We’ve seen worse, but this winter has been particularly tough for birds and mammals in the Lower Susquehanna River Watershed. Due to the dry conditions of late summer and fall in 2024, the wild food crop of seeds, nuts, berries, and other fare has been less than average. The cold temperatures make insects hard to come by. Let’s have a look at how some of our local generalist and specialist species are faring this winter.
House Sparrows (bottom) and House Finches (top) are generalists. To survive and thrive, they are adapted to a variety of habitats and types of food. House Sparrows live almost anywhere man-made structures are found. They are true omnivores and will eat almost anything, especially if they see something else try to eat it first. The House Sparrow’s close association with humans has allowed it to become the most widespread and successful living avian dinosaur. On a cold night, they’ll take shelter either within dense vegetation alongside a building or within the structure itself. Though not nearly as cosmopolitan, the House Finch has successfully colonized much of the eastern United States after escaping from captivity as a cage bird in New York during the middle of the twentieth century. Upon being trans-located here from the arid southwest, they adapted to suburbs and farmlands consuming primarily a granivore diet of seeds supplemented with seasonally available berries. They quickly became accustomed to offerings at bird-feeding stations as well. To survive the harsh winters in the northern sections of their range, eastern populations of House Finches are developing a pattern of migration. These movements are most evident in late fall when dozens or sometimes hundreds can be seen heading south over regional hawk-counting stations.Though they require dead trees for nesting and as places to find the grubs and adult insects upon which they primarily feed, woodpeckers including the Northern Flicker are generalists, seldom passing by a supply of fruits like these Poison Ivy berries as a source of winter food. Flickers regularly visit suburban areas where they’ll drop by at bird-feeding stations for suet. During the warmer months, they are the woodpecker most frequently seen on the ground where swarms of ants garner their full attention.The Pileated Woodpecker is seldom found outside of mature forests where it digs relentlessly to remove grubs and other infestations from dead wood. But it is not a true specialist……it too finds a supply of Poison Ivy berries to be indispensable during a cold winter day.The Ruby-crowned Kinglet is a generalist, feeding mostly on insects, but also consuming small seeds and some berries, including those of Poison Ivy. It nests well to our north in tall spruces and other evergreens. During migration and in winter, the Ruby-crowned Kinglet may be found in deciduous trees, brush, and tall grass in habitats ranging from forests to parks and suburbia. This male is displaying its seldom-seen red crown.The Golden-crowned Kinglet, seen here on a Poison Ivy vine, is more of a specialist than the Ruby-crowned species, though the two will often occur in mixed groups during the winter. The Golden-crowned Kinglet nests in Spruce-Fir forests and in conifers within mixed woodlands. Even during migration, and particularly in winter, these birds are seldom found far from a stand of large evergreens within which they find shelter for the night.The Hermit Thrush’s generalist lifestyle allows it to survive cold season weather in the Lower Susquehanna River Watershed. During summer, it breeds in coniferous and mixed woods from the northern parts of our valley north into Canada and feeds primarily on worms, insects, and other arthropods. During migration and in winter, the Hermit Thrush becomes a regular visitor to deciduous forests, woody parks and suburbs, particularly where a supply of wild berries is available to supplement its diet.Iconic as it pulls earthworms from lawns during the warmer months, the generalist American Robin is fully dependent upon a crop of berries to survive winter conditions in the lower Susquehanna valley. The drought afflicted wild food crop of 2024 has led to fewer robins spending the season here and has delayed the northward push of migrating birds until the ground thaws and the earthworms make the ground rumble once again.It’s insects for the nestlings during spring and summer, then berries through the winter for the cheerful Eastern Bluebirds, another generalist species.American Crows are an excellent example of a generalist species. They’ll go anywhere to find food and they’ll eat almost anything. Like the House Sparrow and several other generalists, they adapt very well to human activity and actually thrive on it. Garbage anyone?Another career generalist is the widely worshiped White-tailed Deity, a species adapted to nearly all man-made landscapes with adequate vegetation upon which to browse. Pushed to the limit during severe weather, some individuals will consume carrion and even resort to cannibalism.You might think the Great Blue Heron is a specialist. Nope, it’s an accomplished generalist. Great blues will live, feed, and breed on almost any body of fresh or brackish water. And their diet includes almost anything that swims. In winter, you’ll even see them in fields hunting mice and voles.The Red-shouldered Hawk is a generalist with a diet ranging from amphibians and reptiles to small rodents and large insects. Mostly regarded as a species of bottomlands, they’ll frequent woodland edges, roadsides, and suburbia during the winter months.During its periodic winter visits to the region, the American Tree Sparrow feeds on seeds among the grasses and forbs of semi-open country with scattered short shrubs and trees. A generalist species, it will show up at backyard bird-feeding stations, particularly during periods of inclement weather. In summer, the American Tree Sparrow nests in tundra with growths of stunted willows and spruce and their diet includes insects as a source of protein for themselves and their young.The White-crowned Sparrow has similar winter habitat preferences to the tree sparrows……it becomes adaptable and something of a generalist when searching for food during bad winter storms.The Savannah Sparrow is an omnivore favoring insects in summer and seeds in winter. Though very closely tied to its grassland habitat year-round, snow cover can push these birds to enter woodier environs to consume fruits like these rose hips.Dusk and dawn during the short days of winter are the prime hunting times of a mammal specialist, the Short-eared Owl. Its presence in the lower Susquehanna valley is dependent on two