get a little exercise too!

If you go, please tread lightly, take your camera, mind your manners, and…

What’s Flying Right Now

Here’s a quick look at just a few of the birds that have been moving northward in recent days…

Gray Catbird
Find a thicket, a garden with plenty of shrubs, or a brushy woodland and you’ll find Gray Catbirds.  Many will stay in the Lower Susquehanna River Watershed to nest.
Orchard Oriole
Orchard Orioles favor old meadows with large shade trees.  Some remain to nest.
Common Yellowthroat
The Common Yellowthroat is one of the more than two dozen species of warblers found regularly during spring migration.   It nests locally in wet thickets.
Warbling Vireo
The Warbling Vireo can be found singing among the branches of large streamside trees like this Green Ash.
Blue Jay
Unlike the preceding species which do the majority of their migrating by night, Blue Jays move north by daylight.  Keep an eye on the sky and you just might see a flock or two go by!

Photo of the Day

Female Red-winged Blackbirds in a cattail wetland.
This year’s unseasonably mild weather has hastened the northward migration of many birds.  Neotropical species including warblers and Broad-winged Hawks are already pushing through the lower Susquehanna region in numbers.  But it seems the spring movement may be a little bit protracted for female Red-winged Blackbirds, which are still rolling through by the hundreds despite the males being here, some of them defending nesting territories, since late February.  No sense of urgency?

Photo of the Day

Potting Trees and Shrubs
Potting small bare root tree and shrub seedlings in a compost mixture allows you to nurture them with some tender loving care in your home nursery at least through the summer.   The extra attention they’ll get allows you to plant them as larger specimens this autumn or thereafter.   We’ve found that giving native species this quick head start in the nursery significantly improves their survival rate in the field.

Photo of the Day

A Reforestation Project
Visible in the background of this image, an infestation of invasive Emerald Ash Borer larvae has killed the trees in a woodlot comprised exclusively of Green Ash.  Left standing, the dead snags provide excellent habitat for a number of animal species including cavity-nesting birds like woodpeckers, known consumers of these destructive larvae.  To reforest the mowed field in the foreground, a variety of native deciduous trees have been planted.  In areas where a diversity of trees are not present to furnish a source of seeds for natural succession, manually planting an array of seedlings provides some insurance against the risk of allowing establishment of a single pioneer species such as the vulnerable Green Ash.  The white plastic tubes on the young trees offer protection from the ever-browsing White-tailed Deity.

Purple Haze Across the Fields

Have you noticed a purple haze across the fields right now?  If so, you may have wondered, “What kind of flowers are they?”

A purple haze of color stretches across fields not already green with cold-season crops like winter wheat.

Say hello to Purple Dead Nettle (Lamium purpureum), a non-native invasive species that has increased its prevalence in recent years by finding an improved niche in no-till cropland.  Purple Dead Nettle, also known as Red Dead Nettle, is native to Asia and Europe.  It has been a familiar early spring “weed” in gardens, along roadsides, and in other disturbed ground for decades.

Purple Dead Nettle owes its new-found success to the timing of its compressed growing season.  Its tiny seeds germinate during the fall and winter, after crops have been harvested and herbicide application has ended for the season.  The plants flower early in the spring and are thus particularly attractive to Honey Bees and other pollinators looking for a source of energy-rich nectar as they ramp up activity after winter lock down.  In many cases, Purple Dead Nettle has already completed its flowering cycle and produced seeds before there is any activity in the field to prepare for planting the summer crop.  The seeds spend the warmer months in dormancy, avoiding the hazards of modern cultivation that expel most other species of native and non-native plants from the agricultural landscape.

