Despite what seemed to be a chilly early spring, the bright green leaves that unfold to close the canopy of our deciduous forests were dense and casting shade by the last days of April. For northbound migrants, this fresh foliage provides the cover they need for foraging, resting, and, for those that will stick around to breed in the lower Susquehanna valley, nesting.
We recall many occasions when sparse foliage during the first days of May seemed to delay the big push of Neotropical species, but the seasonal arrival of these birds in 2025 is thus far mostly ahead of schedule. This absence of delay is due in part to the lushness of the oaks. Some stands have not only leafed out, but are finished flowering and have added up to 12 inches of new branch growth. We spent these early hours of May among the oaks. Here’s a look at the Neotropical migrants and other species we found…
Dozens of species of Neotropical warblers have arrived in the Lower Susquehanna River Watershed. Some will nest locally while others will continue along flight paths that will ultimately take them far to our north. We spotted this Cape May Warbler in the upper reaches of a Northern Red Oak feeding among its spent flowers. Its destination: the spruce forests of northern New England and Canada.Slightly larger than the warblers are the vireos, including this Yellow-throated Vireo that has arrived to nest in an oak-maple bottomland.Always a fan favorite, we found this marvelously tropical Baltimore Oriole among the foliage of a flowering White Oak.This young Red-tailed Hawk has survived its first winter. It appears to have learned from experience how to hunt from advantageous locations such as this Chestnut Oak along a utility right-of-way where prey may include numerous squirrels as well as mice and other small mammals.We were taking a break beneath this Pin Oak when suddenly an American Kestrel arrived to begin scolding a trespasser that was straying a bit too close to its nest;……the intruder, a Northern Harrier, soon took the hint and continued on its way.While among the oaks, it pays to check the understory where Neotropical thrushes including the Veery are arriving to fill the forests with their melodious songs.Following their nocturnal flights, the last of the season’s northbound White-throated Sparrows may presently be found spending the day in the cover of the oak woodland understory.
The movements of our migratory birds typically continue through much of the month of May. And peak numbers of Neotropical species often occur sometime during the second week of the month. But with habitat at the ready, favorable flight conditions could facilitate quick arrival and/or passage of the bulk of the remaining migrants during the coming week. You may want to venture out sooner rather than later—but watch your step!
The Mayapple, a native wildflower also known as the Mandrake, is now in bloom among the oak leaf litter.And the Canada Mayflower (Maianthemum canadense), a native species also known as Wild Lily-of-the-valley, is blooming in moist mixed-oak forests.At the edge of an oak-maple woodland, we found this lingering White-crowned Sparrow foraging among the leaf litter and fresh, shiny foliage of Poison Ivy. Songbirds can spend their days scratching the ground among these leaves of three. If you do it, you’ll be scratching something else, and you may need an ocean of calamine lotion to boot!
During Saturday’s Prescribed Fire Demonstration at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area, we noticed just how fast some species of wildlife return to areas subjected to burns administered to maintain grassland habitat and reduce the risk of high-intensity blazes.
Pennsylvania Game Commission crews ignite a back fire to contain a prescribed burn along its downwind/upslope perimeter during a demonstration at Middle Creek Wildlife Management Area on Saturday.Visitors observe a fire planned to maintain this section of the refuge as warm-season grassland. A species with roots several feet deep, the light-colored vegetation is Indiangrass, a plant adapted to thrive following periodic episodes of wildfire. Prescribed fire can be used to replace naturally occurring infernos with much safer controlled burns that eliminate successional and invasive plants to promote the establishment of Indiangrass and other native warm-season species including Big Bluestem, Little Bluestem, Switchgrass, and a variety of wildflowers as well.Even as the fire reached its brief peak of intensity, we noticed birds already attracted to the site…Dozens of recently arrived Tree Swallows swept in to patrol for flying insects as the burn was in progress.One even stopped by to have a look inside the kestrel nest box as fire approached the dry stand of goldenrod on the slope behind.Red-tailed Hawks and other raptors, including nocturnal owls, are frequently the first visitors attracted to the scene of a prescribed burn or wildfire. In grassland and successional habitats, they come looking for any vulnerable voles or mice that may be moving about looking for cover.
These three Eastern Meadowlarks spent the morning in the grassland areas adjacent to the prescribed fire site, mostly where a burn had been conducted one week prior. During the demonstration, one even perched and sang from the oak trees in the museum/visitor’s center parking lot.
Following the Prescribed Fire Demonstration, we decided to pay a visit to some of the parcels where burns had been administered one week earlier on the north side of Middle Creek’s main impoundment. We found a surprising amount of activity.
Apparently feeding upon slightly heat-treated seeds, sparrows were found by the dozens. White-crowned (left), White-throated (right), Song, and Savannah Sparrows were identified.This Downy Woodpecker was finding something to its liking among the scorched leaves, stems, and twigs.American Robins seem to find areas with lightly burnt vegetation and ash-dusted soil advantageous for finding invertebrates following a fire.We found this flock of Red-winged Blackbirds, Browned-headed Cowbirds, and a few European Starlings feeding throughout a grassland field cleared of early-successional growth by a prescribed fire administered one week ago.They seemed to favor gleaning seeds from among the lightly burned areas of the plot.Nearby, in an island of unburned grass in the same field, we found yet another Eastern Meadowlark, our fourth of the day. High-intensity agriculture, particularly early hay mowing and pesticide treatments, have mostly eliminated this and other grassland species from modern farms. Management practices like prescribed fire and delayed mowing (no spinning blades until at least early August) can maintain ideal grassland habitat for stunningly colorful blackbirds including nesting Eastern Meadowlarks, Bobolinks, and many other species as well.A male American Kestrel at a nest box located among Middle Creek’s warm-season and cool-season grassland habitats, the former maintained by prescribed fire, the latter by delayed mowing.
We’ve seen worse, but this winter has been particularly tough for birds and mammals in the Lower Susquehanna River Watershed. Due to the dry conditions of late summer and fall in 2024, the wild food crop of seeds, nuts, berries, and other fare has been less than average. The cold temperatures make insects hard to come by. Let’s have a look at how some of our local generalist and specialist species are faring this winter.
