Migrating Waterfowl on Lotic Fresh Waters

Lotic vs. Lentic Freshwater Ecosystems
Some comparative attributes of unspoiled lotic vs. lentic freshwater ecosystems.  Low-gradient (slow-moving) lotic waters often create, and remain connected to, accompanying riverine wetlands (a lentic freshwater ecosystem).  These swamps, marshes, and ditches absorb, purify, and infiltrate flood waters while supporting a diverse number of plant and animal species.

We frequently perceive all waterfowl migration to be synchronized with the conspicuous movements of familiar species like Snow Geese, Tundra Swans, and Canada Geese—big flights coming south in October and November, then a return to the north in late February and March.  And we’re all quite aware of the occurrence of large gatherings of some of these migrants while they make stopovers on some of our largest lentic (still) waters—the man-made lakes and reservoirs created by damming local streams.  But did you know that there are populations of colorful waterfowl with dynamic migrations that extend throughout the winter and early spring with movements that are often continuous.  Under favorable conditions, these birds favor the lotic (flowing) waters of the river and its larger tributaries as they transit the lower Susquehanna valley.  That’s because unpolluted lotic freshwater ecosystems support a greater diversity of plants and animals than lentic waters, and therefore offer more opportunities for hungry migrating waterfowl to find food.  Let’s have a look at some of the species that visit the river during their seasonal journeys…

Ice on the Susquehanna
While the urge to head south in the autumn is largely stimulated by the shortening of the photoperiod, it is the presence of ice, particularly on glacial lakes throughout the lands to the north of the lower Susquehanna River basin, that pushes many diving ducks to finally make their way south toward the guarantee of open-water feeding areas along the coast.  This movement may occur at anytime between November and late February or, as we have seen during some of the mild winters of recent decades, it may scarcely be noticed at all.
Common Mergansers
Common Mergansers often lead they way when it comes to migratory diving ducks.  They regularly move south in conspicuous numbers by late November and December and are regularly pushing north as soon as the ice begins to melt.  During a typical year, it is not unusual for some populations of these large diving birds to remain north of us during the winter.  Then, in the days after a sudden rush of frigid polar air, an appreciable increase in ice cover will force a mid-winter movement of birds down the Susquehanna.
Heat Flux Sources Impacting Temperature in Lotic Fresh Waters
Temperatures in lotic fresh waters vary over the length of the stream or river.  They are largely determined by the collective impact of the numerous sources of heat flux depicted in this graphic.  (Environmental Protection Agency image)
Buffleheads
Buffleheads begin passing through the lower Susquehanna region in Novemeber on their way to coastal saltwater bays for the winter.  Lingering populations feed by diving in the river’s pools and riffles for benthic invertebrates including snails and insect larvae.  Lesser quantities of aquatic plant matter supplement their diet.
Common Goldeneye
Many Common Goldeneyes will remain on shallow, ice-free waters of the northern lakes and rivers sculpted by the most recent glacial event, but only until they are forced south into and through the lower Susquehanna valley by the encroachment of freezing conditions.  On the river, they are among the dozen or so species of diving ducks we see visiting or passing through during the typical late fall and early river.

 

Geomorphological Zonation of a Stream or River
During their visits to the lotic (also known as riverine) fresh waters of the Susquehanna and its largest tributaries, benthic-feeding waterfowl make short dives to take advantage of the plants, small fish, invertebrates, and other food sources inhabiting the stream bottom in the riffles and pools of the free-flowing waterway.  Substrates, listed here by size (in descending order), along with other parameters influenced by this flow regime zonation determine the variety and abundance of the forage available to migrating waterfowl and other consumers.  Ice or high water and poor visibility due to flooding can render the riffles and pools of the channel unusable for feeding.  The birds must then choose to either linger and rest without feeding or leave the lotic freshwater habitat to seek sustenance.  During a flood, this may require relocation to a nearby lentic (still) body of fresh water such as a lake or reservoir.  The presence of ice will almost invariably force the birds to fly on to the Atlantic Coastal Plain and the tidal waters of its bays and estuaries.
Hooded Mergansers
Hooded Mergansers are one of the few species of diving ducks likely to utilize flooded shoreline timber and riverine (fluvial) wetlands as refuge from high water on Susquehanna.
Pied-billed Grebe and Canvasbacks
A Pied-billed Grebe and a pair of Canvasbacks on an ice-free stretch of the lower Susquehanna in mid-winter feed and loaf in a riffle-flanked pool where a large mat of American Eelgrass, a submerged aquatic plant also known as Tapegrass or Wild Celery, grows during the summer.  The vast mosaic of riffles and pools in a river this size offers tremendous opportunities for a diverse array of aquatic species to find their niche wherein they can survive and flourish.
Bufflehead
The presence of ice forces Buffleheads and other diving ducks to gather in turbulent open water, often below a riffle or dam.  Another alternative is to continue on toward the salty bays and estuaries of the coast.  High water may push these birds into the shallows among the flooded woods to feed, but they seldom utilize heavily forested riparian wetlands as a refuge due to their need for a running start to get airborne.

 

Riffle and Pool Characteristics of High Gradient (A) and Low Gradient (B) Streams.  This graphic illustrates the change in deposition characteristics of a stream or river as its gradient decreases.  A high-gradient stream (A) has a rapid velocity, often forms falls, and tends to carry away a high volume of all but the largest of particles of potential substrates as they erode from the surrounding landscape.  On a low-gradient stream, the loss in water velocity reduces the water column’s ability to transport even the smallest of substrate particles.  Deposition of this gravel, sand, silt, and clay forms lateral bars that over time create the familiar meandering path of a naturally flowing lowland stream.  (National Park Service image)
Benthic Life in Lotic Freshwaters
Benthic substrates in lotic freshwater pools and riffles support an abundance of life forms ranging from colorful diatoms on rocks and cobble, to invertebrates including snails and insect larvae, to fishes like this young Channel Catfish.  Free of accumulations of sediment, this river bottom not only provides habitat for a healthy fishery, it facilitates the bidirectional exchange of water between the Susquehanna and its underlying aquifer.
Quillback
On the lower Susquehanna, populations of young Quillback suckers are found almost exclusively in clear, high-gradient pools.
Harlequin Duck
The Harlequin Duck winters along the rocky shores and man-made jetties of the Atlantic coast.  In summer, they nest on fast-moving, headwater streams well to our north.  Very rare on the Susquehanna, this is the first of two individuals found during March and April of 2025.  It was observed feeding in the swift waters of the high-gradient riffles and pools where the river cuts through Blue Mountain north of Harrisburg.  During previous weeks, Harlequin Ducks were being seen along the coast as far south as the mouth of the Chesapeake at Cape Charles, Virginia.  It’s very possible that some of these birds traveled north through the bay area and up the Susquehanna on their way north.

 

Fluvial Geomorphology of a Stream.  Many of the Susquehanna’s tributaries pass through each of these three erosional zones.  Along the way, they carry out the process of breaking down the mountains formed by the Allegheny orogeny, the collision of North America and Africa that created the supercontinent Pangea about 325 to 260 million years ago (during portions of both the Carboniferous and Permian periods).  Today’s main stem of the lower Susquehanna passes through a transfer zone (Zone 2), carrying eroded materials to a depositional  zone (Zone 3) located within the ancient Susquehanna canyon stretching from Havre de Grace, Maryland, to Norfolk Canyon on the edge of the continental shelf.  Within this zone, more than a 10,000-year accumulation of post-glacial sediments lies submerged by the rising waters of the Atlantic Ocean and Chesapeake Bay.  Although the present-day lower Susquehanna is largely a transfer zone, some deposition occurs along low-gradient segments of the river, particularly where its course parallels the watershed’s ridges both above and below the high-gradient rapids where its path has eroded passage through the highlands.  (National Park Service image)
Although the present-day lower Susquehanna is largely a transfer zone, some deposition occurs along low-gradient segments of the river, particularly where its course parallels the watershed’s ridges both above and below the high-gradient rapids where its path continues to erode passage through the bedrock.  On this 1908 map of the Susquehanna at Conewago Falls, alluvial terraces of gravel, sand, silt, and clay can readily been seen as pale areas nearly lacking brown contour lines along the shorelines and islands of the river.  Deposits within most of these terraces date back to the melt period following the most recent glacial event and beyond.  The delta shown at the mouth of Conewago Creek (west) includes massive volumes of material deposited by both the creek and the river.  This delta is currently known as Brunner Island, much of it developed as the site of a coal/gas-fired electric generating station.  Terrace deposits along the Susquehanna’s shorelines created extensive perched marshes and swamps (wooded marshes) fed by rains, high river water, small streams, and springs, the latter often seeping from the base of the rocky escarpments carved by the ancient Susquehanna and defining the present-day inland border of the floodplain.  We call these sites “Alluvial Terrace Wetlands”.  Few of these critical components of river morphology survive.  Those not drained for farmland were obliterated by urbanization and canal, railroad, and highway construction.  (United States Geological Survey base image: Middletown, PA, quadrangle, 1908)
Water Willow
Water Willow is a familiar emergent plant that colonizes lateral bars and other alluvial deposits in low-gradient segments of the Susquehanna.
Water Willow Roots
The fine roots of Water Willow collect sediment and absorb nutrients while creating dense cover for young fish and numerous species of invertebrates including the Virginian River Horn Snails seen here.