dominant factors: extensive grassland habitat and an adequate population of the owl’s favored food, the Meadow Vole (Microtus pennsylvanicus). The Short-eared Owl’s requirements as a specialist species make finding a suitable place to live difficult. Unlike the generalist birds and mammals that often adapt to the widespread man-made disturbances in the region, populations of specialists frequently become fragmented, reduced in abundance, and subject to extirpation.The Meadow Vole is a generalist rodent that can be abundant in grasslands, early successional growth, fallow fields, marshlands, and, of course, meadows. They are primarily herbivores, but will occasionally consume insects and other arthropods. Usually nocturnal, some individuals venture out along their surface runways during daylight hours becoming vulnerable to diurnal raptors including kestrels, harriers, and buteos.A Short-eared Owl in near darkness patrolling a grassland for Meadow Voles.The eastward expansion of the Coyote (Canis latrans), a species of western North America’s grasslands and scrublands, and its progressive mixing with the Wolf (Canis lupus) in these eastern extensions of its range, has produced an expanding population of very adaptable generalists we call Eastern Coyotes (Canis latrans var.). These omnivorous canines colonized the Lower Susquehanna River Watershed during the last four decades to replace extirpated wolves as the top-tier natural predator in the region. Their primary diet includes Meadow Voles in grasslands and other small mammals along woodland edges and in successional habitats. Seasonally, they consume the berries of numerous wild plants. Slightly larger than their western ancestors, Eastern Coyotes with the admixture of Wolf genetics can subdue small ungulates. Like other apex predators, they are attracted to vulnerable prey and thus play a crucial role in culling the weak and diseased among their potential quarry species to assure the health and potential of populations of these species as long-term sources of food energy. The benefit to the prey species is however largely diluted in populous areas of the northeast; most venison consumed by Eastern Coyotes here is in the form of road kill. During periods of extended snow cover when small rodents and other foods become inaccessible, Eastern Coyotes, particularly young individuals, will wander into new areas seeking sustenance. Sometimes they venture into cities and suburbs where they explore the neighborhoods in search of garbage and pet foods placed outside the home. (Video clip courtesy of Tyler and Grace Good. Click image to view.)
Wildlife certainly has a tough time making it through the winter in the lower Susquehanna valley. Establishing and/or protecting habitat that includes plenty of year-round cover and sources of food and water can really give generalist species a better chance of survival. But remember, the goal isn’t to create unnatural concentrations of wildlife, it is instead to return the landscape surrounding us into more of a natural state. That’s why we try to use native plants as much as possible. And that’s why we try to attract not only a certain bird, mammal, or other creature, but we try to promote the development of a naturally functioning ecosystem with a food web, a diversity of pollinating plants, pollinating insects, and so on. Through this experience, we stand a better chance of understanding what it takes to graduate to the bigger job at hand—protecting, enhancing, and restoring habitats needed by specialist species. These are efforts worthy of the great resources that are sometimes needed to make them a success. It takes a mindset that goes beyond a focus upon the welfare of each individual animal to instead achieve the discipline to concentrate long-term on the projects and processes necessary to promote the health of the ecosystems within which specialist species live and breed. It sounds easier than it is—the majority of us frequently become distracted.
Being an individual from a population of a very successful generalist species is no guarantee of survival. This Eastern Gray Squirrel fell from a tall tree when the limbs became ice covered during a storm earlier this month. Just a freak accident? Maybe, but mistakes like this are often fatal in the natural world. This squirrel’s passing may seem brutal, but it provides a better opportunity for other squirrels and animals that share its food and cover requirements to make it through the winter. And those survivors that didn’t suffer such a fatal mistake or, more importantly, don’t possess a vulnerability that may have contributed to such a mishap will have a chance to pass those traits on to a new generation. This squirrel as an individual is gone, its species lives on, and may be stronger for its passing.Pennsylvania Game Commission crews maintain a grassland ecosystem for Short-eared Owls and other specialists using prescribed fire to prevent succession beyond its earliest stages. Among the additional specialist species benefiting from this management tool are Monarchs and other butterflies whose host plants survive early-season fire, but not competition with woody vines, shrubs, trees, and invasive herbaceous growth.
On the wider scale, it’s of great importance to identify and protect the existing and potential future habitats necessary for the survival of specialist species. And we’re not saying that solely for their benefit. These protection measures should probably include setting aside areas on higher ground that may become the beach intertidal zone or tidal marsh when the existing ones becomes inundated. And it may mean finally getting out of the wetlands, floodplains, and gullies to let them be the rain-absorbing, storm-buffering, water purifiers they spent millennia becoming. And it may mean it’s time to give up on building stick structures on tinderbox lands, especially hillsides and rocky outcrops with shallow, eroding soils that dry to dust every few years. We need to think ahead and stop living for the view. If you want to enjoy the view from these places, go visit and take plenty of pictures, or a video, that’s always nice, then live somewhere else. Each of these areas includes ecosystems that meet the narrow habitat requirements of many of our specialist species, and we’re building like fools in them. Then we feign victimhood and solicit pity when the calamity strikes: fires, floods, landslides, and washouts—again and again. Wouldn’t it be a whole lot smarter to build somewhere else? It may seem like a lot to do for some specialist animals, but it’s not. Because, you see, we should and can live somewhere else—they can’t.