House Sparrows (bottom) and House Finches (top) are generalists. To survive and thrive, they are adapted to a variety of habitats and types of food. House Sparrows live almost anywhere man-made structures are found. They are true omnivores and will eat almost anything, especially if they see something else try to eat it first. The House Sparrow’s close association with humans has allowed it to become the most widespread and successful living avian dinosaur. On a cold night, they’ll take shelter either within dense vegetation alongside a building or within the structure itself. Though not nearly as cosmopolitan, the House Finch has successfully colonized much of the eastern United States after escaping from captivity as a cage bird in New York during the middle of the twentieth century. Upon being trans-located here from the arid southwest, they adapted to suburbs and farmlands consuming primarily a granivore diet of seeds supplemented with seasonally available berries. They quickly became accustomed to offerings at bird-feeding stations as well. To survive the harsh winters in the northern sections of their range, eastern populations of House Finches are developing a pattern of migration. These movements are most evident in late fall when dozens or sometimes hundreds can be seen heading south over regional hawk-counting stations.Though they require dead trees for nesting and as places to find the grubs and adult insects upon which they primarily feed, woodpeckers including the Northern Flicker are generalists, seldom passing by a supply of fruits like these Poison Ivy berries as a source of winter food. Flickers regularly visit suburban areas where they’ll drop by at bird-feeding stations for suet. During the warmer months, they are the woodpecker most frequently seen on the ground where swarms of ants garner their full attention.The Pileated Woodpecker is seldom found outside of mature forests where it digs relentlessly to remove grubs and other infestations from dead wood. But it is not a true specialist……it too finds a supply of Poison Ivy berries to be indispensable during a cold winter day.The Ruby-crowned Kinglet is a generalist, feeding mostly on insects, but also consuming small seeds and some berries, including those of Poison Ivy. It nests well to our north in tall spruces and other evergreens. During migration and in winter, the Ruby-crowned Kinglet may be found in deciduous trees, brush, and tall grass in habitats ranging from forests to parks and suburbia. This male is displaying its seldom-seen red crown.The Golden-crowned Kinglet, seen here on a Poison Ivy vine, is more of a specialist than the Ruby-crowned species, though the two will often occur in mixed groups during the winter. The Golden-crowned Kinglet nests in Spruce-Fir forests and in conifers within mixed woodlands. Even during migration, and particularly in winter, these birds are seldom found far from a stand of large evergreens within which they find shelter for the night.The Hermit Thrush’s generalist lifestyle allows it to survive cold season weather in the Lower Susquehanna River Watershed. During summer, it breeds in coniferous and mixed woods from the northern parts of our valley north into Canada and feeds primarily on worms, insects, and other arthropods. During migration and in winter, the Hermit Thrush becomes a regular visitor to deciduous forests, woody parks and suburbs, particularly where a supply of wild berries is available to supplement its diet.Iconic as it pulls earthworms from lawns during the warmer months, the generalist American Robin is fully dependent upon a crop of berries to survive winter conditions in the lower Susquehanna valley. The drought afflicted wild food crop of 2024 has led to fewer robins spending the season here and has delayed the northward push of migrating birds until the ground thaws and the earthworms make the ground rumble once again.It’s insects for the nestlings during spring and summer, then berries through the winter for the cheerful Eastern Bluebirds, another generalist species.American Crows are an excellent example of a generalist species. They’ll go anywhere to find food and they’ll eat almost anything. Like the House Sparrow and several other generalists, they adapt very well to human activity and actually thrive on it. Garbage anyone?Another career generalist is the widely worshiped White-tailed Deity, a species adapted to nearly all man-made landscapes with adequate vegetation upon which to browse. Pushed to the limit during severe weather, some individuals will consume carrion and even resort to cannibalism.You might think the Great Blue Heron is a specialist. Nope, it’s an accomplished generalist. Great blues will live, feed, and breed on almost any body of fresh or brackish water. And their diet includes almost anything that swims. In winter, you’ll even see them in fields hunting mice and voles.The Red-shouldered Hawk is a generalist with a diet ranging from amphibians and reptiles to small rodents and large insects. Mostly regarded as a species of bottomlands, they’ll frequent woodland edges, roadsides, and suburbia during the winter months.During its periodic winter visits to the region, the American Tree Sparrow feeds on seeds among the grasses and forbs of semi-open country with scattered short shrubs and trees. A generalist species, it will show up at backyard bird-feeding stations, particularly during periods of inclement weather. In summer, the American Tree Sparrow nests in tundra with growths of stunted willows and spruce and their diet includes insects as a source of protein for themselves and their young.The White-crowned Sparrow has similar winter habitat preferences to the tree sparrows……it becomes adaptable and something of a generalist when searching for food during bad winter storms.The Savannah Sparrow is an omnivore favoring insects in summer and seeds in winter. Though very closely tied to its grassland habitat year-round, snow cover can push these birds to enter woodier environs to consume fruits like these rose hips.Dusk and dawn during the short days of winter are the prime hunting times of a mammal specialist, the Short-eared Owl. Its presence in the lower Susquehanna valley is dependent on two dominant factors: extensive grassland habitat and an adequate population of the owl’s favored food, the Meadow Vole (Microtus pennsylvanicus). The Short-eared Owl’s requirements as a specialist species make finding a suitable place to live difficult. Unlike the generalist birds and mammals that often adapt to the widespread man-made disturbances in the region, populations of specialists frequently become fragmented, reduced in abundance, and subject to extirpation.The Meadow Vole is a generalist rodent that can be abundant in grasslands, early successional growth, fallow fields, marshlands, and, of course, meadows. They are primarily herbivores, but will occasionally consume insects and other arthropods. Usually nocturnal, some individuals venture out along their surface runways during daylight hours becoming vulnerable to diurnal raptors including kestrels, harriers, and buteos.A Short-eared Owl in near darkness patrolling a grassland for Meadow Voles.The eastward expansion of the Coyote (Canis latrans), a species of western North America’s grasslands and scrublands, and its progressive mixing with the Wolf (Canis lupus) in these eastern extensions of its range, has produced an expanding population of very adaptable generalists we call Eastern Coyotes (Canis latrans var.). These omnivorous canines colonized the Lower Susquehanna River Watershed during the last four decades to replace extirpated wolves as the top-tier natural predator in the region. Their primary diet includes Meadow Voles in grasslands and other small mammals along woodland edges and in successional habitats. Seasonally, they consume the berries of numerous wild plants. Slightly larger than their western ancestors, Eastern Coyotes with the admixture of Wolf genetics can subdue small ungulates. Like other apex predators, they are attracted to vulnerable prey and thus play a crucial role in culling the weak and diseased among their potential quarry species to assure the health and potential of populations of these species as long-term sources of food energy. The benefit to the prey species is however largely diluted in populous areas of the northeast; most venison consumed by Eastern Coyotes here is in the form of road kill. During periods of extended snow cover when small rodents and other foods become inaccessible, Eastern Coyotes, particularly young individuals, will wander into new areas seeking sustenance. Sometimes they venture into cities and suburbs where they explore the neighborhoods in search of garbage and pet foods placed outside the home. (Video clip courtesy of Tyler and Grace Good. Click image to view.)
Wildlife certainly has a tough time making it through the winter in the lower Susquehanna valley. Establishing and/or protecting habitat that includes plenty of year-round cover and sources of food and water can really give generalist species a better chance of survival. But remember, the goal isn’t to create unnatural concentrations of wildlife, it is instead to return the landscape surrounding us into more of a natural state. That’s why we try to use native plants as much as possible. And that’s why we try to attract not only a certain bird, mammal, or other creature, but we try to promote the development of a naturally functioning ecosystem with a food web, a diversity of pollinating plants, pollinating insects, and so on. Through this experience, we stand a better chance of understanding what it takes to graduate to the bigger job at hand—protecting, enhancing, and restoring habitats needed by specialist species. These are efforts worthy of the great resources that are sometimes needed to make them a success. It takes a mindset that goes beyond a focus upon the welfare of each individual animal to instead achieve the discipline to concentrate long-term on the projects and processes necessary to promote the health of the ecosystems within which specialist species live and breed. It sounds easier than it is—the majority of us frequently become distracted.
Being an individual from a population of a very successful generalist species is no guarantee of survival. This Eastern Gray Squirrel fell from a tall tree when the limbs became ice covered during a storm earlier this month. Just a freak accident? Maybe, but mistakes like this are often fatal in the natural world. This squirrel’s passing may seem brutal, but it provides a better opportunity for other squirrels and animals that share its food and cover requirements to make it through the winter. And those survivors that didn’t suffer such a fatal mistake or, more importantly, don’t possess a vulnerability that may have contributed to such a mishap will have a chance to pass those traits on to a new generation. This squirrel as an individual is gone, its species lives on, and may be stronger for its passing.Pennsylvania Game Commission crews maintain a grassland ecosystem for Short-eared Owls and other specialists using prescribed fire to prevent succession beyond its earliest stages. Among the additional specialist species benefiting from this management tool are Monarchs and other butterflies whose host plants survive early-season fire, but not competition with woody vines, shrubs, trees, and invasive herbaceous growth.
On the wider scale, it’s of great importance to identify and protect the existing and potential future habitats necessary for the survival of specialist species. And we’re not saying that solely for their benefit. These protection measures should probably include setting aside areas on higher ground that may become the beach intertidal zone or tidal marsh when the existing ones become inundated. And it may mean finally getting out of the wetlands, floodplains, and gullies to let them be the rain-absorbing, storm-buffering, water purifiers they spent millennia becoming. And it may mean it’s time to give up on building stick structures on tinderbox lands, especially hillsides and rocky outcrops with shallow, eroding soils that dry to dust every few years. We need to think ahead and stop living for the view. If you want to enjoy the view from these places, go visit and take plenty of pictures, or a video, that’s always nice—then live somewhere else. Each of these areas includes ecosystems that meet the narrow habitat requirements of many of our specialist species, and we’re building like fools in them. Then we feign victimhood and solicit pity when the calamity strikes: fires, floods, landslides, and washouts—again and again. Wouldn’t it be a whole lot smarter to build somewhere else? It may seem like a lot to do for some specialist animals, but it’s not. Because, you see, we should and can live somewhere else—they can’t.
