The Nighthawks and the Hummers

Chilly nights and shorter days have triggered the autumn migration of Neotropical birds.  You may not have to go far to see these two travelers.  Each is a species you may be able to find migrating through your neighborhood.

Common Nighthawks
Common Nighthawks are large insect-eating nightjars.  Watch for them feeding and migrating overhead during the late hours of the afternoon and continuing through nightfall.  While skies above the Susquehanna and large tracts of forest or grassland offer the best viewing opportunities, even city residents may witness their evening flights during the coming weeks.  Though nighthawk numbers appear to be in decline, as many as a hundred or more are currently being seen nightly at Pine Grove Furnace State Park along South Mountain in Cumberland County, Pennsylvania.  For daytime roosting during their southbound movements, nighthawks seem to be attracted to deep shade in areas of vast forest.  They will often seek sanctuary and become concentrated in “islands of darkness” like Michaux State Forest on South Mountain after passing over light-polluted urban areas such as Harrisburg and the adjoining metroplex of the Great Valley during the previous night.
Male Ruby-throated Hummingbird
Ruby-throated Hummingbirds are now moving south.  It’s time to be extra vigilant about keeping your sugar-water dispensers clean and filled with fresh nectar mixture.  For tips on feeding hummingbirds safely, check out our post from August 5, 2022, “Two Feeders Are Better Than One”.

TOMEX Sounding Rocket Launches

Beginning this evening at about 10:44 PM EDT, and lasting until almost 11 o’clock, the gaseous clouds from two of three TOMEX+ (Turbulent Oxygen Mixing Experiment) sounding rockets launched from NASA’s Wallops Island Flight Facility near Chincoteague, Virginia, were visible in the southern skies of much of the Lower Susquehanna River Watershed.  From susquehannawildlife.net headquarters, we were able to see and photograph the glowing clouds created by these vapor releases.  Within minutes, the contrail-like wisps were swept away by the swift thin air of the mesosphere, the area lying just below the thermosphere and the Kármán Line—the border of outer space 100 kilometers (62 miles) above sea level.  According to NASA, “This mission aims to provide the clearest 3D view yet of turbulence in the region at the edge of space.

TOMEX+ Gas Cloud Dispersing
A gas cloud released in the mesosphere by the first of three rockets launched in quick succession from Wallops Island, Virginia, on the Delmarva Peninsula.  Being the uppermost layer of the atmosphere, the mesosphere functions as an energy conduit into space and can thus be very turbulent.
TOMEX+ Gas Cloud Dispersing
A gaseous cloud created by a vapor tracer release from the second sounding rocket as seen from susquehannawildlife.net headquarters.
TOMEX+ Gas Clouds Dispersing
Faint tracer clouds at the center of the image and in the upper right disperse in mesospheric air currents after release from two sounding rockets.  The mesosphere’s thin air is responsible for creating enough friction to burn up a majority of the meteors that enter the earth’s atmosphere.  A byproduct of the destruction of these meteors is atomic sodium.  As part of the study, the third rocket used a laser to illuminate and excite this sodium in its area of greatest concentration, about 56 miles above sea level.

Migratory Shorebirds on the Freshwater Impoundments at Bombay Hook National Wildlife Refuge

It may seem hard to believe, but the autumn migration of shorebirds and many Neotropical songbirds is now well underway.  To see the former in what we hope will be large numbers in good light, we timed a visit to the man-made freshwater impoundments at Bombay Hook National Wildlife Refuge near Smyrna, Delaware, to coincide with a high-tide during the mid-morning hours.  Come along for a closer look…

