It’s hard to believe, but for almost two months now, sandpipers, plovers, and terns have been filtering south through the Lower Susquehanna River Watershed on their way to the Atlantic coastline as they complete the first leg of their long autumn migration—a journey that will take some species all the way to the far reaches of the South American continent for winter.
Migrating Caspian Terns and an Osprey take a break on a lower Susquehanna valley gravel bar.
As August draws to a close, these early birds are being joined by widespread nocturnal flights of Neotropical migrants—those species, primarily songbirds, on their way to wintering grounds which lie exclusively south of the continental United States.
To catch a glimpse of these night-flying avians, your best bet may be to position yourself on the crest of a ridge or along a linear break in the forest such as a utility right-of-way where waves of warblers, vireos, flycatchers, and other Neotropical passerines sometimes feed on invertebrates after making landfall at daybreak. Pick a place where the trees are bathed in the warm light of the rising sun and be there by 7 A.M. E.D.T. The activity can be tremendous, but it usually ends between 8 and 9.
One of more than half a dozen Red-eyed Vireos seen during a daybreak fallout at Second Mountain Hawkwatch in Lebanon County earlier this week.During sunrise, a just-arrived Blackburnian Warbler checks the foliage of a ridgetop Red Maple for insects.A Least Flycatcher quietly searches the shrubby growth along the forest edge for a morning meal.A Black-and-white Warbler in the first hour of sunlight after a big nocturnal flight.A Scarlet Tanager having a look around. This individual and other migrating birds may be here for a few days before moving on.A Black-throated Green Warbler in a treetop glowing with the light of sunrise.A hungry Chestnut-sided Warbler inspects limbs, leaves, and twigs looking for nourishment after an all-night flight.
Diurnal migrants, birds that make their movements during the daylight hours, are ramping up their flights now as well. Broad-winged Hawks, Bald Eagles, and falcons are currently being tallied at hawk-counting stations throughout the northeast. Many of those lookouts are seeing Ruby-throated Hummingbirds, swallows, and other daytime migrants too.
While many diurnal migrants gather in flocks as a method of defense against predators, Broad-winged Hawks congregate as a reconnaissance measure. Not waiting around for strong autumn winds to be deflected upwards by the region’s numerous ridges, these Neotropical birds migrate early enough in the season to rely upon thermal updrafts from sun-heated surfaces to provide lift, gain altitude, and save energy during their long trip. By traveling in groups, there are collectively able to better locate and utilize rising air columns as they progress southwesterly along their route. Broad-winged Hawks travel to Central and South America for winter. Their numbers in the Lower Susquehanna River Watershed will peak during the third week of September.Many American Kestrels transit our area in August and early September, a time coinciding with flights of migratory dragonflies, insects upon which they and another falcon, the Merlin, frequently prey.Chimney Swifts are fast-flying diurnal migrants. They are beginning to congregate into larger groups in preparation for their departure. Some are already on the move.Cedar Waxwings are currently being seen in roving flocks throughout the region. Their migration is less of a point-to-point flight and more of a continuous wandering in search of berries and other wild foods.Not everyone is yet ready to go. American Goldfinches are still in the midst of their nesting cycle and won’t begin leaving until the young are on their own and colorful adult males like this one are beginning to molt into their drab winter plumage.
Autumn migration flights are an ever-changing process, with different species peaking at different times throughout the season. In these months just after the nesting season, each of these species is more numerous than at any other time of the year. And of course, the more often we as observers get out and have a look, the more of them we’ll see.
Be certain to click the “Birds” tab at the top of this page for a photo guide to the species you’re likely to see passing through the lower Susquehanna valley this fall. Nearly four months of autumn hawk migration flights lie ahead, so don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab to find a hawk-counting station near you, then stop by for a visit or two. See you there!
A large nightfall liftoff of nocturnal migrating birds is shown here on this evening’s State College, Pennsylvania, Doppler Radar base velocity loop. Returns indicating birds moving toward the radar site (the white dot in the center of the color mass) are shown in green, and those indicating birds moving away are indicated in red. This is a big flight headed generally in a south-southwest direction. (NOAA/National Weather Service Doppler Radar image)
Thoughts of October in the Lower Susquehanna River Watershed bring to mind scenes of brilliant fall foliage adorning wooded hillsides and stream courses, frosty mornings bringing an end to the growing season, and geese and other birds flying south for the winter.