The Allegheny Woodrat (Neotoma magister), a threatened species in Pennsylvania and a critically imperiled species in Maryland, is a habitat specialist requiring the forested rocky slopes, talus-flanked ridgetops, and caves of the Ridge and Valley Province for its nest sites and survival. Isolated populations survived within similar environs in the lower Susquehanna River valley’s Piedmont Province and on South Mountain through at least the first half of the twentieth century, but have since been extirpated. Human encroachment that fragments their habitat and promotes exposure to parasite-hosting mammals including the Raccoon (Procyon lotor), carrier of the Raccoon Roundworm (Baylisascarius procyonis), could prove fatal to remaining populations of this native mammal. (National Park Service image by Rick Olsen)The American Oystercatcher (Haematopus palliatus) is a specialist species that uses its highly adapted bill to feed on marine invertebrates including mollusks, few of which are actually oysters. Reliant upon tidal ecosystems for its survival, many of the seashore animals that make up this wader’s diet are themselves specialist species. Oystercatchers spend nearly their entire lives in tidal marshes or within the intertidal zone on beaches. They also frequent rocky jetties, particularly during high tide. This individual was photographed near the mouth of Chesapeake Bay in Northhampton County, Virginia, a location that, when the waters of the Atlantic started rising over 10,000 years ago, was the lower Susquehanna valley about 60 miles from the river’s mouth at present-day Norfolk Canyon along the edge of the continental shelf. Get the drift?
During your foray to view the colorful foliage of the autumn landscape, a little effort will reveal much more than meets the eye of the casual observer.
Don’t just stand there like a bump on a log, go for a stroll, watch your step, and have a look around. There’s a lot to see out there!The dense understory of this forest on Blue Mountain consists almost exclusively of Witch-hazel (Hamamelis virginiana).Witch-hazel is unique among our native flowering shrubs; it blooms in autumn.Until we experience a heavy frost, some of our most beautiful butterflies, including this Common Buckeye, remain active along forest edges.An Eastern Tailed Blue.A Pearl Crescent.A Variegated Fritillary.A Painted Lady.The arrival of large numbers of Yellow-rumped Warblers heralds the transition to movements of migrants that will spend the colder months in temperate climates like ours instead of continuing on to the tropics. In winter, small numbers of “yellow-rumps” sometimes linger in the Lower Susquehanna River Watershed, as long as a supply of wild berries is readily available.The Palm Warbler, a persistent “tail wagger”, is another late-season migrant that occasionally spends the winter. The migration of these drab “Western Palm Warblers” will be followed in coming weeks by the more yellowish “Eastern Palm Warblers” of the eastern third of the species’ breeding range in Canada.White-throated Sparrows are southbound and easily observed right now. Many will remain along our brushy forest edges for winter.Another native sparrow, the familiar Dark-eyed Junco, is now arriving.The Yellow-bellied Sapsucker is a secretive migratory woodpecker that is currently moving through the region on its way to southern pine forests for the winter. Juveniles like this one in the lower right remain close to their parents during migration and continue to beg for food.The Pileated Woodpecker is a resident species in our mature forests and woodlands. We were showered by “sawdust” as we watched this one dismantling a rotted dead tree while searching for beetle grubs and other invertebrates.You may see Eastern Bluebirds year-round in the lower Susquehanna valley, but many are migratory. A southbound push is currently transiting our area.Mix your leaf watching with hawk watching by visiting a ridgetop vista like this one at Boyd Big Tree Preserve Conservation Area on Blue Mountain north of Harrisburg and you can double your pleasure.In addition to spectacular foliage, you stand a good chance of seeing an inquisitive Common Raven……or a migrating Red-tailed Hawk that gives you a good scream as it passes by……or a majestic Bald Eagle.You may even double your fun by seeing two Common Ravens……or a couple of migrating Red-tailed Hawks……or a pair of Bald Eagles passing thru.
You too can experience the joys of walking and chewing gum at the same time, so grab your field glasses, your camera, and your jacket, then spend lots of time outdoors this fall. You can see all of this and much more.
Don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab at the top of this page to help you find a place to see both fall foliage and migrating birds of prey in coming weeks. And click the “Trees, Shrubs, and Woody Vines” tab to find a photo guide that can help you identify the autumn leaves you encounter during your outings.
In the forests and woodlots of the lower Susquehanna watershed, the Black Gum, also known as the Black Tupelo, is always the first of our native trees to pop with autumn color. Its crimson leaves glow in an otherwise green landscape to help lure hungry birds including waxwings, American Robins, and Pileated Woodpeckers to its soon-to-be-ripe berries. The seeds contained in the fruit are thus well-distributed as a by-product of this avian feast. While not among the most massive of our trees, the Black Gum can live for over five hundred years, an age not exceeded by any other non-clonal indigenous flowering plant.
How ’bout a Two-for-the-Price-of-One Father’s Day Special? You got it!
Woodrow the Pileated Woodpecker (left) sure has collected a throat-load of tasty morsels, but he wouldn’t dare keep them all to himself;……he’s forced to share his meal with Little Woody, otherwise the begging will never stop.
As week-old snow and ice slowly disappears from the Lower Susquehanna River Watershed landscape, we ventured out to see what might be lurking in the dense clouds of fog that for more than two days now have accompanied a mid-winter warm spell.
