Wild Senna, a Showy Background Plant for Your Wildflower Garden

Looking for a native wildflower that’s tall, showy, and a great choice for attracting wildlife, especially butterflies and bees?  Then check out Wild Senna (Senna hebecarpa).

Wild Senna in a roadside wildflower garden on Pennsylvania State Gamelands.
Wild Senna currently blooming in a roadside wildflower garden on Pennsylvania State Game Lands.

Wild Senna, also known as American Senna, is a host plant for the larvae of Cloudless Sulphur and Sleepy Orange (Eurema nicippe) butterflies.  It thrives in almost any moist, well-drained soil in habitats including open woodlands, forest edges, meadows, and gardens like yours.  Its height at flowering ranges from three to six feet.  If you prefer, this perennial wildflower can even be cultivated as a shrub-like form.  It is easily grown from seed, which is available from Ernst Conservation Seeds of Meadville, Pennsylvania, as well as numerous other vendors.  And don’t forget to give Wild Senna’s two close relatives, Partridge Pea and Maryland Senna, a try as well.  They attract the same species of butterflies and are just as easy to grow.  You’ll like ’em.

Cloudless Sulphur
Cloudless Sulphur butterflies from populations in the south colonize the Lower Susquehanna River Watershed each summer in varying numbers.
Close-up view of a Wild Senna flower cluster.
Close-up view of a Wild Senna flower cluster.  Typical of members of the pea family, the  seeds of all native Senna species develop within pods after blooming and are sought after by wildfowl (Galliformes), particularly Northern Bobwhite.
Partridge Pea in flower with seed pods.
Partridge Pea with flowers and seed pods in the susquehannawildlife.net garden.  Smaller than the other Senna species, Partridge Pea reaches a height of just two to three feet.

Some Good Reasons to Postpone Mowing Until Mid-August

Here in a series of photographs are just a handful of the reasons why the land stewards at Middle Creek Wildlife Management Area and other properties where conservation and propagation practices are employed delay the mowing of fields composed of cool-season grasses until after August 15 each year.

Eastern Meadowlark
Eastern Meadowlarks, birds of large pastures, hay lots and other meadows of cool-season grasses, build their nests and raise their young on the ground.  In the years since the early twentieth century, loss in the volume of acreage maintained in the lower Susquehanna Valley as grassland habitat types has dramatically reduced the prevalence and abundance of this and other birds with similar nesting requirements.  During the most recent fifty years, early and frequent mowing and other practices introduced as part of agriculture’s Green Revolution have all but eliminated ground-nesting grassland species from the region.
Grasshopper Sparrow
Like the meadowlarks, Grasshopper Sparrows (Ammodramus savannarum) nest on the ground in fields of cool-season grasses.  Mowing prior to the time the young leave the nest and are able to fly away can obliterate a generation of grassland birds.  Because their life span is short, widespread loss of an entire year of reproduction can quickly impact overall populations of native sparrows and other small birds.  Delayed mowing can improve numbers of Grasshopper Sparrows as well as Savannah Sparrows, Vesper Sparrows (Pooecetes gramineus), and the very rare Henslow’s Sparrow (Centronyx henslowii).
Bobolink
The Bobolink, like the meadowlark, is a member of the blackbird family (Icteridae).  It too requires grasslands free of disturbances like mowing for the duration of the nesting season which, for this particular bird, lasts until mid-August in the lower Susquehanna region.  In places lacking their specific habitat requirements, Bobolinks will seldom be detected except as flyovers during migration.
Ring-necked Pheasant
Ring-necked Pheasants were introduced to the lower Susquehanna basin, and their populations were maintained thereafter, by stocking for the purpose of hunting.  But throughout the middle twentieth century, there was a substantial population of ring-necks breeding in fields of cool-season grasses in farmlands throughout the region.  High-intensity agriculture with frequent mowing eliminated not only nesting habitat in grasslands, but winter cover in areas of early successional growth.  Populations of Ring-necked Pheasants, as well as native Northern Bobwhite, crumbled during the late 1970s and early 1980s due to these changes.  For these resident birds that don’t migrate or routinely travel great distances to find new places to live and breed, widespread habitat loss can be particularly catastrophic.  Not surprisingly, the Northern Bobwhite is no longer found in the Lower Susquehanna River Watershed and has been extirpated from all of Pennsylvania.
Blue Grosbeak
At places like Middle Creek Wildlife Management Area where a mix of grasslands, early successional growth, and even some cropland are maintained, the Blue Grosbeak has extended its range well north of the Mason-Dixon and has become a regular nesting species during recent decades.  Good habitat management does pay dividends.

