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LIFE IN THE LOWER SUSQUEHANNA RIVER WATERSHED
A Natural History of Conewago Falls—The Waters of Three Mile Island

Uncle Tyler Dyer wants to know, “What happened to all the butterflies. Man, I’m worried. I’m seeing just one or two of the common species, but not in anywhere near the numbers we usually see by this time of year.” His question is a rhetorical one. He understands the contributing factors that have conspired to diminish, fragment, and even eliminate populations of butterflies and other insects throughout much of the lower Susquehanna valley.
Here at susquehannawildlife.net headquarters, we’ve observed but four or five species this year, and only several individuals of each. Despite our efforts to devote every square inch of available space in our gardens and pools to plants that benefit butterflies and other insects, we can only do so much. Much like other areas of the region, our compact parcel is surrounded by more and more pavement, manicured lawn, and properties planted with commercially produced ornamentals cultivated to look good, grow fast, and not attract “insect pests”. To butterflies, the land beyond our little border is a desert.
Among human habitations where there may sometimes be an effort to provide nectar for pollinators including adult butterflies, the surrounding landscape still remains largely composed mostly of asphalt, cement, mowed turf, and tanbark-covered ground that offers few if any host plants for butterfly eggs and caterpillars, and no winter cover for our resident species. In the farmlands, herbicide-tolerant crops treated with systemic insecticides are grown on ground sterilized against the emergence of any potential butterfly host species—no mustards, milkweeds, native grasses, violets, or indigenous legumes. Just look at that barren ag ground in January, no butterfly survives there.
Application of herbicides and insecticides is more widespread than you might think. Few plots of ground in proximity to human activity experience a year without use of at least one dose of a pesticide. Along the highway, in the field, on the lawn, in the trees, across the forest, in the soil, on the water, around the garden, about the house—it’s pure poison to insects and/or their host plants and they don’t survive it.


Here at susquehannawildlife.net headquarters, our most common butterfly this summer has been the Monarch. Considering the numeric loses this species has experienced in recent years, its good news for the Monarchs but not so great news for the butterflies we typically see in good numbers and have not.
While their wintering habitat in Mexico faces potentially catastrophic loses due to clandestine logging and other criminal activity, Monarchs do have an advantage once they come north to breed. While other butterflies mate and deposit eggs close to where they themselves were hatched and completed their metamorphosis into adults, each in the succession of each year’s several generations of Monarchs will roam widely in search of suitable stands of milkweed upon which they can lay their eggs.

Other than in the few scattered places where habitat is managed for them, butterflies and other insects don’t fare well in proximity to human activity. This is especially true in the Lower Susquehanna River Watershed where an already neat and tidy culture has become obsessed with a cleared landscape and thus compulsively sprays, mows, whacks, and cuts away at the habitat critical to the survival of hundreds, maybe thousands, of species of these indispensable invertebrates. Take a walk or a drive and look around. Other than a hanging basket of flowers that may or may not have been treated with poisonous neonicotinoids, there’s no place for a butterfly, let alone its progeny, to survive. Just listen…that motorized, grinding racket outside is the sound of butterfly habitat being destroyed, April through November, day after day, year after year.
As for Uncle Ty, he’s putting in a little more effort. Scattered sanctuaries for butterflies aren’t going to save them from the wholesale fragmentation of their habitat that has been intensifying in recent years. Less than 1% of landowners actually rewild property or manage at the ecosystem level, so more must get involved. Uncle Ty asked us to pass along this guide to Butterfly Habitat Management from the United States Department of Agriculture’s Natural Resources Conservation Service. He’s used tips from the publication for years and highly recommends the list of host plants it includes.


No, this is not an account chronicling a British invasion, especially during the month we celebrate the country’s 250th anniversary of its separation from commonwealth rule. This is instead a short introduction to some of the numerous regional insects of the order Coleoptera.
Globally, there are somewhere between 300,000 and 2 million or more species of beetles, depending on who’s doing the enumerating. A commonly settled upon number is 400,000, which makes beetles account for about 40% of the known insect species and 25% of the animals in the world. In the United States, there are almost 30,000 species.
Morphological variation among individuals in a population of beetles is easily observed. These differences often manifest into adaptations that provide a survival advantage and speciation. It therefore comes as no surprise that beetles were among the life forms studied by Charles Darwin as he pondered natural selection and preparation of The Origin of Species.