The autumn migration of birds spans a period equaling nearly half the calendar year. Shorebirds and Neotropical perching birds begin moving through as early as late July, just as daylight hours begin decreasing during the weeks following their peak at summer solstice in late June. During the darkest days of the year, those surrounding winter solstice in late December, the last of the southbound migrants, including some hawks, eagles, waterfowl, and gulls, may still be on the move.
The Rough-legged Hawk (Buteo lagopus), a rodent-eating raptor of tundra, grassland, and marsh, is rare as a migrant and winter resident in the lower Susquehanna valley. It may arrive as late as January, if at all.
During October, there is a distinct change in the list of species an observer might find migrating through the lower Susquehanna valley. Reduced hours of daylight and plunges in temperatures—particularly frost and freeze events—impact the food sources available to birds. It is during October that we say goodbye to the Neotropical migrants and hello to those more hardy species that spend their winters in temperate climates like ours.
During several of the first days of October, two hundred Chimney Swifts remained in this roost until temperatures warmed from the low forties at daybreak to the upper fifties at mid-morning; then, at last, the flock ventured out in search of flying insects. When a population of birds loses its food supply or is unable to access it, that population must relocate or perish. Like other insectivorous birds, these swifts must move to warmer climes to be assured a sustained supply of the flying bugs they need to survive. Due to their specialized food source, they can be considered “specialist” feeders in comparison to species with more varied diets, the “generalists”. After returning to this chimney every evening for nearly two months, the swifts departed this roost on October 5 and did not return.A Northern Parula lingers as an October migrant along the Susquehanna. This and other specialist feeders that survive almost entirely on insects found in the forest canopy are largely south of the Susquehanna watershed by the second week of October.The Blackpoll Warbler is among the last of the insectivorous Neotropical warblers to pass through the riparian forests of the lower Susquehanna valley each fall. Through at least mid-October, it is regularly seen searching for crawling insects and larvae among the foliage and bark of Northern Hackberry (Celtis occidentalis) trees near Conewago Falls. Most other warblers, particularly those that feed largely upon flying insects, are, by then, already gone.The Blue-headed Vireo, another insectivore, is the last of the vireo species to pass through the valley. They linger only as long as there are leaves on the trees in which they feed.Brown Creepers begin arriving in early October. They are specialist feeders, well-adapted to finding insect larvae and other invertebrates among the ridges and peeling bark of trees like this hackberry, even through the winter months.Ruby-crowned Kinglets can be abundant migrants in October. They will often behave like cute little flycatchers, but quickly transition to picking insects and other invertebrates from foliage and bark as the weather turns frosty. Some may spend the winter here, particularly in the vicinity of stands of pines, which provide cover and some thermal protection during storms and bitter cold.Beginning in early October, Golden-crowned Kinglets can be seen searching the forest wood for tiny invertebrates. They are the most commonly encountered kinglet in winter.The Yellow-bellied Sapsucker, a woodpecker, is an October migrant that specializes in attracting small insects to tiny seeps of sap it creates by punching horizontal rows of shallow holes through the tree bark. Some remain for winter.The Yellow-rumped Warbler arrives in force during October. It is the most likely of the warblers to be found here in winter. Yellow-rumped Warblers are generalists, feeding upon insects during the warmer months, but able to survive on berries and other foods in late fall and winter. Wild foods like these Poison Ivy berries are crucial for the survival of this and many other generalists.American Robins are most familiar as hunters of earthworms on the suburban lawn, but they are generalist feeders that rely upon fruits like these Eastern Red Cedar (Juniperus virginiana) berries during their southbound migration in late October and early November each year. Robins remain for the winter in areas of the lower Susquehanna valley with ample berries for food and groves of mature pines for roosting.Like other brown woodland thrushes, the Hermit Thrush (Catharus guttatus) is commonly seen scratching through organic matter on the moist forest floor in search of invertebrates. Unlike the other species, it