Right now is a good time to visit Middle Creek Wildlife Management Area to see the effectiveness a delayed mowing schedule can have when applied to fields of cool-season grasses.  If you slowly drive, walk, or bicycle the auto tour route on the north side of the lake, you’ll pass through vast areas maintained as cool-season and warm-season grasses and early successional growth—and you’ll have a chance to see these and other grassland birds raising their young.  It’s like a trip back in time to see farmlands they way they were during the middle years of the twentieth century.

Shorebirds and More at Bombay Hook National Wildlife Refuge

Have you purchased your 2023-2024 Federal Duck Stamp?  Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points.  Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States.  So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.

2023-2024 Federal Duck Stamp. Your Federal Duck Stamp is your free pass to visit the nation's National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.

Still not convinced that a Federal Duck Stamp is worth the money?  Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge.  Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.

Northern Bobwhite
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge.  At Bombay Hook, they don’t waste your money mowing grass.  Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.
Shearness Pool at Bombay Hook N.W.R.
Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools.  Birds there can often be observed at close range.  The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools.  Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.
Great Blue Heron
A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.
Semipalmated Sandpipers
Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.
Semipalmated Sandpipers and Short-billed Dowitcher
More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.
Short-billed Dowitchers
Two more Short-billed Dowitchers on the way in.
Sandpipers, Avocets, Egrets, and Mallards
Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed.  Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge.  Mallards, American Avocets, and egrets are gathered on the shore.
Short-billed Dowitchers
More Short-billed Dowitchers arriving to feed in Raymond Pool.
Semipalmated Sandpipers
Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.
Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.
Semipalmated Sandpipers, Forster's Terns, and a Short-billed Dowitcher
A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).
Short-billed Dowitchers
Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.
Short-billed Dowitchers, American Avocets, and a Snowy Egret
Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool.  A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.
Stilt Sandpiper among Short-billed Dowitchers
Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.
American Avocets
American Avocets probe the muddy bottom of Raymond Pool.
Dunlin and Short-billed Dowitchers
Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early.  Many migrating Dunlin linger at Bombay Hook into October and even November.
Least Sandpiper
This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.
Lesser Yellowlegs
Lesser Yellowlegs at Bear Swamp Pool.
Lesser Yellowlegs
Lesser Yellowlegs at Bear Swamp Pool
Greater Yellowlegs
A Greater Yellowlegs at Bear Swamp Pool.
Caspian Tern
A Caspian Tern patrolling Raymond Pool.
Marsh Wren singing
The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.
Marsh Wren Nest
This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass.  High tide licks at the roots of the cordgrass supporting the temporary domicile.
Seaside Dragonlet
By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice).  It is our only dragonfly able to breed in saltwater.  Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.
Red-winged Blackbird
Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.
Bobolink
Look up!   A migrating Bobolink passes over the dike at Shearness Pool.
Mute Swans and Canada Geese
Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.
Trumpeter Swans
A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool.  Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river.  The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century.  An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers.  Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.
Great Egret
A Great Egret prowling Shearness Pool.
Snowy Egret
A Snowy Egret in Bear Swamp Pool.
A hen Wood Duck (second from right) escorts her young.
Wood Ducks in Bear Swamp Pool.
Black-necked Stilt and young.
A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.

As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes.  Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.

Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.
Atlantic Horseshoe Crab
The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay.  We found just this one during our visit to Bombay Hook.  Man has certainly decimated populations of this ancient crustacean during recent decades.
As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.
Atlantic Marsh Fiddler Crabs
Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).
Atlantic Marsh Fiddler Crabs
Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.
Atlantic Marsh Fiddler Crab
A male Atlantic Marsh Fiddler Crab peers from its den.
Great Egret
Herons and egrets including this Great Egret are quite fond of fiddler crabs.  As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.
Green Heron
A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.
Clapper Rail
A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.
Glossy Ibis
Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.
Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide.
Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.
Ospey
An Osprey patrols the vast tidal areas opposite Shearness Pool.

No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.

Indigo Bunting
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.
Blue Grosbeak
Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook.  This one was in shrubby growth along the dike at the north end of Shearness Pool.
Trumpet Creeper and Poison Ivy
These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.
Pileated Woodpecker in Sweet Gum
The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries.  The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware.  The seed balls are a favorite winter food of goldfinches and siskins.
Red-bellied Slider and Painted Turtle
Finis Pool has no frontage on the tidal marsh but is still worth a visit.  It lies along a spur road on the tour route and is located within a deciduous coastal plain forest.  Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.
White-tailed Deity
The White-tailed Deity is common along the road to Finis Pool.
Fowler's Toad
Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.
Turk's Cap Lily
The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well.  This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.
Wild Turkey
Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.

We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon.  And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.

Forty Years Ago in the Lower Rio Grande Valley: Day One


Back in late May of 1983, four members of the Lancaster County Bird Club—Russ Markert, Harold Morrrin, Steve Santner, and your editor—embarked on an energetic trip to find, observe, and photograph birds in the Lower Rio Grande Valley of Texas.  What follows is a daily account of that two-week-long expedition.  Notes logged by Markert some four decades ago are quoted in italics.  The images are scans of 35 mm color slide photographs taken along the way by your editor.


The trip was scheduled to coincide with breeding season in order to provide a good chance of locating a number of tropical species that happen to reach the northern limit of their range along the Lower Rio Grande.  In addition, there was the possibility of seeing many northbound end-of-season spring migrants funneling through the valley as they followed the Gulf Coast on their way to nesting grounds in eastern and central sections of the United States and Canada.

DAY ONE—May 21, 1983

The trip began at Russ Markert’s home in Lititz, Pennsylvania where I slept the night before departure.  Markert’s van camper would be both our mode of transportation and accommodation for the journey.  Upon arrival in south Texas, we would pick up Harold Morrin and Steve Santner at the Brownsville Airport.  They would join us for a week before returning by air to Pennsylvania.

“Larry slept at my house last night so we could get an early start today — camper packed and read to go.  Arose at 5:30.  We alternated driving, about every two hours.  Heavy rain through Virginia and West Virginia.  Arrived at Sweetwater KOA, Tennessee, at 5:30 P.M.  The weather turned to rain.”

Day One: Lititz, Pennsylvania, to Sweetwater, Tennessee, a distance of 656 miles.  (United States Geological Survey base image)

Northern Bobwhite were among the many birds of farm and field at the Sweetwater KOA campground.  After dinner and before the rain began, I recall going for a swim in the pool, something I craved being able to do during the sweltering evenings we would experience in south Texas during the week to follow.

A Visit to Bombay Hook National Wildlife Refuge

It’s surprising how many millions of people travel the busy coastal routes of Delaware each year to leave the traffic congestion and hectic life of the northeast corridor behind to visit congested hectic shore towns like Rehobeth Beach, Bethany Beach, and Ocean City, Maryland.  They call it a vacation, or a holiday, or a weekend, and it’s exhausting.  What’s amazing is how many of them drive right by a breathtaking national treasure located along Delaware Bay just east of the city of Dover—and never know it.  A short detour on your route will take you there.  It’s Bombay Hook National Wildlife Refuge, a quiet but spectacular place that draws few crowds of tourists, but lots of birds and other wildlife.