If it’s organic, living or dead, a beetle has probably become specially adapted to fill the role of reducing it or consuming it. Beetles are indispensable members of the ecosystems they inhabit. Many function as scavengers, breaking down dead plants or animals into the basic nutrients that renew life. There are those that consume rotting wood, dung, and carrion. Sometimes they eat these things when people would rather they didn’t—such as moist wood that’s part of a house. Then there are species of beetles that are herbivores, consuming parts of the plants that get their energy directly from the sun. There are beetles that eat stems, roots, leaves, buds, flowers, nectar, sap, seeds, and fruit—many specializing in particular plant species or families. Some of these herbivorous primary consumers come to our attention because their populations explode in response to our creation of monocultures of their favored food, but many more act as pollinators for plants critical to human survival. Linnaean Society treasurer Rathvon made these herbivore-plant relationships the focus of many of his nineteenth-century writings, particularly as they affected agriculture. On the upper trophic levels, there are the predatory beetles that feed on herbivores or other carnivores. Many of these species have narrowed their choice of prey to one or fewer species. For example, there is a beetle that feeds only on the subterranean nymphs of cicadas.

Ladies and gentlemen, the beetles…

















Homo sapiens owes much of its success as a species to an acquired knowledge of how to make, control, and utilize fire. Using fire to convert the energy stored in combustible materials into light and heat has enabled humankind to expand its range throughout the globe. Indeed, humans in their furless incomplete mammalian state may have never been able to expand their populations outside of tropical latitudes without mastery of fire. It is fire that has enabled man to exploit more of the earth’s resources than any other species. From cooking otherwise unpalatable foods to powering the modern industrial society, fire has set man apart from the rest of the natural world.
In our modern civilizations, we generally look at the unplanned outbreak of fire as a catastrophe requiring our immediate intercession. A building fire, for example, is extinguished as quickly as possible to save lives and property. And fires detected in fields, brush, and woodlands are promptly controlled to prevent their exponential growth. But has fire gone to our heads? Do we have an anthropocentric view of fire? Aren’t there naturally occurring fires that are essential to the health of some of the world’s ecosystems? And to our own safety? Indeed there are. And many species and the ecosystems they inhabit rely on the periodic occurrence of fire to maintain their health and vigor.

Man has been availed of the direct benefits of fire for possibly 40,000 years or more. Here in the Lower Susquehanna River Watershed, the earliest humans arrived as early as 12,000 years ago—already possessing skills for using fire. Native plants and animals on the other hand, have been part of the ever-changing mix of ecosystems found here for a much longer period of time—millions to tens of millions of years. Many terrestrial native species are adapted to the periodic occurrence of fire. Some, in fact, require it. Most upland ecosystems need an occasional dose of fire, usually ignited by lightning (though volcanism and incoming cosmic projectiles are rare possibilities), to regenerate vegetation, release nutrients, and maintain certain non-climax habitat types.
But much of our region has been deprived of natural-type fires since the time of the clearcutting of the virgin forests during the eighteenth and nineteenth centuries. This absence of a natural fire cycle has contributed to degradation and/or elimination of many forest and non-forest habitats. Without fire, a dangerous stockpile of combustible debris has been collecting, season after season, in some areas for a hundred years or more. Lacking periodic fires or sufficient moisture to sustain prompt decomposition of dead material, wildlands can accumulate enough leaf litter, thatch, dry brush, tinder, and fallen wood to fuel monumentally large forest fires—fires similar to those recently engulfing some areas of the American west. So elimination of natural fire isn’t just a problem for native plants and animals, its a potential problem for humans as well.

To address the habitat ailments caused by a lack of natural fires, federal, state, and local conservation agencies are adopting the practice of “prescribed fire” as a treatment to restore ecosystem health. A prescribed fire is a controlled burn specifically planned to correct one or more vegetative management problems on a given parcel of land. In the Lower Susquehanna River Watershed, prescribed fire is used to…
Let’s look at some examples of prescribed fire being implemented right here in our own neighborhood…

























In Pennsylvania, state law provides landowners and crews conducting prescribed fire burns with reduced legal liability when the latter meet certain educational, planning, and operational requirements. This law may help encourage more widespread application of prescribed fire in the state’s forests and other ecosystems where essential periodic fire has been absent for so very long. Currently in the Lower Susquehanna River Watershed, prescribed fire is most frequently being employed by state agencies on state lands—in particular, the Department of Conservation and Natural Resources on State Forests and the Pennsylvania Game Commission on State Game Lands. Prescribed fire is also part of the vegetation management plan at Fort Indiantown Gap Military Reservation and on the land holdings of the Hershey Trust. Visitors to the nearby Gettysburg National Military Park will also notice prescribed fire being used to maintain the grassland restorations there.
For crews administering prescribed fire burns, late March and early April are a busy time. The relative humidity is often at its lowest level of the year, so the probability of ignition of previous years’ growth is generally at its best. We visited with a crew administering a prescribed fire at Middle Creek Wildlife Management Area last week. Have a look…

