The Allegheny Woodrat (Neotoma magister), a threatened species in Pennsylvania and a critically imperiled species in Maryland, is a habitat specialist requiring the forested rocky slopes, talus-flanked ridgetops, and caves of the Ridge and Valley Province for its nest sites and survival. Isolated populations survived within similar environs in the lower Susquehanna River valley’s Piedmont Province and on South Mountain through at least the first half of the twentieth century, but have since been extirpated. Human encroachment that fragments their habitat and promotes exposure to parasite-hosting mammals including the Raccoon (Procyon lotor), carrier of the Raccoon Roundworm (Baylisascarius procyonis), could prove fatal to remaining populations of this native mammal. (National Park Service image by Rick Olsen)The American Oystercatcher (Haematopus palliatus) is a specialist species that uses its highly adapted bill to feed on marine invertebrates including mollusks, few of which are actually oysters. Reliant upon tidal ecosystems for its survival, many of the seashore animals that make up this wader’s diet are themselves specialist species. Oystercatchers spend nearly their entire lives in tidal marshes or within the intertidal zone on beaches. They also frequent rocky jetties, particularly during high tide. This individual was photographed near the mouth of Chesapeake Bay in Northhampton County, Virginia, a location that, when the waters of the Atlantic started rising over 10,000 years ago, was the lower Susquehanna valley about 60 miles from the river’s mouth at present-day Norfolk Canyon along the edge of the continental shelf. Get the drift?
It’s hard to believe, but for almost two months now, sandpipers, plovers, and terns have been filtering south through the Lower Susquehanna River Watershed on their way to the Atlantic coastline as they complete the first leg of their long autumn migration—a journey that will take some species all the way to the far reaches of the South American continent for winter.
Migrating Caspian Terns and an Osprey take a break on a lower Susquehanna valley gravel bar.
As August draws to a close, these early birds are being joined by widespread nocturnal flights of Neotropical migrants—those species, primarily songbirds, on their way to wintering grounds which lie exclusively south of the continental United States.
To catch a glimpse of these night-flying avians, your best bet may be to position yourself on the crest of a ridge or along a linear break in the forest such as a utility right-of-way where waves of warblers, vireos, flycatchers, and other Neotropical passerines sometimes feed on invertebrates after making landfall at daybreak. Pick a place where the trees are bathed in the warm light of the rising sun and be there by 7 A.M. E.D.T. The activity can be tremendous, but it usually ends between 8 and 9.
One of more than half a dozen Red-eyed Vireos seen during a daybreak fallout at Second Mountain Hawkwatch in Lebanon County earlier this week.During sunrise, a just-arrived Blackburnian Warbler checks the foliage of a ridgetop Red Maple for insects.A Least Flycatcher quietly searches the shrubby growth along the forest edge for a morning meal.A Black-and-white Warbler in the first hour of sunlight after a big nocturnal flight.A Scarlet Tanager having a look around. This individual and other migrating birds may be here for a few days before moving on.A Black-throated Green Warbler in a treetop glowing with the light of sunrise.A hungry Chestnut-sided Warbler inspects limbs, leaves, and twigs looking for nourishment after an all-night flight.
Diurnal migrants, birds that make their movements during the daylight hours, are ramping up their flights now as well. Broad-winged Hawks, Bald Eagles, and falcons are currently being tallied at hawk-counting stations throughout the northeast. Many of those lookouts are seeing Ruby-throated Hummingbirds, swallows, and other daytime migrants too.
While many diurnal migrants gather in flocks as a method of defense against predators, Broad-winged Hawks congregate as a reconnaissance measure. Not waiting around for strong autumn winds to be deflected upwards by the region’s numerous ridges, these Neotropical birds migrate early enough in the season to rely upon thermal updrafts from sun-heated surfaces to provide lift, gain altitude, and save energy during their long trip. By traveling in groups, there are collectively able to better locate and utilize rising air columns as they progress southwesterly along their route. Broad-winged Hawks travel to Central and South America for winter. Their numbers in the Lower Susquehanna River Watershed will peak during the third week of September.Many American Kestrels transit our area in August and early September, a time coinciding with flights of migratory dragonflies, insects upon which they and another falcon, the Merlin, frequently prey.Chimney Swifts are fast-flying diurnal migrants. They are beginning to congregate into larger groups in preparation for their departure. Some are already on the move.Cedar Waxwings are currently being seen in roving flocks throughout the region. Their migration is less of a point-to-point flight and more of a continuous wandering in search of berries and other wild foods.Not everyone is yet ready to go. American Goldfinches are still in the midst of their nesting cycle and won’t begin leaving until the young are on their own and colorful adult males like this one are beginning to molt into their drab winter plumage.
Autumn migration flights are an ever-changing process, with different species peaking at different times throughout the season. In these months just after the nesting season, each of these species is more numerous than at any other time of the year. And of course, the more often we as observers get out and have a look, the more of them we’ll see.
Be certain to click the “Birds” tab at the top of this page for a photo guide to the species you’re likely to see passing through the lower Susquehanna valley this fall. Nearly four months of autumn hawk migration flights lie ahead, so don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab to find a hawk-counting station near you, then stop by for a visit or two. See you there!
A large nightfall liftoff of nocturnal migrating birds is shown here on this evening’s State College, Pennsylvania, Doppler Radar base velocity loop. Returns indicating birds moving toward the radar site (the white dot in the center of the color mass) are shown in green, and those indicating birds moving away are indicated in red. This is a big flight headed generally in a south-southwest direction. (NOAA/National Weather Service Doppler Radar image)
With nearly all of the Neotropical migrants including Broad-winged Hawks gone for the year, observers and counters at eastern hawk watches are busy tallying numbers of the more hardy species of diurnal raptors and other birds. The majority of species now coming through will spend the winter months in temperate and sub-tropical areas of the southern United States and Mexico.
Here is a quick look at the raptors seen this week at two regional counting stations: Kiptopeke Hawk Watch near Cape Charles, Virginia, and Second Mountain Hawk Watch at Fort Indiantown Gap, Pennsylvania.