Bombay Hook National Wildlife Refuge is located just off Route 9, a lightly-traveled coastal road east of Dover, Delaware.  Note the Big Bluestem and other warm season grasses in the background.  Bombay Hook, like other refuges in the system, is managed for the benefit of the wildlife that relies upon it to survive.  Within recent years, most of the mowed grass and tilled ground that once occurred here has been replaced by prairie grasses or successional growth, much to the delight of Northern Bobwhite (Colinus virginianus) and other species.

Let’s join Uncle Tyler Dyer and have a look around Bombay Hook.  He’s got his duck stamp and he’s ready to go.

Uncle Ty’s current United States Fish and Wildlife Service Duck Stamp displayed on his dashboard is free admission to the tour road at Bombay Hook and other National Wildlife Refuges.
The refuge at Bombay Hook includes woodlands, grasslands, and man-made freshwater pools, but it is predominately a protectorate of thousands of acres of tidal salt marsh bordering and purifying the waters of Delaware Bay.  These marshes are renowned wintering areas for an Atlantic population of Snow Goose known as the “Greater Snow Goose” (Anser caerulescens atlanticus).  Witnessing thousands of these birds rising over the marsh and glowing in the amber light of a setting sun is an unforgettable experience.
Trails at various stops along the auto tour route lead to observation towers and other features. This boardwalk meanders into the salt marsh grasses and includes a viewing area alongside a tidal creek.  Our visit coincided with a very high tide induced by east winds and a new moon.
During high tide, an Eastern Cottontail (Sylvilagus floridanus) seeks higher ground near the boardwalk and the wooded edge of the salt marsh.
As the tide rises, fast-flying shorebirds scramble from flooded mudflats in the salt marsh on the east side of the tour road.
When high tide arrives in the salt marshes, shorebirds and waterfowl often concentrate in the man-made freshwater pools on the west side of the tour road.  Glaring afternoon sun is not the best for viewing birds located west of the road.  For ideal light conditions, time your visit for a day when high tide occurs in the morning and recedes to low tide in the afternoon.
A view looking west into Shearness Pool, largest of the freshwater impoundments at Bombay Hook.
Bombay Hook has many secretive birds hiding in its wetlands, but they can often be located by the patient observer.  Here, two Pied-billed Grebes feed in an opening among the vegetation in a freshwater pool.
One of Bombay Hook’s resident Bald Eagles patrols the wetlands.
American Avocets (Recurvirostra americana) gather by the hundreds at Bombay Hook during the fall.  A passing eagle will stir them into flight.
An American Avocet, a delicate wader with a peculiar upturned bill.
As soon as the tide begins receding, shorebirds and waterfowl like these Green-winged Teal begin dispersing into the salt marshes to feed on the exposed mudflats.
The woodlands and forested areas of the refuge host resident songbirds and can be attractive to migrating species like this Yellow-rumped Warbler.
For much of its course, the tour road at Bombay Hook is located atop the dike that creates the man-made freshwater pools on the western edge of the tidal salt marsh.  If you drive slowly and make frequent stops to look and listen, you’ll notice an abundance of birds and other wildlife living along this border between two habitats.  Here, a Swamp Sparrow has a look around.
Savannah Sparrows are common along the tour road where native grasses grow wild.
Bombay Hook is renowned for its rarities. One of the attractions during the late summer and autumn of 2021 was a group of Roseate Spoonbills (Platalea ajaja), vagrants from the southern states, seen here with Great Egrets and Snowy Egrets (Egretta thula).
Roseate Spoonbills and Great Egrets at Bombay Hook.

Remember to go the Post Office and get your duck stamp.  You’ll be supporting habitat acquisition and improvements for the wildlife we cherish.  And if you get the chance, visit a National Wildlife Refuge.  November can be a great time to go, it’s bug-free!  Just take along your warmest clothing and plan to spend the day.  You won’t regret it.