After freezing to a slushy consistency earlier this week, the Susquehanna is already beginning to thaw. Below the York Haven Dam at Conewago Falls, the water is open and ice-free.On frozen man-made lakes and ponds, geese and ducks like these Mallards and American Wigeon are presently concentrated around small pockets of open water.During the past ten days, American Robin numbers have exploded throughout the lower Susquehanna valley. The majority of these birds may be a mix of both those coming south to escape the late onset of wintry conditions to our north and those inching north into our region as early spring migrants.The January thaw has melted the snow from lawns and fields to provide thousands of visiting robins with a chance to forage for earthworms.A visit by this young Cooper’s Hawk to the susquehannnawildlife.net headquarters garden sent songbirds scrambling……but did nothing to unnerve our resident Eastern Gray Squirrels,……which promptly went into tail-waving mode to advertise their presence.But earlier in the week, when heavy snow cover in the rural areas surrounding our urbanized neighborhood made it difficult for rodent-eating raptors to find food, we received brief visits from both a Red-tailed Hawk……and this young Red-shouldered Hawk, an uncommon bird of prey most often found in wet woods and other lowlands.To escape notice during visits by these larger raptors, our squirrels remained motionless and commenced performance of their best bump-on-a-log impressions.Unimpressed, each of our visiting buteos remained for just a few minutes before moving on in search of more favorable hunting grounds and prey.As snow melted and exposed bare ground in fields of early successional growth, we encountered……a flock of White-crowned Sparrows, most in first-winter plumage……and at least a dozen American Tree Sparrows. During the twentieth century, these handsome songbirds were regular winter visitors to the lower Susquehanna region. During recent decades, they’ve become increasingly more difficult to find. Currently, moderate numbers appear to be arriving to escape harsher weather to our north.What could be more appropriate on a foggy, gray evening than finding a “gray ghost” (adult male Northern Harrier) patrolling the fields in search of mice and voles.
If scenes of a January thaw begin to awaken your hopes and aspirations for all things spring, then you’ll appreciate this pair of closing photographs…
The maroon-red flower buds of Silver Maples are beginning to swell. And woodpeckers including Pileated Woodpeckers are beginning to drum, a timber-pounding behavior they use to establish breeding territories in habitats with suitable sites for cavity nesting.In wet soil surrounding spring seeps and streams, Skunk Cabbage is rising through the leaf litter to herald the coming of a new season. Spring must surely be just around the corner.
With the earth at perihelion (its closest approach to the sun) and with our home star just 27 degrees above the horizon at midday, bright low-angle light offered the perfect opportunity for doing some wildlife photography today. We visited a couple of grasslands managed by the Pennsylvania Game Commission to see what we could find…
On this State Game Lands parcel, prescribed fire is used to maintain a mix of grasslands and brushy early successional growth. In nearby areas, both controlled fire and mechanical cutting are used to remove invasive species from hedgerows and the understory of woodlots. Fire tolerant native species then have an opportunity to recolonize the forest and improve wildlife habitat. This management method also reduces the fuel load in areas with the potential for uncontrolled wildfires.The sun-dried fruits of a native Common Persimmon tree found growing in a hedgerow.Just one year ago, mechanical removal of invasive trees and shrubs (including Multiflora Rose) on this State Game Land was followed by a prescribed fire to create this savanna-like grassland.Hundreds of Song Sparrows were found in the grasses and thickets at both locations.White-throated Sparrows were also abundant, but prefer the tangles and shrubs of the thickets.Northern Mockingbirds were vigilantly guarding winter supplies of berries in the woodlots and hedgerows.In grasses and tangles on wetter ground, about a dozen Swamp Sparrows were discovered.The adult White-crowned Sparrow is always a welcome find.And seeing plenty of juvenile White-crowned Sparrows provides some assurance that there will be a steady stream of handsome adult birds arriving to spend the winter during the years to come.Dark-eyed Juncos were encountered only in the vicinity of trees and large shrubs.Several Savannah Sparrows were observed. Though they’re mostly found in treeless country, this particular one happened to pose atop a clump of shrubs located within, you guessed it, the new savanna-like grasslands.A tiny bird, even when compared to a sparrow, the Winter Wren often provides the observer with just a brief glimpse before darting away into the cover of a thicket.Within grasslands, scattered stands of live and dead timber can provide valuable habitat for many species of animals.Woodpeckers and other cavity-nesting birds rely upon an abundance of “snags” (standing dead trees) for breeding sites.This Red-bellied Woodpecker and about a dozen others were found in trees left standing in the project areas.A Yellow-bellied Sapsucker soaks up some sun.This very cooperative Pileated Woodpecker seemed to be preoccupied by insect activity on the sun-drenched bark of the trees. This denizen of mature forests will oft times wander into open country where larger lumber is left intact.
Just as things were really getting fun, some late afternoon clouds arrived to dim the already fading daylight. Just then, this Northern Harrier made a couple of low passes in search of mice and voles hidden in the grasses.It was a fitting end to a very short, but marvelously sunny, early winter day.
Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge. At Bombay Hook, they don’t waste your money mowing grass. Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools. Birds there can often be observed at close range. The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools. Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.Two more Short-billed Dowitchers on the way in.Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed. Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge. Mallards, American Avocets, and egrets are gathered on the shore.More Short-billed Dowitchers arriving to feed in Raymond Pool.Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool. A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.American Avocets probe the muddy bottom of Raymond Pool.Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early. Many migrating Dunlin linger at Bombay Hook into October and even November.This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp PoolA Greater Yellowlegs at Bear Swamp Pool.A Caspian Tern patrolling Raymond Pool.The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass. High tide licks at the roots of the cordgrass supporting the temporary domicile.By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice). It is our only dragonfly able to breed in saltwater. Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.Look up! A migrating Bobolink passes over the dike at Shearness Pool.Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool. Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river. The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century. An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers. Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.A Great Egret prowling Shearness Pool.A Snowy Egret in Bear Swamp Pool.Wood Ducks in Bear Swamp Pool.A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.
As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.
Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay. We found just this one during our visit to Bombay Hook. Man has certainly decimated populations of this ancient crustacean during recent decades.As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.A male Atlantic Marsh Fiddler Crab peers from its den.Herons and egrets including this Great Egret are quite fond of fiddler crabs. As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.An Osprey patrols the vast tidal areas opposite Shearness Pool.
