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LIFE IN THE LOWER SUSQUEHANNA RIVER WATERSHED
A Natural History of Conewago Falls—The Waters of Three Mile Island

Our wildlife has been having a tough winter. The local species not only contend with cold and stormy weather, but they also need to find food and shelter in a landscape that we’ve rendered sterile of these essentials throughout much of the lower Susquehanna valley’s farmlands, suburbs, and cities.
Planting trees, shrubs, flowers, and grasses that benefit our animals can go a long way, often turning a ho-hum parcel of property into a privately owned oasis. Providing places for wildlife to feed, rest, and raise their young can help assure the survival of many of our indigenous species. With a little dedication, you can be liberated from the chore of manicuring a lawn and instead spend your time enjoying the birds, mammals, insects, and other creatures that will visit your custom-made habitat.

Fortunately for us, our local county conservation districts are again conducting springtime tree sales offering a variety of native and beneficial cultivated plants at discount prices. Listed here are links to information on how to pre-order your plants for pickup in April. Click away to check out the species each county is offering in 2025!
Cumberland County Conservation District Annual Tree Seedling Sale—
Orders due by: Friday, March 21, 2025
Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025

Dauphin County Conservation District Seedling Sale—
Orders due by: Monday, March 17, 2025
Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025

Franklin County Conservation District Tree Seedling Sale—
Orders due by: Wednesday, March 19, 2025
Pickup on: Thursday, April 24, 2025

Lancaster County Annual Tree Seedling Sale—
Orders due by: Friday, March 7, 2025
Pickup on: Friday, April 11, 2025

Lebanon County Conservation District Tree and Plant Sale—
Orders due by: Monday, March 3, 2025
Pickup on: Friday, April 18, 2025




York County Conservation District Seedling Sale—
Orders due by: March 15, 2025
Pickup on: Thursday, April 10, 2025


If you live in Adams County, Pennsylvania, you may be eligible to receive free trees and shrubs for your property from the Adams County Planting Partnership (Adams County Conservation District and the Watershed Alliance of Adams County). These trees are provided by the Chesapeake Bay Foundation’s Keystone 10-Million Trees Partnership which aims to close a seven-year project in 2025 by realizing the goal of planting 10 million trees to protect streams by stabilizing soils, taking up nutrients, reducing stormwater runoff, and providing shade. If you own property located outside of Adams County, but still within the Chesapeake Bay Watershed (which includes all of the Susquehanna, Juniata, and Potomac River drainages), you still may have an opportunity to get involved. Contact your local county conservation district office or watershed organization for information.


We hope you’re already shopping. Need help making your selections? Click on the “Trees, Shrubs, and Woody Vines” tab at the top of this page to check out Uncle Tyler Dyer’s leaf collection. He has most of the species labelled with their National Wetland Plant List Indicator Rating. You can consult these ratings to help find species suited to the soil moisture on your planting site(s). For example: if your site has sloped upland ground and/or the soils sometimes dry out in summer, select plants with a rating such as UPL or FACU. If your planting in soils that remain moist or wet, select plants with the OBL or FACW rating. Plants rated FAC are generally adaptable and can usually go either way, but may not thrive or survive under stressful conditions in extremely wet or dry soils.
NATIONAL WETLAND PLANT LIST INDICATOR RATING DEFINITIONS
Using these ratings, you might choose to plant Pin Oaks (FACW) and Swamp White Oaks (FACW) in your riparian buffer along a stream; Northern Red Oaks (FACU) and White Oaks (FACU) in the lawn or along the street, driveway, or parking area; and Chestnut Oaks (UPL) on your really dry hillside with shallow soil. Give it a try.
The deluge of rain that soaked the lower Susquehanna watershed during last week is now just a memory. Streams to the west of the river, where the flooding courtesy of the remnants of Hurricane Debby was most severe, have reached their crest and receded. Sliding away toward the Chesapeake and Atlantic is all that runoff, laden with a brew of pollutants including but not limited to: agricultural nutrients, sediment, petroleum products, sewage, lawn chemicals, tires, dog poop, and all that litter—paper, plastics, glass, Styrofoam, and more. For aquatic organisms including our freshwater fish, these floods, particularly when they occur in summer, can compound the effects of the numerous stressors that already limit their ability to live, thrive, and reproduce.