Purple Dead Nettle in Bloom
Flowering Purple Dead Nettle as a volunteer cover crop among last year’s corn stubble in a no-till field.
Purple Dead Nettle
Like the flowers of orchids, Purple Hedge Nettle blossoms are described as yoke shaped or bilateral (zygomorphic).  Psychedelic experiences are produced only through observation, not by ingestion.  A member of the mint family, its edible young leaves and tops have nutritional value, making a unique addition to salads and soups.
Purple Dead Nettle, Common Dandelion, and Shepherd's Purse
We call them “weeds”, but what do we know?  Purple Dead Nettle, Common Dandelion, and Shepherd’s Purse (Capsella bursa-pastoris), three edible non-native invasive species with similar life cycles seen here flowering along a rural road among fields where intensive farming is practiced.  Shepard’s Purse, like many members of the mustard family, is already producing seeds at the bottom of the flower cluster by the time the uppermost buds open for business.
Spraying Herbicide
In preparation for seeding of a warm-season crop,  herbicide is applied on a no-till field to kill Shepherd’s Purse and other cool-season plants.  To help prevent sediment and nutrient discharge from lands where high-intensity agriculture is practiced, no-till methods are used to reduce runoff from the areas of bare soil that would otherwise be created by traditional plowing.

While modern farming has eliminated a majority of native plant and animal species from agricultural lands of the lower Susquehanna valley, its crop management practices have simultaneously invited vigorous invasion by a select few non-native species.   High-intensity farming devotes its acreage to providing food for a growing population of people—not to providing wildlife habitat.  That’s why it’s so important to minimize our impact on non-farm lands throughout the remainder of the watershed.  If we continue subdividing, paving, and mowing more and more space, we’ll eventually be living in a polluted semi-arid landscape populated by little else but non-native invasive plants and animals.  We can certainly do better than that.

Off To The Races

Trying to get a favorable place to nest before others arrive, the “early birds” are presently racing north through the lower Susquehanna valley.  Check out these sightings from earlier today…

Ring-necked Ducks
A pair of Ring-necked Ducks.
Hooded Mergansers
Hooded Mergansers, two males and a female.
American Wigeons
A pair of American Wigeons.
A male Canvasback.
A male Canvasback.
Eastern Phoebe
During these chilly days of late winter, this hardy Eastern Phoebe finds sustenance by seizing flying insects along the water’s edge.
An American Robin in classic worm-hunting posture.
Possibly our most familiar sign of spring, an American Robin in classic worm-hunting posture.
A Common Grackle in a maple tree that is starting to flower.
An iridescent Common Grackle in a maple tree that is beginning to flower.
A male Red-winged Blackbird singing near a small patch of cattails.
A male Red-winged Blackbird singing from a perch near a small patch of cattails.  During the spring migration, noisy flocks of males compete for a breeding territory at these sites.  Each of the victors defends his spot and awaits the arrival of a female mate while the losers move on to vie for their own breeding location farther north.

Time to get outside and have a look.  The spectacle of spring migration passes quickly.  You don’t want to miss it!

Plantings for Wet Lowlands

This linear grove of mature trees, many of them nearly one hundred years old, is a planting of native White Oaks (Quercus alba) and Swamp White Oaks (Quercus bicolor).

Imagine the benefit of trees like this along that section of stream you’re mowing or grazing right now.  The Swamp White Oak in particular thrives in wet soils and is available now for just a couple of bucks per tree from several of the lower Susquehanna’s County Conservation District Tree Sales.  These and other trees and shrubs planted along creeks and rivers to create a riparian buffer help reduce sediment and nutrient pollution.  In addition, these vegetated borders protect against soil erosion, they provide shade to otherwise sun-scorched waters, and they provide essential wildlife habitat.  What’s not to love?