No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook. This one was in shrubby growth along the dike at the north end of Shearness Pool.These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries. The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware. The seed balls are a favorite winter food of goldfinches and siskins.Finis Pool has no frontage on the tidal marsh but is still worth a visit. It lies along a spur road on the tour route and is located within a deciduous coastal plain forest. Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.The White-tailed Deity is common along the road to Finis Pool.Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well. This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.
We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.
Back in late May of 1983, four members of the Lancaster County Bird Club—Russ Markert, Harold Morrrin, Steve Santner, and your editor—embarked on an energetic trip to find, observe, and photograph birds in the Lower Rio Grande Valley of Texas. What follows is a daily account of that two-week-long expedition. Notes logged by Markert some four decades ago are quoted in italics. The images are scans of 35 mm color slide photographs taken along the way by your editor.
DAY TWO—May 22, 1983
Our goal today was to continue traveling and reach western Louisiana.
“We were on our way at 6:08. Stopped for a quick lunch in the camper and drove to Vinton, Louisiana, KOA. Lots of hard rain through Tennessee, Georgia, and Alabama.
Day Two: Sweetwater, Tennessee, to Vinton, Louisiana, a distance of 786 miles. (United States Geological Survey base image)
As we crossed Mississippi and entered Louisiana, we left the rain and the Appalachians behind. Upon crossing the Mississippi River, we had arrived in the West Gulf Coastal Plain, the physiographic province that extends all the way south along the Texas coast to Mexico and includes the Lower Rio Grande Valley. West of Baton Rouge, we began seeing waders in the picturesque Bald Cypress swamps—Great Egrets, Green Herons (Butorides virescens), Little Blue Herons (Egretta caerulea), and Glossy Ibis (Plegadis falcinellus) were identified. A Pileated Woodpecker was observed as it flew above the roadside treetops.
The rains we endured earlier in the trip had left there mark in much of Louisiana and Texas. Flooding in agricultural fields was widespread and the flat landscape often appeared inundated as far as the eye could see. Along the highway near Vinton, we spotted the first two of the many southern specialties we would find on the trip, a Loggerhead Shrike (Lanius ludovicianus) and a Scissor-tailed Flycatcher (Tyrannus forficatus), both perched on utility wires and searching for a meal.
Near Vinton, Louisiana, a Loggerhead Shrike was on the lookout for either a large insect or small bird upon which it could prey. (Vintage 35 mm image)
County Conservation District Tree Sales are underway throughout the Lower Susquehanna River Watershed. Now is the time to order for pickup in April. The prices are a bargain and the selection is fabulous. For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue). And don’t forget to order bundles of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local birds. They’re only $12.00 for a bundle of 10—can’t beat that deal!
Pickup on: Thursday, April 20, 2023 or Friday, April 21, 2023
Don’t mow it. Plant a meadow or pollinator garden instead.Both Cumberland and Perry Counties are offering a native warm-season grass and wildflower seed mix for planting your own meadow or pollinator garden. Perry County is also taking orders for a seed mix specifically formulated to grow plants for attracting hummingbirds and butterflies.
Pickup on: Thursday, April 20, 2023 or Friday, April 21, 2023
The Eastern Hemlock, Pennsylvania’s official state tree, is an excellent choice for addition to your landscape or reforestation project. It tolerates rocky soils and its cones are an prime source of food for birds ranging from chickadees to finches.
The handsome yet underused Northern Red Oak is a sturdy long-lived native tree that is ideal for street-side, lawn, and reforestation plantings. In spring, it can be a magnet for migrating Neotropical birds when its flowers attract a wide variety of tiny insects to its upper reaches. Unlike many other oaks, this species is a relatively fast grower.
In autumn, even after the bright red foliage is gone, the berries of mature Black Gum (Nyssa sylvatica) trees attract a wide variety of birds like this Pileated Woodpecker. The Lebanon County Conservation District is offering Black Gum, also known as Black Tupelo, during their 2023 tree sale. Why not order and plant a half dozen or more?
The Perry County Conservation District is not only offering plants during this year’s sale, you can also purchase bluebird nest boxes for just $12.00 each!For less than the cost of one year of mowing, this stream corridor in Conewago Township, Dauphin County was reforested by the owner with hundreds of native trees, the majority purchased through County Conservation District Tree Sale events spanning a period of several years. By replacing bare soil and mowed areas, the riparian buffer created by these plantings has significantly reduced the nutrient and sediment loads that were polluting the small stream therein known as Brill’s Run. With determination and not a lot of money, you can do it too.But don’t forget the Eastern White Pines!
A Pileated Woodpecker devours the dark blue berries of a Black Gum (Nyssa sylvatica). The Black Gum, also known as the Black Tupelo, Sour Gum, and Pepperidge, sports crimson leaves in the fall to attract birds to its fruit, thus assuring distribution of its seeds. In 2021, the fruit is so abundant that it has outlasted the foliage, much to the delight of resident and migrating birds.