Prior to the nineteenth century, the low-gradient flow regime of the river both above and below the riffles at Chiques Rock (lower right on map) created prime wildlife habitat.  The natural accumulations of nutrients and substrates carried into and through the lotic waterway’s pools and riffles were cycled into an ideal growing medium for extensive mats of American Eelgrass and other aquatic plants.  This underwater forest hosted a seemingly endless abundance of invertebrates and fishes (both resident and migratory)—supporting a variety of consumer species including various populations of humans.  But soon after the mass clearing of much of the watershed’s land for farming and lumber, the mill ponds created by dams constructed on streams to power saw and grain mills became brimful with sediments eroded from the unprotected ground.  During storm events, torrents of these sediments then flowed full bore toward the Susquehanna, and began accumulating in the low-gradient segments of the river.

Legacy Sediments
Sediments left behind after the removal of mill dams are known as legacy sediments. They disconnect the stream from its historic floodplain and riverine (fluvial) wetlands, thus intensifying the impact of high water in the surrounding landscape.  As these nutrient-charged deposits wash away, they become a source of pollution in the waters of the Susquehanna and Chesapeake.
American Eelgrass
A mat of American Eelgrass growing in the flowing waters of the Susquehanna below Conewago Falls.  Eelgrass and other submerged aquatic plants provide essential habitat for a wide variety of small fish and invertebrates while also consuming nutrients deposited in the cobble, gravel, and sand substrate of the river’s pools.  Excess quantities of smaller particles of silt and clay can clog the substrate and thus inhibit the hyporheic exchange of water between the stream channel and the underlying aquifer, often diminishing the biomass and diversity of organisms inhabiting this benthic habitat.  Buried in these life-choking sediments, the river bottom becomes inhospitable to growth of submerged aquatics including eelgrass.
Susquehanna at Marietta
This low-gradient stretch of the Susquehanna at Marietta flows parallel to the Chickies Quartzite “Hellam Hills” before making a sharp right turn to punch through the ridge as a series of rapids at Chiques Rock.  Formerly a fully functional lotic ecosystem and a paradise for migrating waterfowl, this river segment is now impaired by accumulations of nutrient-laden sediments from agricultural and urban runoff.
Conewago Falls Flood Waters
Nutrient and sediment-loaded flood waters roar across the diabase boulders at the York Haven Dam and Conewago Falls, a high-gradient segment of the Susquehanna.
Flood Waters at Haldeman Riffles
They continue past Brunner Island (left power plant stacks) and through Haldeman Riffles and the Shocks Mills Railroad Bridge…
Flooding on the Marietta Broads
…into the stretch of river known as the “Marietta broads” along the base of the Chickies Quartzite “Hellam Hills”.  The low stream gradient here produces a slower current and increased deposition of sediments.
Susquehanna Flooding at Chiques Rock
As the flood surges through the riffle and pool complex at Chiques Rock, the high stream gradient maintains a velocity in the water column sufficient to keep additional sediments in suspension until they reach the low-gradient river segment just downstream at Washington Boro, site of a naturally occurring lateral bar area known as the Conejohela Flats.  These bars now lie within the man-made depositional zone known as “Lake Clarke”.  Created nearly a century ago by construction of the Safe Harbor Dam, this impoundment is accumulating astounding volumes of nutrient-loaded sediments that continue to encapsulate the flats within a stream-impairing delta.
Redheads, Canvasbacks, and a Horned Grebe
Anytime from November to April on the lower Susquehanna, a group of Redheads, Canvasbacks (3 birds to right of the middle of the picture), and a Horned Grebe (lower left) is a welcome sight in a riverine pool known to have a summertime growth of American Eelgrass.  Noted Dr. Herbert Beck in 1924 when describing the Canvasback, “Like all ducks, …, it stops to feed within the county (Lancaster) less frequently than formerly, principally because the vast beds of wild celery which existed earlier on broads of the Susquehanna, as at Marietta and Washington Borough, have now been almost entirely wiped out by sedimentation of culm.  Prior to 1875 the four or five square miles of quiet water off Marietta were often as abundantly spread with wild fowl as the Susquehanna Flats are now.  Sometimes there were as many as 500,000 ducks of various kinds on the Marietta broad at one time.”
Aythya genus ducks.
Today, seeing just dozens of Aythya genus ducks (Redheads, Canvasbacks, scaup) on the lower Susquehanna is a notable event.  If they happen to be forced down by inclement weather while migrating through, you might get lucky enough to see several hundred.
Aythya Genus Ducks
While the recovery of eelgrass/wild celery beds on the Susquehanna is trivial in scale and offers little support for numbers of waterfowl to return to historic levels, restorations on the upper Chesapeake in the vicinity of the Susquehanna Flats between Havre de Grace and Aberdeen Proving Grounds may have helped refuel a gathering of mostly Aythya genus ducks during the final days of February.  This mass of ducks, many of which were forced south from the frozen Great Lakes during the previous weeks (some by way of the ice-choked Susquehanna) were apparently making an abrupt turn to make their way back north.  Their stay was brief, but estimates by local birders put their numbers as high as one half million.  The vast majority of the concentration consisted of Aythya species: Redheads, Canvasbacks, Ring-necked Ducks, and both Lesser and Greater Scaup.  It probably included a mix of birds including both northbound migrants from further down the coast and the aforementioned refugees that had just arrived to pay a quick visit while escaping the late-season ice before turning around.
Lesser Scaup
During the past two centuries, as food supplies in the Susquehanna grew increasingly compromised for benthic feeders like these Lesser Scaup and other diving ducks, a change in distribution was necessary for survival.
Lesser Scaup in flight.
As individual species, Lesser Scaup and other waterfowl that fail to adapt to natural or man-made changes in their habitats and food supplies may see their overall global numbers falter.

Despite being located in the transfer zone, the lower Susquehanna has become a significant depositional zone along much of its length, mostly courtesy of the placement of sediment-trapping man-made dams.

Following construction of the mill dams and ponds on nearly every mile of the lower Susquehanna’s low-gradient tributary streams, enterprising parties moved on to the river.  The first significant spans were constructed using wide timber cribs filled with large rock.  They were placed to create water deep enough to allow canal boats to cross the Susquehanna at both Clark’s Ferry at the mouth of the Juniata River in Dauphin County and at Columbia/Wrightsville.  These dams also diverted water into the newly excavated canals—the Pennsylvania Eastern Division Canal (completed in 1833) which followed the river’s east shore from Clark’s Ferry to Columbia, and the Susquehanna and Tidewater Canal (completed in 1840) along the west shore from Wrightsville to Havre de Grace, Maryland.  Placement of these sediment-trapping man-made dams began a process of converting vast mileage of the lower Susquehanna from a transfer zone into a deposition zone.  In addition, layout of the canals and locks followed the contours along the base of the riverside ridges, seriously altering most of the alluvial terrace wetlands that the river had created as a feature of its floodplain during the post-glacial period.

Construction of the canal dams was just the beginning.  During the twentieth century, more massive dams would be added to the main stem of the river for hydroelectric energy production at York Haven, Safe Harbor, Holtwood, and Conowingo.  Upon their completion, the days of unassisted anadromous fish migrations were over.  On both the river and its tributaries, smaller dams including dangerous low-head dams maintain water levels for boating and recreation.  They too create current-diminishing, pseudo-lentic waters that blanket the lotic riffle and pool substrates with polluted sediments.

