Golden-crowned Kinglet – LIFE IN THE LOWER SUSQUEHANNA RIVER WATERSHED

February 26, 2025April 28, 2025

Winter Survival: Generalists and Specialists

We’ve seen worse, but this winter has been particularly tough for birds and mammals in the Lower Susquehanna River Watershed. Due to the dry conditions of late summer and fall in 2024, the wild food crop of seeds, nuts, berries, and other fare has been less than average. The cold temperatures make insects hard to come by. Let’s have a look at how some of our local generalist and specialist species are faring this winter.

House Sparrows and a House Finch — House Sparrows (bottom) and House Finches (top) are generalists. To survive and thrive, they are adapted to a variety of habitats and types of food. House Sparrows live almost anywhere man-made structures are found. They are true omnivores and will eat almost anything, especially if they see something else try to eat it first. The House Sparrow’s close association with humans has allowed it to become the most widespread and successful living avian dinosaur. On a cold night, they’ll take shelter either within dense vegetation alongside a building or within the structure itself. Though not nearly as cosmopolitan, the House Finch has successfully colonized much of the eastern United States after escaping from captivity as a cage bird in New York during the middle of the twentieth century. Upon being trans-located here from the arid southwest, they adapted to suburbs and farmlands consuming primarily a granivore diet of seeds supplemented with seasonally available berries. They quickly became accustomed to offerings at bird-feeding stations as well. To survive the harsh winters in the northern sections of their range, eastern populations of House Finches are developing a pattern of migration. These movements are most evident in late fall when dozens or sometimes hundreds can be seen heading south over regional hawk-counting stations.

Northern Flicker feeding on Poison Ivy — Though they require dead trees for nesting and as places to find the grubs and adult insects upon which they primarily feed, woodpeckers including the Northern Flicker are generalists, seldom passing by a supply of fruits like these Poison Ivy berries as a source of winter food. Flickers regularly visit suburban areas where they’ll drop by at bird-feeding stations for suet. During the warmer months, they are the woodpecker most frequently seen on the ground where swarms of ants garner their full attention.

The Pileated Woodpecker is seldom found outside of mature forests where it digs relentlessly to remove grubs and other infestations from dead wood. But it is not a true specialist…

Pileated Woodpecker eating Poison Ivy — …it too finds a supply of Poison Ivy berries to be indispensable during a cold winter day.

The Ruby-crowned Kinglet is a generalist, feeding mostly on insects, but also consuming small seeds and some berries, including those of Poison Ivy. It nests well to our north in tall spruces and other evergreens. During migration and in winter, the Ruby-crowned Kinglet may be found in deciduous trees, brush, and tall grass in habitats ranging from forests to parks and suburbia. This male is displaying its seldom-seen red crown.

The Golden-crowned Kinglet, seen here on a Poison Ivy vine, is more of a specialist than the Ruby-crowned species, though the two will often occur in mixed groups during the winter. The Golden-crowned Kinglet nests in Spruce-Fir forests and in conifers within mixed woodlands. Even during migration, and particularly in winter, these birds are seldom found far from a stand of large evergreens within which they find shelter for the night.

The Hermit Thrush’s generalist lifestyle allows it to survive cold season weather in the Lower Susquehanna River Watershed. During summer, it breeds in coniferous and mixed woods from the northern parts of our valley north into Canada and feeds primarily on worms, insects, and other arthropods. During migration and in winter, the Hermit Thrush becomes a regular visitor to deciduous forests, woody parks and suburbs, particularly where a supply of wild berries is available to supplement its diet.

Iconic as it pulls earthworms from lawns during the warmer months, the generalist American Robin is fully dependent upon a crop of berries to survive winter conditions in the lower Susquehanna valley. The drought afflicted wild food crop of 2024 has led to fewer robins spending the season here and has delayed the northward push of migrating birds until the ground thaws and the earthworms make the ground rumble once again.