is a cold-hardy generalist feeder, often seen eating berries during the southbound October migration. Small numbers of Hermit Thrushes spend the winter in the lower Susquehanna valley, particularly in habitats with a mix of wild foods.Due to their feeding behavior, Cedar Waxwings can easily be mistaken for flycatchers during the nesting season, but by October they’ve transitioned to voracious consumers of small wild fruits. During the remainder of the year, flocks of waxwings wander widely in search of foods like this Fox Grape (Vitis labrusca). An abundance of cedar, holly, Poison Ivy, hackberry, bittersweet , hawthorn, wild grape, and other berries is essential to their survival during the colder months.Red-breasted Nuthatches have moved south in large numbers during the fall of 2020. They were particularly common in the lower Susquehanna region during mid-October. Red-breasted Nuthatches can feed on invertebrates during warm weather, but get forced south from Canada in droves when the cone crops on coniferous trees fail to provide an adequate supply of seeds for the colder fall and winter seasons. In the absence of wild foods, these generalists will visit feeding stations stocked with suet and other provisions.Purple Finches (Haemorhous purpureus) were unusually common as October migrants in 2020. They are often considered seed eaters during cold weather, but will readily consume small fruits like these berries on an invasive Mile-a-minute Weed (Persicaria perfoliata) vine. Purple Finches are quite fond of sunflower seeds at feeding stations, but often shy away if aggressive House Sparrows or House Finches are present.
The need for food and cover is critical for the survival of wildlife during the colder months. If you are a property steward, think about providing places for wildlife in the landscape. Mow less. Plant trees, particularly evergreens. Thickets are good—plant or protect fruit-bearing vines and shrubs, and allow herbaceous native plants to flower and produce seed. And if you’re putting out provisions for songbirds, keep the feeders clean. Remember, even small yards and gardens can provide a life-saving oasis for migrating and wintering birds. With a larger parcel of land, you can do even more.
GOT BERRIES? Common Winterbery (Ilex verticillata) is a native deciduous holly that looks its best in the winter, especially with snow on the ground. It’s slow-growing, and never needs pruning. Birds including bluebirds love the berries and you can plant it in wet ground, even along a stream, in a stormwater basin, or in a rain garden where your downspouts discharge. Because it’s a holly, you’ll need to plant a male and a female to get the berries. Full sun produces the best crop. Fall is a great time to plant, and many garden centers that sell holiday greenery still have winterberry shrubs for sale in November and December. Put a clump of these beauties in your landscape. Gorgeous!
A moderate breeze from the south placed a headwind into the face of migrants trying to wing their way to winter quarters. The urge to reach their destination overwhelmed any inclination a bird or insect may have had to stay put and try again another day.
Blue Jays were joined by increasing numbers of American Robins crossing the river in small groups to continue their migratory voyages. Killdeer (Charadrius vociferous) and a handful of sandpipers headed down the river route. Other migrants today included a Cooper’s Hawk (Astur cooperii), Eastern Bluebirds (Sialia sialis), and a few Common Mergansers (Mergus merganser), House Finches (Haemorhous mexicanus), and Common Grackles (Quiscalus quiscula).
The afternoon belonged to the insects. The warm wind blew scores of Monarchs toward the north as they persistently flapped on a southwest heading. Many may have actually lost ground today. Painted Lady (Vanessa cardui) and Cloudless Sulphur butterflies were observed battling their way south as well. All three of the common migrating dragonflies were seen: Common Green Darner (Anax junius), Wandering Glider (Pantala flavescens), and Black Saddlebags (Tramea lacerata).
The warm weather and summer breeze are expected to continue as the rain and wind from Hurricane Nate, today striking coastal Alabama and Mississippi, progresses toward the Susquehanna River watershed during the coming forty-eight hours.
This Great Blue Heron was joined by numerous other fishermen and a good number of sightseers in the falls today.A colorful young Northern Water Snake (Nerodia sipedon) takes advantage of the sun-heated surface of a Pothole Rock to remain nimble and active. Cooler weather will soon compel this and other reptiles to find shelter for winter hibernation.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
A fresh breeze from the north brought cooler air and a reminder that summer is gone and autumn has arrived.