Bombay Hook NWR tour road at Shearness Pool
Along a large portion of its route, the tour road at Bombay Hook N.W.R. sits atop the man-made dikes that create several sizeable freshwater pools (left) along the inland border of one of the largest remaining salt marsh estuaries in the Mid-Atlantic States (right).
Tidal Creek at High Tide
Twice daily, the rising tide from Delaware Bay flows along the tidal creeks to flood Bombay Hook’s extensive marshes.
High Tide in the Bombay Hook coastal estuary.
At high tide, mudflats in Bombay Hook’s coastal estuary become inundated by salt water, forcing migratory shorebirds to relocate to bay side beaches or other higher ground to rest and feed for several hours.
Shearness Pool
At Bombay Hook, migratory shorebirds, waterfowl, and waders can find refuge from high tide in the freshwater impoundments created by capturing water along the inland west side of a system of earthen dikes.
Water Level Control System
Mechanical or stacked-board gate systems are used to control water depth in the impoundments.  Levels can be adjusted seasonally to manage plant growth and create conditions favorable for use by specific groups of birds and other wildlife.
Shearness Pool
A map at Shearness Pool, the largest impoundment on the refuge, shows the location of other freshwater pools at Bombay Hook.
Shorebirds on Raymod Pool
A mix of mudflats and shallow water on Raymond Pool provides ideal habitat for a variety of shorebirds forced from the vast tidal marshes by the rising tide.  For us, a mid-morning high tide places these birds in perfect light as they feed and loaf in the pools to the west of the tour road located atop the dikes.
Shorebirds arrive on Raymond Pool during high tide.
Migrating shorebirds arrive on Raymond Pool to find refuge from the rising tide to the east.  Showing a single ring around their breast, many Semipalmated Plovers can be seen here among the Semipalmated Sandpipers, the latter the most abundant shorebird presently populating Bombay Hook.  Feeding in deeper water in the background are Short-billed Dowitchers.  All of these birds consume a variety of invertebrates they find both in and on the mud.
Semipalmated Plovers and Semipalmated Sandpipers
Semipalmated Plovers and Semipalmated Sandpipers arriving on a mudflat at Raymond Pool.
Short-billed Dowitcher
A Short-billed Dowitcher glides in from the salt marsh to visit the shallows of Raymond Pool for a couple of hours.
Short-billed Dowitcher and other Shorebirds
A Short-billed Dowitcher feeding among Semipalmated Plovers and Semipalmated Sandpipers.
Short-billed Dowitchers
Short-billed Dowitchers in water almost too deep for the Semipalmated Sandpipers in their company.
Black-bellied Plover and Short-billed Dowitchers
A lone Black-bellied Plover (Pluvialis squatarola) among the abundance of Short-billed Dowitchers.
Short-billed Dowitchers
Short-billed Dowitchers probe the mud with their sewing machine-like feeding style.
Short-billed Dowitchers
Seldom do they take a break long enough for an observer to their bill in its entirety.
Lesser Yellowlegs
A Lesser Yellowlegs arrives at Raymond Pool as a high-tide refugee.
Lesser Yellowlegs
Another Lesser Yellowlegs on a mudflat.
Greater Yellowlegs
A Greater Yellowlegs wades into shallow water to feed.
Semipalmated Plovers and Semipalmated Sandpipers
It’s difficult to estimate just how many Semipalmated Plovers and Semipalmated Sandpipers were seen.  There were at least 500 plovers, and they were very vocal.  The latter species was present in numbers measurable in the thousands.  We don’t think 10,000 Semipalmated Sandpipers is an overestimate.
Semipalmated Sandpipers
Searching through the Semipalmated Sandpipers, one could regularly find a very similar species among them.
Western Sandpiper
We identified the longer-billed and slightly larger Western Sandpiper (Calidris mauri) and found the species to be present possibly by the hundreds among the masses of thousands of Semipalmated Sandpipers.  These seem to be unusually high numbers for this more western species, but who’s complaining?
Western and Semipalmated Sandpipers
As we sifted through these groups of tiny shorebirds known as “peeps”, we found Western Sandpipers (top) regularly distributed among the multitudes of Semipalmated Sandpipers (bottom) we encountered.
Western and Semipalmated Sandpipers
A Western Sandpiper (top) photographed with a Semipalmated Sandpiper (bottom) in Bear Swamp Pool.
Snowy Egret
Of course, shorebirds aren’t all there is to see at Bombay Hook.  More than 100 Snowy Egrets were found on the freshwater pools alongside birds like this Semipalmated Plover.
Great Egret
Great Egrets could be seen stalking small fish in the channels of the pools.
Green Heron
And a few Green Herons were found lurking in the vegetation.
Osprey
This Osprey briefly startled the shorebirds on Raymond Pool until they realized it posed no threat.
Shorebirds Feeding on Mudflat during Receding Tide
By early afternoon, we noticed the tide beginning to retreat from the saltwater marsh and mudflats opposite Shearness Pool.  As shorebirds began returning there to feed, we decided to make our way to Raymond Pool to watch the exodus.
Shorebirds Depart Raymond Pool
With the high tide receding from the coastal estuary back into Delaware Bay, shorebirds promptly departed their concentrated environs on Raymond Pool to spread out over thousands of acres of salt marsh to feed.
Black-bellied Plovers and other Shorebirds Leaving Raymond Pool
More shorebirds exit Raymond Pool en route to the adjacent tidal areas to feed.  This outbound group includes two Black-bellied Plovers (top center and four birds to the right).
Shorebirds Entering the Saltmarsh Estuary at Bombay Hook National Wildlife Refuge
These migrants have just begun their autumn journey from breeding grounds in Canada and Alaska to wintering areas located as far south as southern South America.  For them, Bombay Hook and other refuges are irreplaceable feeding and resting locations to help them refuel for their long journey ahead.  For them, a little vacation along the coast is a matter of life and death.
White-tailed Fawn
A goodwill ambassador bids us farewell at the end of our visit to Bombay Hook.  Remember to support your National Wildlife Refuges by purchasing your annual Federal Duck Stamp.  They’re available right now at your local United States Post Office, at the Bombay Hook visitor’s center, or online at the United States Fish and Wildlife Service website.