Swamp White Oak
Autumn leaf of a Swamp White Oak

The following native species make great companions for Swamp White Oaks in a lowland setting and are available at bargain prices from one or more of the County Conservation District Tree Sales now underway…

Red Maple
The Red Maple is an ideal tree for a stream buffer project. They do so well that you should limit them to 10% or less of the plants in your project so that they don’t overwhelm slower-growing species.
River Birch
The River Birch (Betula nigra) is a multi-trunked tree of lowlands.  Large specimens with arching trunks help shade waterways and provide a source of falling insects for surface-feeding fish.  Its peeling bark is a distinctive feature.
Common Winterberry
The Common Winterberry with its showy red winter-time fruit is a slow-growing shrub of wet soils.  Only female specimens of this deciduous holly produce berries, so you need to plant a bunch to make sure you have both genders for successful pollination.
American Robins feeding on Common Winterberry.
An American Robin feeding on Common Winterberry.
Common Spicebush
Common Spicebush is a shrub of moist lowland soils.  It is the host plant for the Spicebush Swallowtail butterfly and produces small red berries for birds and other wildlife.  Plant it widely among taller trees to provide native vegetation in the understory of your forest.
Common Spicebush foliage and berries.
Common Spicebush foliage and berries in the shade beneath a canopy of tall trees.
Common Pawpaw
The Common Pawpaw a small shade-loving tree of the forest understory.
Common Pawpaw
Common Pawpaw is a colony-forming small tree which produces a fleshy fruit.  It is the host plant for the caterpillars of the Zebra Swallowtail.
Buttonbush
The Buttonbush is a shrub of wet soils.  It produces a round flower cluster, followed by this globular seed cluster.
Eastern Sycamore
And don’t forget the Eastern Sycamore, the giant of the lowlands.  At maturity, the white-and-tan-colored bark on massive specimens makes them a spectacular sight along stream courses and river shores.  Birds ranging from owls, eagles, and herons to smaller species including the Yellow-throated Warbler rely upon them for nesting sites.
Yellow-crowned Night Herons Nesting in an Eastern Sycamore
Yellow-crowned Night Herons, an endangered species in Pennsylvania, nesting in an Eastern Sycamore.

So don’t mow, do something positive and plant a buffer!

Act now to order your plants because deadlines are approaching fast.  For links to the County Conservation District Tree Sales in the Lower Susquehanna River Watershed, see our February 18th post.

Time to Order Your Trees for Spring Planting

County Conservation District Tree Sales are underway throughout the Lower Susquehanna River Watershed.  Now is the time to order for pickup in April.  The prices are a bargain and the selection is fabulous.  For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue).  And don’t forget to order bundles of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local birds.  They’re only $12.00 for a bundle of 10—can’t beat that deal!

Cumberland County Conservation District Annual Tree Seedling Sale—

Orders due by: Friday, March 24, 2023

Pickup on: Thursday, April 20, 2023 or Friday, April 21, 2023

Showy Northeast Meadow Mix
Don’t mow it.  Plant a meadow or pollinator garden instead.
Showy Northeast Meadow Mix
Both Cumberland and Perry Counties are offering a native warm-season grass and wildflower seed mix for planting your own meadow or pollinator garden.  Perry County is also taking orders for a seed mix specifically formulated to grow plants for attracting hummingbirds and butterflies.

Dauphin County Conservation District Seedling Sale—

Orders due by: Monday, March 20, 2023

Pickup on: Thursday, April 20, 2023 or Friday, April 21, 2023

American Goldfinch atop an Eastern Hemlock
The Eastern Hemlock, Pennsylvania’s official state tree, is an excellent choice for addition to your landscape or reforestation project.  It tolerates rocky soils and its cones are an excellent source of food for birds ranging from chickadees to finches.

Lancaster County Annual Tree Seedling Sale—

Orders due by: Friday, March 10, 2023

Pickup on: Thursday, April 13, 2023

Northern Red Oak
The handsome yet underused Northern Red Oak is a sturdy long-lived native tree that is ideal for street-side, lawn, and reforestation plantings.  In spring, it can be a magnet for migrating Neotropical birds when its flowers attract a wide variety of tiny insects to its upper reaches.  Unlike many other oaks, this species is a relatively fast grower.