Many are wont to say that they have no capacity for scientific pursuits, and having no capacity, they consequently have no love for them. I do not believe, that as a general thing, a love for science is necessarily innate in any man. It is the subject of cultivation and is therefore acquired. There are doubtless many, whose love for these and kindred pursuits is hereditary, through the mental biases and preoccupations of their progenitors, but in the masses of mankind it is quite otherwise. In this consists its redeeming qualities, for I do not think the truly scientific mind can either be an idle, a disorderly, or a very wicked one. There may be scientific men, who, forgetful of its teachings, are imperious and ambitious–who may have foregone their fealty to their country and their God, but as a general thing they are humble, social and law-abiding. If, therefore, there is a human being who desires to break off from old and evil associations, and form new and more virtuous ones, I would advise him to turn his attention to some scientific specialty, for the cultivation of a new affection, if there are no other and higher influences more accessible. In this pursuit he will, in time, be enabled to supplant the old and heartfelt affection. The occupation of his mind in the pursuit of scientific lore will wean him from vicious, trivial, and unmanly pursuits, and point out to him a way that is pleasant and instructive to walk in, which will ultimately lead to moral and intellectual usefulness. I wish I was accessible to them, and possessed the ability to impress this truth with sufficient emphasis upon the minds of the rising generation. This fact, that in all moral reformations, a love for the opposite of any besetting evil must be cultivated, before that evil can be surely eradicated, has been too much overlooked and too little valued in moral ethics. But true progress in this direction implies that, under all circumstances, men should “act in freedom according to reason.”
-Simon S. Rathvon
In the cellar of the North Museum on the campus of Franklin and Marshall College in Lancaster, Pennsylvania, is an assemblage of natural history specimens of great antiquity. The core of the collection has its origins in the endeavors of a group of mid-to-late nineteenth-century naturalists whose diligence provided a most thorough study of the plants and animals found within what was at the time America’s most productive farming county.
The members of the Linnaean Society of Lancaster City and County shared a passion for collecting, identifying, classifying, and documenting the flora and fauna of the region. Some members were formally educated and earned a living in the field of science, but the majority were in the process of self-education and balanced their natural history occupation with an unrelated means to provide financially for their families. The latter benefited greatly from their associations with the former, gaining expertise and knowledge while participating in the functions of the group.
On February 24, 1866, Simon S. Rathvon, the society’s Treasurer, read an essay in commemoration of the group’s fourth anniversary. Rathvon earned a living as a tailor, first in Marietta, a thriving river town at the time, then in Lancaster City. In 1840, Rathvon was elected into the Marietta Natural History Lyceum where, as a collections curator, he became associated with principals Judge John J. Libhart, an amateur ornithologist, and Samuel S. Haldeman, a geologist and soon to be widely-known malacologist. Haldeman, in 1842, upon noticing the new member’s interest in beetles and other insects, provided books, guidance, and inspiration, thus intensifying Rathvon’s study of entomology. Rathvon’s steadfast dedication eventually led to his numerous achievements in the field which included the publication of over 30 papers, many on the topic of agricultural entomology. Rathvon’s scientific understanding of insect identification and taxonomy was a foundation for his practical entomology, which moved beyond mere insect collection to focus upon the study of the life histories of insects, particularly the good and bad things they do. He then applied that knowledge to help growers solve pest problems, often stressing the value of beneficial species for maintaining a balance in nature. From 1869 through 1884, Rathvon edited and published Lancaster Farmer, a monthly (quarterly from 1874) agricultural journal in which he educated patrons with his articles on “economic entomology”. Rathvon continued earning a living in the tailor business, seemingly frustrated that his financially prudent advice on insect control in Lancaster Farmer failed to entice more would-be readers to part with the one dollar annual subscription fee. For many years, Rathvon crafted articles for local newspapers and wrote reports for the United States Department of Agriculture. In recognition of his achievements, Simon Rathvon received an honorary Ph.D. from Franklin and Marshall College in 1878.
In Rathvon’s anniversary essay, he details the origins of the Linnaean Society as a natural science committee within the “Lancaster Historical, Mechanical, and Horticultural Society” founded in 1853. The members of the committee, not finding sufficient support within the parent organization for their desired mission, “the cultivation and investigation of the natural history of Lancaster County…”, sought to form an independent natural history society. In February of 1862, the “Linnaean Society of Lancaster City and County” was founded to fulfill these ambitions.
Above all else, the written works by the members of the Linnaean Society and their predecessors have provided us with detailed accounts of the plants and animals found in Lancaster County, and in the lower Susquehanna River valley, using scientific binomial nomenclature, a genus and species name, as opposed to the variable folk and common names which, when used exclusively, often confuse or mislead readers. Consider the number of common names a species could have if just one was assigned by each of the languages of the world. Binomial nomenclature assigns one designation, a genus name and species name, in Latin, to each life-form (such as Homo sapiens for Humans), and it is adopted universally.
Rathvon would say of the naming of the Linnaean Society:
“…the name which the Society has adopted is in honorable commemoration of LINNAEUS, the great Swedish naturalist—one who may be justly regarded as a father in Natural Science. To him belongs the honor of having first promulgated the “binomial system of nomenclature,” a system that has done more to simplify the study of natural science than any light that has been brought to the subject by any man in any age.”
Carl Linnaeus lived from 1707 to 1778, and published his first edition of Systema Naturae in 1735.
The names of a number of the members and corresponding members on the Linnaean Society of Lancaster City and County’s rolls remain familiar. John P. McCaskey (educator) served as Corresponding Secretary. Doctor Abram P. Garber was a prominent Lancaster botanist and society member. Professor Samuel S. Haldeman (naturalist, geologist, and philologist), Professor J. L. LeConte (entomologist), Judge John J. Libhart, Professor Asa Gray (botanist), and the foremost legal egalitarian in the United States House of Representatives, the Honorable Thaddeus Stevens, were listed among the roster of corresponding members.