One of those preexisting stressors, high water temperature, can be either intensified or relieved by summertime precipitation. Runoff from forested or other densely vegetated ground normally has little impact on stream temperature. But segments of waterways receiving significant volumes of runoff from areas of sun-exposed impervious ground will usually see increases during at least the early stages of a rain event. Fortunately, projects implemented to address the negative impacts of stormwater flow and stream impairment can often have the additional benefit of helping to attenuate sudden rises in stream temperature.

Of the fishes inhabiting the Lower Susquehanna River Watershed’s temperate streams, the least tolerant of summer warming are the trouts and sculpins—species often described as “coldwater fishes”. Coldwater fishes require water temperatures below 70° Fahrenheit to thrive and reproduce. The optimal temperature range is 50° to 65° F. In the lower Susquehanna valley, few streams are able to sustain trouts and sculpins through the summer months—largely due to the effects of warm stormwater runoff and other forms of impairment.




Coldwater fishes are generally found in small spring-fed creeks and headwaters runs. Where stream gradient, substrate, dissolved oxygen, and other parameters are favorable, some species may be tolerant of water warmer than the optimal values. In other words, these temperature classifications are not set in stone and nobody ever explained ichthyology to a fish, so there are exceptions. The Brown Trout for example is sometimes listed as a “coldwater transition fish”, able to survive and reproduce in waters where stream quality is exceptionally good but the temperature may periodically reach the mid-seventies.


More tolerant of summer heat than the trouts, sculpins, and daces are the “coolwater fishes”—species able to feed, grow, and reproduce in streams with a temperature of less than 80° F, but higher than 60° F. Coolwater fishes thrive in creeks and rivers that hover in the 65° to 70° F range during summer.


What are the causes of modern-day reductions in coldwater and coolwater fish habitats in the lower Susquehanna River and its hundreds of miles of tributaries? To answer that, let’s take a look at the atmospheric, cosmic, and hydrologic processes that impact water temperature. Technically, these processes could be measured as heat flux—the rate of heat energy transfer per unit area per unit time, frequently expressed as watts per meter squared (W/m²). Without getting too technical, we’ll just take a look at the practical impact these processes have on stream temperatures.
HEAT FLUX PROCESSES IN A SEGMENT OF STREAM























Now that we have a basic understanding of the heat flux processes responsible for determining the water temperatures of our creeks and rivers, let’s venture a look at a few graphics from gauge stations on some of the lower Susquehanna’s tributaries equipped with appropriate United States Geological Survey monitoring devices. While the data from each of these stations is clearly noted to be provisional, it can still be used to generate comparative graphics showing basic trends in easy-to-monitor parameters like temperature and stream flow.
Each image is self-labeled and plots stream temperature in degrees Fahrenheit (bold blue) and stream discharge in cubic feet per second (thin blue).

The daily oscillations in temperature reflect the influence of several heat flux processes. During the day, solar (shortwave) radiation and convection from summer air, especially those hot south winds, are largely responsible for the daily rises of about 5° F. Longwave radiation has a round-the-clock influence—adding heat to the stream during the day and mostly shedding it at night. Atmospheric exchange including evaporative cooling may help moderate the rise in stream temperatures during the day, and certainly plays a role in bringing them back down after sunset. Along its course this summer, the West Conewago Creek absorbed enough heat to render it a warmwater fishery in the area of the gauging station. The West Conewago is a shallow, low gradient stream over almost its entire course. Its waters move very slowly, thus extending their exposure time to radiated heat flux and reducing the benefit of cooling by atmospheric exchange. Fortunately for bass, catfish, and sunfish, these temperatures are in the ideal range for warmwater fishes to feed, grow, and reproduce—generally over 80° F, and ideally in the 70° to 85° F range. Coolwater fishes though, would not find this stream segment favorable. It was consistently above the 80° F maximum and the 60° to 70° F range preferred by these species. And coldwater fishes, well, they wouldn’t be caught dead in this stream segment. Wait, scratch that—the only way they would be caught in this segment is dead. No trouts or sculpins here.

Look closely and you’ll notice that although the temperature pattern on this chart closely resembles that of the West Conewago’s, the readings average about 5 degrees cooler. This may seem surprising when one realizes that the Codorus follows a channelized path through the heart of York City and its urbanized suburbs—a heat island of significance to a stream this size. Before that it passes through numerous impoundments where its waters are exposed to the full energy of the sun. The tempering factor for the Codorus is its baseflow. Despite draining a smaller watershed than its neighbor to the north, the Codorus’s baseflow (low flow between periods of rain) was 96 cubic feet per second on August 5th, nearly twice that of the West Conewago (51.1 cubic feet per second on August 5th). Thus, the incoming heat energy was distributed over a greater mass in the Codorus and had a reduced impact on its temperature. Though the Codorus is certainly a warmwater fishery in its lower reaches, coolwater and transitional fishes could probably inhabit its tributaries in segments located closer to groundwater sources without stress. Several streams in its upper reaches are in fact classified as trout-stocked fisheries.


