Five Favorite Visitors from the Northern Forests

As the autumn bird migration draws to a close for 2024, we’re delighted to be finding five of our favorite visitors from the coniferous and mixed forests of Canada and the northernmost continental United States.

Male Yellow-bellied Sapsucker
Yellow-bellied Sapsuckers drill evenly spaced holes which they repeatedly visit to harvest “sap” and the insects that become trapped therein.  These surgically small excavations penetrate only the bark and the phloem, a shallow, spongy layer of the trunk that carries sugars from the leaves to other parts of the tree.  Seldom do they puncture the thin, underlying cambium layer which protects the true sapwood.  After healing, sapsucker holes leave linear rows of small scars which are sometimes still discernible decades after a bird’s final visit.  This adult male will probably linger in the Lower Susquehanna River Watershed until colder weather renders the tree totally dormant, the flow of “sap” ebbs, and small insects become harder to find.  Then he’ll make his way off toward southern forests of oak and pine for the remainder of the winter.  Sapsuckers work their way north as sugary “sap” begins flowing again in the spring.
Hermit Thrush
The Hermit Thrush is another denizen of coniferous and mixed forests.  In habitats with dense cover and sufficient food sources like fruit-bearing shrubs and vines, some will remain through the winter.  The Hermit Thrush is the only spot-breasted thrush regularly seen in our region from November through March.
Fox Sparrow
Almost the size of a thrush, the largest of our native sparrows, the Fox Sparrow, is currently at the peak of its southbound migration.  While taking a break from their nocturnal flights, some may stray from forest edges to visit suburban feeding stations.  Most leave our area by mid-December, then begin returning north as early as mid-February.
Pine Siskins
Not present every year, flocks of Pine Siskins are beginning to arrive in the lower Susquehanna region.  They are currently most common along the mountain tops of the Ridge and Valley Province where they are feeding on the seeds of the Sweet Birch, a tree also known as Black Birch.  Look for them around niger seed feeders and on the prickly seed balls of cultivated American Sweetgum, but don’t be surprised if they quickly pass us by to spend the winter on the Atlantic Coastal Plain to our south where this native tree grows in abundance.
Female/Juvenile Male Purple Finch
Since late October, Purple Finches have been widespread throughout the region.  These female/juvenile male types are frequently being seen at feeders, particularly near woodlands and other areas with stands of conifers.
Male Purple Finch
Less frequently seen are the adult male Purple Finches in their raspberry-colored plumage.  What a dandy.

While right now is the best time to get out and look for these species from the northern forests, any or all of them could linger into the winter months, particularly where the food supply is sufficient and conifers and other evergreens provide cover from the blustery weather.

Three’s Company

For many species of terrestrial vertebrates living in the lower Susquehanna valley, it’s time to take refuge in a safe place below ground to enter a winter-long slumber.  The numerous frosty nights of the past two weeks have expedited the stragglers’ efforts to find a suitable place to pass our coldest months.

Wood Turtles
Aside from the Eastern Box Turtle (Terrapene carolina carolina), the Wood Turtle (Glyptemys insculpta) is our most terrestrial species of testudine, however it is seldom found far from wet bottomlands.  Last week, we found this pair preparing to burrow into alluvial soils along the edge of a floodplain wetland where they will spend the winter.  In recent years, Wood Turtle numbers have plummeted due to habitat fragmentation and loss, and due to collecting by short-sighted petkeepers and profiteers.
Wood Turtle
A sluggish Wood Turtle catching a few last rays of sunshine before digging in for the winter.   If you encounter these or any turtles now or at any time of year, it’s important that they not be disturbed.  Look and leave them be.

And now, just in time for Halloween, a look at some hibernating species that some of our more squeamish readers consider to be “scary”.

In the mountainous forests of the Ridge and Valley Province, there lives a seldom seen little mouse which, unlike the more familiar species found in the fields, farms, and homes of our valleys and Piedmont, spends almost half the year in true hibernation.

Woodland Jumping Mouse
The Woodland Jumping Mouse (Napaeozapus insignis) is a nocturnal species of rodent found primarily on rocky forest slopes, particularly those with growths of Eastern White Pine and Eastern Hemlock.  Unlike the similar White-footed Mouse, a species that sometimes enters homes in the fall and remains active year-round, the Woodland Jumping Mouse builds a small subterranean nest among rocks and boulders where it will enter a long winter slumber, often not emerging until the milder days of April.  During the long hibernation, these mice sometimes fatally diminish the energy stored in their body fat; up to 75% may perish before spring arrives (Merritt, 1987).

In spring and fall, Eastern (Black) Ratsnakes are frequently seen basking in the sun as they absorb heat and get their metabolism going.  During the hottest days of summer, when ambient air temperature is sufficient for their needs, they become nocturnal, hunting mice and other creatures that are also active at night.  During October, they make their way back to a winter den where they will remain until March or April.

Eastern Ratsnake
A last-minute Eastern (Black) Ratsnake makes a slow late-October trek across a mountain road toward a winter den on the south slope of the ridge.
Eastern Ratsnake
This Eastern Ratsnake is on its way to a cold-season hideout where it may spend the darker months in the company of other communal hibernators including…
Eastern Copperhead
…Eastern Copperheads…
Timber Rattlesnake
…and Timber Rattlesnakes.

That’s right, all three of these snakes are known to cohabitate, not just during hibernation, but at other times of the year as well.  And you thought they were mean and nasty to anything and everything in their path, didn’t you?

Here in the northeastern United States, fear of indigenous wild snakes is a totally irrational anxiety.  It’s a figment of our indoctrination.  Their wicked reputation is almost exclusively the result of a never-ending stream of demonizing propaganda from the pulpit, press, parents, panicked, and picture shows.  All they want from you and I is to be left alone.  Want something to really be afraid of this Halloween?  Try domestic dogs.  That’s right—the “friendly” pooch.  In the United States, they’ll land hundreds of people in the emergency room today.  Don’t like that one.  How ’bout cars and trucks.  They’ll kill and maim people all day long.  And don’t forget about sugar.  Yes friends, that sweet treat is pure poison that kills all day long.  Happy Halloween!

  SOURCES

Merritt, Joseph F.  1987.  Guide to the Mammals of Pennsylvania.  University of Pittsburgh Press.  Pittsburgh, PA.  pp.230-233.

Shaffer, Larry L.  1991.  Pennsylvania Amphibians and Reptiles.  Pennsylvania Fish Commission.  Harrisburg, PA.

This Week’s Enormous Diurnal Flights

For the past several weeks, we’ve given you a look at the fallout of Neotropical songbirds on mornings following significant nocturnal flights of southbound migrants.  Now let’s examine the diurnal (daytime) flights that have developed in recent days over parts of the lower Susquehanna basin and adjacent regions.

For those observing diurnal migrants, particularly at raptor-counting stations, the third week of September is prime time for large flights of tropics-bound Broad-winged Hawks.  Winds from easterly directions during this year’s movements kept the greatest concentrations of these birds in the Ridge and Valley Province and upper areas of the Piedmont as they transited our region along a southwesterly heading.  Counts topped 1,000 birds or more on at least one day at each of these lookouts during the past seven days.  Meanwhile at hawk watches along the Piedmont/Atlantic Coastal Plain border, where flights topped 10,000 or more birds on the best days last fall, observers struggled to see 100 Broad-winged Hawks in a single day.

Broad-winged Hawks
Dozens of migrating Broad-winged Hawks during a morning liftoff to the north of Blue Mountain.

Among the challenges counters faced while enumerating migratory Broad-winged Hawks this week was the often clear blue skies, a glaring sun, and the high altitude at which these raptors fly during the mid-day hours.

"Kettling" Broad-winged Hawks
Large numbers of Broad-winged Hawks climbing on a thermal updraft to gain altitude can become increasingly difficult to spot in a clear blue sky.
"Kettling" Broad-winged Hawks
Higher and higher they go until they practically disappear from view.
Broad-winged Hawks
Cloud cover can help make locating really high-flying birds a bit easier, but they can sometimes climb into the lower portion of the cloud and vanish.

Another factor complicating the hawk counters’ tasks this week, particularly west of the Susquehanna River, was the widespread presence of another group of diurnal fliers.

Blue Jay
Was it the migratory Blue Jays that are beginning to move through the area?   No.
Sharp-shinned Hawk
Was it the increasing numbers of Sharp-shinned Hawks and other raptors?  No.
Hawkwatchers Scanning the Skies
So what was it that complicated the hawkwatchers’ efforts to find high-flying Broad-winged Hawks during this week’s flights?
Spotted Lanternfly
Is it a bird?  Is it a plane?  No.  And it’s nobody’s superhero either.  It’s the non-native Spotted Lanternfly and it is now very numerous west of the Susquehanna and common enough to the east to make even experienced observers take a second look while scanning the skies for Broad-winged Hawks and other migrating raptors.  Just how many do raptor counters have to contend with?  Even where these invasive insects have become less populous in the lower Susquehanna valley, hundreds still fill the skies above forest clearings around hawk-counting stations including Waggoner’s Gap, Second Mountain, and Rocky Ridge County Park.
Spotted Lanternflies on Radar
Spotted Lanternfly populations explode with great vigor as they colonize areas with plenty of tender new growth on native, non-native, and cultivated shrubs, trees, and vines.  Earlier this week, National Weather Service Doppler Radar in Pittsburgh, Pennsylvania, captured enormous mid-afternoon flights of Spotted Lanternflies in recently invaded areas of southwestern Pennsylvania and eastern Ohio.  (NOAA/National Weather Service Doppler Radar image)
Spotted Lanternflies on Radar
A base velocity image consisting primarily of Spotted Lanternflies in southwestern Pennsylvania and eastern Ohio.  (NOAA/National Weather Service Doppler Radar image)

As the Broad-winged Hawk migration draws to a close during the coming ten days, the occurrence of the one dozen other migratory diurnal raptors will be on the upswing.  They generally fly at lower altitudes, often relying upon wind updrafts on the ridges for lift instead of high-rising thermal updrafts.  They therefore present better observation opportunities for visitors at hawk watch lookouts and are less frequently confused with high-flying Spotted Lanternflies.