It’s insects for the nestlings during spring and summer, then berries through the winter for the cheerful Eastern Bluebirds, another generalist species.

American Crows are an excellent example of a generalist species. They’ll go anywhere to find food and they’ll eat almost anything. Like the House Sparrow and several other generalists, they adapt very well to human activity and actually thrive on it. Garbage anyone?

Another career generalist is the widely worshiped White-tailed Deity, a species adapted to nearly all man-made landscapes with adequate vegetation upon which to browse. Pushed to the limit during severe weather, some individuals will consume carrion and even resort to cannibalism.

You might think the Great Blue Heron is a specialist. Nope, it’s an accomplished generalist. Great blues will live, feed, and breed on almost any body of fresh or brackish water. And their diet includes almost anything that swims. In winter, you’ll even see them in fields hunting mice and voles.

The Red-shouldered Hawk is a generalist with a diet ranging from amphibians and reptiles to small rodents and large insects. Mostly regarded as a species of bottomlands, they’ll frequent woodland edges, roadsides, and suburbia during the winter months.

During its periodic winter visits to the region, the American Tree Sparrow feeds on seeds among the grasses and forbs of semi-open country with scattered short shrubs and trees. A generalist species, it will show up at backyard bird-feeding stations, particularly during periods of inclement weather. In summer, the American Tree Sparrow nests in tundra with growths of stunted willows and spruce and their diet includes insects as a source of protein for themselves and their young.

The White-crowned Sparrow has similar winter habitat preferences to the tree sparrows…

White-crowned Sparrows — …it becomes adaptable and something of a generalist when searching for food during bad winter storms.

The Savannah Sparrow is an omnivore favoring insects in summer and seeds in winter. Though very closely tied to its grassland habitat year-round, snow cover can push these birds to enter woodier environs to consume fruits like these rose hips.

Dusk and dawn during the short days of winter are the prime hunting times of a mammal specialist, the Short-eared Owl. Its presence in the lower Susquehanna valley is dependent on two dominant factors: extensive grassland habitat and an adequate population of the owl’s favored food, the Meadow Vole (Microtus pennsylvanicus). The Short-eared Owl’s requirements as a specialist species make finding a suitable place to live difficult. Unlike the generalist birds and mammals that often adapt to the widespread man-made disturbances in the region, populations of specialists frequently become fragmented, reduced in abundance, and subject to extirpation.

The Meadow Vole is a generalist rodent that can be abundant in grasslands, early successional growth, fallow fields, marshlands, and, of course, meadows. They are primarily herbivores, but will occasionally consume insects and other arthropods. Usually nocturnal, some individuals venture out along their surface runways during daylight hours becoming vulnerable to diurnal raptors including kestrels, harriers, and buteos.

Short-eared Owl patrolling a grassland for Meadow Voles — A Short-eared Owl in near darkness patrolling a grassland for Meadow Voles.

The eastward expansion of the Coyote (Canis latrans), a species of western North America’s grasslands and scrublands, and its progressive mixing with the Wolf (Canis lupus) in these eastern extensions of its range, has produced an expanding population of very adaptable generalists we call Eastern Coyotes (Canis latrans var.). These omnivorous canines colonized the Lower Susquehanna River Watershed during the last four decades to replace extirpated wolves as the top-tier natural predator in the region. Their primary diet includes Meadow Voles in grasslands and other small mammals along woodland edges and in successional habitats. Seasonally, they consume the berries of numerous wild plants. Slightly larger than their western ancestors, Eastern Coyotes with the admixture of Wolf genetics can subdue small ungulates. Like other apex predators, they are attracted to vulnerable prey and thus play a crucial role in culling the weak and diseased among their potential quarry species to assure the health and potential of populations of these species as long-term sources of food energy. The benefit to the prey species is however largely diluted in populous areas of the northeast; most venison consumed by Eastern Coyotes here is in the form of road kill. During periods of extended snow cover when small rodents and other foods become inaccessible, Eastern Coyotes, particularly young individuals, will wander into new areas seeking sustenance. Sometimes they venture into cities and suburbs where they explore the neighborhoods in search of garbage and pet foods placed outside the home. (Video clip courtesy of Tyler and Grace Good. Click image to view.)