The Kreutz Creek gauge shows temperature patterns similar to those in the West Conewago and Codorus data sets, but notice the lower overall temperature trend and the flow. Kreutz Creek is a much smaller stream than the other two, with a flow averaging less than one tenth that of the West Conewago and about one twentieth of that in the Codorus. And most of the watershed is cropland or urban/suburban space. Yet, the stream remains below 80° F through most of the summer. The saving graces in Kreutz Creek are reduced exposure time and gradient. The waters of Kreutz Creek tumble their way through a small watershed to enter the Susquehanna within twenty-four hours, barely time to go through a single daily heating and cooling cycle. As a result, their is no chance for water to accumulate radiant and convective heat over multiple summer days. The daily oscillations in temperature are less amplified than we find in the previous streams—a swing of about three degrees compared to five. This indicates a better balance between heat flux processes that raise temperature and those that reduce it. Atmospheric exchange in the stream’s riffles, forest cover, and good hyporheic exchange along its course could all be tempering factors in Kreutz Creek. From a temperature perspective, Kreutz Creek provides suitable waters for coolwater fishes.

Muddy Creek is a trout-stocked fishery, but it cannot sustain coldwater species through the summer heat. Though temperatures in Muddy Creek may be suitable for coolwater fishes, silt, nutrients, low dissolved oxygen, and other factors could easily render it strictly a warmwater fishery, inhabited by species tolerant of significant stream impairment.

A significant number of stream segments in the Chiques watershed have been rehabilitated to eliminate intrusion by grazing livestock, cropland runoff, and other sources of impairment. Through partnerships between a local group of watershed volunteers and landowners, one tributary, Donegal Creek, has seen riparian buffers, exclusion fencing, and other water quality and habitat improvements installed along nearly ever inch of its run from Donegal Springs through high-intensity farmland to its mouth on the main stem of the Chiques just above its confluence with the Susquehanna. The improved water quality parameters in the Donegal support native coldwater sculpins and an introduced population of reproducing Brown Trout. While coldwater habitat is limited to the Donegal, the main stem of the Chiques and its largest tributary, the Little Chiques Creek, both provide suitable temperatures for coolwater fishes.





Despite its meander through and receipt of water from high-intensity farmland, the temperature of the lower Conewago (East) maxes out at about 85° F, making it ideal for warmwater fishes and even those species that are often considered coolwater transition fishes like introduced Smallmouth Bass, Rock Bass, Walleye, and native Margined Madtom. This survivable temperature is a testament to the naturally occurring and planted forest buffers along much of the stream’s course, particularly on its main stem. But the Conewago suffers serious baseflow problems compared to other streams we’ve looked at so far. Just prior to the early August storms, flow was well below 10 cubic feet per second for a drainage area of more than fifty square miles. While some of this reduced flow is the result of evaporation, much of it is anthropogenic in origin as the rate of groundwater removal continues to increase and a recent surge in stream withdraws for irrigation reaches its peak during the hottest days of summer.



A little side note—the flow rate on the Conewago at the Falmouth gauge climbed to about 160 cubic feet per second as a result of the remnants of Hurricane Debby while the gauge on the West Conewago at Manchester skyrocketed to about 20,000 cubic feet per second. Although the West Conewago’s watershed (drainage area) is larger than that of the Conewago on the east shore, it’s larger only by a multiple of two or three, not 125. That’s a dramatic difference in rainfall!

The temperatures at the Bellaire monitoring station, which is located upstream of the Conewago’s halfway point between its headwaters in Mount Gretna and its mouth, are quite comparable to those at the Falmouth gauge. Although a comparison between these two sets of data indicate a low net increase in heat absorption along the stream’s course between the two points, it also suggests sources of significant warming upstream in the areas between the Bellaire gauge and the headwaters.

The waters of the Little Conewago are protected within planted riparian buffers and mature woodland along much of their course to the confluence with the Conewago’s main stem just upstream of Bellaire. This tributary certainly isn’t responsible for raising the temperature of the creek, but is instead probably helping to cool it with what little flow it has.