Merlin
A Merlin seen earlier this week migrating through the Lower Susquehanna River Watershed.  The numbers of migrating Merlins and other falcons will peak sometime in coming weeks.

Heat Flux Processes in Streams and Their Impact on Coldwater and Coolwater Fishes

The deluge of rain that soaked the lower Susquehanna watershed during last week is now just a memory.  Streams to the west of the river, where the flooding courtesy of the remnants of Hurricane Debby was most severe, have reached their crest and receded.  Sliding away toward the Chesapeake and Atlantic is all that runoff, laden with a brew of pollutants including but not limited to: agricultural nutrients, sediment, petroleum products, sewage, lawn chemicals, tires, dog poop, and all that litter—paper, plastics, glass, Styrofoam, and more.  For aquatic organisms including our freshwater fish, these floods, particularly when they occur in summer, can compound the effects of the numerous stressors that already limit their ability to live, thrive, and reproduce.

(Environmental Protection Agency image)

One of those preexisting stressors, high water temperature, can be either intensified or relieved by summertime precipitation.  Runoff from forested or other densely vegetated ground normally has little impact on stream temperature.  But segments of waterways receiving significant volumes of runoff from areas of sun-exposed impervious ground will usually see increases during at least the early stages of a rain event.  Fortunately, projects implemented to address the negative impacts of stormwater flow and stream impairment can often have the additional benefit of helping to attenuate sudden rises in stream temperature.

Stream Subjected to Agricultural Runoff
While a row of trees along a creek can help provide protection from the thermal impact of the sun, a vegetative riparian buffer must be much wider to be effective for absorbing, cooling, and treating runoff from fields, lawns, and paved surfaces.  This buffer is too narrow to prevent surface runoff from polluting the water.

Of the fishes inhabiting the Lower Susquehanna River Watershed’s temperate streams, the least tolerant of summer warming are the trouts and sculpins—species often described as “coldwater fishes”.  Coldwater fishes require water temperatures below 70° Fahrenheit to thrive and reproduce.  The optimal temperature range is 50° to 65° F.  In the lower Susquehanna valley, few streams are able to sustain trouts and sculpins through the summer months—largely due to the effects of warm stormwater runoff and other forms of impairment.

Blue Ridge Sculpin
Sculpins, including the Blue Ridge Sculpin (Cottus caeruleomentum) seen here, are native coldwater fishes which, during the 11,000 years since the last glacial maximum, have had the availability of their favored habitat sharply reduced by warming water temperatures and a rising Atlantic.  During this interval, seawater has inundated the path of the “Late” Pleistocene lower Susquehanna which passed through the section of flooded river watershed we now call Chesapeake Bay and continued across the continental shelf to what was, during the glacial maximum, the river’s mouth at Norfolk Canyon.  Today, cut off from neighboring drainage basins, sculpins survive exclusively in cold headwaters, and only in those where human alterations including pollution, dams, channelization, and reduced base flow haven’t yet eliminated their isolated populations.  Formerly believed to be composed of two widespread North American species, the Slimy Sculpin (Cottus cognatus) and the Mottled Sculpin (Cottus bairdii), study in recent decades is discovering that sculpin populations in the present-day lower Susquehanna and neighboring Potomac headwaters consist of at least three newly delineated species: Blue Ridge Sculpin, Potomac Sculpin (Cottus gerardi), and Checkered Sculpin (Cottus sp.), the latter an as yet undescribed species found only in the refugium of limestone springs in the Potomac drainage in West Virginia; Frederick and Washington Counties, Maryland; and Franklin County, Pennsylvania.  (United States Geological Survey image)
Ice Age Susquehanna
Stare at this for a little while, you’ll figure it out…………More than 11,000 years ago, during the last glacial maximum, when sea level was about 275 feet lower than it is today, there was no Chesapeake Bay, just a great Susquehanna River that flowed to the edge of the continental shelf and its mouth at Norfolk Canyon.  It was a river draining taiga forests of pine, spruce , and fir, and it carried along the waters of all the present-day bay’s tributaries and more.  The section of the river’s watershed we presently call the lower Susquehanna was, at the time, the upper Susquehanna watershed.  Brook Trout and sculpins had the run of the river and its tributaries back then.  And the entire watershed was a coldwater fishery, with limestone and other groundwater springs providing not refuge from summer heat, but a place to escape freezing water.  (United States Geological Survey base image)
Norfolk Canyon, the mouth of the Susquehanna River during the most recent glacial maximum, now lies more than 275 feet below the surface of the ocean and plunges to more than a mile in depth along the finger of out wash from the gorge.  (United States Geological Survey image)
Rainbow. Brown, and Brook Trout
Tens of thousands of trout are raised in state-operated and cooperative nurseries for stocking throughout the lower Susquehanna valley.  These rearing facilities are located on spring-fed headwaters with sufficient flow to assure cold temperatures year round.  While the Rainbow Trout and Brown Trout (Salmo trutta) are the most commonly stocked species, the Brook Trout (Salvelinus fontinalis) is the only one native to American waters.  It is the least tolerant of stream warming and still reproduces in the wild only in a few pristine headwaters streams in the region.  During spring, all three of these species have been observed on rare occasions entering the fish lift facilities at the hydroelectric dams on the river, presumably returning to the Susquehanna as sea-run trout.

Coldwater fishes are generally found in small spring-fed creeks and  headwaters runs. Where stream gradient, substrate, dissolved oxygen, and other parameters are favorable, some species may be tolerant of water warmer than the optimal values.  In other words, these temperature classifications are not set in stone and nobody ever explained ichthyology to a fish, so there are exceptions.  The Brown Trout for example is sometimes listed as a “coldwater transition fish”, able to survive and reproduce in waters where stream quality is exceptionally good but the temperature may periodically reach the mid-seventies.

Eastern Blacknose Dace
The Eastern Blacknose Dace is sometimes classified as a “coldwater transition fish”.   It can be found in headwaters runs as well as in creeks with good water quality.
Longnose Dace
The Longnose Dace is another “coldwater transition fish” known only from clear, clean, flowing waters.

More tolerant of summer heat than the trouts, sculpins, and daces are the “coolwater fishes”—species able to feed, grow, and reproduce in streams with a temperature of less than 80° F, but higher than 60° F.  Coolwater fishes thrive in creeks and rivers that hover in the 65° to 70° F range during summer.

Creek Chubs
The Creek Chub is a familiar species of “coolwater fish” seldom found remaining in waters exceeding 80 degrees Fahrenheit.
The Yellow Perch (Perca flavescens) was perhaps the most frequently targeted coolwater “gamefish” in the Lower Susquehanna River Watershed prior to the introduction of the Northern Pike (Esox lucius) and Muskellunge (Esox masquinongy).  Today’s prevalence of warmwater streams and the dozens of species of non-native predatory fishes now naturalized within them have left the Yellow Perch populations greatly reduced and all but forgotten by anglers.  Out of sight, out of mind.  (National Park Service image)

What are the causes of modern-day reductions in coldwater and coolwater fish habitats in the lower Susquehanna River and its hundreds of miles of tributaries?  To answer that, let’s take a look at the atmospheric, cosmic, and hydrologic processes that impact water temperature.  Technically, these processes could be measured as heat flux—the rate of heat energy transfer per unit area per unit time, frequently expressed as watts per meter squared (W/m²).  Without getting too technical, we’ll just take a look at the practical impact these processes have on stream temperatures.