MAN-MADE DAMS TURN LOTIC WATERS INTO UNFLUSHED TOILETS

Sediment Deposition Behind Lower Susquehanna Dams
The construction of dams on the lower Susquehanna has converted vast mileage of the river from a lotic freshwater system into a series of man-made lentic freshwater lakes.  These areas have lost their function as a lotic transfer zone and are now a sort of dysfunctional series of depositional zones collecting vast volumes of sediment containing nutrients and other pollutants.  Within each impoundment, the reduced velocity of the river causes it to drop suspended sand first, then the finer particles of silt and clay closer to the dam.  The flow regime of riffles and pools is lost and the hyporheic zone that exchanges water between the river and the underlying aquifer is clogged.  These impaired segments of river become ripe for eutrophication: algal blooms followed by die offs that can lead to a fatal reduction of dissolved oxygen in the water column.
Gadwall
Deposits of lateral bars of sediment in low-gradient segments of the Susquehanna can create shallow water feeding habitat for puddle/dabbling ducks like these Gadwall.  Where sediment pollution is severe, benthic foods in these areas often consist mostly of invertebrates and plant matter deposited by the current, the buried substrates devoid of a functioning ecosystem and the waters subject to eutrophication.
Common Goldeneyes
Common Goldeneyes on a patch of open water on an otherwise ice-covered “Lake Clarke”, the impoundment created by Safe Harbor Dam.  While they may find this spot advantageous for loafing, the food supply over the sediment-buried substrate will be limited.
By the end of the twentieth century, accumulations of polluted sediments behind lower Susquehanna dams were nearing capacity.  There is no working plan to attenuate the massive release of these pollutants that may be triggered by a catastrophic flood.  The effort to reduce nutrient and sediment runoff remains the focus so that new loading is kept to a minimum and won’t add to the capacity problems at the dams or continue downriver to the Chesapeake at full strength when the dams are full.  Alleviating the sediment aggregation problem within the river’s impoundments is a tall order and a dilemma not easily solved.  (United States Geological Survey image)
Common Mergansers
Common Mergansers will feed where benthic substrate supports the small fish and invertebrates they prefer. They will, however, gather in extraordinary numbers on the “lakes” created by riverine dams.  Though they can only feed on what floats in with the current, hundreds or sometimes thousands of Common Mergansers will concentrate on “Conowingo Pond” during the late fall or early winter.  There, safety in numbers gives them some guarantee of protection against the multitudes of eagles that simultaneously frequent the vicinity.  Another advantage of staging on “Conowingo Pond” is its close proximity to favorable feeding areas on upper Chesapeake Bay and stretches of the Susquehanna where lotic riffles and pools offer abundant opportunities below the river’s dams.
Rusty Crayfish
Fortunately for everything else living in the benthos, Common Mergansers are big enough to devour invasive, non-native Rusty Crayfish when they find them in our lotic waterways.

TIME TO CLEAN UP OUR ACT

WHERE DOES YOUR STORMWATER GO?

 

Channelized Urban/Suburban Streams Function as Sewers. They have no attached lowlands or floodplains to absorb, purify, and infiltrate runoff from rain events.  Pollutants including litter, pet waste, lawn chemicals, tire-wear particles, hazardous fluids, and sometimes untreated human excrement flush unchecked from the municipal storm drainage system into the waterway.  Thermal shock from summer downpours washing across sun-heated pavements can kill temperature sensitive fishes and other aquatic life.  Nutrient and sediment loads from these impaired tributaries later accumulate downstream in low-gradient segments of the Susquehanna, turning the river into an open-air cesspool.  Aggressively working to implement projects that eliminate these sources of pollution are the only effective way to keep the problem from getting worse.  Making things better requires a lot more dedication and effort.  (United States Geological Survey image by Frank Ippolito)

RIPARIAN BUFFERS MAKE A DIFFERENCE…WIDER IS BETTER

Riparian Buffer
To sequester sediment and cycle nutrients (primarily nitrogen and phosphorus) contained in farm runoff, the U.S.D.A. recommends installing riparian buffers between streams and lands used for grazing and raising crops.  To protect pollinating species including bees and butterflies from pesticide drift and eroding soils rich in fertilizers, they further recommend installing a stand of wind-pollinating plants such as conifers, oaks, and birches between the field and streamside plantings.  These same conservation practices improve water quality and wildlife habitat on waterways located in residential and commercial areas as well.  (United States Department of Agriculture image, click to enlarge)

FLOODPLAIN RESTORATION

Regardless OF HOW LONG YOU’VE BEEN CONDITIONED TO THINK OTHERWISE, THIS IS A DYSFUNCTIONAL, POLLUTED CREEK—AND IT NEEDS HELP
Legacy Sediments
A channelized low-gradient stream eroding a path through deposits of legacy sediments displaces flood waters into previously unaffected areas and provides a continuing source of nutrient and sediment pollution during storm events.  These impaired waters have a diminished capacity for supporting aquatic life including fishes.
RESTORED TO ITS HISTORIC FUNCTIONS
Floodplain and Stream Restoration
On an adjacent segment of the same creek, this legacy sediment removal project restored a braided meandering channel and connected it to its newly liberated, historic floodplain.  In just their second year, the fluvial wetlands are effectively absorbing stormwater and sequestering nutrients as an attached component of the stream’s riffle and pool complex.  During our visit earlier this week, we found American Toads, Northern Leopard Frogs (Lithobates pipiens), and Northern Spring Peepers breeding here.  It’s just as Castor canadensis would have it!

“STOP HEMMING AND HAWING AROUND ALREADY”

“HEY COWBOY, HOW ‘BOUT GETTIN’ THEM FILTHY LITTLE DOGIES OUTTA DAT CRICK?”
Impaired Stream
Here’s a polluted stream in a pasture with grazing livestock.  The site is a former mill pond within which the creek eroded a channel following removal of the dam.  The animals defecate and urinate where access to water is gained at a broken down embankment of the nutrient-loaded legacy sediments deposited in the pond more than a century ago.  It’s a haphazard form of animal husbandry and a reminder that all it takes is just one stubborn jackass to foul up the whole waterway.
DIRECT SOURCES OF NITROGEN (AMMONIA) POLLUTION IN STREAMS

DID YOU KNOW that a dairy cow produces about 80 pounds of waste (excrement and urine) every day?

DID YOU KNOW that a horse produces about 50 pounds of waste (excrement and urine) every day?

DID YOU KNOW that a human produces about 3 to 4 pounds of waste (excrement and urine) every day?  The exceptions, of course, are those who continue to insist that raising farm animals in and alongside a body of water is okey-doke—a harmless practice.  These individuals tend to retain the former constituent of human waste and are thus full of it.

“ATTABOY TEX, THAT’S MORE LIKE IT!”
Stream Fencing and Livestock Crossing
Now that’s better.  Legacy sediments have been removed to reconnect the stream to its floodplain.  A livestock crossing and exclusion fencing has been installed, and a nutrient-consuming riparian buffer has been planted.  This creek segment’s pollution woes have been mitigated.  Do you have a neighbor needing this type of remedial work on their farm?  Encourage them to contact your local county conservation district office or volunteer waterways restoration organization for assistance.  Various programs provide financial assistance covering all or most of the costs of stream improvement as well as monetary incentives for helping to clean up the water.

AND FINALLY

WHEN IT COMES TO BUILDING DAMS ON LOTIC FRESH WATERS…

Sandhill Cranes in Beaver Pond

…LEAVE IT TO THE BEAVERS
North American Beavers
North American Beavers (Castor canadensis) create habitats that connect the riffle and pool regime of a low-gradient stream to a surrounding fluvial wetland that retains sediments, cycles nutrients, and provides essential habitat for hundreds of plant and animal species.  Floodplains are for flooding, and if a beaver floods an area, you can be guaranteed that it was already part of a floodplain.  You see, beavers don’t encroach upon humans, it’s humans engaging in the encroachment upon beavers.  (National Park Service image)
MAD HATTERS

DID YOU KNOW that even before the landscape was cleared for farms and a supply of timber, and before mill dams on local creeks began accumulating soil runoff from the consequently barren hillsides, all the North American Beavers, the keystone species of lower Susquehanna stream ecology, were killed and sold to make hats?  It’s no wonder things are fubar!