Planning a visit?  Here are some upcoming dates with morning high tides to coax the birds out of the tidal estuary and into good light in the freshwater impoundments on the west side of the tour road…

Tuesday, August 19 at approximately 07:00 AM EDT

Wednesday, August 20 at approximately 08:00 AM EDT

Thursday, August 21 at approximately 09:00 AM EDT

Friday, August 22 at approximately 10:00 AM EDT

Saturday, August 23 at approximately 11:00 AM EDT

Photo of the Day

A teneral Eastern Scissor Grinder and exuvia.
Having emerged from the soil overnight, this pale-colored teneral (soft-bodied) Eastern Scissor Grinder has shed its exuvia (left) and is currently pumping blood throughout its extremities to expand its size and unfurl its wings.  During the remainder of the morning, the wings and exoskeleton will darken in color and harden in preparation for flight.  As an adult, this cicada then has just weeks to complete its courtship and breeding cycle before facing inevitable doom.  To see images and listen to sound clips of this and other annual species, visit our cicada page by clicking the “Cicadas” tab at the top of this page.

Disasters: The Big Three

Today’s NOAA/GOES satellite image serves as a little reminder of the big three.  That’s right, it’s the three big “natural” disasters—wildfires, inland flooding, and coastal flooding (lucky for us, our region is at present millions of years removed from severe threats posed by the tectonic disasters—earthquakes, volcanoes, and tsunamis—and is not particularly prone to frequent tornadoes).  Each continues to cause an increasing volume of property damage and threaten a greater number of lives because of where and how we choose to make our homes and erect our structures.

Smoke from Canadian Wildfires and Tropical Storm Dexter
Earlier today, smoke from forest fires in central Canada sweeps through the skies of the Great Lakes, New England, and the Mid-Atlantic States north of the Mason-Dixon Line.  Meanwhile off North Carolina, Tropical Storm Dexter is seen developing over warm ocean waters east of the Gulf Stream.  (NOAA/GOES satellite image)

For all of human existence, the dynamics of the atmosphere have been shaping the topography and the ecosystems of the planet.  In recent times, we’ve had the advantage of satellite technologies to show us spectacular images of atmospheric events as they happen.  And through the various branches of science, we’ve come to understand the impact these events have upon the landscape and the people who live and/or work there.