HEAT FLUX PROCESSES IN A SEGMENT OF STREAM

Heat Flux Processes on Stream and River Segments.  These processes could be measured as heat flux—the rate of heat energy transfer per unit area per unit time.  (Environmental Protection Agency image)
      • INCOMING TEMPERATURE AND FLOW—The baseline temperature of stream water entering a given segment of waterway is obviously the chief factor determining its temperature when exiting that segment.  Incoming temperature and flow also determine the water’s susceptibility to heat absorption or loss while transiting the segment.  Lower flows may subject the given volume of water to a greater loss or gain of heat energy during the time needed to pass through the segment than the same volume at a higher flow.  Lower flows may also reduce stream velocity and extend a given volume of water’s exposure time to the exchange of heat energy while moving through the segment.  Generally speaking…
        1. …the higher the stream flow, the less a given volume of that stream’s  water may be impacted by the effects of the heat flux processes within the segment.
        2. …the lower the stream flow, the more a given volume of that stream’s water may be impacted by the effects of the heat flux processes within that segment.
        3. …the temperature and flow rate of precipitation entering the segment are factors that determine the impact of its heat energy transfer to or from a given volume of the stream’s waters.
        4. …the temperature and flow rate of runoff and point-source discharges entering the segment are factors that determine the impact of their heat energy transfer to or from a given volume of the stream’s waters.
Stormwater Discharge into Channelized Creek
Stormwater from impervious surfaces including roads, parking lots, roofs, and lawns quickly impacts temperatures in small creeks.  Channelized  streams are availed few of the positive attributes provided by many of the heat flux processes we’re about to see.  They therefore suffer from severe impairment and are exposed to temperature extremes that few aquatic organisms can survive.  Runoff from sun-heated pavement during a summer thunderstorm can often exceed 100 degrees Fahrenheit and can, at sufficient flow rate, quickly raise the temperature of a small stream to well over 90 degrees.
Stormwater Runoff
Stormwater runoff not only poses a thermal threat to waterways, its a significant source of a wide variety of pollutants.
      • GROUNDWATER INPUT—In streams connected to the aquifer, the temperature in a flowing segment can be impacted by the influx of cold groundwater.  With temperatures ranging from about 52° to 60° Fahrenheit, groundwater will absorb heat from the stream in summer, and warm it in the winter.  In warmwater streams, coldwater and coolwater fishes will often seek areas of the substrate where groundwater is entering for use as refugium from the summer heat.  Yellow Perch in the lower Susquehanna are known to exhibit this behavior.
Creeks and rivers connected to the aquifer and receiving supplemental flow from it are known as “gaining streams”. These streams frequently feed water into the aquifer as well. (United States Geological Survey image)
When flowing through an area experiencing drought or an excessive removal of groundwater (lots of wells, etc.), a waterway can become a “losing stream”, one that surrenders a portion of its flow to recharge the aquifer.  Further downstream, the reduced flow can make such a creek or river more susceptible to the effects of heat flux processes.  (United States Geological Survey image)
Seriously depleted aquifers can lead to a “disconnected stream”.  Smaller waterways subjected to these conditions will sometimes lose all their flow to the ground, often causing a catastrophic failure of the aquatic ecosystem supported therein.  (United States Geological Survey image)
Urban Flooding and Dry Streambed
Urban runoff overwhelms this small stream with polluted water than can reach temperatures of 100 degrees or more (left), then lets it high and dry with no baseflow during periods of dry weather (right) as the waterway becomes disconnected from the much-depleted aquifer.
Stormwater Retention Basin
Well-designed and properly constructed stormwater retention basins not only recharge groundwater supplies for wells and streams, they can also help prevent thermal pollution in waterways.  Planted with native wetland species and allowed to thrive, they can become treasured wildlife islands in otherwise inhospitable environs.  The benefits don’t stop there; plants also help sequester nutrients contained in the runoff.
      • HYPORHEIC EXCHANGE—Related to groundwater input, hyporheic exchange is the slow movement of water through the rock, sand, gravel, and soils composing the streambed, saturated shoreline, shallow aquifer, and connected floodplain of a creek or river.  As a heat flux process, hyporheic exchange helps moderate extremes in seasonal water temperatures by conducting energy between the solid materials in the zone and the flowing water.  Hyporheic zones are important habitats for many species of aquatic invertebrates and spawning fish.  Natural chemical processes within these zones convert ammonia-producing wastes into nitrite, then nitrate, allowing it to be absorbed as food by plants growing in the stream or in the alluvium within the zone.  Vegetation removal, channelization, legacy sediments, silt deposits, and man-made walls and dams can negate the benefits of hyporheic exchange.
Exchange of surface and ground water within the hyporheic zone is most directly associated with high-gradient (left) and meandering (right) segments of streams. (United States Geological Survey image)
Legacy Sediments and Fill
Very common on streams in the lower Susquehanna valley are these accumulations of legacy sediments at the sites of former mill ponds.  After the dams were removed, the creeks began eroding their way down through the mire as they tried to reestablish their floodplains and find their native substrate.  These trapped waterways are not only cut off from their hyporheic zones, they’re now a major source of nutrient and sediment pollution.  Misguided landowners like this one frequently dump fill into these sites to “save their land” and “control flooding”.  The fill and materials added to “shore up the banks” do nothing to fix what ails the creek, but instead displace more water to make the impact of flooding even more widespread.
Flooplain and Stream Restoration
Rehabilitation projects that remove legacy sediments help restore hyporheic exchange by reconnecting the stream to its underlying geology, its floodplain, and its wetlands.  Rising waters remain in the floodplain where they get a good bio-scrubbing and help replenish the creek and groundwater supply.  As the experts say, “floodplains are for flooding.”
      • ATMOSPHERIC EXCHANGE (CONVECTION, EVAPORATION)—Primarily a process by which a stream loses heat energy and cools its waters, atmospheric exchange is also a means by which a warm air mass can relinquish heat to cooler waters and thus increase their temperature.  This phenomenon can be dramatically enhanced when a stream passes through a so-called urban heat island where air temperatures remain warm through the night.  Convection, the movement of heat energy through a fluid (liquid or gas), causes warmer, less-dense water to rise to the surface of a stream, particularly where there is minimal turbulence.  When the air above is cooler than the water’s surface layer, the stream will conduct heat energy across the water/atmosphere interface causing the warmed air molecules to rise in a convection column.  If the atmospheric relative humidity is less than 100%, some surface water will vaporize—a process that expends more of the stream’s heat energy.  The rate of convective and evaporative cooling in a given stream segment is directly related to the degree of difference between the water temperature and air temperature, and to the relative humidity in the air mass above the lake, creek, or river.  The mechanical action of stream turbulence including rapids, riffles, and falls increases the contact area between air and water to maximize the atmospheric exchange of heat energy.  The convective air current we call surface wind has a turbulent wave-producing effect on water that can also maximize atmospheric exchange; think of a cold autumn wind robbing heat energy from a warm lake or river or a hot summer wind imparting its heat to a cooler creek.  These exchanges are both conductive in nature (air-to-water/water-to-air) and evaporative, the latter being expedited by the movement of dry air over warm water.
Tessellated Darter
Usually classified as one of the coolwater fishes, the bottom-dwelling Tessellated Darter can thrive in the warmer creeks and in the main stem of the Susquehanna by inhabiting riffles where atmospheric exchange in the form of increased evaporation helps reduce temperatures and convective currents carry the cooler, well-oxygenated water to the streambed.
Three mile Island Unit 1 Cooling Towers
Humans utilize the concept of atmospheric exchange, adopting the phenomena of evaporation and convection to cool the hot waters produced during electric generation and other industrial processes before discharge into a lake or river.
      • STREAMBED CONDUCTIVE EXCHANGE—In the lower Susquehanna watershed, there may be no better natural example of streambed conductive exchange than the Triassic-Jurassic diabase pothole bedrocks of Conewago Falls on the river at the south end of Three Mile Island.
During sunny days, the massive diabase pothole rocks at Conewago Falls absorb solar (shortwave) radiation, then conduct that heat energy into the flowing water, often continuing to pass the accumulated warmth into the river during the night.  On cloudy days, the riverbed collects longwave atmospheric radiation, a heat flux process that yields significantly less energy for conduction into the rapids, riffles, and pools of the falls.  During periods of low river flow, the heating effect of streambed conductive exchange can become magnified.  Compared to conditions that prevail when torrents of turbid water are rushing through the falls, partially exposed bedrock surrounded by clear water collects radiated energy much more efficiently, then conducts the heat to a greatly reduced volume of passing water.  During summer and autumn, this process can create a mix of temperature zones within the falls with warmer water lingering in slow-moving pools and cooler water flowing in the deeper fast-moving channels.  Along the falls’ mile-long course, a haven is created for aquatic organisms including warmwater and some coolwater fishes, oft times attracting anglers and a variety of hungry migrating birds as well.
Fallfish
Classified as one of our coolwater fishes, the Fallfish finds favorable conditions for feeding, growing, and spawning in the well-oxygenated waters of Conewago Falls.
Northern Hog Sucker
Though the lower Susquehanna River is classified as a warmwater fishery, the Northern Hog Sucker (Hypentelium nigricans), another of our native coolwater fishes, finds the fast-moving waters of Conewago Falls to its liking.  Northern Hog Suckers are known to inhabit streams cold enough to host trout.  They exhibit remarkable home range fidelity, sometimes spending their entire lives occupying the same several hundred feet of waterway.  Northern Hog Suckers are often designated an indicator of good water quality, intolerant of many stream impairment parameters.  Their presence in Conewago Falls provides testament to the quality of the warmwater fishery there.
Severely Impaired Channelized Stream
An unnatural example.  The reduced base flow in this channelized and severely impaired creek has been rendered vulnerable to the negative impacts of several heat flux processes including streambed conductive exchange.  Urban stormwater/surfacewater inflow, solar (shortwave) radiation, and heat conducted into the stream from the masonry walls, curbs, and raceway can all conspire to cook aquatic organisms with life-quenching summer water temperatures exceeding 90 degrees Fahrenheit.
      • SOLAR (SHORTWAVE) RADIATION—The sun provides the energy that fuels the earth’s complex climate.  The primary heat flux process that heats our planet is the absorption of solar radiation in the shortwave spectrum, which includes ultraviolet, visible, and infrared frequencies at the upper end of the longwave spectrum.  Streams and other bodies of water absorb the greatest amounts of solar (shortwave) radiation during the weeks around summer solstice when the sun at mid-day is closer to zenith than at any other time of the year.  However, the heating impact of the radiation may be greatest when the volume of water in the creek, river, or lake is at its minimum for the year—often during early fall.
The rate, measured in watts per square meter, at which solar (shortwave) energy is directly radiated to a given area on the earth’s surface (including streams and other waters) is determined by: solar activity, the angle of the sun in the sky, aspect (slope) of the receiving surface, the opacity of the overlying atmosphere, and the distance of the earth from the sun.  The former varies with the year’s seasons, the time of day, and the latitude of a given area.  The latter is currently at its annual minimum when earth is at perihelion during the early days of January, thus providing the northern hemisphere with a little bump in radiation during the shortest days of the year when the sun is at its lowest angle in the sky.  (NASA image)
A varying portion of the solar (shortwave) radiation reaching the earth is reflected back into space by clouds.  A smaller share is absorbed by the atmosphere, thus heating it.  An even lesser quantity is reflected back into space by water and land.  The remainder of the energy is absorbed by the planet’s surfaces, its water and land. (NASA image)
      • INCIDENT SHORTWAVE RADIATION—Also known as insolation (incoming solar radiation), incident shortwave radiation is the sum total energy of both the direct solar radiation that travels to the earth’s surface unaffected by the atmosphere and the diffuse radiation, waves that have been weakened and scattered by constituents of the atmosphere before reaching the planet’s surface.  On a cloudy day, the warming of terrestrial surfaces including streams and other bodies of water is the result of diffuse radiation.  On days with any amount of sunshine at all, both direct and diffuse radiation heat our waters and lands.
Pumkinseed
Warmwater fishes such as the native Pumpkinseed (Lepomis gibbosus) thrive in sun-drenched 70-to-85-degree waters as long as other heat flux processes prevent sudden temperature increases and oxygen depletion.
Mowed Stream Bank
Mowed stream banks offer a waterway no protection from incoming solar (shortwave) radiation, nor terrestrial forms of impairment including nutrient-rich stormwater runoff and silt.
      • REFLECTED SHORTWAVE RADIATION—known as albedo, reflected solar (shortwave) radiation is energy directed away from the earth’s surface before being absorbed.  A surface’s albedo value is basically determined by its color, black having little reflective value, white and silvery surfaces reflecting nearly all solar (shortwave) radiation away.  A surface with no reflective properties has an albedo value of 0, while a totally reflective surface has a value of 1.  Clean snow with a value of about 0.85 to 0.9 (85% to 90%) is a highly reflective surface; yellow snow isn’t as good.  A stream, river, or lake blanketed with ice and snow will absorb very little solar energy and will rely upon other heat flux processes to trigger a melt and thaw.  The surface of open water has a varying albedo value determined mostly by the angle of the sun.  Solar radiation striking the water’s surface at a low angle is mostly reflected away, while that originating at an angle closer to zenith is more readily absorbed.
Forested Stream
To avoid the heating effects of solar (shortwave) and atmospheric longwave radiation, coldwater and coolwater fishes require streams offering protection from full exposure to direct sunlight and cloud cover.  Runs and creeks flowing beneath a closed canopy of forest trees are shielded from 25% or more of incoming radiation and are thus able to better maintain thermal stability during the most vulnerable period of the year for temperature-sensitive fishes, May through October.
      • LONGWAVE RADIATION—Radiation in the longwave spectrum is composed of infrared waves at frequencies lower than those of the shortwave spectrum.  Longwave radiation, sometimes just called infrared radiation, is produced by the earth and its atmosphere and is propagated in all directions, day and night.  It warms mostly the lower atmosphere which in turn warms the earth’s surface including its waters.  Some longwave energy can even be radiated into the waterway from its own streambed—and the stream can return the favor.  Other forms of mass surrounding  a stream such as a rocky shoreline or a man-made structure such as bridge pier can trade longwave radiation with a waterway.  The effect of these latter exchanges is largely trivial and never rivals the heat flux transfer of warm to cold provided by  conduction.
Longwave radiation emissions slow as the temperature of the emitting mass decreases, just as they also increase with temperature of the mass.  Longwave radiation emissions therefore decrease with altitude along with the temperature of the water vapor, carbon dioxide, methane, and other gases that produce them.  As such, the highest reaches of the atmosphere have a greatly reduced capability of shedding longwave radiation into space.  At ground level, lakes, creeks, and streams receive their greatest dose of longwave radiation while beneath the cover of low-lying clouds or fog.  (NASA image)
      • CANOPY RADIATION—Trees emit longwave radiation that may have a limited heat flux impact on waterway temperature.  This radiation is diffuse, of scattered effect, and scarcely detectable, particularly beneath multilayered dense canopies.  Some of the infrared energy transmitted by the tree canopy is radiated skyward as well.
      • WATER RADIATION—Water, like all earthly matter composed of vibrating molecules, emits longwave radiation.  This heat flux process provides an ongoing cooling effect to streams, rivers, lakes, and oceans—warmer ones shedding infrared energy at a faster rate than those that are cold.