Wildlife certainly has a tough time making it through the winter in the lower Susquehanna valley. Establishing and/or protecting habitat that includes plenty of year-round cover and sources of food and water can really give generalist species a better chance of survival. But remember, the goal isn’t to create unnatural concentrations of wildlife, it is instead to return the landscape surrounding us into more of a natural state. That’s why we try to use native plants as much as possible. And that’s why we try to attract not only a certain bird, mammal, or other creature, but we try to promote the development of a naturally functioning ecosystem with a food web, a diversity of pollinating plants, pollinating insects, and so on. Through this experience, we stand a better chance of understanding what it takes to graduate to the bigger job at hand—protecting, enhancing, and restoring habitats needed by specialist species. These are efforts worthy of the great resources that are sometimes needed to make them a success. It takes a mindset that goes beyond a focus upon the welfare of each individual animal to instead achieve the discipline to concentrate long-term on the projects and processes necessary to promote the health of the ecosystems within which specialist species live and breed. It sounds easier than it is—the majority of us frequently become distracted.

Being an individual from a population of a very successful generalist species is no guarantee of survival. This Eastern Gray Squirrel fell from a tall tree when the limbs became ice covered during a storm earlier this month. Just a freak accident? Maybe, but mistakes like this are often fatal in the natural world. This squirrel’s passing may seem brutal, but it provides a better opportunity for other squirrels and animals that share its food and cover requirements to make it through the winter. And those survivors that didn’t suffer such a fatal mistake or, more importantly, don’t possess a vulnerability that may have contributed to such a mishap will have a chance to pass those traits on to a new generation. This squirrel as an individual is gone, its species lives on, and may be stronger for its passing.

Prescribed Burn of a Grassland — Pennsylvania Game Commission crews maintain a grassland ecosystem for Short-eared Owls and other specialists using prescribed fire to prevent succession beyond its earliest stages. Among the additional specialist species benefiting from this management tool are Monarchs and other butterflies whose host plants survive early-season fire, but not competition with woody vines, shrubs, trees, and invasive herbaceous growth.

On the wider scale, it’s of great importance to identify and protect the existing and potential future habitats necessary for the survival of specialist species. And we’re not saying that solely for their benefit. These protection measures should probably include setting aside areas on higher ground that may become the beach intertidal zone or tidal marsh when the existing ones become inundated. And it may mean finally getting out of the wetlands, floodplains, and gullies to let them be the rain-absorbing, storm-buffering, water purifiers they spent millennia becoming. And it may mean it’s time to give up on building stick structures on tinderbox lands, especially hillsides and rocky outcrops with shallow, eroding soils that dry to dust every few years. We need to think ahead and stop living for the view. If you want to enjoy the view from these places, go visit and take plenty of pictures, or a video, that’s always nice—then live somewhere else. Each of these areas includes ecosystems that meet the narrow habitat requirements of many of our specialist species, and we’re building like fools in them. Then we feign victimhood and solicit pity when the calamity strikes: fires, floods, landslides, and washouts—again and again. Wouldn’t it be a whole lot smarter to build somewhere else? It may seem like a lot to do for some specialist animals, but it’s not. Because, you see, we should and can live somewhere else—they can’t.