Fast-moving dark clouds provided a perfect backdrop for viewing passing diurnal migrants. Bald Eagles utilized the tail wind to cruise down the Susquehanna toward Chesapeake Bay and points further south. A migrating Merlin began a chase from which a Northern Flicker narrowly escaped by finding shelter among Pothole Rocks and a few small trees. The season’s first American Black Duck (Anas rubripes), Common Loon (Gavia immer), Yellow-bellied Sapsucker (Sphyrapicus varia), and American Pipits (Anthus rubescens) moved through.
Blue Jays continued their hesitant crossings of the river at Conewago Falls. The majority completed the journey by forming groups of a dozen or more birds and following the lead of a lone American Robin, a Northern Flicker, or, odd as it appeared, a small warbler.
By far the most numerous migrants today were swallows. Thousands of Northern Rough-winged Swallows and hundreds of Tree Swallows were on the wing in search of what was suddenly a sparse flying insect supply. To get out of the brisk wind, some of the more resourceful birds landed on the warm rocks. To satisfy their appetite, many were able to pick crawling arthropods from the surface of the boulders. They swallow them whole.
A few of the thousands of swallows seen at Conewago Falls today.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
The Neotropical birds that raised their young in Canada and in the northern United States have now logged many miles on their journey to warmer climates for the coming winter. As their density decreases among the masses of migrating birds, a shift to species with a tolerance for the cooler winter weather of the temperate regions will be evident.
Though it is unusually warm for this late in September, the movement of diurnal migrants continues. This morning at Conewago Falls, five Broad-winged Hawks (Buteo platypterus) lifted from the forested hills to the east, then crossed the river to continue a excursion to the southwest which will eventually lead them and thousands of others that passed through Pennsylvania this week to wintering habitat in South America. Broad-winged Hawks often gather in large migrating groups which swarm in the rising air of thermal updrafts, then, after gaining substantial altitude, glide away to continue their trip. These ever-growing assemblages from all over eastern North America funnel into coastal Texas where they make a turn to south around the Gulf of Mexico, then continue on toward the tropics. In the coming weeks, a migration count at Corpus Christi in Texas could tally 100,000 or more Broad-winged Hawks in a single day as a large portion of the continental population passes by. You can track their movement and that of other diurnal raptors as recorded at sites located all over North America by visiting hawkcount.org on the internet. Check it out. You’ll be glad you did.
Nearly all of the other migrants seen today have a much shorter flight ahead of them. Red-bellied Woodpeckers (Melanerpes carolinus), Red-headed Woodpeckers (Melanerpes erythrocephalus), and Northern Flickers (Colaptes auratus) were on the move. Migrating American Robins (Turdus migratorius) crossed the river early in the day, possibly leftovers from an overnight flight of this primarily nocturnal migrant. The season’s first Great Black-backed Gulls (Larus marinus) arrived. American Goldfinches are easily detected by their calls as they pass overhead. Look carefully at the goldfinches visiting your feeder, the birds of summer are probably gone and are being replaced by migrants currently passing through.
By far, the most conspicuous migrant today was the Blue Jay. Hundreds were seen as they filtered out of the hardwood forests of the diabase ridge to cautiously cross the river and continue to the southwest. Groups of five to fifty birds would noisily congregate in trees along the river’s edge, then begin flying across the falls. Many wary jays abandoned their small crossing parties and turned back. Soon, they would try the trip again in a larger flock.
Sensing that they are being watched, Blue Jays are hesitant to fly across the narrow Susquehanna at Conewago Falls without first assembling into a flock. The local constabulary often penalizes those who freelance and do not move in orderly groups.
A look at this morning’s count reveals few Neotropical migrants. With the exception of the Broad-winged Hawks and warblers, the migratory species seen today will winter in a sub-tropical temperate climate, primarily in the southern United States, but often as far north as the lower Susquehanna River valley. The individual birds observed today will mostly continue to a winter home a bit further south. Those that will winter in the area of Conewago Falls will arrive in October and later.
The Great Blue Heron (Ardea herodius) can be found year-round at Conewago Falls, provided there is open water and adequate food. Migrants from breeding colonies to the north will soon supplement the local population.The Double-crested Cormorant (Phalacrocorax auritus) is a summer resident at Conewago Falls. Migration of the local population and of those from further north will soon begin. All will be gone by the time ice forms on the river. Cormorants are often seen drying their feathers in sunlight following a series of feeding dives.