Now that we have a basic understanding of the heat flux processes responsible for determining the water temperatures of our creeks and rivers, let’s venture a look at a few graphics from gauge stations on some of the lower Susquehanna’s tributaries equipped with appropriate United States Geological Survey monitoring devices.  While the data from each of these stations is clearly noted to be provisional, it can still be used to generate comparative graphics showing basic trends in easy-to-monitor parameters like temperature and stream flow.

Each image is self-labeled and plots stream temperature in degrees Fahrenheit (bold blue) and stream discharge in cubic feet per second (thin blue).

The West Conewago Creek drains much of the Gettysburg Basin’s Triassic redbeds in Adams and northern York Counties in Pennsylvania and includes a small headwaters area in northern Maryland.  The gauge station is located just a over a mile upstream from the waterway’s mouth on the Susquehanna just below Conewago Falls.  Right through the summer heatwave, this 90-day graph shows a consistent daily pattern of daytime rises in temperature and nighttime cooling.  To the right, a rapid cool down can be seen coinciding with two periods of high water, the first from a series of heavy thundershowers, the second from flooding caused by the remnants of Hurricane Debby.  Notice that the early August downpours were so heavy that they cooled the hot surface runoff and waterway quickly, without creating a rise in stream temperature at the gauging station.  Had this monitoring device been located on a small tributary in an area with an abundance of impervious surfaces, there would probably have been a brief rise in stream temperature prior to the cooldown.  (United States Geological Survey image)

The daily oscillations in temperature reflect the influence of several heat flux processes.  During the day, solar (shortwave) radiation and convection from summer air, especially those hot south winds, are largely responsible for the daily rises of about 5° F.  Longwave radiation has a round-the-clock influence—adding heat to the stream during the day and mostly shedding it at night.  Atmospheric exchange including evaporative cooling may help moderate the rise in stream temperatures during the day, and certainly plays a role in bringing them back down after sunset.  Along its course this summer, the West Conewago Creek absorbed enough heat to render it a warmwater fishery in the area of the gauging station.  The West Conewago is a shallow, low gradient stream over almost its entire course.  Its waters move very slowly, thus extending their exposure time to radiated heat flux and reducing the benefit of cooling by atmospheric exchange.  Fortunately for bass, catfish, and sunfish, these temperatures are in the ideal range for warmwater fishes to feed, grow, and reproduce—generally over 80° F, and ideally in the 70° to 85° F range.  Coolwater fishes though, would not find this stream segment favorable.  It was consistently above the 80° F maximum and the 60° to 70° F range preferred by these species.  And coldwater fishes, well, they wouldn’t be caught dead in this stream segment.  Wait, scratch that—the only way they would be caught in this segment is dead.  No trouts or sculpins here.

The Codorus Creek drains primarily the carbonate valleys of York County to the south of the West Conewago watershed.  This gauge station is located about a mile upstream from the creek’s mouth on the Susquehanna just below Haldeman Riffles.  The graphic pattern is very similar to that of the West Conewago’s: daily heating and cooling cycles and a noticeable drop in stream temperature in early August caused by a day of thundershowers followed by the remnants of Hurricane Debby.  (United States Geological Survey image)

Look closely and you’ll notice that although the temperature pattern on this chart closely resembles that of the West Conewago’s, the readings average about 5 degrees cooler.  This may seem surprising when one realizes that the Codorus follows a channelized path through the heart of York City and its urbanized suburbs—a heat island of significance to a stream this size.  Before that it passes through numerous impoundments where its waters are exposed to the full energy of the sun.  The tempering factor for the Codorus is its baseflow.  Despite draining a smaller watershed than its neighbor to the north, the Codorus’s baseflow (low flow between periods of rain) was 96 cubic feet per second on August 5th, nearly twice that of the West Conewago (51.1 cubic feet per second on August 5th).  Thus, the incoming heat energy was distributed over a greater mass in the Codorus and had a reduced impact on its temperature.  Though the Codorus is certainly a warmwater fishery in its lower reaches, coolwater and transitional fishes could probably inhabit its tributaries in segments located closer to groundwater sources without stress.  Several streams in its upper reaches are in fact classified as trout-stocked fisheries.

This is a zoomed-in look at the previous graph showing the impact of a rainfall event on the water temperatures in Codorus Creek.  Unlike the sharp declines accompanying the deluge of flood waters during the two events in early August, these lesser storms in late June generated just enough runoff to capture heat energy from impervious surfaces and warm the creek, temporarily breaking the daily heating/cooling cycle.  Upstream in the immediate area of the runoff, the impact on the stream and/or its tributaries was probably much more dramatic, certainly raising temperatures into the nineties or above.  (United States Geological Survey image)
Kreutz Creek drains a carbonate bedrock area of York County and flows parallel to the Lincoln Highway (US 30) to enter the Susquehanna at Wrightsville.  The gauging station is about one mile upstream from the creek’s mouth.   (United States Geological Survey image)

The Kreutz Creek gauge shows temperature patterns similar to those in the West Conewago and Codorus data sets, but notice the lower overall temperature trend and the flow.  Kreutz Creek is a much smaller stream than the other two, with a flow averaging less than one tenth that of the West Conewago and about one twentieth of that in the Codorus.  And most of the watershed is cropland or urban/suburban space.  Yet, the stream remains below 80° F through most of the summer.  The saving graces in Kreutz Creek are reduced exposure time and gradient.  The waters of Kreutz Creek tumble their way through a small watershed to enter the Susquehanna within twenty-four hours, barely time to go through a single daily heating and cooling cycle.  As a result, their is no chance for water to accumulate radiant and convective heat over multiple summer days.  The daily oscillations in temperature are less amplified than we find in the previous streams—a swing of about three degrees compared to five.  This indicates a better balance between heat flux processes that raise temperature and those that reduce it.  Atmospheric exchange in the stream’s riffles, forest cover, and good hyporheic exchange along its course could all be tempering factors in Kreutz Creek.  From a temperature perspective, Kreutz Creek provides suitable waters for coolwater fishes.

Muddy Creek drains portions of southern York County through rolling farmland and woodlots.  There are no large impoundments or widespread urban impacts in the watershed, which may help explain its slightly lower temperature trends.  (United States Geological Survey image)

Muddy Creek is a trout-stocked fishery, but it cannot sustain coldwater species through the summer heat.  Though temperatures in Muddy Creek may be suitable for coolwater fishes, silt, nutrients, low dissolved oxygen, and other factors could easily render it strictly a warmwater fishery, inhabited by species tolerant of significant stream impairment.