Forestry sciences have helped us to understand how natural fuels, humidity, temperature, rainfall, soil moisture, wind, and human encroachment influence the frequency and severity of wildfires.  These discoveries have led to changes in forest management and implementation of practices such as prescribed burns to reduce accumulated fuel loads.  Because human development typically lowers soil moisture and brings along with it additional sources of ignition, many land managers and fire departments have warned of the ever increasing dangers of wildland-urban interface fires.  These warnings have gone largely unheeded for more than four decades as millions of homes and other combustible structures have been erected within areas prone to fires capable of uncontrollable growth into disastrous conflagrations.  The tinderbox wildlands—they’re a nice place to visit, but we ought not to live there!

Tropical storms and other sources of heavy precipitation bring about quite the hubbub over flooding.  Meteorologists spend a lot of time explaining it all, but it’s almost as if no one pays any mind.  For a people who check the weather forecast several times a day, every day of our adult life, just to get a leg up on how that weather is going to change day by day and hour by hour, you would think we would better anticipate the climatic events that happen over the long term.  In particular, you would think we would have an awareness of our own individual susceptibility to flooding— a grasp of how, where, and why floods occur.  You would think that repeated episodes of flooding would compel society to embrace an ethic that treated water as the valuable commodity it is.  Yet, we all seem to follow the same patterns of behavior.  First, we drain, dump, pipe, curb, channel, ditch, grade, pave, and pump to get the rain that falls upon our property off of our property.  Then, the chump downstream gets really mad that we sent our water his way and flooded him out, so he takes the same measures to send even more water to the next poor slob down the line until finally the now polluted slurry of runoff floods the street, a cellar, a house, a business, or a stream—a stream that has been channelized so it no longer has a floodplain to absorb, hold, purify, and infiltrate the stormwater.  Why was the stream channelized?  So we could fill in the floodplain and build upon it of course.  Two things come to mind here.  First, if we’re going to be selfish enough to flood out our neighbors, then why shouldn’t we be totally selfish and keep for ourselves all the water that falls upon our place.  After all, we’re going to need that water some day.  And second, the floodplain is a nice place to visit, but we ought not to build there.  Floodplains are for flooding; thousands of years of erosion have shaped them that way—it’s a gravity thing!

Next, we look at the lessons from geology, more specifically coastal geomorphology.  Through these disciplines we know that the coastal plain—the flat land that spent most of the last 35.5 million years (the time since the meteor strike at the present-day mouth of Chesapeake Bay) as a beach or a tidal marsh—today stands mostly less than three dozen feet above sea level.  We know that the sands forming barrier islands along the Atlantic seaboard, which are only several feet above sea level, shift their shape and position with the tides.  Over the decades and centuries, these islands migrate and compensate for changes in climate and tidal patterns as well as sea levels.  Behind their shifting dunes, vast tidal marshes are protected from seasonal storms including the periodic nor’easter or hurricane.  Despite the importance of barrier island dynamics to the integrity of the bays and estuaries they protect, and despite their vulnerability to coastal storm surges, winds, and flooding, we choose to build there.  In fact, the greatest population densities in the United States, and in many other countries of the world, are on the beach.  It’s not because these hundreds of millions of people are fishing or loading/unloading ships for a living—it’s mostly for the view.  Despite their importance to fisheries and other coastal life, we continue to alter and destroy the near-tidal areas of the the barrier islands and bays.  We go to great expense to “save” for our uses the lands that should be getting inundated by rising sea levels to create new shallow tidewater zones.  We waste spectacular amounts of money pumping sand back onto beaches to keep naturally migrating sediments from changing their shape and position in response to the tides.  We keep putting more people and more capital at risk by urbanizing these low-lying areas.  Building on the beach is absolute madness.  It’s an ecological catastrophe from day one and a human catastrophe soon after.

All of the lands impacted by these natural events have two things in common.  Each becomes a potential disaster area if people choose to construct their homes or businesses there.  And each, if left in its wild state and given a buffer space from human activity, reacts with natural time-derived mechanisms in response to the same events.  These mechanisms are often essential for provision of the unique ecosystems required by many of our most threatened wildlife species.  Human encroachment into floodplains, wetlands, tidal marshes, beaches, and xeric uplands is a double-edged sword.  It first decimates populations of these uncommon species by destroying and fragmenting their specialized habitats.  Then, it sets the stage for the fires, floods, and other disasters that endanger the lives and property of the people living there.  Considering the ramifications of building in these fire and flood susceptible areas, we can and should live somewhere else, especially when the wildlife requiring these places often can’t.