Prescribed burns aren’t a cure-all for what ails a troubled forest or other ecosystem, but they can be an effective remedy for deficiencies caused by a lack of periodic episodes of naturally occurring fire. They are an important option for modern foresters, wildlife managers, and other conservationists.
If you’re feeling the need to see summertime butterflies and their numbers just don’t seem to be what they used to be in your garden, then plan an afternoon visit to the Boyd Big Tree Preserve along Fishing Creek Valley Road (PA 443) just east of U.S. 22/322 and the Susquehanna River north of Harrisburg. The Pennsylvania Department of Conservation and Natural Resources manages the park’s 1,025 acres mostly as forested land with more than ten miles of trails. While located predominately on the north slope of Blue Mountain, a portion of the preserve straddles the crest of the ridge to include the upper reaches of the southern exposure.

Fortunately, one need not take a strenuous hike up Blue Mountain to observe butterflies. Open space along the park’s quarter-mile-long entrance road is maintained as a rolling meadow of wildflowers and cool-season grasses that provide nectar for adult butterflies and host plants for their larvae.















Do yourself a favor and take a trip to the Boyd Big Tree Preserve Conservation Area. Who knows? It might actually inspire you to convert that lawn or other mowed space into much-needed butterfly/pollinator habitat.
While you’re out, you can identify your sightings using our photographic guide—Butterflies of the Lower Susquehanna River Watershed—by clicking the “Butterflies” tab at the top this page. And while you’re at it, you can brush up on your hawk identification skills ahead of the upcoming migration by clicking the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab. Therein you’ll find a listing and descriptions of hawk watch locations in and around the lower Susquehanna region. Plan to visit one or more this autumn!
Here’s a look at some native plants you can grow in your garden to really help wildlife in late spring and early summer.







This month, the International Union for Conservation of Nature (I.U.C.N.) added the Migratory Monarch Butterfly (Danaus plexippus plexippus) to its “Red List of Threatened Species”, classifying it as endangered. Perhaps there is no better time than the present to have a look at the virtues of replacing areas of mowed and manicured grass with a wildflower garden or meadow that provides essential breeding and feeding habitat for Monarchs and hundreds of other species of animals.

If you’re not quite sure about finally breaking the ties that bind you to the cult of lawn manicuring, then compare the attributes of a parcel maintained as mowed grass with those of a space planted as a wildflower garden or meadow. In our example we’ve mixed native warm season grasses with the wildflowers and thrown in a couple of Eastern Red Cedars to create a more authentic early successional habitat.

Still not ready to take the leap. Think about this: once established, the wildflower planting can be maintained without the use of herbicides or insecticides. There’ll be no pesticide residues leaching into the soil or running off during downpours. Yes friends, it doesn’t matter whether you’re using a private well or a community system, a wildflower meadow is an asset to your water supply. Not only is it free of man-made chemicals, but it also provides stormwater retention to recharge the aquifer by holding precipitation on site and guiding it into the ground. Mowed grass on the other hand, particularly when situated on steep slopes or when the ground is frozen or dry, does little to stop or slow the sheet runoff that floods and pollutes streams during heavy rains.
What if I told you that for less than fifty bucks, you could start a wildflower garden covering 1,000 square feet of space? That’s a nice plot 25′ x 40′ or a strip 10′ wide and 100′ long along a driveway, field margin, roadside, property line, swale, or stream. All you need to do is cast seed evenly across bare soil in a sunny location and you’ll soon have a spectacular wildflower garden. Here at the susquehannawildllife.net headquarters we don’t have that much space, so we just cast the seed along the margins of the driveway and around established trees and shrubs. Look what we get for pennies a plant…

Here’s a closer look…






All this and best of all, we never need to mow.
Around the garden, we’ve used a northeast wildflower mix from American Meadows. It’s a blend of annuals and perennials that’s easy to grow. On their website, you’ll find seeds for individual species as well as mixes and instructions for planting and maintaining your wildflower garden. They even have a mix specifically formulated for hummingbirds and butterflies.