COMING SOON…

Horned Grebe
Horned Grebes are regular migrants and sometimes winter residents on ice-free stretches of the lower Susquehanna.  They spend their time plying the benthic substrate of the river’s clear riffles and pools for a variety of invertebrates and small fish.  Look for them moving north in coming days sporting this beautiful breeding plumage.
Common Loon
April and early May are prime time for observing Common Loons on the Susquehanna as they undertake a journey from the Atlantic surf where they spent the winter to nesting sites on northern lakes.  For this migrant in breeding plumage, clear water for sighting plenty of benthic life in the river’s riffles and pools assures a successful dive in search of energy-replenishing forage.

Eagle Time At Conowingo Dam’s Fisherman’s Park

Wintering Bald Eagles are again congregating on the lower Susquehanna River, particularly in the area of Conowingo Dam near Rising Sun, Maryland.  To catch a glimpse of the action earlier this week, we took a drive on U.S. Route 1 atop Conowingo’s impounding structure to reach Fisherman’s Park on the river’s west shore below the powerhouse.

Eagle Watching at Conowingo Dam
Scores of dedicated eagle watchers and photographers brave the raw weather to see and document the concentration of eagles that gather to feed and roost in the vicinity of Exelon Energy’s Fisherman’s Park.
Susquehanna at Conowingo Dam
The panoramic view of the Susquehanna from Fisherman’s Park offers excellent opportunities to witness Bald Eagle activity.
Immature Bald Eagles
When you arrive, it’s not unusual to hear the sounds of squabbling eagles immediately upon exiting the shelter of your vehicle.  During our visit, we sighted probably 60 to 80 individuals of various age classes among the rocks and trees along the river shorelines below the dam.
Second-year (Basic I) Bald Eagle
Soon enough, we experienced a close fly-by from this second-year Bald Eagle.
Second-year (Basic I) Bald Eagle
Another of the many second-year Bald Eagles seen on the Susquehanna at Conowingo Dam.
Hatch-year (Juvenile) Bald Eagle
We were a little bit concerned to see only one hatch-year (juvenile) Bald Eagle among the birds at Fisherman’s Park.  Perhaps the aggressive behavior of the large number of older and more experienced eagles in the area has these first-year individuals shying away.  We discerned no third-year birds either, though they may certainly have been present.
Fourth-year (Basic III) Bald Eagle
A probable fourth-year Bald Eagle shows a white head with the remains of a dark line through the eye, a trait often more conspicuous in third-year birds often known as  “osprey face” eagles.
Fifth-year (Basic IV) Bald Eagle
This probable fifth-year Bald Eagle has nearly lost the dark markings on the head and tail that differentiate immature birds from adults.  A molt during the coming year will yield adult plumage and mark the completion of this bird’s sexual maturity.
Adult Bald Eagle
An adult Bald Eagle.
Eagle Photographers at Fisherman's Park
Meanwhile, a little action gets the shutters clicking,…
Eagle Dogfight in the Incipient Stage
…a fourth-year Bald Eagle (top) is drawing the ire of an adult bird,…
Eagle Dogfight
…necessitating a reprisal for the taunting behavior.

To the delight of photographers at Conowingo, some of the eagles can be seen grabbing fish, mostly Gizzard Shad, from the tailrace area of the river below the powerhouse.  But Bald Eagles are opportunistic feeders, and their feeding habits are similar to those of numerous other birds found in the vicinity of the dam at this time of year—they’re scavengers.  Here’s a glimpse of some of the other scavengers found in the midst of this Bald Eagle realm…

Fish Crows are recognized by their nasal call.  They’ll eat almost anything they can find including garbage, fish remains, discarded bait, lunch scraps, road kill, and more.
Black Vulture
Visitors to Fisherman’s Park are warned to keep out of sight any food they may have stored in their cars.  Black Vultures are known to peel rubber away from windows as they search for something to eat, a habit they possibly learned during productive forays to landfills where the edges of rubber coverings sometimes hide a freshly dumped buffet of potential sustenance.
Turkey Vulture
During our stop at Conowingo Dam earlier this week, we saw only one Turkey Vulture, though more are certainly in the vicinity feeding on road kill and other carrion.
Ring-billed Gulls
Like eagles, Ring-billed Gulls are opportunistic feeders, seen here looking for disoriented Gizzard Shad and other fish,…
Ring-billed Gull
…then quickly changing focus to check the humans along the shoreline for discarded bait or fumbled snack foods.
Juvenile Ring-billed Gull
Even young Ring-billed Gulls learn the value of watching people for activities that provide an opportunity to scavenge food.
Double-crested Cormorants
While Ring-billed Gulls and other scavengers aren’t particularly fussy about what they eat, Double-crested Cormorants are;…
Double-crested Cormorant
…they’re targeting Gizzard Shad and other fish in the waters below the dam.  Thus, we would categorize cormorants as predators, eating mollusks and other aquatic organisms as well.
Common Mergansers
And while you’re on the lower Susquehanna, keep an eye on the sky.  Common Mergansers winter on ice-free sections of the waterway and are now arriving in the vicinity of Conowingo and elsewhere.
Bald Eagles at Heron Rookery
But if perhaps winter isn’t your thing, don’t despair.  These Bald Eagles came upon last year’s Great Blue Heron rookery on the island below the dam and it seems to be giving them some ideas.  If you think like an eagle, spring is just weeks away!

Seven Reasons to Visit Middle Creek’s Willow Point Right Now

Here are 7 reasons why you, during the coming week or so, should consider spending some time at Willow Point overlooking the lake at the Pennsylvania Game Commission’s Middle Creek Wildlife Management Area.

REASON NUMBER ONE— Wildlife is at close quarters along the trail leading from the parking lot to Willow Point…

Eastern Gray Squirrel
Eastern Gray Squirrels are common and easily seen along the edge of the woods.
Eastern Bluebirds
Eastern Bluebirds are investigating nest boxes and may presently be using them as communal roost sites during cold, windy nights.
Hermit Thrush
A Hermit Thrush was just one of the songbirds we found foraging along the edge of Willow Point Trail.
Yellow-bellied Sapsucker
Look carefully and you may see one or more species of woodpeckers in the mature trees.  We found this Yellow-bellied Sapsucker in a maple near the trail’s terminus at the Willow Point viewing area.

REASON NUMBER TWO— A variety of waterfowl species are lingering on ice-free sections of the lake surrounding Willow Point…

Tundra Swans and Canada Geese
Noisy flocks of Tundra Swans and Canada Geese on open water at Willow Point.
American Black Ducks
American Black Ducks on a fly by.
Mallards
Mallard drakes near the point.
Green-winged Teal
A small flock of Green-winged Teal feeding among the Mallards.
Northern Shovelers
A flock of Northern Shovelers has been frequenting the shallows along the south side of Willow Point for at least two weeks.
Common Mergansers
Common Mergansers diving for benthic fare.

REASON NUMBER THREE— Bald Eagles are conspicuous, easily seen and heard…

Bald Eagles
Bald Eagles on tree stumps in the lake.
Bald Eagle
A hatch-year/juvenile Bald Eagle on a glide over Willow Point.

REASON NUMBER FOUR— Northern Harriers have been making close passes over Willow Point as they patrol Middle Creek’s grasslands while hunting voles…

Northern Harrier
A female Northern Harrier over Willow Point.
Northern Harrier
A hatch-year/juvenile Northern Harrier gazing over a field of goldenrod adjacent to Willow Point.
Northern Harrier
Hatch-year/juvenile Northern Harrier patrolling a field of goldenrod and preparing to pounce.
Northern Harrier
A Northern Harrier buoyantly flying past Willow Point just prior to sunset.

REASON NUMBER FIVE— The annual observance of the White-tailed Deity holidays may be drawing to a close for the gasoline and gunpowder gang, but for the supreme ungulates, the rituals that lead to consummation of their unions are still ongoing…

White-tailed Deity
Mystical White-tailed Deities hiding in plain sight near Willow Point.
White-tailed Deity
Having so far survived the ceremonies of sacrifice practiced by worshipers clad in vibrant orange attire, these divine idols agree to a more civilized ritual, a gentlemanly duel.

White-tailed Deity

White-tailed Deity

White-tailed Deity

White-tailed Deity

White-tailed Deity

White-tailed Deity
…then it’s off to find the fair maidens.

REASON NUMBER SIX— Sandhill Cranes are still being seen from Willow Point…

Sandhill Cranes
Sandhill Cranes have been spending time on dry portions of the lake bed and in grasslands and croplands to the north.