Chiques Creek drains mostly limestone farmland in northwestern Lancaster County.  The gauging station is located near the stream’s mouth on the Susquehanna at Chiques (Chickies) Rock.  Oscillations in temperature again resemble the other waterways, but daily highs remain almost entirely below 80 degrees.  (United States Geological Survey image)

A significant number of stream segments in the Chiques watershed have been rehabilitated to eliminate intrusion by grazing livestock, cropland runoff, and other sources of impairment.  Through partnerships between a local group of watershed volunteers and landowners, one tributary, Donegal Creek, has seen riparian buffers, exclusion fencing, and other water quality and habitat improvements installed along nearly ever inch of its run from Donegal Springs through high-intensity farmland to its mouth on the main stem of the Chiques just above its confluence with the Susquehanna.  The improved water quality parameters in the Donegal support native coldwater sculpins and an introduced population of reproducing Brown Trout.  While coldwater habitat is limited to the Donegal, the main stem of the Chiques and its largest tributary, the Little Chiques Creek, both provide suitable temperatures for coolwater fishes.

Limestone Formation on Little Chiques Creek
Streams in the Chiques Creek and similar limestone watersheds often pass through areas with significant bedrock formations.  Heat flux processes including groundwater input, hyporheic exchange, and streambed conductive exchange can have a greater influence on water temperature along these segments.
Eastern Blacknose Dace
A breeding condition Eastern Balcknose Dace, one of the coldwater transition fishes found in the Chiques and its tributaries.
Common Shiner
The Common Shiner (Luxilus cornutus), a fish tolerant of warmwater streams, prefers cool, clear waters for spawning.  For protection from late-spring and summer heat, breeding males may seek a section of creek with a streambed inflow of limestone groundwater to defend as their nesting territory.
A closeup of the Chiques Creek graph showing what appears to be a little bump in temperature caused by surface runoff during a couple of late-May showers.  Stream rehabilitation is an ongoing process and the pressures of land disturbances both old and new present challenges to those who make it their passion to fix the wrongs that have been inflicted upon our local waters.  Even the  exemplary Donegal Creek faces new threats from urbanization in one of its headwater areas several miles to the northwest of the historic springs.  (United States Geological Survey image)
Conewago Creek (East) drains primarily Triassic redbed farmlands in Dauphin, Lancaster, and Lebanon Counties.  Much of the headwaters area is forested but is experiencing an increasing rate of encroachment by housing and some commercial development.  Conewago Creek (East) enters the Susquehanna on the east side of Conewago Falls at Three Mile Island.  The watershed is equipped with three U.S.G.S. gauge stations capable of providing temperature data.  This first one is located just over a mile upstream of the creek’s mouth.  (United States Geological Survey image)

Despite its meander through and receipt of water from high-intensity farmland, the temperature of the lower Conewago (East) maxes out at about 85° F, making it ideal for warmwater fishes and even those species that are often considered coolwater transition fishes like introduced Smallmouth Bass, Rock Bass, Walleye, and native Margined Madtom.  This survivable temperature is a testament to the naturally occurring and planted forest buffers along much of the stream’s course, particularly on its main stem.  But the Conewago suffers serious baseflow problems compared to other streams we’ve looked at so far.  Just prior to the early August storms, flow was well below 10 cubic feet per second for a drainage area of more than fifty square miles.  While some of this reduced flow is the result of evaporation, much of it is anthropogenic in origin as the rate of groundwater removal continues to increase  and a recent surge in stream withdraws for irrigation reaches its peak during the hottest days of summer.

Juvenile Rock Bass
A juvenile Rock Bass.
A juvenile Margined Madtom.
A juvenile Margined Madtom.
A closer look at the Conewago Creek (East) graphic shows the temperature drop associated with a series of thundershowers and the remnants of Hurricane Debby in early August.  Despite the baseflow being below five cubic feet per second, the cooling effect of the downpours as measured in the area of the gauge was significant enough to overwhelm any heating of runoff that may have occurred as precipitation drained across hardened soils or man-made impervious surfaces.  (United States Geological Survey image)

A little side note—the flow rate on the Conewago at the Falmouth gauge climbed to about 160 cubic feet per second as a result of the remnants of Hurricane Debby while the gauge on the West Conewago at Manchester skyrocketed to about 20,000 cubic feet per second.  Although the West Conewago’s watershed (drainage area) is larger than that of the Conewago on the east shore, it’s larger only by a multiple of two or three, not 125.  That’s a dramatic difference in rainfall!

The Bellaire monitoring station on Conewago Creek (East) is located on the stream’s main stem just downstream from the mouth of Little Conewago Creek, a tributary with its origins in farmland and woodlots.  (United States Geological Survey image)

The temperatures at the Bellaire monitoring station, which is located upstream of the Conewago’s halfway point between its headwaters in Mount Gretna and its mouth, are quite comparable to those at the Falmouth gauge.  Although a comparison between these two sets of data indicate a low net increase in heat absorption along the stream’s course between the two points, it also suggests sources of significant warming upstream in the areas between the Bellaire gauge and the headwaters.

Data from the gauge site on the Little Conewago Creek shows a temperature averaging about five degrees cooler than the gauge several miles downstream on the main stem of the Conewago at Bellaire.  (United States Geological Survey image)

The waters of the Little Conewago are protected within planted riparian buffers and mature woodland along much of their course to the confluence with the Conewago’s main stem just upstream of Bellaire.  This tributary certainly isn’t responsible for raising the temperature of the creek, but is instead probably helping to cool it with what little flow it has.

Juvenile Eastern Blacknose Dace (top) and a juvenile Longnose Dace.
A stream like the Little Conewago Creek with daily temperatures that remain mostly below 80 degrees and retreat to 75 degrees or less during the night can be suitable for coldwater transition fishes like these juvenile Eastern Blacknose Dace (top) and Longnose Dace.

Though mostly passing through natural and planted forest buffers above its confluence with the Little Conewago, the main stem’s critically low baseflow makes it particularly susceptible to heat flux processes that raise stream temperatures in segments within the two or three large agricultural properties where owners have opted not to participate in partnerships to rehabilitate the waterway.  The headwaters area, while largely within Pennsylvania State Game Lands, is interspersed with growing residential communities where potable water is sourced from hundreds of private and community wells—every one of them removing groundwater and contributing to the diminishing baseflow of the creek.  Some of that water is discharged into the stream after treatment at the two municipal sewer plants in the upper Conewago.  This effluent can become quite warm during processing and may have significant thermal impact when the stream is at a reduced rate of flow.  A sizeable headwaters lake is seasonally flooded for recreation in Mount Gretna.  Such lakes can function as effective mid-day collectors of solar (shortwave) radiation that both warms the water and expedites atmospheric exchange.

The Conewago Creek (East) Watershed from the Bellaire U.S.G.S. Gauging Station (lower left) upstream to the headwaters in Mount Gretna.  (United States Geological Survey image)

Though Conewago Creek (East) is classified as a trout-stocked fishery in its upper reaches in Lebanon County, its low baseflow and susceptibility to warming render it inhospitable to these coldwater fishes by late-spring/early summer.

River Chub
Despite being considered a warmwater fish, the River Chub (Nocomis micropogon) will ascend streams like the Conewago to seek cooler, gravel-bottomed waters for spawning.  Reduced baseflow has probably rendered the stream currently too small for this species on Pennsylvania State Game Lands in Colebrook where this specimen was photographed in 2018.
Juvenile Golden Shiner
The Golden Shiner, another warmwater fish, often ascends streams to enter cooler water. Juvenile Golden Shiners like this one will move into shallower headwaters not only to seek reduced temperatures, but to escape large predatory fishes as well.
Irrigation using stream water.
Irrigation of agricultural fields using a large portion of the already diminished baseflow in the Conewago Creek (East) just downstream of the Bellaire gauging station.  Despite millions of dollars in investment to rehabilitate this Susquehanna valley stream, the riparian buffers and other practices can have little effect when the creek gets sucked down to just a trickle.  Low baseflow is a hard nut to crack.  It’s best prevented, not corrected.
Hammer Creek, a trout-stocked fishery, originates, in part, within Triassic conglomerate in the Furnace Hills of Lebanon County, then flows north into the limestone Lebanon Valley where it picks up significant flow from other tributaries before working its way south back through the Furnace Hills into the limestone farmlands of Lancaster County.  From there the stream merges with the Cocalico Creek, then the Conestoga River, and at last the Susquehanna.  Note the tremendous daily temperature oscillations on this headwaters stream as it surges about 15 degrees each day before recovering back close to groundwater temperature by sunrise the next day.  (United States Geological Survey image)
Headwaters of Hammer Creek including Buffalo Springs, a significant source of cold groundwater feeding the western leg of the stream.  The large dams on this section that created the Lebanon and Rexmont Reservoirs have been removed.  (United States Geological Survey base image)

The removal of two water supply dams on the headwaters of Hammer Creek at Rexmont eliminated a large source of temperature fluctuation on the waterway, but did little to address the stream’s exposure to radiant and convective heat flux processes as it meanders largely unprotected out of the forest cover of Pennsylvania State Game Lands and through high-intensity farmlands in the Lebanon Valley.  Moderating the temperature to a large degree is the influx of karst water from Buffalo Springs, located about two miles upstream from this gauging station, and other limestone springs that feed tributaries which enter the Hammer from the east and north.  Despite the cold water, the impact of the stream’s nearly total exposure to radiative and other warming heat flux processes can readily be seen in the graphic.  Though still a coldwater fishery by temperature standards, it is rather obvious that rapid heating and other forms of impairment await these waters as they continue flowing through segments with few best management practices in place for mitigating pollutants.  By the time Hammer Creek passes back through the Furnace Hills and Pennsylvania State Game Lands, it is leaning toward classification as a coolwater fishery with significant accumulations of sediment and nutrients.  But this creek has a lot going for it—mainly, sources of cold water.  A core group of enthusiastic landowners could begin implementing the best management practices and undertaking the necessary water quality improvement projects that could turn this stream around and make it a coldwater treasure.  An organized effort is currently underway to do just that.  Visit Trout Unlimited’s Don Fritchey Chapter and Donegal Chapter to learn more.  Better yet, join them as a volunteer or cooperating landowner!