Sunset over the Susquehanna at Chiques Rock
This evening’s smoky sunset over the Susquehanna at Chiques Rock was courtesy of Canadian wildfires.

A Sixth Species of Annual Cicada in the Lower Susquehanna Basin: Neotibicen robinsonianus

Just as we were very pleased last month to have the opportunity to hear the sounds of the rarest of the Periodical Cicadas—the Little Seventeen-year Cicada—in the Conewago Hills of York County to thus provide our only record of the species during the Brood XIV emergence in the Lower Susquehanna River Watershed, we were this week delighted to find and record a population of what may be the valley’s rarest cicada experiencing annual flights—the Robinson’s Cicada—just a few miles away at Gifford Pinchot State Park.

Like the Little Seventeen-year Cicada, Robinson’s Cicada (Neotibicen robinsonianus) is a species found more commonly in the southern United States, occurring with scattered distribution in a range that extends west into Missouri, Kansas, and Texas.  They are of rare occurrence in the lands of the Chesapeake drainage basin in Virginia and Maryland.

Swamp Cicada
Populations of Swamp Cicadas, also known as Morning Cicadas, are presently in the midst of their yearly courtship rituals.  Their songs are easily heard in woods, edge habitat, and suburbia throughout the region during the AM hours.  It was while recording the sounds of these common insects in the Conewago Day-use Area at Gifford Pinchot State Park that we noticed a very unique song from high in the oaks, hickories, and other hardwoods along the lake.  Listen to this sound clip featuring a group of serenading male Swamp Cicadas.  In the background, a Robinson’s Cicada’s song consisting of a pulsing series of raspy buzzes, each about one second in duration, can be heard, particularly starting at 00:40.
Robinson's Cicada
Dozen’s of Robinson’s Cicadas were heard this week in the large trees of the lakeside picnic grove in the Conewago Day-use Area at Gifford Pinchot State Park.  They remained high in the canopy and were glimpsed only when flying to a new perch, so we’ll have to settle for a photo of an individual from the more southerly portions of the species’ range (provided courtesy of AmaryllisGardener, under license: CC BY-SA 4.0).  We recorded these two sound clips of males singing in the forested area west of the picnic grove.  The first was nearer the park campground.  The second was in the vicinity of the nature center building and includes a Wood Thrush and another Robinson’s Cicada in the background.

The Lower Susquehanna River Watershed, and Gifford Pinchot State Park in particular, may currently represent the northern limit of the geographic range of the Robinson’s Cicada.  If you’re in the area during the coming weeks, drop by the park and have a listen.  And don’t forget to check out our “Cicadas” page for sound clips of all the species found in our area.

Sunny Swarms of Insects

It begins on a sunny morning in spring each year, just as the ground temperature reaches sixty degrees or more…