Though mostly passing through natural and planted forest buffers above its confluence with the Little Conewago, the main stem’s critically low baseflow makes it particularly susceptible to heat flux processes that raise stream temperatures in segments within the two or three large agricultural properties where owners have opted not to participate in partnerships to rehabilitate the waterway. The headwaters area, while largely within Pennsylvania State Game Lands, is interspersed with growing residential communities where potable water is sourced from hundreds of private and community wells—every one of them removing groundwater and contributing to the diminishing baseflow of the creek. Some of that water is discharged into the stream after treatment at the two municipal sewer plants in the upper Conewago. This effluent can become quite warm during processing and may have significant thermal impact when the stream is at a reduced rate of flow. A sizeable headwaters lake is seasonally flooded for recreation in Mount Gretna. Such lakes can function as effective mid-day collectors of solar (shortwave) radiation that both warms the water and expedites atmospheric exchange.

Though Conewago Creek (East) is classified as a trout-stocked fishery in its upper reaches in Lebanon County, its low baseflow and susceptibility to warming render it inhospitable to these coldwater fishes by late-spring/early summer.





The removal of two water supply dams on the headwaters of Hammer Creek at Rexmont eliminated a large source of temperature fluctuation on the waterway, but did little to address the stream’s exposure to radiant and convective heat flux processes as it meanders largely unprotected out of the forest cover of Pennsylvania State Game Lands and through high-intensity farmlands in the Lebanon Valley. Moderating the temperature to a large degree is the influx of karst water from Buffalo Springs, located about two miles upstream from this gauging station, and other limestone springs that feed tributaries which enter the Hammer from the east and north. Despite the cold water, the impact of the stream’s nearly total exposure to radiative and other warming heat flux processes can readily be seen in the graphic. Though still a coldwater fishery by temperature standards, it is rather obvious that rapid heating and other forms of impairment await these waters as they continue flowing through segments with few best management practices in place for mitigating pollutants. By the time Hammer Creek passes back through the Furnace Hills and Pennsylvania State Game Lands, it is leaning toward classification as a coolwater fishery with significant accumulations of sediment and nutrients. But this creek has a lot going for it—mainly, sources of cold water. A core group of enthusiastic landowners could begin implementing the best management practices and undertaking the necessary water quality improvement projects that could turn this stream around and make it a coldwater treasure. An organized effort is currently underway to do just that. Visit Trout Unlimited’s Don Fritchey Chapter and Donegal Chapter to learn more. Better yet, join them as a volunteer or cooperating landowner!



For coldwater fishes, the thousands of years since the most recent glacial maximum have seen their range slowly contract from nearly the entirety of the once much larger Susquehanna watershed to the headwaters of only our most pristine streams. Through no fault of their own, they had the misfortune of bad timing—humans arrived and found coldwater streams and the groundwater that feeds them to their liking. Some of the later arrivals even built their houses right on top of the best-flowing springs. Today, populations of these fishes in the region we presently call the Lower Susquehanna River Watershed are seriously disconnected and the prospect for survival of these species here is not good. Stream rehabilitation, groundwater management, and better civil planning and land/water stewardship are the only way coldwater fishes, and very possibly coolwater fishes as well, will survive. For some streams like Hammer Creek, it’s not too late to make spectacular things happen. It mostly requires a cadre of citizens, local government, project specialists, and especially stakeholders to step up and be willing to remain focused upon project goals so that the many years of work required to turn a failing stream around can lead to success.




You’re probably glad this look at heat flux processes in streams has at last come to an end. That’s good, because we’ve got a lot of work to do.


The rain and clouds have at last departed. With blue skies and sunshine to remind us just how wonderful a spring afternoon can be, we took a stroll at Memorial Lake State Park in Lebanon County, Pennsylvania, to look for some migratory birds.








What? You thought we were gonna drop in on Maryland’s largest city for a couple of ball games and some oysters, clams, and crab cakes—not likely.
It’s that time of year. Your local county conservation district is taking orders for their annual tree sale and it’s a deal that can’t be beat. Order now for pickup in April.
The prices are a bargain and the selection includes the varieties you need to improve wildlife habitat and water quality on your property. For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue). And don’t forget to order packs of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local native birds. They’re only $12.00 for a bundle of 10.