Sandhill Cranes

Sandhill Cranes

Sandhill Cranes
These Sandhill Cranes could depart from Middle Creek’s refuge at any time, particularly if the deep freeze returns to make feeding more difficult.  You’ll want to visit soon if you want to see them.

REASON NUMBER SEVEN— The crowds that will accompany the arrival of thousands of Snow Geese in early 2025 can make visiting Willow Point a stressful experience.  Visit now to see these birds and mammals at Willow Point and you might just have the place all to yourself.  Then you can spend your time looking through the flocks of waterfowl and other birds for unusual new arrivals instead of wading through a sea of humanity.

White-crowned Sparrows
White-crowned Sparrows at Middle Creek Wildlife Management Area’s Willow Point.

Chesapeake Bay Maritime Accidents and Their Impact on Susquehanna Wildlife

Tuesday’s collision of the container ship Dali into Baltimore’s Francis Scott Key Bridge and the nearly immediate collapse of the span into the chilly waters below reminds us just how unforgiving and deadly maritime accidents can be.  Upon termination of rescue and recovery operations, salvage and cleanup will be prioritized as the next steps in the long-term process of reopening the navigable waters to ship traffic and construction of a new bridge.  Part of the effort will include monitoring for leaks of fuels and other hazardous materials from the ship, its damaged cargo containers, and vehicles and equipment that were on the bridge when it failed.

Damage to the hull of the Dali and to the cargo containers on her deck could lead to leaks of hazardous liquids or other materials into Chesapeake Bay.  (United States Army Corps of Engineers Baltimore image)

On the waters and shores of today’s Chesapeake, numerous county, state, and federal agencies, including the United States Coast Guard, monitor and inspect looking for conditions and situations that could lead to point-source or accidental discharges of petroleum products and other hazardous materials into the bay.  Many are trained, equipped, and organized for emergency response to contain and mitigate spills upon detection.  But this was not always the case.

Through much of the twentieth century, maritime spills of oil and other chemicals magnified the effects of routine discharges of hazardous materials and sanitary sewer effluent into the Chesapeake and its tributaries.  The cumulative effect of these pollutants progressively impaired fisheries and bay ecosystems leading to noticeable declines in numbers of many aquatic species.  Rather frequently, spills or discharges resulted in conspicuous fish and/or bird kills.

One of the worst spills occurred near the mouth of the Potomac River on February 2, 1976, when a barge carrying 250,000 gallons of number 6 oil sank in a storm and lost its cargo into the bay.  During a month-long cleanup, the United States Coast Guard recovered approximately 167,000 gallons of the spilled oil, the remainder dispersed into the environment.  A survey counted 8,469 “sea ducks” killed.  Of the total number, the great majority were Horned Grebes (4,347 or 51.3%) and Long-tailed Ducks (2,959 or 34.9%).  Other species included Surf Scoter (Melanitta perspicillata) (405 or 4.8%), Common Loon (195 or 2.3%), Bufflehead (166 or 2.0%), Ruddy Duck (107 or 1.3%), Common Goldeneye (78 or 0.9%), Tundra Swan (46 or 0.5%), Greater Scaup (19 or 0.2%), American Black Duck (12 or 0.2%), Common Merganser (11 or 0.1%), Canvasback (10 or 0.1%), Double-crested Cormorant (10 or 0.1%), Canada Goose (8 or 0.1%), White-winged Scoter (Melanitta deglandi) (7 or 0.1%), Redhead (5 or 0.1%), gull species (10 or 0.1%), miscellaneous ducks and herons (13 or 0.2%) and unidentified (61 or 0.7%).  During the spring migration, a majority of these birds would have made their way north and passed through the lower Susquehanna valley.  The accident certainly impacted the occurrence of the listed species during that spring in 1976, and possibly for a number of years after.

Horned Grebe during migration on the Susquehanna near Haldeman Riffles.
Of the 8,469 birds killed by the February 2, 1976, oil spill on the Chesapeake, 51.3% (4,347) were Horned Grebes.  Many of them would have migrated north through the Lower Susquehanna River Watershed during the coming spring.

The Federal Water Pollution Control Act Amendments of 1972, commonly known as the Clean Water Act, put teeth into the original FWCPCA of 1948 and began reversing the accumulation of pollutants in the bay and other bodies of water around the nation.  Additional amendments in 1977 and 1987 have strengthened protections and changed the culture of “dump-and-run” disposal and “dilution-is-the-solution” treatment of hazardous wastes.  During the late nineteen-seventies and early nineteen-eighties, emergency response teams and agencies began organizing to control and mitigate spill events.  The result has been a greater awareness and competency for handling accidental discharges of fuels and other chemicals into Chesapeake Bay and other waterways.  These improvements can help minimize the environmental impact of the Dali’s collision with the Francis Scott Key Bridge in Baltimore.

Hickory Shad
Oil spills and other pollution in the Chesapeake can impact populations of migratory fish including the anadromous Hickory Shad which are presently transiting the bay on their way to the waters of the Susquehanna below Conowingo Dam.

SOURCES

Roland, John V., Moore, Glenn E., and Bellanca, Michael A.  1977.  “The Chesapeake Bay Oil Spill—February 2, 1976: A Case History”.  International Oil Spill Conference Proceedings (1977).  1977 (1): 523-527.

Waterfowl on a Rainy Day

Though soggy, windy, and rainy, it happened to be a delightfully mild day to search for migrating waterfowl in the Lower Susquehanna River Watershed.  Here’s a look at what we found temporarily grounded by the poor flight conditions…

Northern Pintails
It’s prime time for northbound Northern Pintails.
American Wigeons and Ring-necked Ducks
This raft of distant waterfowl appears to include mostly American Wigeons and Ring-necked Ducks.
Ring-necked Duck
A much more cooperative drake Ring-necked Duck.
American Black Ducks
A couple of American Black Ducks take a break during an afternoon shower.
Northern Shovelers
A pair of Northern Shovelers in the rain.
Common Mergansers
And a pair of blissful Common Mergansers.
Common Mergansers
Uh oh!…must be something he said.

Migrating Ducks Find Emergency Refuge on the Susquehanna

As anticipated, lakes and ponds throughout the lower Susquehanna basin are beginning to freeze.  Fortunately for the waterfowl thereon, particularly diving ducks, the rain-swollen river is slowly receding and water clarity is improving to provide a suitable alternative to life on the man-made impoundments.

Ice-free Susquehanna at Harrisburg, Pennsylvania
The Susquehanna, ice-free and receding from near flood stage levels last week, can presently provide suitable habitat for diving ducks and other wintering waterfowl.
A Common Merganser on the Susquehanna in Dauphin County, Pennsylvania.
A feeding Common Merganser takes a breather between dives on the Susquehanna in Dauphin County, Pennsylvania.

The deep freeze is not only impacting ponds and lakes in the lower Susquehanna valley, but is evidently affecting the larger bodies of water to our north and northwest.  During Tuesday’s snow event, thousands of diving ducks arrived on the main stem of the river—apparently forced down by the inclement weather while en route to the Atlantic Coast from the Great Lakes and its connected waterways, which are currently beginning to freeze.