Eastern Subterranean Termites
Eastern Subterranean Termites emerge in unison from a nest located in the soil beneath a log.  Each of these swarming “alates” is a potential king or queen seeking to find a location with an ample supply of fallen timber to provide food for establishment of a new colony.  They are escorted to the surface by a soldier (lower center) equipped with powerful jaws for protecting the existing nest.  Similar-looking worker termites tend the nest, the queen, her eggs, and their siblings, but usually remain hidden from view.  The workers feed upon wood, hosting cellulose-digesting protozoa and bacteria in their guts to break down the fibers.  This symbiotic relationship is an important mode of decomposition in the forest, the process that turns wood into the organic matter that enriches soil and helps it to retain more moisture.
Fungi on Fallen Log
Termites are among the numerous arthropods that join fungi and a variety of microbes to decompose dead wood and other plant matter into the nutrients and organic materials used by living plants to thrive and grow.
Eastern Subterranean Termites
Within moments of emerging, swarming “alates” ascend a tree trunk or other vertical surface from which they can take flight.
Eastern Subterranean Termites
Eastern Subterranean Termite “alates” gather atop a stump before launching skyward.
Eastern Subterranean Termites
Termites swarm in massive numbers in an attempt to overwhelm the predators that are inevitably attracted to their sudden appearance.  The few “alates” that survive to find a source of rotting wood in which to begin a new colony are the only hope for continuing their king and queen’s legacy.
Green Frog Stalking a Eastern Subterranean Termite
Soon after lift off, the majority of swarming termites are consumed by swallows, swifts, and other birds, but some are discovered at ground level.
Green Frog Eating Termites
A lightning-fast strike with its tongue and this Green frog has snatched up yet another termite.  Those that slip by the dragnet of terrestrial and aerial predators can sometimes start a new colony in the ground beneath a dead tree or in a vulnerable house or other wooden structure.  To keep a small clan from invading your home, be certain the wood elements of your building(s) are kept dry and are not in contact with dirt, soil, bark mulch, etc.  Regular inspections for evidence of their presence can head off the long-term damage termites can inflict on the stuff we construct with tree skeletons.
Green Frog Eating Termites
A Green Frog wearing its breakfast.  Its next chance for a termite feast may come during the autumn when the Drywood Termites (Kalotermitidae) swarm.

Of course, termites aren’t the only groups of insects to swarm.  As heated runoff from slow-moving thundershowers has increased stream temperatures during the past couple of weeks, there have occurred a number of seasonal mayfly “hatches” on the Susquehanna and its tributaries.  These “hatches” are actually the nuptial flights of newly emerged imago and adult mayflies.  The most conspicuous of these is the Great Brown Drake.

Great Brown Drake
Seen here with a much smaller and more typical regional mayfly to its left, the Great Brown Drake was for several years infamous for swarming the lights and creating traffic hazards on bridges spanning the Susquehanna.  During the past two weeks, nighttime flights of these giants have ventured out to gather at well lit locations in housing and business districts more than a dozen miles from the river.  Earlier this century, this proclivity to wander probably led the Great Brown Drake to first invade silty segments of the Susquehanna as a colonizer from its native range in the Mississippi watershed.

Swarms of another storm-related visitor are being seen throughout the lower Susquehanna valley right now.  Have you noticed the Wandering Gliders?

Wandering Glider
Throughout the month, swarms of Wandering Gliders, the most widespread dragonflies in the world, descended on areas hit by localized slow-moving thundershowers.  Large numbers of these global travelers are known to get swept up within the thermal air masses that lead to these storms.  In suitable terrain within the path of the downpours, they linger to search for flooded places where they can mate and deposit eggs.  Wandering Gliders frequently mistake large parking lots at shopping malls, grocery stores, etc. for wetlands and will be seen in these areas depositing eggs upon the hoods and roofs of shiny motor vehicles, surfaces which appear puddle-like in their eyes.
Mating Common Green Darners
More dragonfly swarms are yet to come.  Adult Common Green Darners are presently in the Lower Susquehanna River Watershed mating and depositing eggs within vegetated ponds, lakes, and wetlands.  Beginning in August, adult Common Green Darners and other migratory species including Black Saddlebags, Carolina Saddlebags, Wandering Gliders, and Twelve-spotted Skimmers will begin swarming as they feed on flying insects (including lots of mosquitos and gnats) and start working their way toward the Atlantic Coastal Plain.  Along the barrier islands by September, concentrations of southbound dragonflies can reach the thousands, particularly at choke points like Cape May, New Jersey, and Cape Charles, Virginia.  So be sure to keep an eye on the sky for swarms of dragonflies during coming weeks.  And don’t forget to check out our “Damselflies and Dragonflies” page by clicking the tab at the top of this page.

It’s Prime Time for Hummingbirds

Ruby-throated Hummingbird at Scarlet Bee Balm
Now that the nesting season is drawing to a close, Ruby-throated Hummingbirds are a bit less fussy about where they spend their time.  Even in urban settings, gardens with an abundance of nectar-producing flowers like this Scarlet Bee Balm (Monarda didyma) have a good chance of attracting them.  Hummingbirds will be wandering the landscape and starting to drift south during the coming two months, so keep your feeders clean and filled with a fresh blend of sugar water to keep them energized and happy.  If you’re feeding hummingbirds, or thinking about feeding hummingbirds, be sure to review the helpful tips contained in our post from August 5, 2022, “Two Feeders Are Better Than One”.  Their health and your peace of mind may depend on it.