The hawk-watching platform at Kiptopeke State Park is located along Chesapeake Bay near the southern tip of Delmarva Peninsula. In autumn, thousands of raptors and other birds migrate through the Atlantic Coastal Plain Province. Those that follow the shorelines south frequently concentrate in spectacular numbers before crossing the mouths of the bays they encounter. This phenomenon makes both Cape May, New Jersey, on Delaware Bay and Kiptopeke, Virginia, on Chesapeake Bay exceptional places to experience fall flights of migrating birds.A Sharp-shinned Hawk is counted as it swoops by the owl decoy at Second Mountain Hawk Watch at Fort Indiantown Gap Military Reservation. Migrating raptors save energy by riding updrafts of air created by winds blowing against the slopes of the mountainsides in the Ridge and Valley Province.A Sharp-shinned Hawk passes the lookout at Second Mountain Hawk Watch. “Sharp-shins” are currently the most numerous migrants both on the coast and at inland counting stations.A Sharp-shinned Hawk nearly passes observers unnoticed as it skims the treetops.A Sharp-shinned Hawk eyes up an owl decoy. Under cover of darkness, nocturnal owls could rather easily prey upon young and small adult hawks and falcons, both on the nest or at roost. Accordingly, many diurnal raptors instinctively harass owls to drive them from their presence. An owl decoy at the lookout helps attract migrating birds for a closer look.An adult Cooper’s Hawk flaps its way past a counting station. Like the similar Sharp-shinned Hawk, the larger Cooper’s Hawk is a member of the genus Accipiter. As a proportion of the annual fall Accipiter flight, the Cooper’s Hawk is more numerous at coastal hawk watches than at inland sites. (Editor’s Update: As of autumn, 2024, the Cooper’s Hawk and the American Goshawk have been placed in the genus Astur. The Sharp-shinned Hawk remains in the genus Accipiter. Though the Cooper’s Hawk and Sharp-shinned Hawk are oft times confused by observers due to their like appearance, it turns out that the two are not as closely related as originally believed. Neither is an offshoot of the other, nor do they descend from an immediate common ancestor. Their resemblance may instead be a case of convergent evolution, most readily characterized by acquisition of similar physical traits due to shared demands for survival within their environment.)The majority of Osprey migrate along the coast, but a few are still being seen at inland hawk watches.Bald Eagles are commonly seen at both coastal and inland lookouts. Their movements continue well into late fall.A Northern Harrier illuminated by a setting sun. Northern Harriers are often still flying when many other species have gone to roost for the day.An adult male Northern Harrier, the “gray ghost”, flying in misty weather, at a time when few other birds were in the air.The American Kestrel, like our other falcons, is seen in greatest concentrations at coastal counting stations. It is our most numerous falcon.The Merlin provides only a brief observation opportunity as it passes the lookout. These falcons are dark, speedy, and easily missed as they fly by.While moving south, Merlins often accompany flights of migrating Tree Swallows, a potential food source.A Merlin consumes a dragonfly. Eating is no reason to stop moving.The “Tundra Peregrine” is an arctic-breeding Peregrine Falcon that travels a distance of over 6,000 miles to southern South America for winter. It is strictly a migratory species in our region with numbers peaking during the first two weeks of October each year. These strong fliers have little need for the updrafts from mountain ridges, inland birds often observed flying in a north to south direction. The majority of “Tundra Peregrines” are observed following coastlines, with some migrating offshore to make landfall at points as far south as Florida and the Caribbean islands before continuing across water again to reach the northern shores of Central and South America. This “Tundra Peregrine” is a juvenile bird on its first southbound trip.
During coming days, fewer and fewer of these birds will be counted at our local hawk watches. Soon, the larger raptors—Red-tailed Hawks, Red-shouldered Hawks, and Golden Eagles—will be thrilling observers. Cooler weather will bring several flights of these spectacular species. Why not plan a visit to a lookout near you? Click on the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab at the top of this page for site information and a photo guide to identification. See you at the hawk watch!
It’s not all hawks at the hawk watch. Even the coastal sites are now seeing fun birds like the playful Common Raven on a regular basis.Coastal locations are renowned places to see migrating songbirds in places outside of their typical habitat. Here a flock of Eastern Meadowlarks has set down in the top of a Loblolly Pine (Pinus taeda) in downtown Cape Charles, Virginia, not far from Kiptopeke Hawk Watch.
Grasshoppers are perhaps best known for the occasions throughout history when an enormous congregation of these insects—a “plague of locusts”—would assemble and rove a region to feed. These swarms, which sometimes covered tens of thousands of square miles or more, often decimated crops, darkened the sky, and, on occasion, resulted in catastrophic famine among human settlements in various parts of the world.
The largest “plague of locusts” in the United States occurred during the mid-1870s in the Great Plains. The Rocky Mountain Locust (Melanoplus spretus), a grasshopper of prairies in the American west, had a range that extended east into New England, possibly settling there on lands cleared for farming. Rocky Mountain Locusts, aside from their native habitat on grasslands, apparently thrived on fields planted with warm-season crops. Like most grasshoppers, they fed and developed most vigorously during periods of dry, hot weather. With plenty of vegetative matter to consume during periods of scorching temperatures, the stage was set for populations of these insects to explode in agricultural areas, then take wing in search of more forage. Plagues struck parts of northern New England as early as the mid-1700s and were numerous in various states in the Great Plains through the middle of the 1800s. The big ones hit between 1873 and 1877 when swarms numbering as many as trillions of grasshoppers did $200 million in crop damage and caused a famine so severe that many farmers abandoned the westward migration. To prevent recurrent outbreaks of locust plagues and famine, experts suggested planting more cool-season grains like winter wheat, a crop which could mature and be harvested before the grasshoppers had a chance to cause any significant damage. In the years that followed, and as prairies gave way to the expansive agricultural lands that presently cover most of the Rocky Mountain Locust’s former range, the grasshopper began to disappear. By the early years of the twentieth century, the species was extinct. No one was quite certain why, and the precise cause is still a topic of debate to this day. Conversion of nearly all of its native habitat to cropland and grazing acreage seems to be the most likely culprit.
The critically endangered Eskimo Curlew (Numenius borealis), a species not photographed since 1962 and not confirmed since 1963, fed on Rocky Mountain Locusts during its spring migration through the Great Plains. Excessive hunting and conversion of grasslands to agriculture are believed responsible for the bird’s demise. (United States Fish and Wildlife Service image by Christina Nelson)
In the Mid-Atlantic States, the mosaic of the landscape—farmland interspersed with a mix of forest and disturbed urban/suburban lots—prevents grasshoppers from reaching the densities from which swarms arise. In the years since the implementation of “Green Revolution” farming practices, numbers of grasshoppers in our region have declined. Systemic insecticides including neonicotinoids keep grasshoppers and other insects from munching on warm-season crops like corn and soybeans. And herbicides including 2,4-D (2,4-Dichlorophenoxyacetic acid) have, in effect, become the equivalent of insecticides, eliminating broadleaf food plants from the pasturelands and hayfields where grasshoppers once fed and reproduced in abundance. As a result, few of the approximately three dozen species of grasshoppers with ranges that include the Lower Susquehanna River Watershed are common here. Those that still thrive are largely adapted to roadsides, waste ground, and small clearings where native and some non-native plants make up their diet.
Here’s a look at four species of grasshoppers you’re likely to find in disturbed habitats throughout our region. Each remains common in relatively pesticide-free spaces with stands of dense grasses and broadleaf plants nearby.
CAROLINA GRASSHOPPER
Dissosteira carolina
The Carolina Grasshopper, also known as the Carolina Locust or Quaker, is one of the band-winged grasshoppers. It is commonly found along roadsides and on other bare ground near stands of tall grass and broadleaf plants.The Carolina Grasshopper is variable in color, ranging from very dark brown……to a rich tan or khaki shade. These earth-tone colors provide the insect with effective camouflage while spending time on the ground.The Carolina Grasshopper is most readily detected and identified when it flies. The colors of the wings resemble those of the Mourning Cloak butterfly.Carolina Grasshoppers are among the preferred victims of Great Black Wasps (Sphex pensylvanicus). A female wasp stings the grasshopper to paralyze it, then drags it away to one of numerous cells in an underground burrow where she lays an egg on it. The body of the disabled grasshopper then provides nourishment for the larval wasp.
DIFFERENTIAL GRASSHOPPER
Melanoplus differentialis
Differential Grasshopper nymph with small “fairy wings”.An adult female Differential Grasshopper with fully developed wings.An adult female Differential Grasshopper
TWO-STRIPED GRASSHOPPER
Melanoplus bivittatus
An early-stage Two-striped Grasshopper nymph.A Two-striped Grasshopper nymph in a later stage.An adult female Two-striped Grasshopper.An adult female Two-striped Grasshopper. Note the pale stripe originating at each eye and joining near the posterior end of the wings to form a V-shaped pattern.An adult female Two-striped Grasshopper.
RED-LEGGED GRASSHOPPER
Melanoplus femurrubrum
An adult male Red-legged Grasshopper hiding in dense urban vegetation.The Red-legged Grasshopper may currently be our most abundant and widespread species.An adult male Red-legged Grasshopper.An adult female Red-legged Grasshopper.
