We’ve seen worse, but this winter has been particularly tough for birds and mammals in the Lower Susquehanna River Watershed. Due to the dry conditions of late summer and fall in 2024, the wild food crop of seeds, nuts, berries, and other fare has been less than average. The cold temperatures make insects hard to come by. Let’s have a look at how some of our local generalist and specialist species are faring this winter.
House Sparrows (bottom) and House Finches (top) are generalists. To survive and thrive, they are adapted to a variety of habitats and types of food. House Sparrows live almost anywhere man-made structures are found. They are true omnivores and will eat almost anything, especially if they see something else try to eat it first. The House Sparrow’s close association with humans has allowed it to become the most widespread and successful living avian dinosaur. On a cold night, they’ll take shelter either within dense vegetation alongside a building or within the structure itself. Though not nearly as cosmopolitan, the House Finch has successfully colonized much of the eastern United States after escaping from captivity as a cage bird in New York during the middle of the twentieth century. Upon being trans-located here from the arid southwest, they adapted to suburbs and farmlands consuming primarily a granivore diet of seeds supplemented with seasonally available berries. They quickly became accustomed to offerings at bird-feeding stations as well. To survive the harsh winters in the northern sections of their range, eastern populations of House Finches are developing a pattern of migration. These movements are most evident in late fall when dozens or sometimes hundreds can be seen heading south over regional hawk-counting stations.Though they require dead trees for nesting and as places to find the grubs and adult insects upon which they primarily feed, woodpeckers including the Northern Flicker are generalists, seldom passing by a supply of fruits like these Poison Ivy berries as a source of winter food. Flickers regularly visit suburban areas where they’ll drop by at bird-feeding stations for suet. During the warmer months, they are the woodpecker most frequently seen on the ground where swarms of ants garner their full attention.The Pileated Woodpecker is seldom found outside of mature forests where it digs relentlessly to remove grubs and other infestations from dead wood. But it is not a true specialist……it too finds a supply of Poison Ivy berries to be indispensable during a cold winter day.The Ruby-crowned Kinglet is a generalist, feeding mostly on insects, but also consuming small seeds and some berries, including those of Poison Ivy. It nests well to our north in tall spruces and other evergreens. During migration and in winter, the Ruby-crowned Kinglet may be found in deciduous trees, brush, and tall grass in habitats ranging from forests to parks and suburbia. This male is displaying its seldom-seen red crown.The Golden-crowned Kinglet, seen here on a Poison Ivy vine, is more of a specialist than the Ruby-crowned species, though the two will often occur in mixed groups during the winter. The Golden-crowned Kinglet nests in Spruce-Fir forests and in conifers within mixed woodlands. Even during migration, and particularly in winter, these birds are seldom found far from a stand of large evergreens within which they find shelter for the night.The Hermit Thrush’s generalist lifestyle allows it to survive cold season weather in the Lower Susquehanna River Watershed. During summer, it breeds in coniferous and mixed woods from the northern parts of our valley north into Canada and feeds primarily on worms, insects, and other arthropods. During migration and in winter, the Hermit Thrush becomes a regular visitor to deciduous forests, woody parks and suburbs, particularly where a supply of wild berries is available to supplement its diet.Iconic as it pulls earthworms from lawns during the warmer months, the generalist American Robin is fully dependent upon a crop of berries to survive winter conditions in the lower Susquehanna valley. The drought afflicted wild food crop of 2024 has led to fewer robins spending the season here and has delayed the northward push of migrating birds until the ground thaws and the earthworms make the ground rumble once again.It’s insects for the nestlings during spring and summer, then berries through the winter for the cheerful Eastern Bluebirds, another generalist species.American Crows are an excellent example of a generalist species. They’ll go anywhere to find food and they’ll eat almost anything. Like the House Sparrow and several other generalists, they adapt very well to human activity and actually thrive on it. Garbage anyone?Another career generalist is the widely worshiped White-tailed Deity, a species adapted to nearly all man-made landscapes with adequate vegetation upon which to browse. Pushed to the limit during severe weather, some individuals will consume carrion and even resort to cannibalism.You might think the Great Blue Heron is a specialist. Nope, it’s an accomplished generalist. Great blues will live, feed, and breed on almost any body of fresh or brackish water. And their diet includes almost anything that swims. In winter, you’ll even see them in fields hunting mice and voles.The Red-shouldered Hawk is a generalist with a diet ranging from amphibians and reptiles to small rodents and large insects. Mostly regarded as a species of bottomlands, they’ll frequent woodland edges, roadsides, and suburbia during the winter months.During its periodic winter visits to the region, the American Tree Sparrow feeds on seeds among the grasses and forbs of semi-open country with scattered short shrubs and trees. A generalist species, it will show up at backyard bird-feeding stations, particularly during periods of inclement weather. In summer, the American Tree Sparrow nests in tundra with growths of stunted willows and spruce and their diet includes insects as a source of protein for themselves and their young.The White-crowned Sparrow has similar winter habitat preferences to the tree sparrows……it becomes adaptable and something of a generalist when searching for food during bad winter storms.The Savannah Sparrow is an omnivore favoring insects in summer and seeds in winter. Though very closely tied to its grassland habitat year-round, snow cover can push these birds to enter woodier environs to consume fruits like these rose hips.Dusk and dawn during the short days of winter are the prime hunting times of a mammal specialist, the Short-eared Owl. Its presence in the lower Susquehanna valley is dependent on two dominant factors: extensive grassland habitat and an adequate population of the owl’s favored food, the Meadow Vole (Microtus pennsylvanicus). The Short-eared Owl’s requirements as a specialist species make finding a suitable place to live difficult. Unlike the generalist birds and mammals that often adapt to the widespread man-made disturbances in the region, populations of specialists frequently become fragmented, reduced in abundance, and subject to extirpation.The Meadow Vole is a generalist rodent that can be abundant in grasslands, early successional growth, fallow fields, marshlands, and, of course, meadows. They are primarily herbivores, but will occasionally consume insects and other arthropods. Usually nocturnal, some individuals venture out along their surface runways during daylight hours becoming vulnerable to diurnal raptors including kestrels, harriers, and buteos.A Short-eared Owl in near darkness patrolling a grassland for Meadow Voles.The eastward expansion of the Coyote (Canis latrans), a species of western North America’s grasslands and scrublands, and its progressive mixing with the Wolf (Canis lupus) in these eastern extensions of its range, has produced an expanding population of very adaptable generalists we call Eastern Coyotes (Canis latrans var.). These omnivorous canines colonized the Lower Susquehanna River Watershed during the last four decades to replace extirpated wolves as the top-tier natural predator in the region. Their primary diet includes Meadow Voles in grasslands and other small mammals along woodland edges and in successional habitats. Seasonally, they consume the berries of numerous wild plants. Slightly larger than their western ancestors, Eastern Coyotes with the admixture of Wolf genetics can subdue small ungulates. Like other apex predators, they are attracted to vulnerable prey and thus play a crucial role in culling the weak and diseased among their potential quarry species to assure the health and potential of populations of these species as long-term sources of food energy. The benefit to the prey species is however largely diluted in populous areas of the northeast; most venison consumed by Eastern Coyotes here is in the form of road kill. During periods of extended snow cover when small rodents and other foods become inaccessible, Eastern Coyotes, particularly young individuals, will wander into new areas seeking sustenance. Sometimes they venture into cities and suburbs where they explore the neighborhoods in search of garbage and pet foods placed outside the home. (Video clip courtesy of Tyler and Grace Good. Click image to view.)