Ice-free Susquehanna
Tuesday’s snowfall not only blanketed the landscape with a coating of white,…
Bay Ducks on the Susquehanna
…it prompted thousands of migrating “bay ducks” including Canvasbacks, Redheads, scaup, and other diving species to seek open water and make a forced landing.
Scaup on the Susquehanna
Scaup were by far the most numerous of the birds in the grounded flight.  The majority appeared to be Lesser Scaup.
Scaup in flight.
Scaup in flight on the Susquehanna.
Scaup, Buffleheads, and Long-tailed Duck
A mixed raft of scaup and Buffleheads seen one day after a snowstorm-related fallout of late-season migrants.  A single Long-tailed Duck, a species formerly known as Oldsquaw, can be seen to the lower left.
Scaup, Bufflehead, and Long-tailed Duck
Scaup and a Bufflehead (center) fly past a Long-tailed Duck.  Because they winter primarily in coastal waters, both of the latter species are sometimes categorized as “sea ducks”.
Ruddy Ducks
A small flock of Ruddy Ducks.
Common Mergansers
While Common Mergansers on the Susquehanna are fish eaters (piscivores), other diving ducks observed during this fallout event are primarily benthic feeders, eating plant matter and invertebrate animals collected from the river bottom.
Common Goldeneyes
A small flock of Common Goldeneyes. They, like the Long-tailed Duck and Bufflehead, are sometimes known as a species of “sea duck”.
Scaup and Buffleheads
Another mixed raft of scaup and Buffleheads loafing on the Susquehanna.
Bald Eagle
Large numbers of waterfowl attract the attention of the river’s ever-vigilant Bald Eagles.
Adult Bald Eagle
An adult Bald Eagle patrolling the area of a fallout in search of dead, sick, or injured ducks.  In addition to the victims of naturally occurring ailments, eagles find birds and mammals wounded or killed by hunters to be particularly attractive sources of food.   They can, therefore, quite easily ingest pieces of shot.  Because eagles in the lower Susquehanna valley feed as frequently in upland habitats as they do in riverine environs, use of alternatives to toxic lead shot is prudent practice in all habitat types.
Fourth-year Bald Eagle
A Bald Eagle in the first month of its fourth calendar year.   Though not yet matured to breeding age, this bird is nevertheless smart enough to be on the lookout for vulnerable or deceased waterfowl during a post-storm fallout.
Seeing Some Waterfowl
A fallout of some migrating waterfowl seldom escapes notice by members of the gasoline and gunpowder gang…
Taking Aim at Some Waterfowl
…who find their very presence an irresistible temptation to arouse their adolescent urges…
Harrassing Some Waterfowl
…to get an adrenaline junkie’s fix.

With more snow on the way for tomorrow, you may be wondering if another fallout like this could be in the works.  The only way to find out is to get out there and have a look.  Good luck!  And be good!

Piscivorous Waterfowl Visiting Lakes and Ponds

Heavy rains and snow melt have turned the main stem of the Susquehanna and its larger tributaries into a muddy torrent.  For fish-eating (piscivorous) ducks, the poor visibility in fast-flowing turbid waters forces them to seek better places to dive for food.  With man-made lakes and ponds throughout most of the region still ice-free, waterfowl are taking to these sources of open water until the rivers and streams recede and clear.

Common Mergansers
The Common Merganser is a species of diving duck with a primary winter range that, along the Atlantic Coast, reaches its southern extreme in the lower Susquehanna and Potomac watersheds.  Recently, many have left the main stem of the muddy rivers to congregate on waters with better visibility at some of the area’s larger man-made lakes.
Common Mergansers Feeding
Common Mergansers dive to locate and capture prey, primarily small fish.  During this century, their numbers have declined along the southern edge of their winter range, possibly due to birds remaining to the north on open water, particularly on the Great Lakes.  In the lower Susquehanna valley, some of these cavity-nesting ducks can now be found year-round in areas where heavy timber again provides breeding sites in riparian forests.  After nesting, females lead their young to wander widely along our many miles of larger rivers and streams to feed.
Several Common Mergansers Intimidating a Male with a Freshly Caught Fish
The behavior of these mergansers demonstrates the stiff competition for food that can result when predators are forced away from ideal habitat and become compressed into less favorable space.  On the river, piscivores can feed on the widespread abundance of small fish including different species of minnows, shiners, darters, and more.  In man-made lakes stocked for recreational anglers with sunfish, bass, and other predators (many of them non-native), small forage species are usually nonexistent.  As a result, fish-eating birds can catch larger fish, but are successful far less often.  Seen here are several mergansers resorting to intimidation in an effort to steal a young bass away from the male bird that just surfaced with it.  While being charged by the aggressors, he must quickly swallow his oversize catch or risk losing it.

With a hard freeze on the way, the fight for life will get even more desperate in the coming weeks.  Lakes will ice over and the struggle for food will intensify.  Fortunately for mergansers and other piscivorous waterfowl, high water on the Susquehanna is expected to recede and clarify, allowing them to return to their traditional environs.  Those with the most suitable skills and adaptations to survive until spring will have a chance to breed and pass their vigor on to a new generation of these amazing birds.

An Encore of the Susquehanna Seawatch

In late March and early April, a rainy night and fog at daybreak can lead to an ideal morning for spotting migratory waterfowl and seabirds during their layover on the lower Susquehanna.  Visibility was just good enough to spot these birds at Harrisburg, Pennsylvania, most of them feeding at midriver.

Northern Shovelers are regular migrants, more often seen on ponds and in wetlands than on the river.
A pair of American Wigeons head upriver.
A Horned Grebe.
A small flock of northbound Buffleheads.
Ring-necked Ducks.
Lesser Scaup, eight of the more than 100 seen along Front Street in Harrisburg at the Pennsylvania Governor’s Residence.  Note how the white bar on the wing’s secondaries becomes diffused and dusky in the primaries.
Lesser Scaup spend the winter on bays and lakes to our south.
More Scaup, the lead bird with bright white extending through the secondaries into the primaries is possibly a Greater Scaup (Aythya marila).
Long-tailed Ducks, formerly known as Oldsquaw, are a diving duck that winters on the Great Lakes and on bays along the Atlantic Coast.  They nest on freshwater ponds and lakes in the tundra of Canada and Alaska.
A male Common Merganser.
This pair of Hooded Mergansers may be nesting in a tree cavity nearby.
The local Peregrine Falcon grabbed a passing Common Grackle…
…prompting the more than 100 Bonaparte’s Gulls in the vicinity to quickly depart and fly upstream.
Birds of Conewago Falls in the Lower Susquehanna River Watershed: Northern Flicker
It pays to keep an eye on the trees along the shoreline too.  Migrants like this Northern Flicker are beginning to come through in numbers.

Common Goldeneyes, Buffleheads, and Migrating Canada Geese

Spring migration is underway and waterfowl are on the move along the lower Susquehanna River.  Here is a sample of sightings collected during a walk across the Veteran’s Memorial Bridge at Columbia-Wrightsville this morning.

At the Veteran’s Memorial Bridge, the Susquehanna was cresting this morning after recent rains and accompanying snow melt.
An overnight breeze from the southwest and calm winds during the morning hours created ideal conditions for flocks of Canada Geese to begin migrating north from Chesapeake Bay through the lower Susquehanna valley.
These three Common Mergansers and a Common Goldeneye (right) are some of the hundreds of diving ducks presently gathered on the river in the vicinity of the Veteran’s Memorial Bridge.
Common Goldeneyes and a first-winter male Bufflehead (upper right).  All of the ducks seen today in the waters surrounding the bridge are benthic feeders, diving to the river bottom to pluck invertebrates from the substrate.
A male Common Goldeneye (Bucephala clangula).
A female Common Goldeneye.
A pair of Common Goldeneyes in flight.
A pair of Buffleheads in flight.
Common Goldeneyes repositioning for another series of feeding dives.
During the morning flight, thousands of Canada Geese were seen moving north in flocks numbering about 50 to 100 birds each.  The bird in the lower right was one of thousands of Ring-billed Gulls seen headed upriver as well.
As they pass over the lower Susquehanna region, migrating Canada Geese are typically observed flying much higher than flocks from the local resident populations, often reaching the cruising altitudes of aircraft.  Aviators are always alert for flights of resident geese around airfields.  But to prevent bird strikes during days like today when thousands of migratory geese traverse the airspace, air traffic controllers can become extra busy relaying the location and altitude of potential targets to pilots flying aircraft entering areas where birds have been reported.

This is, of course, just the beginning of the great spring migration.  Do make a point of getting out to observe the spectacle.  And remember, keep looking up—you wouldn’t want to miss anything.

Snow Geese Arriving

With plenty of open water on the main lake and no snow cover on the fields where they graze, Snow Geese have begun arriving at Middle Creek Wildlife Management Area in Lancaster/Lebanon Counties.  As long as our mild winter weather continues, more can be expected to begin moving inland from coastal areas to prepare for their spring migration and a return to arctic breeding grounds.

You probably need a break from being indoors all month, so why not get out and have a look?