Protein-rich grasshoppers are an important late-summer, early-fall food source for birds. The absence of these insects has forced many species of breeding birds to abandon farmland or, in some cases, disappear altogether.
Beginning in the early 1930s, the Western Cattle Egret (Bubulcus ibis), a notoriously nomadic species, transited the Atlantic from Africa to colonize the Americas…and they did it without any direct assistance from humans. During the 1970s and early 1980s, a nesting population of Western Cattle Egrets on river islands adjacent to the Susquehanna’s Conejohela Flats off Washington Boro was the largest inland rookery in the northeastern United States. The Lancaster County Bird Club censused the birds each August and found peak numbers in 1981 (7,580). During their years of abundance, V-shaped flocks of cattle egrets from the rookery islands ventured into grazing lands throughout portions of Lancaster, York, Dauphin, and Lebanon Counties to hunt grasshoppers. These daily flights were a familiar summertime sight for nearly two decades. Then, in the early 1980s, reductions in pastureland acreage and plummeting grasshopper numbers quickly took their toll. By 1988, the rookery was abandoned. The cattle egrets had moved on. (Vintage 33 mm image)During the summer and early fall, juvenile and adult Ring-necked Pheasants feed heavily on grasshoppers. Earlier and more frequent mowing along with declining numbers of grasshoppers on farmlands due to an increase in pesticide use were factors contributing to the crash of the pheasant population in the early 1980s.To the delight of Wild Turkeys, each of the four species of grasshoppers shown above frequents clearings and roadsides adjacent to forest areas. While changes in grasshopper distribution have been detrimental to populations of birds like pheasants, they’ve created a feeding bonanza for turkeys.Wild Turkeys feeding on an abundance of grasshoppers along a forest road.An American Kestrel feeds on a grasshopper while ignoring the abundance of Spotted Lanternflies swarming the adjacent utility pole. In Susquehanna valley farmlands, grasshopper and kestrel numbers are down. Lanternflies, on the other hand, have got it made.Maintaining areas bordering roads, forests, wetlands, farmlands, and human development in a state of early succession can provide and ideal mix of mature grasses and broadleaf plants for grasshoppers, pollinators, birds, and other wildlife.
It’s that time of year—Snow Geese on their northbound migration, more than 100,000 of them, have arrived for a stopover at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area in Lancaster and Lebanon Counties. Get there now to see scenes like these in person…
Thousands of Snow Geese gather on the main impoundment at Middle Creek. Presently, over 100,000 are estimated to be visiting the refuge.More than two thousand Tundra Swans are visiting Middle Creek right now too.A passing American Kestrel doesn’t seem to ruffle any feathers among the multitudes of Snow Geese on the lake……but when the local pair of Bald Eagles gets close, such as when they’re escorting a third party away from their nesting territory, ……Snow Geese lift off in a panic.Snow Geese rise above the shoreline treetops.Feeding and roosting Snow Geese take flight in the presence of a potential predator to not only make a direct escape, but also to seek a safer position toward the center of the flock, keeping away from the vulnerable areas on the periphery.Thousands of Snow Geese rise behind the placid group of Tundra Swans in the foreground.Observers along Willow Point Trail with swarming Snow Geese just ahead.In the midst of the thunderous cackle of thousands of Snow Geese.Don’t forget your camera……for an unforgettable photo op at Willow Point.
Can it be that time already? Most Neotropical birds have passed through the Lower Susquehanna River Watershed on their way south and the hardier species that will spend our winter in the more temperate climes of the eastern United States are beginning to arrive.
Here’s a gallery of sightings from recent days…
During the past two weeks, thousands of Broad-winged Hawks, including this adult bird, crossed the skies of the lower Susquehanna valley on their way to Central and South America for our winter.A juvenile Broad-winged Hawk passes into the sunset during its first autumn migration.Blackpoll Warblers are among the last of the Neotropical species to transit the region. They’ll continue to be seen locally through at least early October.Blue-headed Vireos are the October vireo during the fall, the other species having already continued toward tropical forests for a winter vacation.The lower Susquehanna region lies just on the northern edge of the wintering range of the Blue-gray Gnatcatcher, a species found nesting locally among treetops in deciduous woods. Look for their numbers to swell in coming days as birds from further north begin rolling through the region on their way south.Sharp-shinned Hawks delight visitors at ridgetop hawk watches during breezy late-September and early-October days. They allow closer observation than high-flying Broad-winged Hawks due to their habit of cruising just above the treetops while migrating.A Sharp-shinned Hawk glides over a lookout.Late September/early October is falcon time at area hawk-counting stations, the Peregrine Falcon often being the most anticipated species.Pale “Tundra Peregrines”, a subspecies that nests in the arctic, are strictly migratory birds in the Mid-Atlantic States. They are presently passing through on their way to South America. Like Neotropical songbirds, their long flights provide them with the luxury of never experiencing a winter season.This Carolina Saddlebags and other migratory dragonflies, which normally leave the area by mid-September, are still lingering in the lower Susquehanna region, much to the pleasure of the falcons that feed upon them.An male American Kestrel in pursuit of dragonflies found swarming around the lookout at Second Mountain Hawk Watch in Lebanon County, Pennsylvania.A male American Kestrel stooping on a dragonfly.Osprey will be among the birds of prey passing hawk watch sites during the coming two weeks. The first week of October often provides the best opportunity for seeing the maximum variety of raptors at a given site. On a good day, a dozen species are possible.Seeing cinnamon-colored juvenile Northern Harriers is symbolic of the October migration flights.Bald Eagles always thrill the crowd.In addition to raptors, resident Common Ravens are regularly sighted by observers at hawk watches and elsewhere during the fall season.Hawk-counting stations sometimes log movements of Red-bellied Woodpeckers during late September and early October. This species has extended its range into the Lower Susquehanna River Watershed only during the past one hundred years, making these seasonal migration movements a recent local phenomenon.Blue Jays are currently on the move with breeding birds from the forests of Canada and the northern United States moving south. Hundreds can be seen passing a given observation point during an ideal morning.Blue Jays find a pile of peanuts to be an irresistible treat. Provide the unsalted variety and watch the show!
Be sure to click on these tabs at the top of this page to find image guides to help you identify the dragonflies, birds, and raptors you see in the Lower Susquehanna River Watershed…
Damselflies and Dragonflies
Birds of Conewago Falls
Hawkwatcher’s Helper: Identifying Bald Eagles and other Raptors
Late August and early September is prime time to see migrating shorebirds as they pass through the lower Susquehanna valley during their autumn migration, which, believe it or not, can begin as early as late June. These species that are often assumed to spend their lives only near the seashore are regular visitors each fall as they make their way from breeding grounds in the interior of Canada to wintering sites in seacoast wetlands—many traveling as far south as Central and South America.
Low water levels on the Susquehanna River often coincide with the shorebird migration each year, exposing gravel and sand bars as well as vast expanses of muddy shorelines as feeding and resting areas for these traveling birds. This week though, rain from the remnants of Tropical Storm Fred arrived to increase the flow in the Susquehanna and inundate most of the natural habitat for shorebirds. Those on the move must either continue through the area without stopping or find alternate locations to loaf and find food.
The draining and filling of wetlands along the river and elsewhere in the region has left few naturally-occurring options. The Conejohela Flats south of Columbia offer refuge to many migrating sandpipers and their allies, the river level there being controlled by releases from the Safe Harbor Dam during all but the severest of floods. Shorebirds will sometimes visit flooded fields, but wide-open puddles and farmland resembling mudflats is more of springtime occurrence—preceding the planting and growth of crops. Well-designed stormwater holding facilities can function as habitat for sandpipers and other wildlife. They are worth checking on a regular basis—you never know what might drop in.
Right now, there is a new shorebird hot spot in the Lower Susquehanna River Watershed—Middle Creek Wildlife Management Area. The water level in the main impoundment there has been drawn down during recent weeks to expose mudflats along the periphery of nearly the entire lake. Viewing from “Stop 1” (the roadside section of the lake in front of the refuge museum) is best. The variety of species and their numbers can change throughout the day as birds filter in and out—at times traveling to other mudflats around the lake where they are hidden from view. The birds at “Stop 1” are backlit in the morning with favorable illumination developing in the afternoon.