By the end of its fourth year, Rathvon enumerated the specimens in the collections of the society to exceed 32,000. These included all the species of mosses and plants known in the county, 200 bird specimens, an enormous insect collection with nearly 12,000 Coleoptera (Beetles), and more than 1,400 mollusk shells. The work of the society had already provided a thorough baseline of the flora and fauna of the lower Susquehanna River valley and Lancaster County.
Rathvon would continue as Treasurer and primary curator through the group’s first twenty-five years, their most active. By 1887, their library contained over 1,000 volumes, they possessed over 40,000 specimens, and more than 600 scientific papers had been read at their meetings.
Many of the society’s specimens were moved to the custody of Franklin and Marshall College following the group’s dissolution. In 1953, the collection found a home on the F&M campus at the newly constructed North Museum, named for benefactor Hugh M. North, where many of the specimens, particularly the birds, are on prominent display.
Among the mounted specimens in the North Museum collection is a Heath Hen, once a numerous coastal plain bird which was also of limited abundance in the Piedmont Province areas of southeast Pennsylvania prior to its rapid decline during the first half of the nineteenth century. In southern Lancaster County, the burned grasslands of the serpentine barrens in Fulton Township may have provided suitable Heath Hen habitat prior to the bird’s demise. Curiously, Judge John J. Libhart did not note the Heath Hen in his enumeration of the birds of Lancaster County in either 1844 or 1869, indicating it was seriously imperiled or may have already been extirpated.
The Heath Hen (Tympanuchus cupido cupido) became extinct in 1932. While the collection of this particular specimen had little significant impact on the population of this subspecies as a whole, prolonged hunting pressure was largely responsible for decimating the numbers of Heath Hens on the mainland of the Atlantic Coastal Plain. According to the museum tag, this specimen was “probably taken in southern Lancaster County prior to 1850”, and was part of the collection belonging to the Linnaean Society of Lancaster City and County. It is among hundreds of bird specimens on display in antique wood and glass cabinets in the North Museum.
The Heath Hen was extirpated from its entire Atlantic Coastal Plain mainland range by the mid-1860s. The last remaining population was restricted to Martha’s Vineyard where, for the first time, a conservation effort was initiated to try to save a species. After some promising rebounds, the Heath Hen’s recovery failed for a variety of reasons including: the population’s isolation on an island, severe winter storms, feral cat predation, and a flawed understanding of methods for conducting mosaic burns to maintain the bird’s scrub habitat and prevent large catastrophic fires. A large fire in 1906 reduced the island population to just 80 birds, then there was a strong rebound to an estimated 2,000 birds (800 counted) by April, 1916. One month later, a fire burned twenty percent of Martha’s Vineyard, striking while females were on the nest, and leaving mostly males as survivors. A downward spiral in numbers followed for another decade. Finally, from 1929 until his death in 1932, “Booming Ben”, the last Heath Hen, searched the island every spring for a mate that wasn’t there.
Based on life history and the morphology of specimens, the Heath Hen has long been considered to be a subspecies of the Greater Prairie Chicken (Tympanuchus cupido pinnatus), a bird of the tallgrass prairies. However, for more than a decade now, modern DNA analysis has kept taxonomists busy reclassifying and reworking the “tree of life”. For certain species, genetic discoveries often disqualify the long-trusted practice of determining a binomial name based on the visual appearance of specimens. Molecular study is making Linnaean classification more scientific, and is gradually untangling a web of names that man has been weaving for 200 years, often with scant evidence, in an effort to better understand the world around him. In the case of the Heath Hen, DNA research has thus far failed to conclusively determine its relationship to other species of prairie chickens. The lack of a sufficient pool of genetic material, particularly from mainland Heath Hens, reduces the ability of researchers to draw conclusions on this group of birds. There remains the possibility that the Heath Hen was genetically distinct from the Greater Prairie Chickens of the mid-western United States. This would be bad news for organizations studying the possibility of introducing the latter into the former’s historic range as a restoration program.
The Carolina Parakeet (Conuropsis carolinensis) specimen on display at the North Museum was collected by John C. Jenkins in Nanchez, Mississippi in 1835. The specimen was remounted by conservator H. Justin Roddy.
The last Carolina Parakeet (the only parrot species native to the eastern United States) died in captivity in the Cincinnati Zoo on February 21, 1918, one hundred years ago this past week. It was a species inhabiting primarily the lowland forests of the southeastern United States
In Lancaster County, Judge John J. Libhart wrote of the species in 1869, “…Carolina Parrot, Accidental; a flock seen near Manheim by Mr. G. W. Hensel.” Libhart did not mention the species in his earlier ornithological writings (1844). Therefore, the Hensel sighting probably occurred sometime between 1844 and 1869. The fate of a specimen reported to have been collected in the town of Willow Street sometime during the nineteenth century is unknown, the written details lack the date of its origin and other particulars that may clarify the authenticity of the sighting.
McKinley (1979) researched numerous historical sight records of Carolina Parakeets, but found no specimen from Lancaster County, or from Pennsylvania, New Jersey, Delaware, the District of Columbia, or Maryland to substantiate any of the reports in the Mid-Atlantic states. In the days prior to high-speed photography, verification and documentation of the presence of an animal species relied on what seems today to be a brutal and excessive method of nature study, killing. Lacking a specimen, the historical status of Carolina Parakeets in Pennsylvania, an area often considered to be within the bird’s former range, may be considered by many authorities to be hypothetical.
The Passenger Pigeon (Ectopistes migratorius) was abundant in the lower Susquehanna River valley through the early nineteenth century. Specimens in the North Museum collection include colorful males in breeding plumage. Several are from the original Linnaean Society of Lancaster City and County collection.