The Allegheny Woodrat (Neotoma magister), a threatened species in Pennsylvania and a critically imperiled species in Maryland, is a habitat specialist requiring the forested rocky slopes, talus-flanked ridgetops, and caves of the Ridge and Valley Province for its nest sites and survival. Isolated populations survived within similar environs in the lower Susquehanna River valley’s Piedmont Province and on South Mountain through at least the first half of the twentieth century, but have since been extirpated. Human encroachment that fragments their habitat and promotes exposure to parasite-hosting mammals including the Raccoon (*Procyon lotor)*, carrier of the Raccoon Roundworm (Baylisascarius procyonis), could prove fatal to remaining populations of this native mammal. (National Park Service image by Rick Olsen)

The American Oystercatcher (Haematopus palliatus) is a specialist species that uses its highly adapted bill to feed on marine invertebrates including mollusks, few of which are actually oysters. Reliant upon tidal ecosystems for its survival, many of the seashore animals that make up this wader’s diet are themselves specialist species. Oystercatchers spend nearly their entire lives in tidal marshes or within the intertidal zone on beaches. They also frequent rocky jetties, particularly during high tide. This individual was photographed near the mouth of Chesapeake Bay in Northhampton County, Virginia, a location that, when the waters of the Atlantic started rising over 10,000 years ago, was the lower Susquehanna valley about 60 miles from the river’s mouth at present-day Norfolk Canyon along the edge of the continental shelf. Get the drift?

January 22, 2024January 22, 2024

Birds Along the River’s Edge

Just as bare ground along a plowed road attracts birds in an otherwise snow-covered landscape, a receding river or large stream can provide the same benefit to hungry avians looking for food following a winter storm.

Here is a small sample of some of the species seen during a brief stop along the Susquehanna earlier this week.

Along vegetated edges of the Susquehanna and its tributaries, the Song Sparrow is ubiquitous in its search for small seeds and other foods. As the river recedes from the effects of this month’s rains, the shoreline is left bare of more recently deposited snow cover. Song Sparrows and other birds are attracted to streamside corridors of frost-free ground to find sufficient consumables for supplying enough energy to survive the long cold nights of winter.

This Golden-crowned Kinglet was observed searching the trees and shrubs along the Susquehanna shoreline for tiny insects and spiders. Temperatures above the bare ground along the receding river can be a few degrees higher than in surrounding snow-covered areas, thus improving the chances of finding active prey among the trunks and limbs of the riparian forest.

Not far from the kinglet, a Brown Creeper is seen searching the bark of a Silver Maple (Acer saccharinum) for wintering insects, as well as their eggs and larvae. Spiders in all their life stages are a favorite too.

American Pipits not only inhabit farm fields during the winter months, they are quite fond of bare ground along the Susquehanna. Seen quite easily along a strip of pebbly shoreline exposed by receding water, these birds will often escape notice when spending time on mid-river gravel and sand bars during periods of low flow.

An American Pipit on a bitterly cold afternoon along the Susquehanna.

October 31, 2020August 17, 2022

October Transition

Thoughts of October in the Lower Susquehanna River Watershed bring to mind scenes of brilliant fall foliage adorning wooded hillsides and stream courses, frosty mornings bringing an end to the growing season, and geese and other birds flying south for the winter.

The autumn migration of birds spans a period equaling nearly half the calendar year. Shorebirds and Neotropical perching birds begin moving through as early as late July, just as daylight hours begin decreasing during the weeks following their peak at summer solstice in late June. During the darkest days of the year, those surrounding winter solstice in late December, the last of the southbound migrants, including some hawks, eagles, waterfowl, and gulls, may still be on the move.

The Rough-legged Hawk (Buteo lagopus), a rodent-eating raptor of tundra, grassland, and marsh, is rare as a migrant and winter resident in the lower Susquehanna valley. It may arrive as late as January, if at all.

During October, there is a distinct change in the list of species an observer might find migrating through the lower Susquehanna valley. Reduced hours of daylight and plunges in temperatures—particularly frost and freeze events—impact the food sources available to birds. It is during October that we say goodbye to the Neotropical migrants and hello to those more hardy species that spend their winters in temperate climates like ours.