Hundreds of high-flying Snow Geese descended onto the main lake at Middle Creek Wildlife Management Area this afternoon.
Snow Geese and a few Ring-necked Ducks as seen from the Willow Point observation area at Middle Creek this afternoon.
The sound of calling Tundra Swans is music to the ears on an otherwise quiet winter day.
Dozens of Common Mergansers are at Middle Creek W.M.A. right now.
Look carefully and you might see American Black Ducks among the waterfowl in the refuge’s impoundments.
Northern Shovelers have been at Middle Creek since late fall.  If they can continue to access the muddy bottom of the refuge’s lake for food, they’ll stay until the spring migration.
Check those flocks of Canada Geese carefully, sometimes you’ll find something different among them,…
…like this noticeably smaller bird, possibly either a Lesser Canada Goose (Branta canadensis parvipes) or a Richardson’s Cackling Goose (Branta hutchinsii hutchinsii), the latter being more likely.

Don’t just sit there—don your coat, grab a pair of binoculars, and get out and have a gander!

Looking Up

One can get a stiff neck looking up at the flurry of bird activity in the treetops at this time of year.  Many of the Neotropical migrants favor rich forests as daytime resting sites after flying through the night.  For others, these forests are a destination where they will nest and raise their young.

The Veery (Catharus fuscescens) is a Neotropical thrush that breeds in extensive mature forest on the dampest slopes of the Diabase ridges in the Gettysburg Basin. Their rolling flute-like songs echo through the understory as newly arrived birds establish nesting territories.
The whistled song of the Baltimore Oriole is often heard long before this colorful Neotropical is seen among the foliage of a treetop.  Some dead branches allow us a glimpse of this curious beauty.
The “Pee-a-wee……..Pee-urr” song of the Eastern Wood-Pewee (Contopus virens), a small flycatcher, is presently heard in the Riparian Woodlands at Conewago Falls.  It breeds in forested tracts throughout the lower Susquehanna valley. The vocalizations often continue through the summer, ending only when the birds depart to return to the tropics for the winter.
While constructing a nest beneath a tree canopy, an Eastern Wood-Pewee form-fits the cup where eggs will soon be laid.
The Yellow-billed Cuckoo (Coccyzus americana) nests in the treetops of Riparian Woodlands along the Susquehanna and its tributaries.  Most arrive during the second half of May for their summer stay.  It is a renowned consumer of caterpillars.
The Cedar Waxwing is a notorious wanderer.  Though not a Neotropical migrant, it is a very late nester.  Flocks may continue moving for another month before pairs settle on a place to raise young.
Of the more than twenty species of warblers which regularly migrate through the lower Susquehanna Valley, the Common Yellowthroat (Geothlypis trichas) is among those which breeds here.  It is particularly fond of streamside thickets.

For the birds that arrive earlier in spring than the Neotropical migrants, the breeding season is well underway.  The wet weather may be impacting the success of the early nests.

Northern Rough-winged Swallows arrived back in April.  At traditional nest sites, including the York Haven Dam and local creek bridges, small groups of adults were seen actively feeding and at times perching in dead treetops during recent days.  There was an absence of visits to the actual nest cavities where they should be feeding and fledging young by now.  It’s very possible that these nests failed due to the wet weather and flooding.  Another nest attempt may follow if drier conditions allow stream levels to subside and there is an increase in the mass of flying insects available for the adults to feed to their young..
A Carolina Chickadee, a resident species, is seen atop a hollow stump where it and a mate are constructing a new nest for a second brood.  Did the first brood fail?  Not sure.
Common Mergansers are an uncommon but regular nesting species of waterfowl on the lower Susquehanna River.  They nest in cavities, requiring very large trees to accommodate their needs.  It was therefore encouraging to see this pair on a forested stream in northern Lancaster County during the weekend.  However, a little while after this photograph was taken the pair flew away, indicating that they are not caring for young which by now should be out of the nest and on the move under the watchful care of the female.

So long for now, if you’ll excuse me please, I have a sore neck to tend to.

A Flock of Seagulls?

At the moment there is a heavy snow falling, not an unusual occurrence for mid-February, nevertheless, it is a change in weather.  Forty-eight hours ago we were in the midst of a steady rain and temperatures were in the sixties.  The snow and ice had melted away and a touch of spring was in the air.

Big Bluestem in the Riverine Grasslands is inundated by the rising waters of the Susquehanna at Conewago Falls.   The river ice has been dispersed by the recent mild temperatures and rains.

Anyone casually looking about while outdoors during these last several days may have noticed that birds are indeed beginning to migrate north in the lower Susquehanna valley.  Killdeer, American Robins, Eastern Bluebirds, Red-winged Blackbirds, and Common Grackles are easily seen or heard in most of the area now.

Just hours ago, between nine o’clock this morning and one o’clock this afternoon, there was a spectacular flight of birds following the river north, their spring migration well underway.  In the blue skies above Conewago Falls, a steady parade of Ring-billed Gulls was utilizing thermals and riding a tailwind from the south-southeast to cruise high overhead on a course toward their breeding range.

Ring-billed Gulls swarm in a thermal updraft above Conewago Falls to gain altitude prior to streaming off to the north and continuing their journey.
Ring-billed Gulls climbing to heights sometimes exceeding 1,000 feet before breaking off and gliding away to the north.

The swirling hoards of Ring-billed Gulls attracted other migrants to take advantage of the thermals and glide paths on the breeze.  Right among them were 44 American Herring Gulls, 3 Great Black-backed Gulls, 12 Tundra Swans (Cygnus columbianus), 10 Canada Geese, 3 Northern Pintails (Anas acuta), 6 Common Mergansers, 3 Red-tailed Hawks, a Red-shouldered Hawk, 6 Bald Eagles (non-adults), 8 Black Vultures, and 5 Turkey Vultures.

A first-winter American Herring Gull (top center) is a standout in a “kettle” of Ring-billed Gulls.
How many Ring-billed Gulls passed by today?  More than 18,000…with emphasis on MORE THAN.  You see, early this afternoon, the handy-dandy clicker-counter used to tick off and tally the big flights of birds as they pass by quit clicking and counting.  Therefore, 18,000 is the absolute minimum number of Ring-billed Gulls seen migrating north today.  Hopefully the trusty old oil can will get the clicker working again soon.

In the afternoon, the clouds closed in quickly, the flight ended, and by dusk more than an inch of snow was on the ground.  Looks like spring to me.

Culinary Reminder

It was a crisp clear morning with birdless blue skies.  The migration has mostly drawn to a close; very little was seen despite a suitable northwest breeze to support a flight.   There were no robins and no blackbirds.  Not even a starling was seen today.  The only highlights were a Bufflehead (Bucephala albeola) and a couple of Swamp Sparrows.

A Swamp Sparrow is coaxed from the dense leafy cover of the Riverine Grasslands of Conewago Falls.
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And now ladies and gentlemen, boys and girls, it’s time for a Thanksgiving Day culinary reminder from the local Conewago Falls Turkey…

“That’s the Conewago Falls Turkey Vulture if you don’t mind.  And I’m here to remind you that this Thanksgiving, you don’t have to fight your way in there to get first pickings of the feast.  Take it from me, let those flavors and aromas intensify over time.  And juices, who needs juices?  Let that stuff congeal so everything sticks to your ribs.  Are you paying attention?  I’m telling you, just back off, let the crowd die down, then get in there and devour those leftovers.  They’re the best.  You’ll thank me later!”

Feathered Fallout

The NOAA National Weather Service radar images from last evening provided an indication that there may be a good fallout of birds at daybreak in the lower Susquehanna valley.  The moon was bright, nearly full, and there was a gentle breeze from the north to move the nocturnal migrants along.  The conditions were ideal.

Rising from daytime roosts in New York and Pennsylvania, then streaming south in moonlit skies, migrating birds are recorded as echoes on this post-sunset composite NEXRAD loop from last evening.  (NOAA/National Weather Service image)

The Riparian Woodlands at Conewago Falls were alive with migrants this morning.  American Robins and White-throated Sparrows were joined by new arrivals for the season: Brown Creeper (Certhia americana), Ruby-crowned Kinglets (Regulus calendula), Golden-crowned Kinglets (Regulus satrapa), Dark-eyed Junco (Junco hyemalis), and Yellow-rumped Warbler (Setophaga coronata).  These are the perching birds one would expect to have comprised the overnight flight.  While the individuals that will remain may not yet be among them, these are the species we will see wintering in the Mid-Atlantic states.  No trip to the tropics for these hardy passerines.