The Passenger Pigeon, too, has been extinct for more than a century. In Lancaster County, Judge John J. Libhart listed the Passenger Pigeon by the common name “Wild Pigeon” and wrote of the species in 1869, “Migratory; spring and autumn; feeds on grain, oak and beach, mostly on berries; stragglers sometimes remain and breed in the county.” There are numerous accounts of their precipitous decline both locally and throughout their former range, each illustrating the tragic loss of another portion of the North American natural legacy.
The North Museum specimen label describes the precipitous decline of the Passenger Pigeon in the lower Susquehanna River valley.
Martha, the last surviving Passenger Pigeon, died on September 1, 1914, in the Cincinnati Zoo. Ironically, the last Carolina Parakeet would die in the same enclosure just three-and-one-half years later. In the wild, the final three records of Passenger Pigeons were all of birds that were shot for taxidermy mounts in 1900, 1901, and 1902—an embarrassing human legacy.
By the early twentieth century, concerned citizens were beginning to realize the danger posed to many species of flora and fauna by man’s activities. In the eastern United States, the vast forests had been logged, the wetlands drained, and the streams and rivers dammed. Nearly all of the landscape had been altered in some way. Animals were harvested with little concern for the sustenance of their populations. Nearly unnoticed, the seemingly endless abundance and diversity of wildlife found in the early days of European colonization had dwindled critically.
In 1844, Judge John J. Libhart noted the “Log-Cock” among the birds found in Lancaster County. Fortunately, he included the scientific name “Picus pileatus”, the binomial nomenclature then recognized for the Pileated Woodpecker (specimens to right) among taxonomists. A record of “Log-Cock” could confuse researchers, leaving them to guess whether Libhart was referring to a woodpecker, a woodcock, a grouse, or any number of other birds including the long-extinct(?) Ivory-billed Woodpecker (Campephilus principalis). Of the Pileated Woodpecker (Dryocopus pileatus today), Libhart wrote in 1869, “…now become rare and is only met with in old and extensive woods; breeds in the county.” The Ivory-billed Woodpecker (specimen to left), a species of vast forests of large timber, living and dead, was restricted to the southeastern United States and Cuba. Logging following the American Civil War and, to a lesser degree, shooting impacted both species detrimentally. The Pileated Woodpecker recovered, the larger Ivory-billed Woodpecker, which has never been documented in the northeastern United States, has not. These specimens are in the North Museum collection.
The movement to conserve and protect threatened species from relentless persecution owes its start to the Linnaean taxonomists, the specimen collectors who gave uniformly recognizable names to nearly all of North America’s plants and animals. Significant too were John James Audubon and many others who used specimens as models to create accurate artwork which allowed scientists and citizens alike to learn to identify and name the living things they were seeing and, as time went by, not seeing.
Binomial nomenclature enabled the new conservationists to communicate accurately, reducing misunderstandings resulting from the use of many different names for one species or a shared name for multiple species. Discussions on the status of Columba migratorius (the binomial name for Passenger Pigeon in the nineteenth century) could occur without using the confusing local names for the Passenger Pigeon such as Wood Pigeon or, here in Pennsylvania, Wild Pigeon, a term which could describe any number of free-ranging pigeon or dove species. A binomial name, genus and species, makes the identity of a particular plant or animal, for lack of a more fitting term, specific.
Appreciation for the work completed by taxonomists who killed thousands of animals so each could be classified and assigned a name particular to its lineage is what finally motivated some to seek a cessation of the unchecked catastrophic killing of living things. It’s the paradox of late nineteenth-century conservation. The combined realization that a species is unique among other life-forms and that continuing to kill it for specimens, “style”, “sport”, or just an adrenaline thrill could eliminate it forever became an intolerable revelation. The blood would be on the hands of an audacious mankind, and it was unthinkable. Something had to be done. Unfortunately for the Passenger Pigeon, the Carolina Parakeet, and the Heath Hen, help came too late.
SOURCES
Greenburg, Joel. 2014. A Feathered River Across the Sky: The Passenger Pigeon’s Flight to Extinction. Bloomsbury Publishing. New York.
Libhart, John J. 1844. “Birds of Lancaster County”. I. Daniel Rupp’s History of Lancaster County. Gilbert Hills. Lancaster, PA.
Libhart, John J. 1869. “Ornithology”. J. I. Mombert’s An Authentic History of Lancaster County. J. E. Barr and Company. Lancaster, PA.
McKinley, Daniel. 1979. “History of the Carolina Parakeet in Pennsylvania, New Jersey, Delaware, Maryland, and the District of Columbia”. Maryland Birdlife. 35(1):1-10.
Palkovacs, Eric P.; Oppenheimer, Adam J.; Gladyshev, Eugene; Toepfer, John E.; Amato, George; Chase, Thomas; Caccone, Adalgesia. 2004. “Genetic Evaluation of a Proposed Introduction: The Case of the Greater Prairie Chicken and the Extinct Heath Hen”. Molecular Ecology. 13(7):1759-1769.
Rathvon, S. S. 1866. An Essay on the Origin of the Linnaean Society of Lancaster City and County, Its Objects and Progress. Pearsol and Geist. Lancaster, Pennsylvania.
Wheeler, Alfred G., Jr. and Miller, Gary L. 2006. “Simon Snyder Rathvon: Popularizer of Agricultural Entomology in Mid-19th Century America”. American Entomologist. 52(1):36-47.
Winpenny, Thomas R. 1990. “The Triumphs and Anguish of a Self-Made Man: 19th Century Naturalist S. S. Rathvon”. Pennsylvania History. 57(2):136-149.