The long-distance migrating insect so beloved among butterfly enthusiasts shows signs of improving numbers. Today, more than two dozen Monarchs were seen crossing the falls and slowly flapping and gliding their way to Mexico.
CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
A few nocturnal migrants flew through the moonlit night to arrive at Conewago Falls for a sunrise showing this morning. A dozen warblers were in the treetops and a Wood Thrush (Hylocichla mustelina) chattered away in the understory of the Riparian Woodlands. Three species of shorebirds were in the falls and on the Pothole Rocks: Least Sandpiper (Calidris minutilla), Lesser Yellowlegs (Tringa flavipes), and Greater Yellowlegs (Tringa melanoleuca).
A Greater Yellowlegs (right) and two Lesser Yellowlegs sandpipers dropped by for breakfast.
The diurnal migration was highlighted by a Merlin (Falco columbarius), an Osprey, and a Bald Eagle, each flying down the river. Most of the other birds in the falls seemed content to linger and feed. There’s no need to hurry folks, only trouble lurks down there in paradise at the moment.
A light to moderate flight of nocturnal migrants in the eastern United States is displayed on NOAA National Weather Service NEXRAD radar at 4:58 AM EDT. The eye of Hurricane Irma can be seen approaching the Florida Keys. (NOAA/National Weather Service image)CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
A couple of inches of rain this week caused a small increase in the flow of the river, just a burp, nothing major. This higher water coincided with some breezy days that kicked up some chop on the open waters of the Susquehanna upstream of Conewago Falls. Apparently it was just enough turbulence to uproot some aquatic plants and send them floating into the falls.
Piled against and upon the upstream side of many of the Pothole Rocks were thousands of two to three feet-long flat ribbon-like opaque green leaves of Tapegrass, also called Wild Celery, but better known as American Eelgrass (Vallisneria americana). Some leaves were still attached to a short set of clustered roots. It appears that most of the plants broke free from creeping rootstock along the edge of one of this species’ spreading masses which happened to thrive during the second half of the summer. You’ll recall that persistent high water through much of the growing season kept aquatic plants beneath a blanket of muddy current. The American Eelgrass colonies from which these specimens originated must have grown vigorously during the favorable conditions in the month of August. A few plants bore the long thread-like pistillate flower stems with a fruit cluster still intact. During the recent few weeks, there have been mats of American Eelgrass visible, the tops of their leaves floating on the shallow river surface, near the east and west shorelines of the Susquehanna where it begins its pass through the Gettysburg Basin near the Pennsylvania Turnpike bridge at Highspire. This location is a probable source of the plants found in the falls today.
Uprooted American Eelgrass floating into the Pothole Rocks under the power of a north wind. Note the white thread-like pistillate flower stem to the left and the small rooted specimen to the upper right. The latter is likely a plant from the creeping rootstock on the edge of a colony. As a native aquatic species, American Eelgrass is a critical link in the Susquehanna River and Chesapeake Bay food chain. Its decimation by pollution during the twentieth century led to migration pattern alterations and severe population losses for the Canvasback (Aythya valisineria) duck.American Eelgrass, a very small specimen, found growing in a low-lying Pothole Rock alongside the accumulations of freshly arriving material from upstream. Note that the creeping rootstock has leaves growing from at least three nodes on this plant. Eelgrass dislocations are regular occurrences which sometimes begin new colonies, like the small one seen here in this Diabase Pothole Rock Microhabitat.
The cool breeze from the north was a perfect fit for today’s migration count. Nocturnal migrants settling down for the day in the Riparian Woodlands at sunrise included more than a dozen warblers and some Gray Catbirds (Dumetella carolinensis). Diurnal migration was underway shortly thereafter.
A moderate flight of nocturnal migrants is indicated around NEXRAD sites in the northeastern states at 3:18 AM EDT. The outer rain bands of Hurricane Irma can be seen approaching the Florida Keys as the storm closes in on the peninsula. (NOAA/National Weather Service image)
Four Bald Eagles were counted as migrants this morning. Based on plumage, two were first-year eagles (Juvenile) seen up high and flying the river downstream, one was a second-year bird (Basic I) with a jagged-looking wing molt, and a third was probably a fourth year (Basic III) eagle looking much like an adult with the exception of a black terminal band on the tail. These birds were the only ones which could safely be differentiated from the seven or more Bald Eagles of varying ages found within the past few weeks to be lingering at Conewago Falls. There were as many as a dozen eagles which appeared to be moving through the falls area that may have been migrating, but the four counted were the only ones readily separable from the locals.