Have a look at a few of the shorebirds currently being seen at Middle Creek…
The Killdeer is familiar as a breeding bird in the lower Susquehanna region. Large numbers can congregate ahead of and during migration on mudflats and gravel bars.The Least Sandpiper is one of the “peeps”, a group of very small shorebirds. This species is quite common at Middle Creek right now. Note the plants beginning to grow in the mud. Later in the fall, after the shorebirds are gone, raising the water level in the lake will flood these newly vegetated areas to provide an abundance of food for migrating waterfowl. This cycle can be repeated annually to support transient birds during what is often the most vulnerable time of their lives…fall migration.The Baird’s Sandpiper (Calidris bairdii) is an uncommon “peep” along the east coast during autumn migration. On the lower Susquehanna it is most frequently encountered on the vegetated gravel bars in mid-river during the last days of August or first days of September each year. The mudflats and shallows at Middle Creek are providing a suitable alternative for this juvenile bird.Numbers of Lesser Yellowlegs are increasing as flocks drop by for a rest and refueling. Bring your binoculars and your spotting scope to see the oddities that may be hiding among these groups of newly-arriving migrants.
The aquatic environs at Middle Creek attract other species as well. Here are some of the most photogenic…
Wood Ducks atop the dam.The migration of Caspian Terns coincides with that of shorebirds. Just look at that blood-red bill; it’s unmistakable. Two of these big terns are currently patrolling Middle Creek’s lake and shoreline.A female American Kestrel creates a stir among the “peeps” as it passes by. The larger falcons (the Merlin and Peregrine) can be expected to more readily take advantage of concentrations of shorebirds as a food supply.Osprey migration is underway, and many will stop at Middle Creek while in transit.Even if shorebirds aren’t your thing, there are almost always Bald Eagles to be seen at Middle Creek. See you there!
It’s been more than a week since Tropical Storm Isaias moved swiftly up the Atlantic seaboard leaving wind and flood damage in its wake. Here in the Lower Susquehanna River Watershed, the brevity of its presence minimized the effects.
Tropical Storm Isaias moves north-northeast across the Delmarva Peninsula.
You may have noticed some summertime visitors flying about during these hot humid days that followed Isaias’ passing. They’re the dragonflies.
Our familiar friend the Wandering Glider is widespread throughout the valley right now—dropping eggs on shiny automobile hoods that look to them like a nice quiet puddle of water.
The Wandering Glider is a global traveler. Here in the lower Susquehanna valley, it is currently abundant around still water and in large parking lots.
Each of the other common migratory species is here too. Look for them patrolling the skies over large bodies of water and over adjacent fallow land and meadows where tiny flying insects abound. Did these dragonflies arrive on the winds associated with the tropical storm, or did they move in with the waves of warm air that followed it? Probably a little of both.
The Common Green Darner, a large dragonfly, can be the most abundant of the migratory species. Watch for high-flying swarms in the coming weeks.The Black Saddlebags is recognized in flight by the black base of each hindwing. These patches give the appearance of a pair of saddlebags draped across the dragonfly’s thorax.The Twelve-spotted Skimmer(Libellula pulchella) is a regular migrant.The Prince Baskettail (Epitheca princeps) can occur among mixed groups of dragonflies. Despite it rarely being mentioned as a migrant, its proclivity for non-stop day-long flight makes it a likely sighting among sizeable swarms.
Big swarms of dragonflies don’t go unnoticed by predators—particularly birds. The southbound migration of kites, Broad-winged Hawks, American Kestrels, and Merlins often coincides with the swarming of migratory dragonflies in late summer. Each of these raptors will grab and feed upon these insects while on the wing—so keep an eye on the sky.
This Peregrine Falcon found the congregations of hundreds of dragonflies worthy of a closer look……and an acrobatic fly-by to disrupt the swarms. Fun, fun, fun, fun, fun.
You need to get outside and go for a walk. You’ll be sorry if you don’t. It’s prime time to see wildlife in all its glory. The songs and colors of spring are upon us!
Flooding that resulted from mid-week rains is subsiding. The muddy torrents of Conewago Falls are seen here racing by the powerhouse at the York Haven Dam.Receding waters will soon leave the parking area at Falmouth and other access points along the river high and dry.Migrating Yellow-rumped Warblers are currently very common in the riparian woodlands near Conewago Falls. They and all the Neotropical warblers, thrushes, vireos, flycatchers are moving through the Susquehanna watershed right now.A Baltimore Oriole feeds in a riverside maple tree.Ruby-crowned Kinglets are migrating through the Susquehanna valley. These tiny birds may be encountered among the foliage of trees and shrubs as they feed upon insects .Gray Catbirds are arriving. Many will stay to nest in shrubby thickets and in suburban gardens.American Robins and other birds take advantage of rising flood waters to feed upon earthworms and other invertebrates that are forced to the soil’s surface along the inundated river shoreline.Spotted Sandpipers are a familiar sight as they feed along water’s edge.The Yellow Warbler (Setophaga petechia) is a Neotropical migrant that nests locally in wet shrubby thickets. Let your streamside vegetation grow and in a few years you just might have these “wild canaries” singing their chorus of “sweet-sweet-sweet-I’m-so-sweet” on your property.
If you’re not up to a walk and you just want to go for a slow drive, why not take a trip to Middle Creek Wildlife Management Area and visit the managed grasslands on the north side of the refuge. To those of us over fifty, it’s a reminder of how Susquehanna valley farmlands were before the advent of high-intensity agriculture. Take a look at the birds found there right now.
Red-winged Blackbirds commonly nest in cattail marshes, but are very fond of untreated hayfields, lightly-grazed pastures, and fallow ground too. These habitats are becoming increasingly rare in the lower Susquehanna region. Farmers have little choice, they either engage in intensive agriculture or go broke.Nest boxes are provided for Tree Swallows at the refuge.Numbers of American Kestrels have tumbled with the loss of grassy agricultural habitats that provide large insects and small rodents for them to feed upon.White-crowned Sparrows (Zonotrichia leucophrys) are a migrant and winter resident species that favors small clumps of shrubby cover in pastures and fallow land.When was the last time you saw an Eastern Meadowlark (Sturnella magna) singing “spring-of-the-year” in a pasture near your home?And yes, the grasslands at Middle Creek do support nesting Ring-necked Pheasants (Phasianus colcichus). If you stop for a while and listen, you’ll hear the calls of “kowk-kuk” and a whir of wings. Go check it out.
And remember, if you happen to own land and aren’t growing crops on it, put it to good use. Mow less, live more. Mow less, more lives.
Second Mountain Hawk Watch is located on a ridge top along the northern edge of the Fort Indiantown Gap Military Reservation and the southern edge of State Game Lands 211 in Lebanon County, Pennsylvania. The valley on the north side of the ridge, also known as St. Anthony’s Wilderness, is drained to the Susquehanna by Stony Creek. The valley to the south is drained toward the river by Indiantown Run, a tributary of Swatara Creek.
The hawk watch is able to operate at this prime location for observing the autumn migration of birds, butterflies, dragonflies, and bats through the courtesy of the Pennsylvania Game Commission and the Garrison Commander at Fort Indiantown Gap. The Second Mountain Hawk Watch Association is a non-profit organization that staffs the count site daily throughout the season and reports data to the North American Hawk Watch Association (posted daily at hawkcount.org).
Today, Second Mountain Hawk Watch was populated by observers who enjoyed today’s break in the rainy weather with a visit to the lookout to see what birds might be on the move. All were anxiously awaiting a big flight of Broad-winged Hawks, a forest-dwelling Neotropical species that often travels back to its wintering grounds in groups exceeding one hundred birds. Each autumn, many inland hawk watches in the northeast experience at least one day in mid-September with a Broad-winged Hawk count exceeding 1,000 birds. They are an early-season migrant and today’s southeast winds ahead of the remnants of Hurricane Florence (currently in the Carolinas) could push southwest-heading “Broad-wings” out of the Piedmont Province and into the Ridge and Valley Province for a pass by the Second Mountain lookout.