Cumberland County Conservation District Annual Tree Seedling Sale—
Orders due by: Friday, March 22, 2024
Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024



Dauphin County Conservation District Seedling Sale—
Orders due by: Monday, March 18, 2024
Pickup on: Thursday, April 18, 2024 or Friday, April 19, 2024

Lancaster County Annual Tree Seedling Sale—
Orders due by: Friday, March 8, 2024
Pickup on: Friday, April 12, 2024






Lebanon County Conservation District Tree and Plant Sale—
Orders due by: Friday, March 8, 2024
Pickup on: Friday, April 19, 2024




Perry County Conservation District Tree Sale—
Orders due by: Sunday, March 24, 2024
Pickup on: Thursday, April 11, 2024

Again this year, Perry County is offering bluebird nest boxes for sale. The price?—just $12.00.

York County Conservation District Seedling Sale—
Orders due by: Friday, March 15, 2024
Pickup on: Thursday, April 11, 2024

To get your deciduous trees like gums, maples, oaks, birches, and poplars off to a safe start, conservation district tree sales in Cumberland, Dauphin, Lancaster, and Perry Counties are offering protective tree shelters. Consider purchasing these plastic tubes and supporting stakes for each of your hardwoods, especially if you have hungry deer in your neighborhood.

There you have it. Be sure to check out each tree sale’s web page to find the selections you like, then get your order placed. The deadlines will be here before you know it and you wouldn’t want to miss values like these!
I’m worried about the beaver. Here’s why.
Imagine a network of brooks and rivulets meandering through a mosaic of shrubby, sometimes boggy, marshland, purifying water and absorbing high volumes of flow during storm events. This was a typical low-gradient stream in the valleys of the Lower Susquehanna River Watershed in the days prior to the arrival of the waves of trans-Atlantic human migrants that started to inundate the area during the seventeenth century. Then, a frenzy of trapping, tree chopping, mill building, and stream channelization accompanied the east to west surge of settlement across the region. The first casualty: the indispensable lowlands manager, the North American Beaver (Castor canadensis).

Without the widespread presence of beavers, stream ecology quickly collapsed. Pristine waterways were all at once gone, as were many of their floral and faunal inhabitants. It was a streams-to-sewers saga completed in just one generation. So, if we really want to restore our creeks and rivers, maybe we need to give the North American Beaver some space and respect. After all, we as a species have yet to build an environmentally friendly dam and have yet to fully restore a wetland to its natural state. The beaver is nature’s irreplaceable silt deposition engineer and could be called the 007 of wetland construction—doomed upon discovery, it must do its work without being noticed, but nobody does it better.










Few landowners are receptive to the arrival of North American Beavers as guests or neighbors. This is indeed unfortunate. Upon discovery, beavers, like wolves, coyotes, sharks, spiders, snakes, and so many other animals, evoke an irrational negative response from the majority of people. This too is quite unfortunate, and foolish.
North American Beavers spend their lives and construct their dams, pools, and lodges exclusively within floodplains—lands that are going to flood. Their existence should create no conflict with the day to day business of human beings. But humans can’t resist encroachment into beaver territory. Because they lack any basic understanding of floodplain function, people look at these indispensable lowlands as something that must be eliminated in the name of progress. They’ll fill them with soil, stone, rock, asphalt, concrete, and all kinds of debris. You name it, they’ll dump it. It’s an ill-fated effort to eliminate these vital areas and the high waters that occasionally inundate them. Having the audacity to believe that the threat of flooding has been mitigated, buildings and poorly engineered roads and bridges are constructed in these “reclaimed lands”. Much of the Lower Susquehanna River Watershed has now been subjected to over three hundred years-worth of these “improvements” within spaces that are and will remain—floodplains. Face it folks, they’re going to flood, no matter what we do to try to stop it. And as a matter of fact, the more junk we put into them, the more we displace flood waters into areas that otherwise would not have been impacted! It’s absolute madness.
By now we should know that floodplains are going to flood. And by now we should know that the impacts of flooding are costly where poor municipal planning and negligent civil engineering have been the norm for decades and decades. So aren’t we tired of hearing the endless squawking that goes on every time we get more than an inch of rain? Imagine the difference it would make if we backed out and turned over just one quarter or, better yet, one half of the mileage along streams in the Lower Susquehanna River Watershed to North American Beavers. No more mowing, plowing, grazing, dumping, paving, spraying, or building—just leave it to the beavers. Think of the improvements they would make to floodplain function, water quality, and much-needed wildlife habitat. Could you do it? Could you overcome the typical emotional response to beavers arriving on your property and instead of issuing a death warrant, welcome them as the talented engineers they are? I’ll bet you could.

We’ve got the summertime blues for you, right here at susquehannawildlife.net…





…so don’t let the summertime blues get you down. Grab a pair of binoculars and/or a camera and go for a stroll!