Wildlife certainly has a tough time making it through the winter in the lower Susquehanna valley. Establishing and/or protecting habitat that includes plenty of year-round cover and sources of food and water can really give generalist species a better chance of survival. But remember, the goal isn’t to create unnatural concentrations of wildlife, it is instead to return the landscape surrounding us into more of a natural state. That’s why we try to use native plants as much as possible. And that’s why we try to attract not only a certain bird, mammal, or other creature, but we try to promote the development of a naturally functioning ecosystem with a food web, a diversity of pollinating plants, pollinating insects, and so on. Through this experience, we stand a better chance of understanding what it takes to graduate to the bigger job at hand—protecting, enhancing, and restoring habitats needed by specialist species. These are efforts worthy of the great resources that are sometimes needed to make them a success. It takes a mindset that goes beyond a focus upon the welfare of each individual animal to instead achieve the discipline to concentrate long-term on the projects and processes necessary to promote the health of the ecosystems within which specialist species live and breed. It sounds easier than it is—the majority of us frequently become distracted.
Being an individual from a population of a very successful generalist species is no guarantee of survival. This Eastern Gray Squirrel fell from a tall tree when the limbs became ice covered during a storm earlier this month. Just a freak accident? Maybe, but mistakes like this are often fatal in the natural world. This squirrel’s passing may seem brutal, but it provides a better opportunity for other squirrels and animals that share its food and cover requirements to make it through the winter. And those survivors that didn’t suffer such a fatal mistake or, more importantly, don’t possess a vulnerability that may have contributed to such a mishap will have a chance to pass those traits on to a new generation. This squirrel as an individual is gone, its species lives on, and may be stronger for its passing.Pennsylvania Game Commission crews maintain a grassland ecosystem for Short-eared Owls and other specialists using prescribed fire to prevent succession beyond its earliest stages. Among the additional specialist species benefiting from this management tool are Monarchs and other butterflies whose host plants survive early-season fire, but not competition with woody vines, shrubs, trees, and invasive herbaceous growth.
On the wider scale, it’s of great importance to identify and protect the existing and potential future habitats necessary for the survival of specialist species. And we’re not saying that solely for their benefit. These protection measures should probably include setting aside areas on higher ground that may become the beach intertidal zone or tidal marsh when the existing ones becomes inundated. And it may mean finally getting out of the wetlands, floodplains, and gullies to let them be the rain-absorbing, storm-buffering, water purifiers they spent millennia becoming. And it may mean it’s time to give up on building stick structures on tinderbox lands, especially hillsides and rocky outcrops with shallow, eroding soils that dry to dust every few years. We need to think ahead and stop living for the view. If you want to enjoy the view from these places, go visit and take plenty of pictures, or a video, that’s always nice, then live somewhere else. Each of these areas includes ecosystems that meet the narrow habitat requirements of many of our specialist species, and we’re building like fools in them. Then we feign victimhood and solicit pity when the calamity strikes: fires, floods, landslides, and washouts—again and again. Wouldn’t it be a whole lot smarter to build somewhere else? It may seem like a lot to do for some specialist animals, but it’s not. Because, you see, we should and can live somewhere else—they can’t.
The Allegheny Woodrat (Neotoma magister), a threatened species in Pennsylvania and a critically imperiled species in Maryland, is a habitat specialist requiring the forested rocky slopes, talus-flanked ridgetops, and caves of the Ridge and Valley Province for its nest sites and survival. Isolated populations survived within similar environs in the lower Susquehanna River valley’s Piedmont Province and on South Mountain through at least the first half of the twentieth century, but have since been extirpated. Human encroachment that fragments their habitat and promotes exposure to parasite-hosting mammals including the Raccoon (Procyon lotor), carrier of the Raccoon Roundworm (Baylisascarius procyonis), could prove fatal to remaining populations of this native mammal. (National Park Service image by Rick Olsen)The American Oystercatcher (Haematopus palliatus) is a specialist species that uses its highly adapted bill to feed on marine invertebrates including mollusks, few of which are actually oysters. Reliant upon tidal ecosystems for its survival, many of the seashore animals that make up this wader’s diet are themselves specialist species. Oystercatchers spend nearly their entire lives in tidal marshes or within the intertidal zone on beaches. They also frequent rocky jetties, particularly during high tide. This individual was photographed near the mouth of Chesapeake Bay in Northhampton County, Virginia, a location that, when the waters of the Atlantic started rising over 10,000 years ago, was the lower Susquehanna valley about 60 miles from the river’s mouth at present-day Norfolk Canyon along the edge of the continental shelf. Get the drift?