Red-tailed Hawks (Buteo jamaicensis) were observed riding the wind to journey not on a course following the river, but flying across it and riding the updraft on the York Haven Diabase ridge from northeast to southwest.
Bank Swallows (Riparia riparia) seem to have moved on. None were discovered among the swarms of other species today.
Ruby-throated Hummingbirds, Caspian Terns, Cedar Waxwings (Bombycilla cedrorum), and Chimney Swifts (Chaetura pelagica) were migrating today, as were Monarch butterflies.
Not migrating, but always fun to have around, all four wise guys were here today. I’m referring to the four members of the Corvid family regularly found in the Mid-Atlantic states: Blue Jay (Cyanocitta cristata), American Crow (Corvus brachyrhynchos), Fish Crow (Corvus ossifragus), and Common Raven (Corvus corax).
It looks like a big Blue Jay, but it’s not. This Belted Kingfisher (Megaceryle alcyon) takes a break after flying around the falls trying to shake a marauding Ruby-throated Hummingbird off its tail.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
SOURCES
Klots, Elsie B. 1966. The New Field Book of Freshwater Life. G. P. Putnam’s Sons. New York, NY.
We all know that birds (and many other animals) migrate. It’s a survival phenomenon which, above all, allows them to utilize their mobility to translocate to a climate which provides an advantage for obtaining food, enduring seasonal weather, and raising offspring.
In the northern hemisphere, most migratory birds fly north in the spring to latitudes with progressively greater hours of daylight to breed, nest, and provide for their young. In the southern hemisphere there are similar movements, these to the south during their spring (our autumn). The goal is the same, procreation, though the landmass offering sustenance for species other than seabirds is limited “down under”. Interestingly, there are some seabirds that breed in the southern hemisphere during our winter and spend our summer (their winter) feeding on the abundant food sources of the northern oceans.
Each autumn, migratory breeding birds leave their nesting grounds as the hours of sunlight slowly recede with each passing day. They fly to lower latitudes where the nights aren’t so long and the climate is less brutal. There, they pass their winter season.
Food supply, weather, the start/finish of the nesting cycle, and other factors motivate some birds to begin their spring and autumn journeys. But overall, the hours of daylight and the angle of the sun prompt most species to get going.
But what happens after birds begin their trips to favorable habitats? Do they follow true north and south routes? Do they fly continuously, day and night? Do they ease their way from point to point, stopping to feed along the way? Do they all migrate in flocks? Well, the tactics of migration differ widely from bird species to species, from population to population, and sometimes from individual to individual. The variables encountered when examining the dynamics of bird migration are seemingly endless, but fascinatingly so. Bird migration is well-studied, but most of its intricacies and details remain a mystery.
Consider for a moment that just 10,000 years ago, an Ice Age was coming to an end, with the southernmost edge of the most recent glaciers already withdrawn into present-day Canada from points as near as the upper Susquehanna River watershed. Back then, the birds migrating to the lower portion of the drainage basin each spring probably weren’t forest-dwelling tropical warblers, orioles, and other songbirds. The migratory birds that nested in the lower Susquehanna River valley tens of millennia ago were probably those species found nesting today in taiga and tundra much closer to the Arctic Circle. And the ancestors of most of the tropical migrants that nest here now surely spent their entire lives much closer to the Equator, finding no advantage by journeying to the frigid Susquehanna valley to nest. It’s safe to say that since those times, and probably prior to them, migration patterns have been in a state of flux.
During the intervening years since the great ice sheets, birds have been able to adapt to the shifts in their environment on a gradual basis, often using their unmatched mobility to exploit new opportunities. Migration patterns change slowly, but continuously, resulting in differences that can be substantial over time. If the natural transformations of habitat and climate have kept bird migration evolving, then man’s impact on the planet shows great potential to expedite future changes, for better or worse.