With Periodical Cicadas Gone, Here Come the Annual Cicadas…Right on Schedule!

Eastern Scissor Grinder
As the choruses of Brood XIV Periodical Cicadas fall silent for another seventeen years, the sounds of the more widespread annual Neotibicen cicadas are starting to be heard throughout the lower Susquehanna valley.  These latter insects have a more abbreviated life cycle, spending just two to five years in the subterranean nymphal stage before appearing during mid-summer to breed.  The emergence of these annual insects is not synchronized into broods, so some adults are found taking flight, singing, and mating every year.  Over the centuries, male annual cicadas that emerged and commenced courtship songs earlier than July have certainly failed to successfully reproduce during Periodical Cicada years.  So to avoid competition with the overwhelming drone of the seventeen-year cicadas that may emerge along with them, natural selection has delayed the maturation of the annual species until just after the periodicals have gone quiet.  To see pictures and hear the sounds of our five (now 6 as of July 24th) species of annual cicadas, click the “Cicadas” tab at the top of this page.

Sorry, But We Have No Steamed Crabs Today

Goldenrod Crab Spider
Earlier today, we collected some mint leaves from the garden to make a batch of iced tea.  Just prior to plunging them into boiling water, we noticed this Goldenrod Crab Spider (Misumena vatia) hiding in the foliage.
Goldenrod Crab Spider
Goldenrod Crab Spiders live among the stems and leaves of flowering plants.  They are particularly fond of goldenrods, milkweeds, and other species that attract an abundance of flying insects.  When the plants bloom, the spiders will use their white or yellow coloration to hide among the petals and disks of the flowers.  From there, they ambush the visitors that stop by for a sample of nectar or pollen.  In an effort to lure prey directly into their clutches, these acrobatic arachnids will even dangle among the clusters of blossoms with their legs spread like petals surrounding their disk-like body.  This behavior helps inspire their other common name: Flower Crab Spider. 
Goldenrod Crab Spider
Needless to say, this crab was spared the pot and instead returned to the garden to help keep the plants healthy and ecosystem in balance in our wildflower patch.  So we’ll just be having tea, thank you.

Photo of the Day

Independence Day Safety Equipment
The gasoline and gunpowder gang’s biggest holiday of the year is once again upon us.  This Independence Day weekend, don’t let the celebration turn to tragedy.  Keep a garden hose or fire extinguisher at the ready to dowse any hot embers or other potential troublemakers.  Fill a bucket with water for use as a trash receptacle for your hot sparkler rods and other pyrotechnic waste.  Have non-combustible lids or covers handy to smother any flare ups while you’re grilling.  Don’t forget, they’ll be plenty of testosterone and adrenaline circulating to keep the festivities exciting, so skip the alcohol and energy drinks if you’re driving or lighting off fireworks.  Things don’t need to be any more explosive.  And remember, the anthem’s lyrics say, “…the rockets red glare, bombs bursting in air…”, not burning your neighbor’s garage down, so maintain a safe distance with your grills, camp fires, and July 4th displays, won’t you please?  Have an exciting weekend!

Brood XIV Periodical Cicada Wrap Up

While the heat and humidity of early summer blankets the region, Brood XIV Periodical Cicadas are wrapping up their courtship and breeding cycle for 2025.  We’ve spent the past week visiting additional sites in and near the Lower Susquehanna River Watershed where their emergence is evident.

We begin in York County just to the west of the river and Conewago Falls in mostly forested terrain located just southeast of Gifford Pinchot State Park.  Within this area, often called the Conewago Hills, a very localized population of cicadas could be heard in the woodlands surrounding the scattered homes along Bull Road.  Despite the dominant drone of an abundance of singing Pharaoh Periodical Cicadas, we were able to hear and record the courtship song of a small number of the rare Little Seventeen-year Cicadas.  Their lawn sprinkler-like pulsating songs help mate-seeking males penetrate the otherwise overwhelming chorus of the Pharaoh cicadas in the area.