Back on March 24th, we took a detailed look at the process involved in administering prescribed fire as a tool for managing grassland and early successional habitat. Today we’re going turn back the hands of time to give you a glimpse of how the treated site fared during the five months since the controlled burn. Let’s go back to Middle Creek Wildlife Management Area for a photo tour to see how things have come along…
Pennsylvania Game Commission crews administering prescribed fire on a grassland demonstration site back on March 16.By late May, native herbaceous perennial wildflowers including Joe-pye Weed had re-greened the site. One of the goals of the burn was to kill fire-sensitive woody plants, thus preventing the process of succession from reforesting the site.The scorched, lifeless remains of small trees and shrubs indicate that that goal was met.Because prescribed fire is administered in a mosaic pattern that permits some early successional growth to remain until the next burn, birds including this thicket-nesting Yellow-breasted Chat are able to take advantage of the mixed habitat during their breeding season in May and June.By August, the site is a haven for native plants and animals.The burn has promoted the growth and late-summer bloom of fire-tolerant native wildflowers and warm-season grasses……including Indiangrass,……Big Bluestem……Thin-leaved Coneflower (Rudbeckia triloba), a plant also known as Brown-eyed Susan,……and Joe-pye Weed, a plant butterflies find irresistible.Eastern Tiger Swallowtail collecting nectar from Joe-pye Weed.A black-morph Eastern Tiger Swallowtail collecting nectar from Spotted Knapweed (Centaurea steobe micranthos), a non-native invasive plant found growing in an area of the burn site missed by this year’s fire. While many non-native plants are unable to survive the flames and heat produced by prescribed fire, it isn’t an absolute cure-all. It doesn’t eliminate all invasive plants, it just keeps them from dominating a landscape by out-competing native species. Left unmanaged, Spotted Knapweed is a tough perennial invasive that can easily become one the species able to overtake a vulnerable grassland. It can be a stubborn survivor of some prescribed burns. On the plus side, butterflies really like it.By August, native grassland plants in the prescribed fire area were already providing an abundance of seeds for birds including this American Goldfinch.For larger birds like turkeys and pheasants, an abundance of Carolina Grasshoppers are providing a protein-dense food source in managed grasslands.And tiny flying insects, a nuisance to us as we take a stroll alongside the grasslands, are a meal taken on the wing by dragonflies including this Black Saddlebags.
Elsewhere around the refuge at Middle Creek, prescribed fire and other management techniques are providing high-quality grassland habitat for numerous species of nesting birds…
Bobolinks nested both in areas subjected to controlled burns……and in hay fields where mowing was delayed until the nesting season, including the fledging process, was completed earlier this month.As advertised, Grasshopper Sparrows nested in these fields as well.
We hope you enjoyed this short photo tour of grassland management practices. Now, we’d like to leave you with one last set of pictures—a set you may find as interesting as we found them. Each is of a different Eastern Cottontail, a species we found to be particularly common on prescribed fire sites when we took these images in late May. The first two are of the individuals we happened to be able to photograph in areas subjected to fire two months earlier in March. The latter two are of cottontails we happened to photograph elsewhere on the refuge in areas not in proximity to ground treated with a prescribed burn or exposed to accidental fire in recent years.
Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.
These first two rabbits are living the good life in a warm-season grass wonderland.
Eastern Cottontail at a site not subjected to any recent fire activity.Eastern Cottontail at a site not subjected to any recent fire activity.
Oh Deer! Oh Deer! These last two rabbits have no clock to track the time; they have only ticks. Better not go for a stroll with them Alice—that’s no wonderland! I know, I know, it’s time to go. See ya later.
Homo sapiens owes much of its success as a species to an acquired knowledge of how to make, control, and utilize fire. Using fire to convert the energy stored in combustible materials into light and heat has enabled humankind to expand its range throughout the globe. Indeed, humans in their furless incomplete mammalian state may have never been able to expand their populations outside of tropical latitudes without mastery of fire. It is fire that has enabled man to exploit more of the earth’s resources than any other species. From cooking otherwise unpalatable foods to powering the modern industrial society, fire has set man apart from the rest of the natural world.
In our modern civilizations, we generally look at the unplanned outbreak of fire as a catastrophe requiring our immediate intercession. A building fire, for example, is extinguished as quickly as possible to save lives and property. And fires detected in fields, brush, and woodlands are promptly controlled to prevent their exponential growth. But has fire gone to our heads? Do we have an anthropocentric view of fire? Aren’t there naturally occurring fires that are essential to the health of some of the world’s ecosystems? And to our own safety? Indeed there are. And many species and the ecosystems they inhabit rely on the periodic occurrence of fire to maintain their health and vigor.
For the war effort- The campaign to reduce the frequency of forest fires got its start during World War II with distribution of this poster in 1942. The goal was to protect the nation’s timber resources from accidental or malicious loss due to fire caused by man-made ignition sources. The release of the Walt Disney film “Bambi” during the same year and the adoption of the Smokey the Bear mascot in 1944 softened the message’s delivery, but the public relations outreach continued to be a key element of a no-fire policy to save trees for lumber. Protection and management of healthy forest ecosystems in their entirety has only recently become a priority. (National Archives image)
Man has been availed of the direct benefits of fire for possibly 40,000 years or more. Here in the Lower Susquehanna River Watershed, the earliest humans arrived as early as 12,000 years ago—already possessing skills for using fire. Native plants and animals on the other hand, have been part of the ever-changing mix of ecosystems found here for a much longer period of time—millions to tens of millions of years. Many terrestrial native species are adapted to the periodic occurrence of fire. Some, in fact, require it. Most upland ecosystems need an occasional dose of fire, usually ignited by lightning (though volcanism and incoming cosmic projectiles are rare possibilities), to regenerate vegetation, release nutrients, and maintain certain non-climax habitat types.
But much of our region has been deprived of natural-type fires since the time of the clearcutting of the virgin forests during the eighteenth and nineteenth centuries. This absence of a natural fire cycle has contributed to degradation and/or elimination of many forest and non-forest habitats. Without fire, a dangerous stockpile of combustible debris has been collecting, season after season, in some areas for a hundred years or more. Lacking periodic fires or sufficient moisture to sustain prompt decomposition of dead material, wildlands can accumulate enough leaf litter, thatch, dry brush, tinder, and fallen wood to fuel monumentally large forest fires—fires similar to those recently engulfing some areas of the American west. So elimination of natural fire isn’t just a problem for native plants and animals, its a potential problem for humans as well.