Now, let’s look at two different bird migration strategies, that of day-fliers or diurnal migrants, and that of night-fliers, the nocturnal migrants.
Diurnal migrants are the most familiar to people who notice birds on the move. The majority of these species have one thing in common, some form of defense to lessen the threat of becoming the victim of a predator while flying in daylight. Of course the vultures, hawks, and eagles fly during the day. Swallows and swifts employ speed and agility on the wing to avoid becoming prey, as do hummingbirds. Finches have an undulating flight, never flying on a horizontal plane, which makes their capture more difficult. Other songbirds seen migrating by day, Red-winged Blackbirds for example, congregate into flocks soon after breeding season to avoid being alone. Defense flocks change shape constantly as birds position themselves toward the center and away from the vulnerable fringes of the swarm. The larger the flock, the safer the individual. For a lone bird, large size can be a form of protection against all but the biggest of predators. Among the more unusual defenses is that of birds like Indigo Buntings and other tropical migrants that fly across the Gulf of Mexico each autumn (often completing a portion of the flight during the day), risking exhaustion at sea to avoid the daylight hazards, including numerous predators, found in the coastal and arid lands of south Texas. Above all, diurnal migrants capture our attention and provide a spectacle which fascinates us. Perhaps diurnal migrants attract our favor because we can just stand or sit somewhere and watch them go by. We can see, identify, and even count them. It’s fantastic.
What about a bird like the Canada Goose (Branta canadensis)? It is often seen migrating in flocks during the day (the truly migratory ones flying much higher than the local year-round resident “transplants”), but then, during the big peak movements of spring and fall, they can be heard overhead all through the night. Perhaps the Canada Goose and related waterfowl bridge the gap between day and night, introducing us to the secretive starlight and moonshine commuters, the nocturnal migrants.
The high-flying migratory Canada Goose can be seen during the daytime and heard at night when passing over the lower Susquehanna River valley. A large flight exiting from Chesapeake Bay in late February or early March often results in a 12 to 24 hour-long stream of northbound flocks.
The skies are sometimes filled with thousands of them, mostly small perching birds and waders. These strangers in the night fly inconspicuously in small groups or individually, and most can be detected when passing above us only when heard making short calls to remain in contact with their travel partners. They need not worry about predators, but instead must have a method of finding their way. Many, like the Indigo Bunting, can navigate by the stars, a capability which certainly required many generations to refine. The nocturnal migrants begin moving just after darkness falls and ascend without delay to establish a safe flight path void of obstacles (though lights and tall structures can create a deadly counter to this tactic). Often, the only clue we have that a big overnight flight has occurred is the sudden appearance of new bird species or individuals, on occasion in great numbers, in a place where we observe regularly. Just days ago, the arrival of various warbler species at Conewago Falls indicated that there was at least a small to moderate movement of these birds during previous nights.
In recent years, the availability of National Weather Service radar has brought the capability to observe nocturnal migrants into easy reach. Through the night, you can log on to your local National Oceanic and Atmospheric Administration’s National Weather Service radar page (State College for the Conewago Falls area) and watch on the map as the masses of migrating bird pass through the sweep of the radar beam. As they lift off just after nightfall, rising birds will create an echo as they enter the sweeping beam close to the radar site. Then, due to the incline of the transmitted signal and the curvature of the earth, migrants will be displayed as an expanding donut-like ring around the radar’s map location as returns from climbing birds are received from progressively higher altitudes at increasing distances from the center of the site’s coverage area. On a night with a local or regional flight, several radar locations may show signs of birds in the air. On nights with a widespread flight, an exodus of sorts, the entire eastern half of the United States may display birds around the sites. You’ll find the terrain in the east allows it to be well-covered while radars in the west are less effective due to the large mountains. At daybreak, the donut-shaped displays around each radar site location on the map contract as birds descend out of the transmitted beam and are no longer detected.
Weather systems sometimes seem to motivate some flights and stifle others. The first example seen below is a northbound spring exodus, the majority of which is probably migrants from the tropics, the Neotropical migrants, including our two dozen species of warblers. A cold front passing into the northeastern United States appears to have stifled any flight behind it, while favorable winds from the southwest are motivating a heavy concentration ahead of the front.