During several of the first days of October, two hundred Chimney Swifts remained in this roost until temperatures warmed from the low forties at daybreak to the upper fifties at mid-morning; then, at last, the flock ventured out in search of flying insects. When a population of birds loses its food supply or is unable to access it, that population must relocate or perish. Like other insectivorous birds, these swifts must move to warmer climes to be assured a sustained supply of the flying bugs they need to survive. Due to their specialized food source, they can be considered “specialist” feeders in comparison to species with more varied diets, the “generalists”. After returning to this chimney every evening for nearly two months, the swifts departed this roost on October 5 and did not return.

A Northern Parula lingers as an October migrant along the Susquehanna. This and other specialist feeders that survive almost entirely on insects found in the forest canopy are largely south of the Susquehanna watershed by the second week of October.

The Blackpoll Warbler is among the last of the insectivorous Neotropical warblers to pass through the riparian forests of the lower Susquehanna valley each fall. Through at least mid-October, it is regularly seen searching for crawling insects and larvae among the foliage and bark of Northern Hackberry (Celtis occidentalis) trees near Conewago Falls. Most other warblers, particularly those that feed largely upon flying insects, are, by then, already gone.

The Blue-headed Vireo, another insectivore, is the last of the vireo species to pass through the valley. They linger only as long as there are leaves on the trees in which they feed.

Brown Creepers begin arriving in early October. They are specialist feeders, well-adapted to finding insect larvae and other invertebrates among the ridges and peeling bark of trees like this hackberry, even through the winter months.

Ruby-crowned Kinglets can be abundant migrants in October. They will often behave like cute little flycatchers, but quickly transition to picking insects and other invertebrates from foliage and bark as the weather turns frosty. Some may spend the winter here, particularly in the vicinity of stands of pines, which provide cover and some thermal protection during storms and bitter cold.

Beginning in early October, Golden-crowned Kinglets can be seen searching the forest wood for tiny invertebrates. They are the most commonly encountered kinglet in winter.

The Yellow-bellied Sapsucker, a woodpecker, is an October migrant that specializes in attracting small insects to tiny seeps of sap it creates by punching horizontal rows of shallow holes through the tree bark. Some remain for winter.

The Yellow-rumped Warbler arrives in force during October. It is the most likely of the warblers to be found here in winter. Yellow-rumped Warblers are generalists, feeding upon insects during the warmer months, but able to survive on berries and other foods in late fall and winter. Wild foods like these Poison Ivy berries are crucial for the survival of this and many other generalists.

American Robins are most familiar as hunters of earthworms on the suburban lawn, but they are generalist feeders that rely upon fruits like these Eastern Red Cedar (Juniperus virginiana) berries during their southbound migration in late October and early November each year. Robins remain for the winter in areas of the lower Susquehanna valley with ample berries for food and groves of mature pines for roosting.

Like other brown woodland thrushes, the Hermit Thrush (Catharus guttatus) is commonly seen scratching through organic matter on the moist forest floor in search of invertebrates. Unlike the other species, it is a cold-hardy generalist feeder, often seen eating berries during the southbound October migration. Small numbers of Hermit Thrushes spend the winter in the lower Susquehanna valley, particularly in habitats with a mix of wild foods.

Due to their feeding behavior, Cedar Waxwings can easily be mistaken for flycatchers during the nesting season, but by October they’ve transitioned to voracious consumers of small wild fruits. During the remainder of the year, flocks of waxwings wander widely in search of foods like this Fox Grape (Vitis labrusca). An abundance of cedar, holly, Poison Ivy, hackberry, bittersweet , hawthorn, wild grape, and other berries is essential to their survival during the colder months.

Red-breasted Nuthatches have moved south in large numbers during the fall of 2020. They were particularly common in the lower Susquehanna region during mid-October. Red-breasted Nuthatches can feed on invertebrates during warm weather, but get forced south from Canada in droves when the cone crops on coniferous trees fail to provide an adequate supply of seeds for the colder fall and winter seasons. In the absence of wild foods, these generalists will visit feeding stations stocked with suet and other provisions.