Indiangrass (seen here), Switchgrass, Big Bluestem, and Little Bluestem are native species requiring periodic forms of disturbance to eliminate competition by woody plants. These warm-season grasses develop roots that penetrate deep into the soil, sometimes to depths of six feet or more, allowing them to survive severe drought and flash fire events. In the tall grass prairies, these extensive root systems allow these grasses to return following heavy grazing by roaming herds of American Bison (Bison bison). Without these habitat disturbances, warm season grasslands succumb to succession in about seven years. With their periodic occurrence, the plants thrive and provide excellent wildlife habitat, erosion control, and grazing forage.
To address the habitat ailments caused by a lack of natural fires, federal, state, and local conservation agencies are adopting the practice of “prescribed fire” as a treatment to restore ecosystem health. A prescribed fire is a controlled burn specifically planned to correct one or more vegetative management problems on a given parcel of land. In the Lower Susquehanna River Watershed, prescribed fire is used to…
Eliminate dangerous accumulations of combustible fuels in woodlands.
Reduce accumulations of dead plant material that may harbor disease.
Provide top kill to promote oak regeneration.
Regenerate other targeted species of trees, wildflowers, grasses, and vegetation.
Kill non-native plants and promote growth of native plants.
Prevent succession.
Remove woody growth and thatch from grasslands.
Promote fire tolerant species of plants and animals.
Improve habitat for rare species (Regal Fritillary, etc.)
Recycle nutrients and minerals contained in dead plant material.
Let’s look at some examples of prescribed fire being implemented right here in our own neighborhood…
Prescribed fires are typically planned for the dormant season extending from late fall into early spring with burns best conducted on days when the relative humidity is low.Prescribed fire is used regularly at Fort Indiantown Gap Military Reservation in Lebanon County, Pennsylvania, to keep accumulations of woody and herbaceous fuels from accumulating on and around the training range areas where live ordinance and other sources of ignition could otherwise spark large, hard-to-control wildfires.Prescribed fires replace the periodic natural burns that would normally reduce the fuel load in forested areas. Where these fuels are allowed to accumulate, south-facing slopes are particularly susceptible to extreme fires due to their exposure to the drying effects of intense sunlight for much of the year. The majority of small oaks subjected to treatment by the prescribed fire shown here will have the chance to regenerate without immediate competition from other species including invasive plants. The larger trees are mostly unaffected by the quick exposure to the flames. Note too that these fires don’t completely burn everything on the forest floor, they burn that which is most combustible. There are still plenty of fallen logs for salamanders, skinks, and other animals to live beneath and within.
A prescribed fire in late winter prevents this grassland consisting of Big Bluestem and native wildflowers from being overtaken by woody growth and invasive species. Fires such as this that are intended to interrupt the process of succession are repeated at least every three to five years.In its wildlife food plots, prescribed fire is used by the Pennsylvania Game Commission to prevent succession and control invasive species such as Multiflora Rose, instead promoting the growth of native plants.An example of a woodlot understory choked with combustible fuels and dense tangles of invasive Multiflora Rose. A forester has the option of prescribing a dose of dormant-season fire for a site like this to reduce the fuel load, top kill non-native vegetation, and regenerate native plants.A dose of prescribed fire was administered on this grassland to kill the woody growth of small trees beginning to overtake the habitat by succession.The Pennsylvania Game Commission employs prescribed fire at Middle Creek Wildlife Management Area and on many of their other holdings to maintain grasslands.Prescribed fire is used to eliminate invasive species including Multiflora Rose from grasslands at Middle Creek W.M.A. Annual burns on the property are conducted in a mosaic pattern so that each individual area of the grassland is exposed to the effects of fire only once every two to five years. Without fire or some type of mechanical or chemical intervention, succession by woody trees and shrubs would take hold after about seven years.Prescribed fire is planned for a fraction of total grassland acreage at Middle Creek W.M.A. each year. Another section of the mosaic is targeted in the following year and yet another in the year that follows that. Because burns are conducted in the spring, grassland cover is available for wildlife throughout the winter. And because each year’s fire burns only a portion of the total grassland acreage, wildlife still has plenty of standing grass in which to take shelter during and after the prescribed fire.Prescribed fire at Middle Creek W.M.A. provides grassland habitat for dozens of species of birds and mammals including the not-so-common Grasshopper Sparrow……and stocked Ring-necked Pheasants that do nest and raise young there.On a few sites in the Lower Susquehanna River Watershed , prescribed fire is being used to establish and maintain savanna-like grasslands. This one, located on a dry, south-facing slope near numerous man-made sources of ignition, can easily be dosed with periodic prescribed burns to both prevent succession and reduce fuel accumulations that may lead to a devastating extreme fire.One year following a prescribed burn, this is the autumn appearance of a savanna-like habitat with fire-tolerant Pitch Pine (Pinus rigida), Bear Oak, warm-season grasses, and a variety of nectar-producing wildflowers for pollinators. These ecosystems are magnets for wildlife and may prove to be a manageable fit on sun-drenched sites adjacent to man-made land disturbances and their sources of ignition.Savanna-like grasslands with oaks and other scattered large trees, some of them dead, make attractive nesting habitat for the uncommon Red-headed Woodpecker.Prescribed fire can benefit hungry Wild Turkeys by maintaining savanna-like grasslands for an abundance of grasshoppers and other insects in summer and improving the success of mast-producing oaks for winter.In the Lower Susquehanna River Watershed, the caterpillar of the rare Eastern Buck Moth feeds on the foliage of the Bear Oak, also known as the Scrub Oak, a shrubby species that relies upon periodic fire to eliminate competition from larger trees in its early successional habitat.Leaves of the Bear Oak in fall. The Bear Oak regenerates readily from top kill caused by fire.Reed Canary Grass (Phalaris arundinacea) is a native cool-season grass with a colorful inflorescence in spring. But given the right situation, it can aggressively overtake other species to create a pure stand lacking biodiversity. It is one of the few native species which is sometimes labelled “invasive”.Prescribed fire can be used to reduce an overabundance of Reed Canary Grass and its thatch in wetlands. Periodic burning can help restore species diversity in these habitats for plants and animals including rare species such as the endangered Bog Turtle (Glyptemys muhlenbergii).On the range areas at Fort Indiantown Gap in Lebanon County, Pennsylvania, disturbances by armored vehicles mimic the effects of large mammals such as the American Bison which periodically trampled grasses to prevent succession and the establishment of woody plants on its prairie habitat. To supplement the activity of the heavy vehicles and to provide suitable habitat for the very rare Regal Fritillary (Speyeria idalia) butterflies found there, prescribed fire is periodically employed to maintain the grasslands on the range. These burns are planned to encourage the growth of “Fort Indiantown Gap Little Bluestem” grass as well as the violets used as host plants by the Regal Fritillary caterpillars. These fires also promote growth of a variety of native summer-blooming wildflowers to provide nectar for the adults butterflies.A last record of a wild American Bison killed in Pennsylvania was an animal taken in the Susquehanna watershed in Union County in 1801. The species is thereafter considered extirpated from the state. Since that time, natural disturbances needed to regenerate warm-season grasses have been limited primarily to fires and riverine ice scour. The waning occurrence of both has reduced the range of these grasses and their prairie-like ecosystems in the commonwealth. (Exhibit: State Museum of Pennsylvania, Harrisburg)A male Regal Fritillary on the range at Fort Indiantown Gap, where armored vehicles and prescribed fire provide suitable prairie-like habitat for this vulnerable species.Prescribed fires return the nutrients and minerals contained in dead plant material to the soil. Following these controlled burns, insects like this Honey Bee can often be seen collecting minerals from the ashes.A Greenbottle Fly gathering minerals from the ash following a prescribed burn.