Male Creek Chub
The male Creek Chub, one of our coolwater fishes, develops head tubercles and becomes flushed with color during spawning season.  Hammer Creek not only provides a home for the Creek Chub, its cold headwaters provide refuge for a population of native Brook Trout too.
Like no other example we’ve looked at so far, this closeup of the Hammer Creek graphic shows temperature bumps correlating with the stormwater runoff from early August’s rains.  Because the stream flow is small and the precipitation rate was not as great at this location, the effect of heat flux from runoff is more readily apparent.  (United States Geological Survey image)
Brook Trout adult and juvenile.  (United States Fish and Wildlife Service image by Ryan Hagerty)

For coldwater fishes, the thousands of years since the most recent glacial maximum have seen their range slowly contract from nearly the entirety of the once much larger Susquehanna watershed to the headwaters of only our most pristine streams.  Through no fault of their own, they had the misfortune of bad timing—humans arrived and found coldwater streams and the groundwater that feeds them to their liking.  Some of the later arrivals even built their houses right on top of the best-flowing springs.  Today, populations of these fishes in the region we presently call the Lower Susquehanna River Watershed are seriously disconnected and the prospect for survival of these species here is not good.  Stream rehabilitation, groundwater management, and better civil planning and land/water stewardship are the only way coldwater fishes, and very possibly coolwater fishes as well, will survive.  For some streams like Hammer Creek, it’s not too late to make spectacular things happen.  It mostly requires a cadre of citizens, local government, project specialists, and especially stakeholders to step up and be willing to remain focused upon project goals so that the many years of work required to turn a failing stream around can lead to success.

Riparian Buffer
Riparian buffers with fences to exclude livestock can immediately begin improving water quality.  With establishment of such vegetative buffers, the effects of stressors that otherwise eliminate coldwater and coolwater fishes from these segments will begin to diminish.
Riparian Buffer
Within five to ten years, a riparian buffer planted with native trees is not only helping to reduce nutrient and sediment loads in the stream, it is also shielding the waters from heat flux processes including the solar (shortwave) radiation that raises water temperatures to levels not tolerated by coldwater and coolwater fishes.
Riparian Buffer
A well-established riparian buffer.
Forested Stream
A forested stream.

You’re probably glad this look at heat flux processes in streams has at last come to an end.  That’s good, because we’ve got a lot of work to do.

Add one more benefit to the wildflower meadow, it infiltrates stormwater to recharge the aquifer much better than mowed grass.  And another related plus, it reduces runoff and its thermal pollution.  Besides, you don’t have time to mow grass, because we have work to do!
Potomac Sculpin
Our native coldwater fishes including the Potomac Sculpin will survive only if we protect and expand the scattered few habitats where they have taken refuge.  They have no choice but to live in these seriously threatened places, but we do.  So let’s give ’em some space.  How ’bout it?  (United States Fish and Wildlife Service image by Ryan Hagerty)

Time to Order Trees and Shrubs for Spring

It’s that time of year.  Your local county conservation district is taking orders for their annual tree sale and it’s a deal that can’t be beat.  Order now for pickup in April.

The prices are a bargain and the selection includes the varieties you need to improve wildlife habitat and water quality on your property.  For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue).  And don’t forget to order packs of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local native birds.  They’re only $12.00 for a bundle of 10.

Mature Trees in a Suburban Neighborhood
It’s the most desirable block in town, not because the houses are any different from others built during the post-war years of the mid-twentieth century, but because the first owners of these domiciles had the good taste and foresight to plant long-lived trees on their lots, the majority of them native species.  Pin Oak, Northern Red Oak, Yellow Poplar, Flowering Dogwood (Cornus florida), Eastern Red Cedar, Eastern White Pine, Eastern Hemlock, Norway Spruce, and American Holly dominate the landscape and create excellent habitat for birds and other wildlife.  These 75-year-old plantings provide an abundance of shade in summer and thermal stability in winter, making it a “cool” place to live or take a stroll at any time of the year.

Cumberland County Conservation District Annual Tree Seedling Sale—

Orders due by: Friday, March 22, 2024

Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024

Common Winterberry
Cumberland County Conservation District is taking orders for Common Winterberry, the ideal small shrub for wet soil anywhere on your property.  To get berries, you’ll need both males and females, so buy a bunch and plant them in a clump or scattered group.
Pin Oak
To live for a century or more like this towering giant, a Pin Oak needs to grow in well-drained soils with adequate moisture.  These sturdy shade providers do well along streams and on low ground receiving clean runoff from hillsides, roofs, streets, and parking areas.  As they age, Pin Oaks can fail to thrive and may become vulnerable to disease in locations where rainfall is not adequately infiltrated into the soil.  Therefore, in drier areas such as raised ground or slopes, avoid the Pin Oak and select the more durable Northern Red Oak for planting.  This year, Pin Oaks are available from the Cumberland and Lancaster County Conservation Districts, while Dauphin, Lancaster, Lebanon, and York Counties are taking orders for Northern Red Oaks.
Purple Coneflower
The Cumberland County Conservation District is again offering a “Showy Northeast Native Wildflower and Grass Mix” for seeding your own pollinator meadow or garden.  It consists of more than twenty species including this perennial favorite, Purple Coneflower.

Dauphin County Conservation District Seedling Sale—

Orders due by: Monday, March 18, 2024

Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024

Eastern Redbud
The Eastern Redbud is small tree native to our forest edges, particularly in areas of the Piedmont Province with Triassic geology (Furnace Hills, Conewago Hills, Gettysburg/Hammer Creek Formations, etc.)  Also known as the Judas Tree, the redbud’s brilliant flowers are followed by heart-shaped leaves.  As seen here, it is suitable for planting near houses and other buildings.  Eastern Redbud seedlings are being offered through tree sales in Dauphin, Cumberland, and Lancaster Counties.

Lancaster County Annual Tree Seedling Sale—

Orders due by: Friday, March 8, 2024

Pickup on: Friday, April 12, 2024

Yellow Poplar
The Yellow Poplar, often called Tuliptree or Tulip Poplar for its showy flowers, is a sturdy, fast-growing deciduous tree native to forests throughout the Lower Susquehanna River Watershed.  Its pole-straight growth habit in shady woodlands becomes more spreading and picturesque when the plant is grown as a specimen or shade tree in an urban or suburban setting.  The Yellow Poplar can live for hundreds of years and is a host plant for the Eastern Tiger Swallowtail butterfly.  It is available this year from the Lancaster County Conservation District.
The American Sweetgum, also known as Sweet Gum, is a large, long-lived tree adorned with a mix of vibrant colors in autumn.
American Goldfinches and Pine Siskin on Sweet Gum
Ever wonder where all the American Goldfinches and particularly the Pine Siskins go after passing through our region in fall?  Well, many are headed to the lowland forests of the Atlantic Coastal Plain where they feed on an abundance of seeds contained in spiky American Sweetgum fruits.  In the Piedmont and Ridge and Valley Provinces of the Lower Susquehanna River Watershed, American Sweetgum transplants can provide enough sustenance to sometimes lure our friendly finches into lingering through the winter.
Sweet Gum in a Beaver Pond
The American Sweetgum is a versatile tree.  It can be planted on upland sites as well as in wet ground along streams, lakes, and rivers.  In the beaver pond seen here it is the dominate tree species.  This year, you can buy the American Sweetgum from the Lancaster County Conservation District.
"Red-twig Dogwood"
“Red-twig Dogwood” is a group of similar native shrubs that, in our region, includes Silky Dogwood and the more northerly Red-Osier Dogwood (Cornus sericea).  Both have clusters of white flowers in spring and showy red twigs in winter.  They are an excellent choice for wet soils.  Landscapers often ruin these plants by shearing them off horizontally a foot or two from the ground each year.  To produce flowers and fruit, and to preserve winter attractiveness, trim them during dormancy by removing three-year-old and older canes at ground level, letting younger growth untouched.
Silky Dogwood Stream Buffer
“Red-twig Dogwoods” make ideal mass plantings for streamside buffers and remain showy through winter, even on a gloomy day.  They not only mitigate nutrient and sediment pollution, they provide excellent food and cover for birds and other wildlife.  Both Silky and Red-osier Dogwoods are available for sale through the Lancaster County Conservation District as part of their special multi-species offers, the former is included in its “Beauty Pack” and the latter in its “Wildlife Pack”.  The similar Gray Dogwood (Cornus racemosa) is being offered for sale by the York County Conservation District.

Lebanon County Conservation District Tree and Plant Sale—

Orders due by: Friday, March 8, 2024

Pickup on: Friday, April 19, 2024

Common Pawpaw flower
The unique maroon flowers of the Common Pawpaw produce banana-like fruits in summer.  These small native trees grow best in damp, well-drained soils on slopes along waterways, where they often form clonal understory patches.  To get fruit, plant a small grove to increase the probability of pollination.  The Common Pawpaw is a host plant for the Zebra Swallowtail butterfly.  It is available through both the Lebanon and Lancaster County sales.
Eastern Red Cedar
The Eastern Red Cedar provides excellent food, cover, and nesting sites for numerous songbirds.  Planted in clumps of dozens or groves of hundreds of trees, they can provide winter shelter for larger animals including deer and owls.  The Eastern Red Cedar is being offered for purchase through both the Lebanon and Lancaster County Conservation Districts.
Hybrid American Chestnut
Care to try your hand at raising some chestnuts?  Lebanon County Conservation District has hybrid American Chestnut seedlings for sale.
Common Winterberry
Lebanon County Conservation District is offering Common Winterberry and Eastern White Pine during their 2024 Tree and Plant Sale.  Plant them both for striking color during the colder months.  Eastern White Pine is also available from the Cumberland, Dauphin, Lancaster, and York County sales.