National Oceanic and Atmospheric Administration/National Weather Service radar image from May 5, 2010, at 11:18 PM EDT, shows rain associated with a cold front moving east from the border of Ontario, Canada, and the United States into New England and the Mid-Atlantic region. The heavy blue and green reflections surrounding the radar locations ahead of the front are nocturnal migrating birds taking advantage of favorable conditions for flight including a tail wind from the southwest. Note the lighter migration behind the advancing front. Heavy radar echoes on the gulf coast, particularly in Texas, indicate dense bird concentrations exiting the tropics to fan out into North American breeding areas. The westward progression of expanding echoes surrounding individual radar sites indicates birds rising into the radar beam at local nightfall.
The second and third examples seen below are an autumn nocturnal migration movement, probably composed of many of the same tropics-bound species which were on the way north in the previous example. Note that during autumn, the cold front seems to motivate the flight following its passage. Ahead of the front, there is a reduced and, in places, undetectable volume of birds. The two images below are separated by about 42 hours.
National Oceanic and Atmospheric Administration/National Weather Service radar image (still) from September 5, 2017, at 2:38 AM EDT, shows rain in the northeast associated with a slow-moving cold front stretching from Maine southwest to New Mexico. Heaviest nocturnal bird flights can be seen behind the advancing front where there are favorable tail winds from the north or northwest.Nearly two full days later, the slow-moving cold front from the previous image has crossed Pennsylvania. As nightfall progresses from east to west, ascending nocturnal migrants enter NEXRAD radar beams, their echoes creating expanding rings around individual sites. Concentrations of southbound birds can be seen along the gulf coast. Many will follow the Texas coastline into Mexico. The Neotropical migrants that try to cross the Gulf of Mexico this night could be in for a perilous voyage. Hurricane Katia is churning in the southern gulf and a much stronger storm, Hurricane Irma, is rolling toward the Bahamas and Florida from the southeast. Masses of birds that follow learned routes or instinct to venture offshore and cross seas under such circumstances could suffer catastrophic losses. (NOAA/National Weather Service image)
You can easily learn much more about birds (and insects and bats) on radar, including both diurnal and nocturnal migrants, by visiting the Clemson University Radar Ornithology Laboratory (CUROL) website. There you’ll find information on using the various mode settings on NEXRAD (Next-Generation Radar) to differentiate between birds, other flying animals, and inanimate airborne or grounded objects. It’s superbly done and you’ll be glad you gave it a try.
SOURCES
Clemson University Radar Ornithology Laboratory (CUROL) website: http://virtual.clemson.edu/groups/birdrad/ as accessed September 6, 2017.
Rain from the remnants of Hurricane/Tropical Storm Harvey ended just after daybreak this morning. Locally, the precipitation was mostly absorbed into the soil. There was little runoff and no flooding. The river level at Conewago Falls is presently as low as it has been all summer. Among the pools and rapids of the Pothole Rocks, numbers of migrating birds are building.
Mist and a low cloud ceiling created poor visibility while trying to see early morning birds, but they’re here. The warblers are moving south and a small wave of them was filtering through the foliage on the edge of the Riparian Woodlands. One must bend backwards to have a look, and most could not be identified due to the poor lighting in the crowns of the trees where they were zipping about. Five species of warblers and two species of vireos were discerned.
There are increasing concentrations of swallows feeding on insects over the falls. Hundreds were here today, mostly Tree Swallows (Tachycineta bicolor). Bank Swallows (Stelgidopteryx riparia) numbered in the hundreds, far below the thousands, often 10,000, which staged here for migration and peaked during the first week of September annually during the 1980s and 1990s. Their numbers have been falling steadily. Loss of nesting locations in embankments near water may be impacting the entire population. A reduction in the abundance of late-summer flying insects here on the lower Susquehanna River may be cause for them to abandon this area as a migration staging point.
Bank and Tree Swallows by the hundreds were feeding upon flying insects above the waters of Conewago Falls today. Lesser numbers of Northern Rough-winged Swallows (Stelgidopteryx serripennis) and Barn Swallows (Hirundo rustica) joined the swarm.
Clear weather in the coming nights and days may get the migrants up and flying in large numbers. For those species headed to the tropics for winter, the time to get moving has arrived.
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