In Pennsylvania, state law provides landowners and crews conducting prescribed fire burns with reduced legal liability when the latter meet certain educational, planning, and operational requirements. This law may help encourage more widespread application of prescribed fire in the state’s forests and other ecosystems where essential periodic fire has been absent for so very long. Currently in the Lower Susquehanna River Watershed, prescribed fire is most frequently being employed by state agencies on state lands—in particular, the Department of Conservation and Natural Resources on State Forests and the Pennsylvania Game Commission on State Game Lands. Prescribed fire is also part of the vegetation management plan at Fort Indiantown Gap Military Reservation and on the land holdings of the Hershey Trust. Visitors to the nearby Gettysburg National Military Park will also notice prescribed fire being used to maintain the grassland restorations there.
For crews administering prescribed fire burns, late March and early April are a busy time. The relative humidity is often at its lowest level of the year, so the probability of ignition of previous years’ growth is generally at its best. We visited with a crew administering a prescribed fire at Middle Creek Wildlife Management Area last week. Have a look…
Members of a Pennsylvania Game Commission burn crew provide visitors to Middle Creek Wildlife Management Area with an overview of prescribed fire and the equipment and techniques they use to conduct a burn.Pennsylvania Game Commission Southeast Region Forester Andy Weaver will fulfill the role of Burn Boss for administering this day’s dose of fire. His responsibilities include assessing the weather before the burn and calculating a probability of ignition.The Burn Boss briefs personnel with information on site layout, water supply location(s), places of refuge, emergency procedures, the event’s goals and plan of action, crew assignments, and the results of the weather check: wind from the northwest at 5 miles per hour, temperature 48 degrees, and the relative humidity 63%. Today’s patient is a parcel of warm-season grasses receiving a dose of fire to eliminate invasive non-native plants, woody growth, and thatch. The probability of ignition is 20%, but improving by the minute.To begin the burn, a test fire is started in the downwind corner of the parcel, which also happens to be the bottom of the slope. Fuel ignition is good. The burn can proceed.Crews proceed uphill from the location of the test fire while igniting combustibles along both flanks of the area being treated.A drip torch is used to ignite the dried stems and leaves of warm-season grasses and wildflowers. Each member of the burn crew wears Nomex fire-resistant clothing and carries safety equipment including a two-way radio, a hydration pack, and a cocoon-like emergency fire shelter.An all-terrain vehicle equipped with various tools, a fire pump, hose, and a small water tank accompanies the crew on each flank of the fire.A mowed strip of cool-season grasses along the perimeter of the burn area is already green and functions as an ideal fire break. While the drip torch is perfect for lighting combustibles along the fire’s perimeter, the paintball gun-looking device is an effective tool used to lob incendiaries into the center areas of the burn zone for ignition.With green cool-season grasses already growing on the trails surrounding the burn zone, very little water was used to contain this prescribed fire. Where such convenient fire breaks don’t already exist, crews carry tools including chain saws, shovels, and leaf blowers to create their own. They also carry flame swatters, backpack water pumps, shovels, and other tools to extinguish fires if necessary. None of these items were needed to control this particular fire.This fast-burning fire provides enough heat to damage the cambium layer of the woody tree and shrub saplings in this parcel being maintained as a grassland/wildflower plot, thus the process of succession is forestalled. Burns conducted during previous years on this and adjacent fields have also controlled aggressive growth of invasive Multiflora Rose and Olives (Elaeagnus species).Crews proceed up the slope while maintaining the perimeter by igniting dry plant material along the flanks of the burn zone.Ignition complete, the crews monitor the fire.The Burn Boss surveys the final stages of a safe and successful prescribed fire. The fire has left behind a mosaic of burned and unburned areas, just as a naturally occurring event may have done. Wildlife dodging the flames may be taking refuge in the standing grasses, so there is no remedial attempt to go back and ignite these areas. They’ll be burned during prescribed fires in coming years.By June, this grassland will again be lush and green with warm-season grasses and blooming wildflowers like this Common Milkweed being visited by a Great Spangled Fritillary.And later in the summer, Eastern Tiger Swallowtails on Joe-pye Weed.Indiangrass in flower in mid-summer.Bobolinks glow in the late August sun while taking flight from a stand of warm-season grasses maintained using springtime prescribed fire. The small dots on the dark background at the top of the image are multitudes of flying insects, many of them pollinators. The vegetation is predominately Indiangrass, excellent winter cover for birds, mammals, and other wildlife.
Prescribed burns aren’t a cure-all for what ails a troubled forest or other ecosystem, but they can be an effective remedy for deficiencies caused by a lack of periodic episodes of naturally occurring fire. They are an important option for modern foresters, wildlife managers, and other conservationists.