American Robins continued migratory flight into the first hour of daylight this morning.  Their calls are commonly heard at night as migrating individuals pass overhead.
White-throated Sparrows are nocturnal migrants, and are a familiar find on woodland edges and at suburban feeding stations through the winter.
Dark-eyed Juncos, also nocturnal migrants, are common winter residents in the Lower Susquehanna River Watershed, frequently visiting bird feeders.
Heavy rain earlier this week in the Susquehanna River drainage basin has flooded most of the Pothole Rocks; the rapids of Conewago Falls have returned.
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A Quick Getaway

It was a placid morning on Conewago Falls with blue skies dotted every now and then by a small flock of migrating robins or blackbirds.  The jumbled notes of a singing Winter Wren (Troglodytes hiemalis) in the Riparian Woodland softly mixed with the sounds of water spilling over the dam.  The season’s first Wood Ducks (Aix sponsa), Blue-winged Teal (Spatula discors), American Herring Gull (Larus smithsonianus), Horned Larks (Eremophila alpestris), and White-throated Sparrows (Zonotrichia albicollis) were seen.

There was a small ruckus when one of the adult Bald Eagles from a local pair spotted an Osprey passing through carrying a fish.  This eagle’s effort to steal the Osprey’s catch was soon interrupted when an adult eagle from a second pair that has been lingering in the area joined the pursuit.  Two eagles are certainly better than one when it’s time to hustle a skinny little Osprey, don’t you think?

But you see, this just won’t do.  It’s a breach of eagle etiquette, don’t you know?  Soon both pairs of adult eagles were engaged in a noisy dogfight.  It was fussing and cackling and the four eagles going in every direction overhead.  Things calmed down after about five minutes, then a staring match commenced on the crest of the dam with the two pairs of eagles, the “home team” and the “visiting team”, perched about 100 feet from each other.  Soon the pair which seems to be visiting gave up and moved out of the falls for the remainder of the day.  The Osprey, in the meantime, was able to slip away.

In recent weeks, the “home team” pair of Bald Eagles, seen regularly defending territory at Conewago Falls, has been hanging sticks and branched tree limbs on the cross members of the power line tower where they often perch.  They seem only to collect and display these would-be nest materials when the “visiting team” pair is perched in the nearby tower just several hundred yards away…an attempt to intimidate by homesteading.  It appears that with winter and breeding time approaching, territorial behavior is on the increase.

The second migrating Osprey of the day ran the gauntlet of marauding eagles without incident.

In the afternoon, a fresh breeze from the south sent ripples across the waters among the Pothole Rocks.  The updraft on the south face of the diabase ridge on the east shore was like a highway for some migrating hawks, falcons, and vultures.  Black Vultures (Coragyps atratus) and Turkey Vultures streamed off to the south headlong into the wind after leaving the ridge and crossing the river.  A male and female Northern Harrier (Circus hudsonius), ten Red-tailed Hawks, two Red-shouldered Hawks (Buteo lineatus), six Sharp-shinned Hawks, and two Merlins crossed the river and continued along the diabase ridge on the west shore, accessing a strong updraft along its slope to propel their journey further to the southwest.  Four high-flying Bald Eagles migrated through, each following the east river shore downstream and making little use of the ridge except to gain a little altitude while passing by.

(Top and Middle) Turkey Vultures riding the fresh breeze and teetering to-and-fro on up-tilted wings.  This wing posture is known as a dihedral.  (Bottom) More than 100 migrating Black Vultures climbed high on the afternoon breeze to make an oblique crossing of the river and maintain a southbound course.

Late in the afternoon, the local Bald Eagles were again airborne and cackling up a storm.  This time they intercepted an eagle coming down the ridge toward the river and immediately forced the bird to climb if it intended to pass.  It turned out to be the best sighting of the day, and these “home team” eagles found it first.  It was a Golden Eagle (Aquila chrysaetos) in crisp juvenile plumage.  On its first southward voyage, it seemed to linger after climbing high enough for the Bald Eagles to loose concern, then finally selected the ridge route and crossed the river to head off to the southwest.

Ring-billed Gulls began feeding during the afternoon as clouds preceding stormy weather approached.
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State of Confusion

The humid rainy remains of Hurricane Nate have long since passed by Pennsylvania, yet mild wet weather lingers to confuse one’s sense of the seasons.  This gloomy misty day was less than spectacular for watching migrating birds and insects, but some did pass by.  Many resident animals of the falls are availing themselves of the opportunity to continue active behavior before the cold winds of autumn and winter force a change of lifestyle.

Warm drizzle at daybreak prompted several Northern Spring Peepers (Pseudacris crucifer crucifer) to begin calling from the wetlands in the Riparian Woodlands of Conewago Falls.  An enormous chorus of these calls normally begins with the first warm rains of early spring to usher in this tiny frog’s mating season.  Today, it was just a few “peeps” among anxious friends.

The tiny Northern Spring Peeper is recognized by the dark “X” across its back.  Soon, shelter must be found among loose bark and fallen logs to commence hibernation.  Emergence, often prompted by warm spring rains, will quickly be followed by a growing chorus of breeding calls as sometimes hundreds of these frogs assemble in vernal pools where mating will then occur.

Any additional river flow that resulted from the rains of the previous week is scarcely noticeable among the Pothole Rocks.  The water level remains low, the water column is fairly clear, and the water temperatures are in the 60s Fahrenheit.

It’s no real surprise then to see aquatic turtles climbing onto the boulders in the falls to enjoy a little warmth, if not from the sun, then from the stored heat in the rocks.  As usual, they’re quick to slide into the depths soon after sensing someone approaching or moving nearby.  Seldom found anywhere but on the river, these skilled divers are Common Map Turtles (Graptemys geographica), also known as Northern Map Turtles.  Their paddle-like feet are well adapted to swimming in strong current.  They are benthic feeders, feasting upon a wide variety of invertebrates found among the stone and substrate of the river bottom.

Adult Common Map Turtles hibernate communally on the river bottom in a location protected from ice scour and turbulent flow, often using boulders, logs, or other structures as shelter from strong current.  The oxygenation of waters tumbling through Conewago Falls may be critical to the survival of the turtles overwintering downstream.  Dissolved oxygen in the water is absorbed by the nearly inactive turtles as they remain submerged at their hideout through the winter.  Though Common Map Turtles, particularly males, may occasionally move about in their hibernation location, they are not seen coming to the surface to breathe.

The Common Map Turtles in the Susquehanna River basin are a population disconnected from that found in the main range of the species in the Great Lakes and upper Mississippi basin.  Another isolated population exists in the Delaware River.

Common Map Turtles, including this recently hatched young seen in August, are often observed climbing onto rocks in the river.
Note the oversize swimming fin adaptations of the feet on this adult Common Map Turtle found among the Pothole Rocks in Conewago Falls.  Young and adults are capable of navigating some strong current to feed and escape danger.
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SOURCES

Committee on the Status of Endangered Wildlife in Canada.  2002.  Status Report of the Northern Map Turtle.  Canadian Wildlife Service.  Ottawa, Ontario.

Summer Breeze

A moderate breeze from the south placed a headwind into the face of migrants trying to wing their way to winter quarters.  The urge to reach their destination overwhelmed any inclination a bird or insect may have had to stay put and try again another day.

Blue Jays were joined by increasing numbers of American Robins crossing the river in small groups to continue their migratory voyages.  Killdeer (Charadrius vociferous) and a handful of sandpipers headed down the river route.  Other migrants today included a Cooper’s Hawk (Astur cooperii), Eastern Bluebirds (Sialia sialis), and a few Common Mergansers (Mergus merganser), House Finches (Haemorhous mexicanus), and Common Grackles (Quiscalus quiscula).

The afternoon belonged to the insects.  The warm wind blew scores of Monarchs toward the north as they persistently flapped on a southwest heading.  Many may have actually lost ground today.  Painted Lady (Vanessa cardui) and Cloudless Sulphur butterflies were observed battling their way south as well.  All three of the common migrating dragonflies were seen: Common Green Darner (Anax junius), Wandering Glider (Pantala flavescens), and Black Saddlebags (Tramea lacerata).

The warm weather and summer breeze are expected to continue as the rain and wind from Hurricane Nate, today striking coastal Alabama and Mississippi, progresses toward the Susquehanna River watershed during the coming forty-eight hours.

This Great Blue Heron was joined by numerous other fishermen and a good number of sightseers in the falls today.
A colorful young Northern Water Snake (Nerodia sipedon) takes advantage of the sun-heated surface of a Pothole Rock to remain nimble and active.  Cooler weather will soon compel this and other reptiles to find shelter for winter hibernation.
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