The flight turned out to be steady through the day with over three hundred Broad-winged Hawks sighted. The largest group consisted of several dozen birds. We would hope there are probably many more yet to come after the Florence rains pass through the northeast and out to sea by mid-week. Also seen today were Bald Eagles, Ospreys, American Kestrels, and a migrating Red-headed Woodpecker.
Migrating Broad-winged Hawks circle on a thermal updraft above Second Mountain Hawk Watch to gain altitude before gliding away to the southwest.
Migrating insects included Monarch butterflies, and the three commonest species of migratory dragonflies: Wandering Glider, Black Saddlebags, and Common Green Darner. The Common Green Darners swarmed the lookout by the dozens late in the afternoon and attracted a couple of American Kestrels, which had apparently set down from a day of migration. American Kestrels and Broad-winged Hawks feed upon dragonflies and often migrate in tandem with them for at least a portion of their journey.
Still later, as the last of the Broad-winged Hawks descended from great heights and began passing by just above the trees looking for a place to settle down, a most unwelcome visitor arrived at the lookout. It glided in from the St. Anthony’s Wilderness side of the ridge on showy crimson-red wings, then became nearly indiscernible from gray tree bark when it landed on a limb. It was the dreaded and potentially invasive Spotted Lanternfly (Lycorma delicatula). This large leafhopper is native to Asia and was first discovered in North America in the Oley Valley of eastern Berks County, Pennsylvania in 2014. The larval stage is exceptionally damaging to cultivated grape and orchard crops. It poses a threat to forest trees as well. Despite efforts to contain the species through quarantine and other methods, it’s obviously spreading quickly. Here on the Second Mountain lookout, we know that wind has a huge influence on the movement of birds and insects. The east and southeast winds we’ve experienced for nearly a week may be carrying Spotted Lanternflies well out of their most recent range and into the forests of the Ridge and Valley Province. We do know for certain that the Spotted Lanternfly has found its way into the Lower Susquehanna River Watershed.
This adult Spotted Lanternfly landed in a birch tree behind the observers at the Second Mountain Hawk Watch late this afternoon. It was first recognized by its bright red wings as it glided from treetops on the north side of the lookout.
The humid rainy remains of Hurricane Nate have long since passed by Pennsylvania, yet mild wet weather lingers to confuse one’s sense of the seasons. This gloomy misty day was less than spectacular for watching migrating birds and insects, but some did pass by. Many resident animals of the falls are availing themselves of the opportunity to continue active behavior before the cold winds of autumn and winter force a change of lifestyle.
Warm drizzle at daybreak prompted several Northern Spring Peepers (Pseudacris crucifer crucifer) to begin calling from the wetlands in the Riparian Woodlands of Conewago Falls. An enormous chorus of these calls normally begins with the first warm rains of early spring to usher in this tiny frog’s mating season. Today, it was just a few “peeps” among anxious friends.
The tiny Northern Spring Peeper is recognized by the dark “X” across its back. Soon, shelter must be found among loose bark and fallen logs to commence hibernation. Emergence, often prompted by warm spring rains, will quickly be followed by a growing chorus of breeding calls as sometimes hundreds of these frogs assemble in vernal pools where mating will then occur.
Any additional river flow that resulted from the rains of the previous week is scarcely noticeable among the Pothole Rocks. The water level remains low, the water column is fairly clear, and the water temperatures are in the 60s Fahrenheit.
It’s no real surprise then to see aquatic turtles climbing onto the boulders in the falls to enjoy a little warmth, if not from the sun, then from the stored heat in the rocks. As usual, they’re quick to slide into the depths soon after sensing someone approaching or moving nearby. Seldom found anywhere but on the river, these skilled divers are Common Map Turtles (Graptemys geographica), also known as Northern Map Turtles. Their paddle-like feet are well adapted to swimming in strong current. They are benthic feeders, feasting upon a wide variety of invertebrates found among the stone and substrate of the river bottom.
Adult Common Map Turtles hibernate communally on the river bottom in a location protected from ice scour and turbulent flow, often using boulders, logs, or other structures as shelter from strong current. The oxygenation of waters tumbling through Conewago Falls may be critical to the survival of the turtles overwintering downstream. Dissolved oxygen in the water is absorbed by the nearly inactive turtles as they remain submerged at their hideout through the winter. Though Common Map Turtles, particularly males, may occasionally move about in their hibernation location, they are not seen coming to the surface to breathe.
The Common Map Turtles in the Susquehanna River basin are a population disconnected from that found in the main range of the species in the Great Lakes and upper Mississippi basin. Another isolated population exists in the Delaware River.
Common Map Turtles, including this recently hatched young seen in August, are often observed climbing onto rocks in the river.Note the oversize swimming fin adaptations of the feet on this adult Common Map Turtle found among the Pothole Rocks in Conewago Falls. Young and adults are capable of navigating some strong current to feed and escape danger.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
SOURCES
Committee on the Status of Endangered Wildlife in Canada. 2002. Status Report of the Northern Map Turtle. Canadian Wildlife Service. Ottawa, Ontario.
A moderate breeze from the south placed a headwind into the face of migrants trying to wing their way to winter quarters. The urge to reach their destination overwhelmed any inclination a bird or insect may have had to stay put and try again another day.
Blue Jays were joined by increasing numbers of American Robins crossing the river in small groups to continue their migratory voyages. Killdeer (Charadrius vociferous) and a handful of sandpipers headed down the river route. Other migrants today included a Cooper’s Hawk (Astur cooperii), Eastern Bluebirds (Sialia sialis), and a few Common Mergansers (Mergus merganser), House Finches (Haemorhous mexicanus), and Common Grackles (Quiscalus quiscula).
The afternoon belonged to the insects. The warm wind blew scores of Monarchs toward the north as they persistently flapped on a southwest heading. Many may have actually lost ground today. Painted Lady (Vanessa cardui) and Cloudless Sulphur butterflies were observed battling their way south as well. All three of the common migrating dragonflies were seen: Common Green Darner (Anax junius), Wandering Glider (Pantala flavescens), and Black Saddlebags (Tramea lacerata).
The warm weather and summer breeze are expected to continue as the rain and wind from Hurricane Nate, today striking coastal Alabama and Mississippi, progresses toward the Susquehanna River watershed during the coming forty-eight hours.
This Great Blue Heron was joined by numerous other fishermen and a good number of sightseers in the falls today.A colorful young Northern Water Snake (Nerodia sipedon) takes advantage of the sun-heated surface of a Pothole Rock to remain nimble and active. Cooler weather will soon compel this and other reptiles to find shelter for winter hibernation.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
A fresh breeze from the north brought cooler air and a reminder that summer is gone and autumn has arrived.
Fast-moving dark clouds provided a perfect backdrop for viewing passing diurnal migrants. Bald Eagles utilized the tail wind to cruise down the Susquehanna toward Chesapeake Bay and points further south. A migrating Merlin began a chase from which a Northern Flicker narrowly escaped by finding shelter among Pothole Rocks and a few small trees. The season’s first American Black Duck (Anas rubripes), Common Loon (Gavia immer), Yellow-bellied Sapsucker (Sphyrapicus varia), and American Pipits (Anthus rubescens) moved through.
Blue Jays continued their hesitant crossings of the river at Conewago Falls. The majority completed the journey by forming groups of a dozen or more birds and following the lead of a lone American Robin, a Northern Flicker, or, odd as it appeared, a small warbler.
By far the most numerous migrants today were swallows. Thousands of Northern Rough-winged Swallows and hundreds of Tree Swallows were on the wing in search of what was suddenly a sparse flying insect supply. To get out of the brisk wind, some of the more resourceful birds landed on the warm rocks. To satisfy their appetite, many were able to pick crawling arthropods from the surface of the boulders. They swallow them whole.