As week-old snow and ice slowly disappears from the Lower Susquehanna River Watershed landscape, we ventured out to see what might be lurking in the dense clouds of fog that for more than two days now have accompanied a mid-winter warm spell.
After freezing to a slushy consistency earlier this week, the Susquehanna is already beginning to thaw. Below the York Haven Dam at Conewago Falls, the water is open and ice-free.On frozen man-made lakes and ponds, geese and ducks like these Mallards and American Wigeon are presently concentrated around small pockets of open water.During the past ten days, American Robin numbers have exploded throughout the lower Susquehanna valley. The majority of these birds may be a mix of both those coming south to escape the late onset of wintry conditions to our north and those inching north into our region as early spring migrants.The January thaw has melted the snow from lawns and fields to provide thousands of visiting robins with a chance to forage for earthworms.A visit by this young Cooper’s Hawk to the susquehannnawildlife.net headquarters garden sent songbirds scrambling……but did nothing to unnerve our resident Eastern Gray Squirrels,……which promptly went into tail-waving mode to advertise their presence.But earlier in the week, when heavy snow cover in the rural areas surrounding our urbanized neighborhood made it difficult for rodent-eating raptors to find food, we received brief visits from both a Red-tailed Hawk……and this young Red-shouldered Hawk, an uncommon bird of prey most often found in wet woods and other lowlands.To escape notice during visits by these larger raptors, our squirrels remained motionless and commenced performance of their best bump-on-a-log impressions.Unimpressed, each of our visiting buteos remained for just a few minutes before moving on in search of more favorable hunting grounds and prey.As snow melted and exposed bare ground in fields of early successional growth, we encountered……a flock of White-crowned Sparrows, most in first-winter plumage……and at least a dozen American Tree Sparrows. During the twentieth century, these handsome songbirds were regular winter visitors to the lower Susquehanna region. During recent decades, they’ve become increasingly more difficult to find. Currently, moderate numbers appear to be arriving to escape harsher weather to our north.What could be more appropriate on a foggy, gray evening than finding a “gray ghost” (adult male Northern Harrier) patrolling the fields in search of mice and voles.
If scenes of a January thaw begin to awaken your hopes and aspirations for all things spring, then you’ll appreciate this pair of closing photographs…
The maroon-red flower buds of Silver Maples are beginning to swell. And woodpeckers including Pileated Woodpeckers are beginning to drum, a timber-pounding behavior they use to establish breeding territories in habitats with suitable sites for cavity nesting.In wet soil surrounding spring seeps and streams, Skunk Cabbage is rising through the leaf litter to herald the coming of a new season. Spring must surely be just around the corner.
With the earth at perihelion (its closest approach to the sun) and with our home star just 27 degrees above the horizon at midday, bright low-angle light offered the perfect opportunity for doing some wildlife photography today. We visited a couple of grasslands managed by the Pennsylvania Game Commission to see what we could find…
On this State Game Lands parcel, prescribed fire is used to maintain a mix of grasslands and brushy early successional growth. In nearby areas, both controlled fire and mechanical cutting are used to remove invasive species from hedgerows and the understory of woodlots. Fire tolerant native species then have an opportunity to recolonize the forest and improve wildlife habitat. This management method also reduces the fuel load in areas with the potential for uncontrolled wildfires.The sun-dried fruits of a native Common Persimmon tree found growing in a hedgerow.Just one year ago, mechanical removal of invasive trees and shrubs (including Multiflora Rose) on this State Game Land was followed by a prescribed fire to create this savanna-like grassland.Hundreds of Song Sparrows were found in the grasses and thickets at both locations.White-throated Sparrows were also abundant, but prefer the tangles and shrubs of the thickets.Northern Mockingbirds were vigilantly guarding winter supplies of berries in the woodlots and hedgerows.In grasses and tangles on wetter ground, about a dozen Swamp Sparrows were discovered.The adult White-crowned Sparrow is always a welcome find.And seeing plenty of juvenile White-crowned Sparrows provides some assurance that there will be a steady stream of handsome adult birds arriving to spend the winter during the years to come.Dark-eyed Juncos were encountered only in the vicinity of trees and large shrubs.Several Savannah Sparrows were observed. Though they’re mostly found in treeless country, this particular one happened to pose atop a clump of shrubs located within, you guessed it, the new savanna-like grasslands.A tiny bird, even when compared to a sparrow, the Winter Wren often provides the observer with just a brief glimpse before darting away into the cover of a thicket.Within grasslands, scattered stands of live and dead timber can provide valuable habitat for many species of animals.Woodpeckers and other cavity-nesting birds rely upon an abundance of “snags” (standing dead trees) for breeding sites.This Red-bellied Woodpecker and about a dozen others were found in trees left standing in the project areas.A Yellow-bellied Sapsucker soaks up some sun.This very cooperative Pileated Woodpecker seemed to be preoccupied by insect activity on the sun-drenched bark of the trees. This denizen of mature forests will oft times wander into open country where larger lumber is left intact.
Just as things were really getting fun, some late afternoon clouds arrived to dim the already fading daylight. Just then, this Northern Harrier made a couple of low passes in search of mice and voles hidden in the grasses.It was a fitting end to a very short, but marvelously sunny, early winter day.
As the annual autumn songbird migration begins to reach its end, native sparrows can be found concentrating in fallow fields, early successional thickets, and brushy margins along forest edges throughout the Lower Susquehanna River Watershed.
A streamside thicket composed of seed-producing grasses and wildflowers as well as fruit-bearing shrubs and vines can be ideal habitat for migrating and wintering native sparrows.
Visit native sparrow habitat during mid-to-late November and you have a good chance of seeing these species and more…
The Song Sparrow can be found in woody brush and grassy margins from the shores of the Susquehanna all the way up to the ridgetops of the Appalachians.During the colder months of the year, the Dark-eyed Junco is a familiar visitor to bird-feeding stations. Where suitable natural cover is present, they regularly venture into suburban and urban settings.The White-throated Sparrow is commonly found in the company of juncos, but is generally less adventurous, being more likely in weedy fields near young woodlands than in suburban gardens.The Eastern Towhee is a large native sparrow most often found in early successional growth near woodlands. Look for them in utility right-of-ways.The elusive Fox Sparrow is a regular late-fall migrant. Few stay for the winter, but northbound birds can be seen as early as mid-February each year.