Purple Finches (Haemorhous purpureus) were unusually common as October migrants in 2020. They are often considered seed eaters during cold weather, but will readily consume small fruits like these berries on an invasive Mile-a-minute Weed (Persicaria perfoliata) vine. Purple Finches are quite fond of sunflower seeds at feeding stations, but often shy away if aggressive House Sparrows or House Finches are present.

The need for food and cover is critical for the survival of wildlife during the colder months. If you are a property steward, think about providing places for wildlife in the landscape. Mow less. Plant trees, particularly evergreens. Thickets are good—plant or protect fruit-bearing vines and shrubs, and allow herbaceous native plants to flower and produce seed. And if you’re putting out provisions for songbirds, keep the feeders clean. Remember, even small yards and gardens can provide a life-saving oasis for migrating and wintering birds. With a larger parcel of land, you can do even more.

**GOT BERRIES?** Common Winterbery (Ilex verticillata) is a native deciduous holly that looks its best in the winter, especially with snow on the ground. It’s slow-growing, and never needs pruning. Birds including bluebirds love the berries and you can plant it in wet ground, even along a stream, in a stormwater basin, or in a rain garden where your downspouts discharge. Because it’s a holly, you’ll need to plant a male and a female to get the berries. Full sun produces the best crop. Fall is a great time to plant, and many garden centers that sell holiday greenery still have winterberry shrubs for sale in November and December. Put a clump of these beauties in your landscape. Gorgeous!

October 11, 2020November 27, 2020

A Visit to Waggoner’s Gap

Nothing beats spending a day at a hawk watch lookout—except of course spending a day at a hawk watch lookout when the birds are parading through nonstop for hours on end.

Check out Waggoner’s Gap, a hawk count site located on the border of Cumberland and Perry Counties atop Blue Mountain just north of Carlisle, Pennsylvania. It is by far the Lower Susquehanna River Watershed’s best location for observing large numbers of migrating raptors during the October and November flights.

Waggoner’s Gap is located where Route 74 crosses Blue Mountain north of Carlisle.

The entrance to a parking area for hawk watch visitors is designated by this sign located along Route 74 several hundred yards north of the summit of Blue Mountain.

Since acquiring the site in 2000, Audubon Pennsylvania has added improvements to expand the function of Waggoner’s Gap Hawk Watch to include education for both formal students and the public at large.

The site is named in honor of the late conservationist Clifford L. Jones, a business leader, a former Chairman of the Pennsylvania Public Utilities Commission, a cabinet secretary for six Pennsylvania governors (both major parties), a director on the boards of numerous conservation organizations, and an active birder.

Orange falcon silhouettes function as blazes for the trails that lead from the parking area to the lookout. The trail and the lookout consist entirely of boulders. Some of these move when stepped upon. Others may be slick. Use caution at all times.

The lookout at Waggoner’s Gap is staffed by official counters from August through December each year. They are tasked with enumerating every migratory raptor’s passage during that period.

Sure-footed observers climb into a comfortable position among the Tuscarora quartzite boulders and begin watching the flight.

The view from the lookout is spectacular. To the east, downtown Harrisburg can be seen in the distance.

During a recent afternoon with breezes from the “southwesterlies”, a steady stream of migrating Sharp-shinned Hawks, including this juvenile, passed by the lookout.

Sharp-shinned Hawks were ready subjects for photography as they sailed on updrafts along the south side of the ridge.

A Sharp-shinned Hawk below eye level. Over 400 Sharp-shinned Hawks migrated past Waggoner’s Gap Hawk Watch on this particular early October day.

The local Turkey Vultures at Waggoner’s Gap seem ubiquitous at times. They’re on the radio towers, they’re flying overhead, and a few are cruising the slopes below the crest. But on the day of our recent visit, their numbers were eclipsed by the more than 300 “T.V.s” that migrated down the ridge.

Black Vultures, both migrants and local birds, are seen from the lookout.