Perry County Conservation District Tree Sale—

Orders due by: Sunday, March 24, 2024

Pickup on: Thursday, April 11, 2024

Pollinator Garden
In addition to a selection of trees and shrubs, the Perry County Conservation District is again selling wildflower seed mixes for starting your own pollinator meadow or garden.  For 2024, they have both a “Northeast Perennials and Annuals Mix” and a “Butterfly and Hummingbird Seed Mix” available.  Give them a try so you can give up the mower!

Again this year, Perry County is offering bluebird nest boxes for sale.  The price?—just $12.00.

Eastern Bluebird
Wait, what?,…twelve bucks,…that’s cheaper than renting!

York County Conservation District Seedling Sale—

Orders due by: Friday, March 15, 2024

Pickup on: Thursday, April 11, 2024

Buttonbush flower
The Buttonbush, a shrub of wet soils, produces a cosmic-looking flower.  It grows well in wetlands, along streams, and in rain gardens.  Buttonbush seedlings are for sale from both the York and Lancaster County Conservation Districts.

To get your deciduous trees like gums, maples, oaks, birches, and poplars off to a safe start, conservation district tree sales in Cumberland, Dauphin, Lancaster, and Perry Counties are offering protective tree shelters.  Consider purchasing these plastic tubes and supporting stakes for each of your hardwoods, especially if you have hungry deer in your neighborhood.

Deciduous Tree Planting Protected by Shelters
Tree shelters protect newly transplanted seedlings from browsing deer, klutzy hikers, visually impaired mower operators, and other hazards.

There you have it.  Be sure to check out each tree sale’s web page to find the selections you like, then get your order placed.  The deadlines will be here before you know it and you wouldn’t want to miss values like these!

Visit South Mountain for the Brightest Foliage Display You’ve Ever Seen

Where should you go this weekend to see vibrantly colored foliage in our region?  Where are there eye-popping displays of reds, oranges, yellows, and greens without so much brown and gray?  The answer is Michaux State Forest on South Mountain in Adams, Cumberland, and Franklin Counties.

South Mountain is the northern extension of the Blue Ridge Section of the Ridge and Valley Province in Pennsylvania.  Michaux State Forest includes much of the wooded land on South Mountain.  Within or adjacent to its borders are located four state parks: King’s Gap Environmental Education Center, Pine Grove Furnace State Park, Caledonia State Park, and Mont Alto State Park.  The vast network of trails on these state lands includes the Appalachian Trail, which remains in the mountainous Blue Ridge Section all the way to its southern terminus in Georgia.

Pennsylvania State Parks on South Mounatin
In Pennsylvania, the forested highlands of the Blue Ridge Section of the Ridge and Valley Province are known as South Mountain.  Much of South Mountain lies within the boundaries of Michaux State Forest.  Stars indicate the locations of 1) King’s Gap Environmental Education Center, 2) Pine Grove Furnace State Park, 3) Caledonia State Park, and 4) Mont Alto State Park.  A drive on US 30 between Gettysburg and Chambersburg will take you right through Michaux State Forest along an east to west axis while a scenic northbound or southbound trip along PA 233 will bring you in proximity to each of the state parks located therein.  (Base image from NASA Earth Observatory Collection)

If you want a closeup look at the many species of trees found in Michaux State Forest, and you want them to be labeled so you know what they are, a stop at the Pennsylvania State University’s Mont Alto arboretum is a must.  Located next door to Mont Alto State Park along PA 233, the Arboretum at Penn State Mont Alto covers the entire campus.  Planting began on Arbor Day in 1905 shortly after establishment of the Pennsylvania State Forest Academy at the site in 1903.  Back then, the state’s “forests” were in the process of regeneration after nineteenth-century clear cutting.  These harvests balded the landscape and left behind the combustible waste which fueled the frequent wildfires that plagued reforestation efforts for more than half a century.  The academy educated future foresters on the skills needed to regrow and manage the state’s woodlands.

Online resources can help you plan your visit to the Arboretum at Penn State Mont Alto.  More than 800 trees on the campus are numbered with small blue tags.  The “List of arboretum trees by Tag Number” can be downloaded to tell you the species or variety of each.  The interactive map provides the locations of individual trees plotted by tag number while the Grove Map displays the locations of groups of trees on the campus categorized by region of origin.  A Founder’s Tree Map will help you find some of the oldest specimens in the collection and a Commemorative Tree Map will help you find dedicated trees.  There is also a species list of the common and scientific tree names.

Yellow Buckeye
The Yellow Buckeye (Aesculus flava) is a tree found in the forests of the Blue Ridge Section of the Ridge and Valley Province in West Virginia, Virginia, North Carolina and Georgia.  You can see it in Pennsylvania by visiting the collection of trees in the Arboretum at Penn State Mont Alto.
American Chestnut
The American Chestnut can be difficult to find due to the impact of chestnut blight, but you can see it in the Arboretum at Penn State Mont Alto.
Shagbark Hickory
Shagbark Hickory is a common tree in the forests of South Mountain.
Sweet Cherry
The Sweet Cherry (Prunus avium), a native of Europe, is naturalized throughout eastern and south-central Pennsylvania and is one of the more than 150 species of trees in the arboretum’s collection.
Sweet Birch
Sweet Birch (Betula lenta) foliage is particularly bright yellow on South Mountain this autumn.  It really “pops” against the backdrop of the evergreen Eastern White Pines and Eastern Hemlocks.  During less-than-ideal years, Sweet Birch leaves can subtly transition from green to drab brown without much fanfare before falling.
Common Persimmon
You might have a difficult time finding a Common Persimmon (Diospyros virginiana) growing wild in Pennsylvania, but you can find it in the Arboretum at Penn State Mont Alto.

The autumn leaves will be falling fast, so make it a point this weekend to check out the show on South Mountain.

This Week at Regional Hawk Watches

With nearly all of the Neotropical migrants including Broad-winged Hawks gone for the year, observers and counters at eastern hawk watches are busy tallying numbers of the more hardy species of diurnal raptors and other birds.  The majority of species now coming through will spend the winter months in temperate and sub-tropical areas of the southern United States and Mexico.

Here is a quick look at the raptors seen this week at two regional counting stations: Kiptopeke Hawk Watch near Cape Charles, Virginia, and Second Mountain Hawk Watch at Fort Indiantown Gap, Pennsylvania.

Kiptopeke Hawk Watch
The hawk-watching platform at Kiptopeke State Park is located along Chesapeake Bay near the southern tip of Delmarva Peninsula.  In autumn, thousands of raptors and other birds migrate through the Atlantic Coastal Plain Province.  Those that follow the shorelines south frequently concentrate in spectacular numbers before crossing the mouths of the bays they encounter.  This phenomenon makes both Cape May, New Jersey, on Delaware Bay and Kiptopeke, Virginia, on Chesapeake Bay exceptional places to experience fall flights of migrating birds.
Second Mounatin Hawk Watch
A Sharp-shinned Hawk is counted as it swoops by the owl decoy at Second Mountain Hawk Watch at Fort Indiantown Gap Military Reservation.  Migrating raptors save energy by riding updrafts of air created by winds blowing against the slopes of the mountainsides in the Ridge and Valley Province.
Sharp-shinned Hawk
A Sharp-shinned Hawk passes the lookout at Second Mountain Hawk Watch.  “Sharp-shins” are currently the most numerous migrants both on the coast and at inland counting stations.
Sharp-shinned Hawk
A Sharp-shinned Hawk nearly passes observers unnoticed as it skims the treetops.
Sharp-shinned Hawk at owl decoy.
A Sharp-shinned Hawk eyes up an owl decoy.  Under cover of darkness, nocturnal owls could rather easily prey upon young and small adult hawks and falcons, both on the nest or at roost.  Accordingly, many diurnal raptors instinctively harass owls to drive them from their presence.  An owl decoy at the lookout helps attract migrating birds for a closer look.
Cooper's Hawk
An adult Cooper’s Hawk flaps its way past a counting station.  Like the similar Sharp-shinned Hawk, the larger Cooper’s Hawk is a member of the genus Accipiter.  As a proportion of the annual fall Accipiter flight, the Cooper’s Hawk is more numerous at coastal hawk watches than at inland sites.  (Editor’s Update:  As of autumn, 2024, the Cooper’s Hawk and the American Goshawk have been placed in the genus Astur.  The Sharp-shinned Hawk remains in the genus Accipiter.  Though the Cooper’s Hawk and Sharp-shinned Hawk are oft times confused by observers due to their like appearance, it turns out that the two are not as closely related as originally believed.  Neither is an offshoot of the other, nor do they descend from an immediate common ancestor.  Their resemblance may instead be a case of convergent evolution, most readily characterized by acquisition of similar physical traits due to shared demands for survival within their environment.)
Osprey
The majority of Osprey migrate along the coast, but a few are still being seen at inland hawk watches.
Bald Eagle
Bald Eagles are commonly seen at both coastal and inland lookouts.  Their movements continue well into late fall.
Northern Harrier
A Northern Harrier illuminated by a setting sun.  Northern Harriers are often still flying when many other species have gone to roost for the day.
An adult male Northern Harrier flying in misty weather.
An adult male Northern Harrier, the “gray ghost”, flying in misty weather, at a time when few other birds were in the air.
American Kestrel
The American Kestrel, like our other falcons, is seen in greatest concentrations at coastal counting stations.  It is our most numerous falcon.
Merlin
The Merlin provides only a brief observation opportunity as it passes the lookout.  These falcons are dark, speedy, and easily missed as they fly by.
Tree Swallow
While moving south, Merlins often accompany flights of migrating Tree Swallows, a potential food source.
Merlin with Dragonfly
A Merlin consumes a dragonfly.  Eating is no reason to stop moving.
Juvenile "Tundra Peregrine"
The “Tundra Peregrine” is an arctic-breeding Peregrine Falcon that travels a distance of over 6,000 miles to southern South America for winter.  It is strictly a migratory species in our region with numbers peaking during the first two weeks of October each year.  These strong fliers have little need for the updrafts from mountain ridges, inland birds often observed flying in a north to south direction.  The majority of “Tundra Peregrines” are observed following coastlines, with some migrating offshore to make landfall at points as far south as Florida and the Caribbean islands before continuing across water again to reach the northern shores of Central and South America.  This “Tundra Peregrine” is a juvenile bird on its first southbound trip.