If you’re lucky enough to live where non-native House Sparrows won’t overrun your bird feeders, you can offer white millet as a supplement to the wild foods these beautiful sparrows might find in your garden sanctuary. Give it a try!
Grasshoppers are perhaps best known for the occasions throughout history when an enormous congregation of these insects—a “plague of locusts”—would assemble and rove a region to feed. These swarms, which sometimes covered tens of thousands of square miles or more, often decimated crops, darkened the sky, and, on occasion, resulted in catastrophic famine among human settlements in various parts of the world.
The largest “plague of locusts” in the United States occurred during the mid-1870s in the Great Plains. The Rocky Mountain Locust (Melanoplus spretus), a grasshopper of prairies in the American west, had a range that extended east into New England, possibly settling there on lands cleared for farming. Rocky Mountain Locusts, aside from their native habitat on grasslands, apparently thrived on fields planted with warm-season crops. Like most grasshoppers, they fed and developed most vigorously during periods of dry, hot weather. With plenty of vegetative matter to consume during periods of scorching temperatures, the stage was set for populations of these insects to explode in agricultural areas, then take wing in search of more forage. Plagues struck parts of northern New England as early as the mid-1700s and were numerous in various states in the Great Plains through the middle of the 1800s. The big ones hit between 1873 and 1877 when swarms numbering as many as trillions of grasshoppers did $200 million in crop damage and caused a famine so severe that many farmers abandoned the westward migration. To prevent recurrent outbreaks of locust plagues and famine, experts suggested planting more cool-season grains like winter wheat, a crop which could mature and be harvested before the grasshoppers had a chance to cause any significant damage. In the years that followed, and as prairies gave way to the expansive agricultural lands that presently cover most of the Rocky Mountain Locust’s former range, the grasshopper began to disappear. By the early years of the twentieth century, the species was extinct. No one was quite certain why, and the precise cause is still a topic of debate to this day. Conversion of nearly all of its native habitat to cropland and grazing acreage seems to be the most likely culprit.
The critically endangered Eskimo Curlew (Numenius borealis), a species not photographed since 1962 and not confirmed since 1963, fed on Rocky Mountain Locusts during its spring migration through the Great Plains. Excessive hunting and conversion of grasslands to agriculture are believed responsible for the bird’s demise. (United States Fish and Wildlife Service image by Christina Nelson)
In the Mid-Atlantic States, the mosaic of the landscape—farmland interspersed with a mix of forest and disturbed urban/suburban lots—prevents grasshoppers from reaching the densities from which swarms arise. In the years since the implementation of “Green Revolution” farming practices, numbers of grasshoppers in our region have declined. Systemic insecticides including neonicotinoids keep grasshoppers and other insects from munching on warm-season crops like corn and soybeans. And herbicides including 2,4-D (2,4-Dichlorophenoxyacetic acid) have, in effect, become the equivalent of insecticides, eliminating broadleaf food plants from the pasturelands and hayfields where grasshoppers once fed and reproduced in abundance. As a result, few of the approximately three dozen species of grasshoppers with ranges that include the Lower Susquehanna River Watershed are common here. Those that still thrive are largely adapted to roadsides, waste ground, and small clearings where native and some non-native plants make up their diet.
Here’s a look at four species of grasshoppers you’re likely to find in disturbed habitats throughout our region. Each remains common in relatively pesticide-free spaces with stands of dense grasses and broadleaf plants nearby.
CAROLINA GRASSHOPPER
Dissosteira carolina
The Carolina Grasshopper, also known as the Carolina Locust or Quaker, is one of the band-winged grasshoppers. It is commonly found along roadsides and on other bare ground near stands of tall grass and broadleaf plants.The Carolina Grasshopper is variable in color, ranging from very dark brown……to a rich tan or khaki shade. These earth-tone colors provide the insect with effective camouflage while spending time on the ground.The Carolina Grasshopper is most readily detected and identified when it flies. The colors of the wings resemble those of the Mourning Cloak butterfly.Carolina Grasshoppers are among the preferred victims of Great Black Wasps (Sphex pensylvanicus). A female wasp stings the grasshopper to paralyze it, then drags it away to one of numerous cells in an underground burrow where she lays an egg on it. The body of the disabled grasshopper then provides nourishment for the larval wasp.
DIFFERENTIAL GRASSHOPPER
Melanoplus differentialis
Differential Grasshopper nymph with small “fairy wings”.An adult female Differential Grasshopper with fully developed wings.An adult female Differential Grasshopper
TWO-STRIPED GRASSHOPPER
Melanoplus bivittatus
An early-stage Two-striped Grasshopper nymph.A Two-striped Grasshopper nymph in a later stage.An adult female Two-striped Grasshopper.An adult female Two-striped Grasshopper. Note the pale stripe originating at each eye and joining near the posterior end of the wings to form a V-shaped pattern.An adult female Two-striped Grasshopper.
RED-LEGGED GRASSHOPPER
Melanoplus femurrubrum
An adult male Red-legged Grasshopper hiding in dense urban vegetation.The Red-legged Grasshopper may currently be our most abundant and widespread species.An adult male Red-legged Grasshopper.An adult female Red-legged Grasshopper.
Protein-rich grasshoppers are an important late-summer, early-fall food source for birds. The absence of these insects has forced many species of breeding birds to abandon farmland or, in some cases, disappear altogether.