Northern Harriers are a hawk watch favorite. Their long uptilted wings, long tail, and white rump make them easy to identify, even for beginners. Their plummeting numbers make them a treasured sighting for everyone.

A distant Red-shouldered Hawk. Numbers of these migrants peak later in the season.

A Peregrine Falcon darts past the lookout. Note the white forehead, throat, and breast. This bird is probably a “Tundra Peregrine” (Falco peregrinus tundrius). In the lower Susquehanna valley, this subspecies is strictly migratory, a transient in spring and fall. “Tundra Peregrines” breed in the arctic and winter as far south as South America.

An immature Bald Eagle. Waggoner’s Gap is a superb place for sighting eagles, especially on a breezy day.

Hundreds of Blue Jays filtered through as their southbound exodus continues. Other songbirds of interest included Blue-headed Vireos (Vireo solitarius), Winter Wrens, Red-breasted Nuthatches, Pine Siskins, and both Ruby-crowned and Golden-crowned Kinglets.

Waggoner’s Gap is a hardy birder’s paradise. During the latter portion of the season, excellent flights often occur on days that follow the passage of a cold front and have strong northwest winds. But be prepared, it can be brutal on those rocks during a gusty late-October or early-November day after the leaves fall—so dress appropriately.

To see the daily totals for the raptor count at Waggoner’s Gap Hawk Watch and other hawk watches in North America, and to learn more about each site, be sure to visit hawkcount.org

November 25, 2017November 22, 2024

Loading Up For Winter

A very light fog lifted quickly at sunrise. Afterward, there was a minor movement of migrants: forty-nine Ring-billed Gulls, a few American Herring Gulls, a Red-shouldered Hawk following the river to the southeast, and small flocks totaling nine Cedar Waxwings and twenty-eight Red-winged Blackbirds.

A Ring-billed Gull calls as active migrants pass overhead on their way downriver.

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In the Riparian Woodland, small mixed flocks of winter resident and year-round resident birds were actively feeding. They must build and maintain a layer of body fat to survive blustery cold nights and the possible lack of access to food during snowstorms. There’s no time to waste; nasty weather could bring fatal hardship to these birds soon.

A Carolina Chickadee *(Poecile carolinensis)* feeds on the seeds of an Eastern Sycamore *(Platanus occidentalis)*, also known as American Sycamore. Chickadees are generalist feeders, eating invertebrates and suet at feeding stations in addition to the seeds of many plants. Carolina Chickadees are year-round residents at Conewago Falls.

A fast-moving Golden-crowned Kinglet zips from limb to limb to grab tiny insects and other invertebrates. During the winter, these petite birds will carefully probe the bark and crevices of trees to glean enough food to survive. Golden-crowned Kinglets are winter residents at Conewago Falls. In spring, they will depart to nest in coniferous forests.

A Downy Woodpecker *(Picoides pubescens)* searches an infected tree for insects. They are year-round residents.

Carolina Wrens *(Thryothorus ludovicianus)* are considered year-round residents at Conewago Falls, though they may withdraw to the south during severe winters. Carolina Wrens sing year-round. Today, their loud melody echoed through the Riparian Woodland all morning.

The tiny bob-tailed Winter Wren is an elusive ground-dwelling winter resident at the falls. You may hear their scolding chatter from rocky areas and tree logs where they climb around mouse-like in search of small invertebrates. Their song is a fast jumble of dainty musical trills that can sometimes be heard echoing through the Riparian Woodland in winter. In spring, they’ll depart to nest in damp coniferous forests.

November 12, 2017July 1, 2018

Anthropoavians

Temperatures plummeted to well below freezing during the past two nights, but there was little sign of it in Conewago Falls this morning. The fast current in the rapids and swirling waters in flooded Pothole Rocks did not freeze. Ice coated the standing water in potholes only in those rocks lacking a favorable orientation to the sun for collecting solar heat during the day to conduct into the water during the cold nights.