During coming days, fewer and fewer of these birds will be counted at our local hawk watches.  Soon, the larger raptors—Red-tailed Hawks, Red-shouldered Hawks, and Golden Eagles—will be thrilling observers.  Cooler weather will bring several flights of these spectacular species.  Why not plan a visit to a lookout near you?  Click on the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab at the top of this page for site information and a photo guide to identification.  See you at the hawk watch!

Common Raven
It’s not all hawks at the hawk watch.  Even the coastal sites are now seeing fun birds like the playful Common Raven on a regular basis.
Eastern Meadowlarks in a Loblolly Pine.
Coastal locations are renowned places to see migrating songbirds in places outside of their typical habitat.  Here a flock of Eastern Meadowlarks has set down in the top of a Loblolly Pine (Pinus taeda) in downtown Cape Charles, Virginia, not far from Kiptopeke Hawk Watch.

Blooming in Early July: Great Rhododendron

With the gasoline and gunpowder gang’s biggest holiday of the year now upon us, wouldn’t it be nice to get away from the noise and the enduring adolescence for just a little while to see something spectacular that isn’t exploding or on fire?  Well, here’s a suggestion: head for the hills to check out the flowers of our native rhododendron, the Great Rhododendron (Rhododendron maximum), also known as Rosebay.

Great Rhododendron
The Great Rhododendron is an evergreen shrub found growing in the forest understory on slopes with consistently moist (mesic) soils.  The large, thick leaves make it easy to identify.  During really cold weather, they may droop and curl, but they still remain green and attached to the plant.

Thickets composed of our native heathers/heaths (Ericaceae) including Great Rhododendron, Mountain Laurel, and Pinxter Flower (Rhododendron periclymenoides), particularly when growing in association with Eastern Hemlock and/or Eastern White Pine, provide critical winter shelter for forest wildlife.  The flowers of native heathers/heaths attract bees and other pollinating insects and those of the deciduous Pinxter Flower, which blooms in May, are a favorite of butterflies and Ruby-throated Hummingbirds.

Pinxter Flower in bloom
A close relative of the Great Rhododendron is the Pinxter Flower, also known as the Pink Azalea.

Forests with understories that include Great Rhododendrons do not respond well to logging.  Although many Great Rhododendrons regenerate after cutting, the loss of consistent moisture levels in the soil due to the absence of a forest canopy during the sunny summertime can, over time, decimate an entire population of plants.  In addition, few rhododendrons are produced by seed, even under optimal conditions.  Great Rhododendron seeds and seedlings are very sensitive to the physical composition of forest substrate and its moisture content during both germination and growth.  A lack of humus, the damp organic matter in soil, nullifies the chances of successful recolonization of a rhododendron understory by seed.  In locations where moisture levels are adequate for their survival and regeneration after logging, impenetrable Great Rhododendron thickets will sometimes come to dominate a site.  These monocultures can, at least in the short term, cause problems for foresters by interrupting the cycle of succession and excluding the reestablishment of native trees.  In the case of forests harboring stands of Great Rhododendron, it can take a long time for a balanced ecological state to return following a disturbance as significant as logging.

Birds of Conewago Falls in the Lower Susquehanna River Watershed: Ruffed Grouse
Ruffed Grouse (Bonasa umbellus) may be particularly sensitive to the loss of winter shelter and travel lanes provided by thickets of Great Rhododendron and other members of the heather/heath family.  (Vintage 35 mm image)

In the lower Susquehanna region, the Great Rhododendron blooms from late June through the middle of July, much later than the ornamental rhododendrons and azaleas found in our gardens.   Set against a backdrop of deep green foliage, the enormous clusters of white flowers are hard to miss.

Great Rhododendron Flower Cluster
Great Rhododendrons sport an attractive blossom cluster.  The colors of the flower, especially the markings found only on the uppermost petal, guide pollinators to the stamens (male organs) and pistil (female organ).
Bumble Bee Pollinating a Great Rhododendron Flower
To this Bumble Bee (Bombus species), the yellowish spots on the uppermost petal of the Great Rhododendron may appear to be clumps of pollen and are thus an irresistible lure.  

In the Lower Susquehanna River Watershed, there are but a few remaining stands of Great Rhododendron.  One of the most extensive populations is in the Ridge and Valley Province on the north side of Second Mountain along Swatara Creek near Ravine (just off Interstate 81) in Schuylkill County, Pennsylvania.  Smaller groves are found in the Piedmont Province in the resort town of Mount Gretna in Lebanon County and in stream ravines along the lower river gorge at the Lancaster Conservancy’s Ferncliff and Wissler’s Run Preserves.  Go have a look.  You’ll be glad you did.

Great Rhododendron along Route 125 near Ravine
Great Rhododendron along Route 125 along the base of the north slope of Second Mountain north of Ravine, Schuylkill County, Pennsylvania.
Great Rhododendron along Swatara Creek
Great Rhododendrons beginning to bloom during the second week of July along Swatara Creek north of Ravine, Schuylkill County, Pennsylvania.  Note how acid mine drainage has stained the rocks in the upper reaches of this tributary of the lower Susquehanna.  Mitigation of leachate pollutants from legacy mines has facilitated the return of over two dozen species of fish to this waterway, but the telltale discoloration of the stream substrate will remain for some time.

Photo of the Day

A hatch-year/juvenile Red-tailed Hawk on its first southbound migration glides on an updraft created as northwest winds strike the slope of a hillside in the Valley and Ridge Province.  It’s a free ride, and this bird could easily cover one hundred miles or more in just one day of flying.

Spotted Lanternfly in the Lower Susquehanna River Watershed

Second Mountain Hawk Watch is located on a ridge top along the northern edge of the Fort Indiantown Gap Military Reservation and the southern edge of State Game Lands 211 in Lebanon County, Pennsylvania.  The valley on the north side of the ridge, also known as St. Anthony’s Wilderness, is drained to the Susquehanna by Stony Creek.  The valley to the south is drained toward the river by Indiantown Run, a tributary of Swatara Creek.

The hawk watch is able to operate at this prime location for observing the autumn migration of birds, butterflies, dragonflies, and bats through the courtesy of the Pennsylvania Game Commission and the Garrison Commander at Fort Indiantown Gap.  The Second Mountain Hawk Watch Association is a non-profit organization that staffs the count site daily throughout the season and reports data to the North American Hawk Watch Association (posted daily at hawkcount.org).

Today, Second Mountain Hawk Watch was populated by observers who enjoyed today’s break in the rainy weather with a visit to the lookout to see what birds might be on the move.  All were anxiously awaiting a big flight of Broad-winged Hawks, a forest-dwelling Neotropical species that often travels back to its wintering grounds in groups exceeding one hundred birds.  Each autumn, many inland hawk watches in the northeast experience at least one day in mid-September with a Broad-winged Hawk count exceeding 1,000 birds.  They are an early-season migrant and today’s southeast winds ahead of the remnants of Hurricane Florence (currently in the Carolinas) could push southwest-heading “Broad-wings” out of the Piedmont Province and into the Ridge and Valley Province for a pass by the Second Mountain lookout.

The flight turned out to be steady through the day with over three hundred Broad-winged Hawks sighted.  The largest group consisted of several dozen birds.  We would hope there are probably many more yet to come after the Florence rains pass through the northeast and out to sea by mid-week.  Also seen today were Bald Eagles, Ospreys, American Kestrels, and a migrating Red-headed Woodpecker.

Migrating Broad-winged Hawks circle on a thermal updraft above Second Mountain Hawk Watch to gain altitude before gliding away to the southwest.

Migrating insects included Monarch butterflies, and the three commonest species of migratory dragonflies: Wandering Glider, Black Saddlebags, and Common Green Darner.  The Common Green Darners swarmed the lookout by the dozens late in the afternoon and attracted a couple of American Kestrels, which had apparently set down from a day of migration.  American Kestrels and Broad-winged Hawks feed upon dragonflies and often migrate in tandem with them for at least a portion of their journey.

Still later, as the last of the Broad-winged Hawks descended from great heights and began passing by just above the trees looking for a place to settle down, a most unwelcome visitor arrived at the lookout.  It glided in from the St. Anthony’s Wilderness side of the ridge on showy crimson-red wings, then became nearly indiscernible from gray tree bark when it landed on a limb.  It was the dreaded and potentially invasive Spotted Lanternfly (Lycorma delicatula).  This large leafhopper is native to Asia and was first discovered in North America in the Oley Valley of eastern Berks County, Pennsylvania in 2014.  The larval stage is exceptionally damaging to cultivated grape and orchard crops.  It poses a threat to forest trees as well.  Despite efforts to contain the species through quarantine and other methods, it’s obviously spreading quickly.  Here on the Second Mountain lookout, we know that wind has a huge influence on the movement of birds and insects.  The east and southeast winds we’ve experienced for nearly a week may be carrying Spotted Lanternflies well out of their most recent range and into the forests of the Ridge and Valley Province.  We do know for certain that the Spotted Lanternfly has found its way into the Lower Susquehanna River Watershed.

This adult Spotted Lanternfly landed in a birch tree behind the observers at the Second Mountain Hawk Watch late this afternoon.  It was first recognized by its bright red wings as it glided from treetops on the north side of the lookout.