Beginning in the early 1930s, the Western Cattle Egret (Bubulcus ibis), a notoriously nomadic species, transited the Atlantic from Africa to colonize the Americas…and they did it without any direct assistance from humans. During the 1970s and early 1980s, a nesting population of Western Cattle Egrets on river islands adjacent to the Susquehanna’s Conejohela Flats off Washington Boro was the largest inland rookery in the northeastern United States. The Lancaster County Bird Club censused the birds each August and found peak numbers in 1981 (7,580). During their years of abundance, V-shaped flocks of cattle egrets from the rookery islands ventured into grazing lands throughout portions of Lancaster, York, Dauphin, and Lebanon Counties to hunt grasshoppers. These daily flights were a familiar summertime sight for nearly two decades. Then, in the early 1980s, reductions in pastureland acreage and plummeting grasshopper numbers quickly took their toll. By 1988, the rookery was abandoned. The cattle egrets had moved on. (Vintage 33 mm image)During the summer and early fall, juvenile and adult Ring-necked Pheasants feed heavily on grasshoppers. Earlier and more frequent mowing along with declining numbers of grasshoppers on farmlands due to an increase in pesticide use were factors contributing to the crash of the pheasant population in the early 1980s.To the delight of Wild Turkeys, each of the four species of grasshoppers shown above frequents clearings and roadsides adjacent to forest areas. While changes in grasshopper distribution have been detrimental to populations of birds like pheasants, they’ve created a feeding bonanza for turkeys.Wild Turkeys feeding on an abundance of grasshoppers along a forest road.An American Kestrel feeds on a grasshopper while ignoring the abundance of Spotted Lanternflies swarming the adjacent utility pole. In Susquehanna valley farmlands, grasshopper and kestrel numbers are down. Lanternflies, on the other hand, have got it made.Maintaining areas bordering roads, forests, wetlands, farmlands, and human development in a state of early succession can provide and ideal mix of mature grasses and broadleaf plants for grasshoppers, pollinators, birds, and other wildlife.
Here in a series of photographs are just a handful of the reasons why the land stewards at Middle Creek Wildlife Management Area and other properties where conservation and propagation practices are employed delay the mowing of fields composed of cool-season grasses until after August 15 each year.
Eastern Meadowlarks, birds of large pastures, hay lots and other meadows of cool-season grasses, build their nests and raise their young on the ground. In the years since the early twentieth century, loss in the volume of acreage maintained in the lower Susquehanna Valley as grassland habitat types has dramatically reduced the prevalence and abundance of this and other birds with similar nesting requirements. During the most recent fifty years, early and frequent mowing and other practices introduced as part of agriculture’s Green Revolution have all but eliminated ground-nesting grassland species from the region.Like the meadowlarks, Grasshopper Sparrows (Ammodramus savannarum) nest on the ground in fields of cool-season grasses. Mowing prior to the time the young leave the nest and are able to fly away can obliterate a generation of grassland birds. Because their life span is short, widespread loss of an entire year of reproduction can quickly impact overall populations of native sparrows and other small birds. Delayed mowing can improve numbers of Grasshopper Sparrows as well as Savannah Sparrows, Vesper Sparrows (Pooecetes gramineus), and the very rare Henslow’s Sparrow (Centronyx henslowii).The Bobolink, like the meadowlark, is a member of the blackbird family (Icteridae). It too requires grasslands free of disturbances like mowing for the duration of the nesting season which, for this particular bird, lasts until mid-August in the lower Susquehanna region. In places lacking their specific habitat requirements, Bobolinks will seldom be detected except as flyovers during migration.Ring-necked Pheasants were introduced to the lower Susquehanna basin, and their populations were maintained thereafter, by stocking for the purpose of hunting. But throughout the middle twentieth century, there was a substantial population of ring-necks breeding in fields of cool-season grasses in farmlands throughout the region. High-intensity agriculture with frequent mowing eliminated not only nesting habitat in grasslands, but winter cover in areas of early successional growth. Populations of Ring-necked Pheasants, as well as native Northern Bobwhite, crumbled during the late 1970s and early 1980s due to these changes. For these resident birds that don’t migrate or routinely travel great distances to find new places to live and breed, widespread habitat loss can be particularly catastrophic. Not surprisingly, the Northern Bobwhite is no longer found in the Lower Susquehanna River Watershed and has been extirpated from all of Pennsylvania.At places like Middle Creek Wildlife Management Area where a mix of grasslands, early successional growth, and even some cropland are maintained, the Blue Grosbeak has extended its range well north of the Mason-Dixon and has become a regular nesting species during recent decades. Good habitat management does pay dividends.
Right now is a good time to visit Middle Creek Wildlife Management Area to see the effectiveness a delayed mowing schedule can have when applied to fields of cool-season grasses. If you slowly drive, walk, or bicycle the auto tour route on the north side of the lake, you’ll pass through vast areas maintained as cool-season and warm-season grasses and early successional growth—and you’ll have a chance to see these and other grassland birds raising their young. It’s like a trip back in time to see farmlands they way they were during the middle years of the twentieth century.
Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge. At Bombay Hook, they don’t waste your money mowing grass. Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools. Birds there can often be observed at close range. The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools. Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.Two more Short-billed Dowitchers on the way in.Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed. Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge. Mallards, American Avocets, and egrets are gathered on the shore.More Short-billed Dowitchers arriving to feed in Raymond Pool.Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool. A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.American Avocets probe the muddy bottom of Raymond Pool.Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early. Many migrating Dunlin linger at Bombay Hook into October and even November.This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp Pool.Lesser Yellowlegs at Bear Swamp PoolA Greater Yellowlegs at Bear Swamp Pool.A Caspian Tern patrolling Raymond Pool.The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass. High tide licks at the roots of the cordgrass supporting the temporary domicile.By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice). It is our only dragonfly able to breed in saltwater. Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.Look up! A migrating Bobolink passes over the dike at Shearness Pool.Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool. Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river. The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century. An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers. Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.A Great Egret prowling Shearness Pool.A Snowy Egret in Bear Swamp Pool.Wood Ducks in Bear Swamp Pool.A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.
As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.
Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay. We found just this one during our visit to Bombay Hook. Man has certainly decimated populations of this ancient crustacean during recent decades.As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.A male Atlantic Marsh Fiddler Crab peers from its den.Herons and egrets including this Great Egret are quite fond of fiddler crabs. As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.An Osprey patrols the vast tidal areas opposite Shearness Pool.
No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook. This one was in shrubby growth along the dike at the north end of Shearness Pool.These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries. The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware. The seed balls are a favorite winter food of goldfinches and siskins.Finis Pool has no frontage on the tidal marsh but is still worth a visit. It lies along a spur road on the tour route and is located within a deciduous coastal plain forest. Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.The White-tailed Deity is common along the road to Finis Pool.Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well. This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.
We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.