A few of the thousands of swallows seen at Conewago Falls today.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
The Neotropical birds that raised their young in Canada and in the northern United States have now logged many miles on their journey to warmer climates for the coming winter. As their density decreases among the masses of migrating birds, a shift to species with a tolerance for the cooler winter weather of the temperate regions will be evident.
Though it is unusually warm for this late in September, the movement of diurnal migrants continues. This morning at Conewago Falls, five Broad-winged Hawks (Buteo platypterus) lifted from the forested hills to the east, then crossed the river to continue a excursion to the southwest which will eventually lead them and thousands of others that passed through Pennsylvania this week to wintering habitat in South America. Broad-winged Hawks often gather in large migrating groups which swarm in the rising air of thermal updrafts, then, after gaining substantial altitude, glide away to continue their trip. These ever-growing assemblages from all over eastern North America funnel into coastal Texas where they make a turn to south around the Gulf of Mexico, then continue on toward the tropics. In the coming weeks, a migration count at Corpus Christi in Texas could tally 100,000 or more Broad-winged Hawks in a single day as a large portion of the continental population passes by. You can track their movement and that of other diurnal raptors as recorded at sites located all over North America by visiting hawkcount.org on the internet. Check it out. You’ll be glad you did.
Nearly all of the other migrants seen today have a much shorter flight ahead of them. Red-bellied Woodpeckers (Melanerpes carolinus), Red-headed Woodpeckers (Melanerpes erythrocephalus), and Northern Flickers (Colaptes auratus) were on the move. Migrating American Robins (Turdus migratorius) crossed the river early in the day, possibly leftovers from an overnight flight of this primarily nocturnal migrant. The season’s first Great Black-backed Gulls (Larus marinus) arrived. American Goldfinches are easily detected by their calls as they pass overhead. Look carefully at the goldfinches visiting your feeder, the birds of summer are probably gone and are being replaced by migrants currently passing through.
By far, the most conspicuous migrant today was the Blue Jay. Hundreds were seen as they filtered out of the hardwood forests of the diabase ridge to cautiously cross the river and continue to the southwest. Groups of five to fifty birds would noisily congregate in trees along the river’s edge, then begin flying across the falls. Many wary jays abandoned their small crossing parties and turned back. Soon, they would try the trip again in a larger flock.
Sensing that they are being watched, Blue Jays are hesitant to fly across the narrow Susquehanna at Conewago Falls without first assembling into a flock. The local constabulary often penalizes those who freelance and do not move in orderly groups.
A look at this morning’s count reveals few Neotropical migrants. With the exception of the Broad-winged Hawks and warblers, the migratory species seen today will winter in a sub-tropical temperate climate, primarily in the southern United States, but often as far north as the lower Susquehanna River valley. The individual birds observed today will mostly continue to a winter home a bit further south. Those that will winter in the area of Conewago Falls will arrive in October and later.
The Great Blue Heron (Ardea herodius) can be found year-round at Conewago Falls, provided there is open water and adequate food. Migrants from breeding colonies to the north will soon supplement the local population.The Double-crested Cormorant (Phalacrocorax auritus) is a summer resident at Conewago Falls. Migration of the local population and of those from further north will soon begin. All will be gone by the time ice forms on the river. Cormorants are often seen drying their feathers in sunlight following a series of feeding dives.
The long-distance migrating insect so beloved among butterfly enthusiasts shows signs of improving numbers. Today, more than two dozen Monarchs were seen crossing the falls and slowly flapping and gliding their way to Mexico.
CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
A couple of inches of rain this week caused a small increase in the flow of the river, just a burp, nothing major. This higher water coincided with some breezy days that kicked up some chop on the open waters of the Susquehanna upstream of Conewago Falls. Apparently it was just enough turbulence to uproot some aquatic plants and send them floating into the falls.
Piled against and upon the upstream side of many of the Pothole Rocks were thousands of two to three feet-long flat ribbon-like opaque green leaves of Tapegrass, also called Wild Celery, but better known as American Eelgrass (Vallisneria americana). Some leaves were still attached to a short set of clustered roots. It appears that most of the plants broke free from creeping rootstock along the edge of one of this species’ spreading masses which happened to thrive during the second half of the summer. You’ll recall that persistent high water through much of the growing season kept aquatic plants beneath a blanket of muddy current. The American Eelgrass colonies from which these specimens originated must have grown vigorously during the favorable conditions in the month of August. A few plants bore the long thread-like pistillate flower stems with a fruit cluster still intact. During the recent few weeks, there have been mats of American Eelgrass visible, the tops of their leaves floating on the shallow river surface, near the east and west shorelines of the Susquehanna where it begins its pass through the Gettysburg Basin near the Pennsylvania Turnpike bridge at Highspire. This location is a probable source of the plants found in the falls today.
Uprooted American Eelgrass floating into the Pothole Rocks under the power of a north wind. Note the white thread-like pistillate flower stem to the left and the small rooted specimen to the upper right. The latter is likely a plant from the creeping rootstock on the edge of a colony. As a native aquatic species, American Eelgrass is a critical link in the Susquehanna River and Chesapeake Bay food chain. Its decimation by pollution during the twentieth century led to migration pattern alterations and severe population losses for the Canvasback (Aythya valisineria) duck.American Eelgrass, a very small specimen, found growing in a low-lying Pothole Rock alongside the accumulations of freshly arriving material from upstream. Note that the creeping rootstock has leaves growing from at least three nodes on this plant. Eelgrass dislocations are regular occurrences which sometimes begin new colonies, like the small one seen here in this Diabase Pothole Rock Microhabitat.
The cool breeze from the north was a perfect fit for today’s migration count. Nocturnal migrants settling down for the day in the Riparian Woodlands at sunrise included more than a dozen warblers and some Gray Catbirds (Dumetella carolinensis). Diurnal migration was underway shortly thereafter.
A moderate flight of nocturnal migrants is indicated around NEXRAD sites in the northeastern states at 3:18 AM EDT. The outer rain bands of Hurricane Irma can be seen approaching the Florida Keys as the storm closes in on the peninsula. (NOAA/National Weather Service image)
Four Bald Eagles were counted as migrants this morning. Based on plumage, two were first-year eagles (Juvenile) seen up high and flying the river downstream, one was a second-year bird (Basic I) with a jagged-looking wing molt, and a third was probably a fourth year (Basic III) eagle looking much like an adult with the exception of a black terminal band on the tail. These birds were the only ones which could safely be differentiated from the seven or more Bald Eagles of varying ages found within the past few weeks to be lingering at Conewago Falls. There were as many as a dozen eagles which appeared to be moving through the falls area that may have been migrating, but the four counted were the only ones readily separable from the locals.
Red-tailed Hawks (Buteo jamaicensis) were observed riding the wind to journey not on a course following the river, but flying across it and riding the updraft on the York Haven Diabase ridge from northeast to southwest.
Bank Swallows (Riparia riparia) seem to have moved on. None were discovered among the swarms of other species today.
Ruby-throated Hummingbirds, Caspian Terns, Cedar Waxwings (Bombycilla cedrorum), and Chimney Swifts (Chaetura pelagica) were migrating today, as were Monarch butterflies.
Not migrating, but always fun to have around, all four wise guys were here today. I’m referring to the four members of the Corvid family regularly found in the Mid-Atlantic states: Blue Jay (Cyanocitta cristata), American Crow (Corvus brachyrhynchos), Fish Crow (Corvus ossifragus), and Common Raven (Corvus corax).
It looks like a big Blue Jay, but it’s not. This Belted Kingfisher (Megaceryle alcyon) takes a break after flying around the falls trying to shake a marauding Ruby-throated Hummingbird off its tail.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
SOURCES
Klots, Elsie B. 1966. The New Field Book of Freshwater Life. G. P. Putnam’s Sons. New York, NY.