On the shoreline, the cold snap has left its mark. Ice covers the still waters of the wetlands. Frost on exposed vegetation lasted until nearly noontime in shady areas. Insect activity is now grounded and out of sight. The leaves of the trees tumble and fall to cover the evidence of a lively summer.

The nocturnal bird flight is narrowing down to just a few species. White-throated Sparrows, a Swamp Sparrow (Melospiza georgiana), and Song Sparrows are still on the move. Though their numbers are not included in the migration count, hundreds of the latter are along the shoreline and in edge habitat around the falls right now. Song Sparrows are present year-round, migrate at night, and are not seen far from cover in daylight, so migratory movements are difficult to detect. It is certain that many, if not all of the Song Sparrows here today have migrated and arrived here recently. The breeding population from spring and summer has probably moved further south. And many of the birds here now may remain for the winter. Defining the moment of this dynamic, yet discrete, population change and logging it in a count would certainly require different methods.

Song Sparrows are now abundant in the brushy edges of fields and woodlands. They may even break into song on sunny days.

Diurnal migration was foiled today by winds from southerly directions and moderating temperatures. The only highlight was an American Robin flight that extended into the morning for a couple of hours after daybreak and totaled over 800 birds. This flight was peppered with an occasional flock of blackbirds. Then too, there were the villains.

They’re dastardly, devious, selfish, opportunistic, and abundant. Today, they were the most numerous diurnal migrant. Their numbers made this one of the biggest migration days of the season, but they are not recorded on the count sheet. It’s no landmark day. They excite no one. For the most part, they are not recognized as migrants because of their nearly complete occupation of North America south of the taiga. If people build on it or alter it, these birds will be there. They’re everywhere people are. If the rotten attributes of man were wrapped up into one bird, an “anthropoavian”, this would be it.

Meet the European Starling (Sturnus vulgaris). Introduced into North America in 1890, the species has spread across the entire continent. It nests in cavities in buildings and in trees. Starlings are aggressive, particularly when nesting, and have had detrimental impacts on the populations of native cavity nesting birds, particularly Red-headed Woodpeckers, Purple Martins (Progne subis), and Eastern Bluebirds. They commonly terrorize these and other native species to evict them from their nest sites. European Starlings are one of the earlier of the scores of introduced plants and animals we have come to call invasive species.

Noisy flocks of European Starlings are right at home on man-made structures in city and country.

Today, thousands of European Starlings were on the move, working their way down the river shoreline and raiding berries from the vines and trees of the Riparian Woodlands. My estimate is between three and five thousand migrated through during the morning. But don’t worry, thousands more will be around for the winter.

European Starlings mob a Sharp-shinned Hawk from above, a common behavior.

An Eastern Bluebird feeds on the few berries left untouched by passing European Starlings.

November 4, 2017September 4, 2019

Feathered Fallout

The NOAA National Weather Service radar images from last evening provided an indication that there may be a good fallout of birds at daybreak in the lower Susquehanna valley. The moon was bright, nearly full, and there was a gentle breeze from the north to move the nocturnal migrants along. The conditions were ideal.

Rising from daytime roosts in New York and Pennsylvania, then streaming south in moonlit skies, migrating birds are recorded as echoes on this post-sunset composite NEXRAD loop from last evening. (NOAA/National Weather Service image)

The Riparian Woodlands at Conewago Falls were alive with migrants this morning. American Robins and White-throated Sparrows were joined by new arrivals for the season: Brown Creeper (Certhia americana), Ruby-crowned Kinglets (Regulus calendula), Golden-crowned Kinglets (Regulus satrapa), Dark-eyed Junco (Junco hyemalis), and Yellow-rumped Warbler (Setophaga coronata). These are the perching birds one would expect to have comprised the overnight flight. While the individuals that will remain may not yet be among them, these are the species we will see wintering in the Mid-Atlantic states. No trip to the tropics for these hardy passerines.