Nothing does more to promote the spread and abundance of non-native plants, including invasive species, than repetitive mowing. One of the big advantages of planting a wildflower garden or meadow is the opportunity to promote the growth of a community of diverse native plants on your property. A single mowing is done only during the dormant season to reseed annuals and to maintain the meadow in an early successional stage—preventing reversion to forest.
For wildflower mixes containing native species, including ecotypes from locations in and near the Lower Susquehanna River Watershed, nobody beats Ernst Conservation Seeds of Meadville, Pennsylvania. Their selection of grass and wildflower seed mixes could keep you planting new projects for a lifetime. They craft blends for specific regions, states, physiographic provinces, habitats, soils, and uses. Check out these examples of some of the scores of mixes offered at Ernst Conservation Seeds…
We’ve used their “Showy Northeast Native Wildflower and Grass Mix” on streambank renewal projects with great success. For Monarchs, we really recommend the “Butterfly and Hummingbird Garden Mix”. It includes many of the species pictured above plus “Fort Indiantown Gap” Little Bluestem, a warm-season grass native to Lebanon County, Pennsylvania, and milkweeds (Asclepias), which are not included in their northeast native wildflower blends. More than a dozen of the flowers and grasses currently included in this mix are derived from Pennsylvania ecotypes, so you can expect them to thrive in the Lower Susquehanna River Watershed.




In addition to the milkweeds, you’ll find these attractive plants included in Ernst Conservation Seed’s “Butterfly and Hummingbird Garden Mix”, as well as in some of their other blends.







Why not give the Monarchs and other wildlife living around you a little help? Plant a wildflower garden or meadow. It’s so easy, a child can do it.



It’s sprayed with herbicides. It’s mowed and mangled. It’s ground to shreds with noisy weed-trimmers. It’s scorned and maligned. It’s been targeted for elimination by some governments because it’s undesirable and “noxious”. And it has that four letter word in its name which dooms the fate of any plant that possesses it. It’s the Common Milkweed, and it’s the center of activity in our garden at this time of year. Yep, we said milk-WEED.
Now, you need to understand that our garden is small—less than 2,500 square feet. There is no lawn, and there will be no lawn. We’ll have nothing to do with the lawn nonsense. Those of you who know us, know that the lawn, or anything that looks like lawn, are through.
Anyway, most of the plants in the garden are native species. There are trees, numerous shrubs, some water features with aquatic plants, and filling the sunny margins is a mix of native grassland plants including Common Milkweed. The unusually wet growing season in 2018 has been very kind to these plants. They are still very green and lush. And the animals that rely on them are having a banner year. Have a look…





We’ve planted a variety of native grassland species to help support the milkweed structurally and to provide a more complete habitat for Monarch butterflies and other native insects. This year, these plants are exceptionally colorful for late-August due to the abundance of rain. The warm season grasses shown below are the four primary species found in the American tall-grass prairies and elsewhere.






There was Monarch activity in the garden today like we’ve never seen before—and it revolved around milkweed and the companion plants.











SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York.
She ate only toaster pastries…that’s it…nothing else. Every now and then, on special occasions, when a big dinner was served, she’d have a small helping of mashed potatoes, no gravy, just plain, thank you. She received all her nutrition from several meals a week of macaroni and cheese assembled from processed ingredients found in a cardboard box. It contains eight essential vitamins and minerals, don’t you know? You remember her, don’t you?
Adult female butterflies must lay their eggs where the hatched larvae will promptly find the precise food needed to fuel their growth. These caterpillars are fussy eaters, with some able to feed upon only one particular species or genus of plant to grow through the five stages, the instars, of larval life. The energy for their fifth molt into a pupa, known as a chrysalis, and metamorphosis into an adult butterfly requires mass consumption of the required plant matter. Their life cycle causes most butterflies to be very habitat specific. These splendid insects may visit the urban or suburban garden as adults to feed on nectar plants, however, successful reproduction relies upon environs which include suitable, thriving, pesticide-free host plants for the caterpillars. Their survival depends upon more than the vegetation surrounding the typical lawn will provide.
The Monarch (Danaus plexippus), a butterfly familiar in North America for its conspicuous autumn migrations to forests in Mexico, uses the milkweeds (Asclepias) almost exclusively as a host plant. Here at Conewago Falls, wetlands with Swamp Milkweed (Asclepias incarnata) and unsprayed clearings with Common Milkweed (A. syriaca) are essential to the successful reproduction of the species. Human disturbance, including liberal use of herbicides, and invasive plant species can diminish the biomass of the Monarch’s favored nourishment, thus reducing significantly the abundance of the migratory late-season generation.



Butterflies are good indicators of the ecological health of a given environment. A diversity of butterfly species in a given area requires a wide array of mostly indigenous plants to provide food for reproduction. Let’s have a look at some of the species seen around Conewago Falls this week…








The spectacularly colorful butterflies are a real treat on a hot summer day. Their affinity for showy plants doubles the pleasure.
By the way, I’m certain by now you’ve recalled that fussy eater…and how beautiful she grew up to be.
SOURCES
Brock, Jim P., and Kaufman, Kenn. 2003. Butterflies of North America. Houghton Mifflin Company. New York, NY.