Lebanon County Conservation District Tree and Plant Sale—

Orders due by: Wednesday, March 8, 2023

Pickup on: Friday, April 7, 2023

Pileated Woodpecker feeding on Black Gum berries.
In autumn, even after the bright red foliage is gone, the berries of mature Black Gum (Nyssa sylvatica) trees attract a wide variety of birds like this Pileated Woodpecker.  The Lebanon County Conservation District is offering Black Gum, also known as Black Tupelo, during their 2023 tree sale.  Why not order and plant a half dozen or more?

Perry County Conservation District Tree Sale—(click on 2023 Tree Sale Brochure tab when it scrolls across the page)

Orders due by: March 22, 2023

Pickup on: Thursday, April 13, 2023

Female Eastern Bluebird with Food for Young
The Perry County Conservation District is not only offering plants during this year’s sale, you can also purchase bluebird nest boxes for just $12.00 each!
Riparian Buffer at 15 Years
For less than the cost of one year of mowing, this stream corridor in Conewago Township, Dauphin County was reforested by the owner with hundreds of native trees, the majority purchased through County Conservation District Tree Sale events spanning a period of several years.  By replacing bare soil and mowed areas, the riparian buffer created by these plantings has significantly reduced the nutrient and sediment loads that were polluting the small stream therein known as Brill’s Run.  With determination and not a lot of money, you can do it too.
Maples, Pin Oaks, Eastern White Pines, and other trees in the Brill's Run riparian buffer.
But don’t forget the Eastern White Pines!

Berries and More on a Bitter Cold Morning

The annual arrival of hoards of American Robins to devour the fruits found on the various berry-producing shrubs and trees in the garden at susquehannawildlife.net headquarters happened to coincide with this morning’s bitter cold temperatures.  Here are photos of some of those hungry robins—plus shots of the handful of other songbirds that joined them for a frosty feeding frenzy.

American Robins consuming Common Winterberry fruits.
American Robins consuming Common Winterberry fruits.
American Robin
One of between one hundred and two hundred American Robins seen feeding on berries at susquehannawildife.net headquarters this morning.
Dark-eyed junco
A Dark-eyed Junco searching the ground for seeds.
American Robin in a "Hollywood Juniper".
An American Robin in the boughs of a “Hollywood Juniper”, a cultivar of the Chinese Juniper (Juniperus chinensis ‘Torulosa’, also known as J. c. ‘Kaizuka’).
A Carolina Wren on the peanut feeder.
A Carolina Wren on the peanut feeder.
American Robin
An American Robin searching for fallen berries beneath a holly.
A female Eastern Bluebird.
A female Eastern Bluebird.
American Robin
An American Robin takes a break from the buffet.
A Carolina Chickadee preparing to pluck a sunflower seed from a tube feeder.
A Carolina Chickadee preparing to pluck a sunflower seed from a tube feeder.
American Robins feeding on "Hollywood Juniper" berries.
American Robins feeding on “Hollywood Juniper” berries.
Red-breasted Nuthatch
One of two Red-breasted Nuthatches spending the week at susquehannawildlife.net headquarters.
Eastern Bluebird. American Robin, and Tufted Titmouse.
A male Eastern Bluebird among the crowd in the garden’s trees.
American Robin and Common Winterberry, a native deciduous holly.
An American Robin and Common Winterberry, a native deciduous holly.
A Carolina Wren investigates a tree cavity, a potential nest site.
A Carolina Wren investigates a tree cavity, a potential nest site in coming weeks.

Photo of the Day

American Robin
We don’t have a resident groundhog at susquehannawildlife.net headquarters, but the arrival of an American Robin to begin cleaning the abundance of berries from our holly trees and shrubs gives us the idea that spring is just around the corner.  We could hardly be happier.

Photo of the Day

Northern Mockingbird
From a lookout atop an Eastern Hemlock, a Northern Mockingbird maintains a vigil over a garden full of berry-producing plants. To assure their survival during cold weather, these bold birds will vigorously defend winter food supplies on hollies, viburnums, poison-ivy, bittersweet, and other fruit-producing trees, shrubs, and vines, often swooping in to startle and flush birds and other animals that approach its stores too closely.