Little Seventeen-year Cicada
The Little Seventeen-year Cicada’s (Magicicada septendecula) thorax is black between the eye and the origin of the wings.  It is the rarest of the three species of seventeen-year cicadas.
Little Seventeen-year Cicada
The underside of male (left) and female (right) Little Seventeen-year Cicadas shows narrow orange edges on the abdominal segments.

From the Conewago Hills we moved northwest into the section of southern Cumberland County known as South Mountain.  Here, Pharaoh Periodical Cicadas were widespread in ridgetop forests along the Appalachian Trail, particularly in the area extending from Long Mountain in the east through Mount Holly to forests south of King’s Gap Environmental Education Center in the west.

Pharaoh Periodical Cicadas
Pharaoh Periodical Cicadas along the Appalachian Trail on South Mountain, Cumberland County, Pennsylvania.  Like the cicadas we visited last week on the east side of the Susquehanna, this population is surviving on lands with a history of timber harvest and charcoal production to fuel nearby iron furnaces during the nineteenth century.

While on South Mountain, we opted for a side trip into the neighboring Potomac watershed of Frederick County, Maryland, where these hills ascend to greater altitude and are known as the Blue Ridge Mountains, a name that sticks with them all the way through Shenandoah National Park, the Great Smoky Mountains, and to their southern terminus in northwestern Georgia.  We found a fragmented emergence of Pharaoh Periodical Cicadas atop the Catoctin Mountain section of the Blue Ridge just above the remains of Catoctin Furnace, again on lands that had been timbered to make charcoal to fuel iron production prior to their protection as vast expanses of forest.

Pharaoh Periodical Cicada
A Pharaoh Periodical Cicada on Catoctin Mountain near Catoctin Furnace south of Thurmont, Maryland.  These cicadas are not part of a Brood XIV emergence, but are instead a population of Brood X (2021) stragglers.
Pharaoh Periodical Cicada
A female Brood X Pharaoh Periodical Cicada straggler on Catoctin Mountain.  The website “cicadamania.com” notes, “Experts (Gaye Williams, State Entomologist of Maryland, John Cooley of UCONN) have confirmed that there will be no Brood XIV cicadas for Maryland.”

Back in Pennsylvania, we’re on our way to the watersheds of the northernmost tributaries of the lower Susquehanna’s largest tributary, the Juniata River.  There, we found Brood XIV cicadas more widespread and in larger numbers than occurred at previous sites.  Both Pharaoh and Cassin’s Periodical Cicadas were seen and heard along Jack’s Mountain and the Kishacoquillas Creek north of Lewistown/Burnham in Mifflin County.  To the north of the Kishacoquillas Valley and Stone Mountain in northernmost Huntingdon County, the choruses of the two species were again widespread, particularly along the forest edges in Greenwood Furnace State Park, Rothrock State Forest, and adjacent areas of the Standing Stone Creek watershed.

Tymbal on the male Pharaoh Periodical Cicada
A view of the sound-generating tymbal on a male Pharaoh Periodical Cicada at Greenwood Furnace State Park.  Rapid vibration of the tymbals by a set of specialized muscles generates the distinctive calls and courtship songs of the various cicada species.  When handled, these tymbals can produce a harsh “panic call”.  This distress sound could startle a would-be predator and provide the cicada with an opportunity to escape.
Tymbal on a Cassin's Periodical Cicada
The sound organs comprised of ribbed tymbals and specialized muscles on the male Cassin’s Periodical Cicada generate a “panic call” as well as the distinctive calls and songs used to penetrate the droning choruses of the more numerous Pharaoh Periodical Cicadas with which it shares a seventeen-year flight.
Cassin's Periodical Cicadas
Using their specialized sound organs, Cassin’s Periodical Cicadas generate a courtship song that usually includes buzzy phrases and ticking notes (first sound clip).  The buzzing and ticking helps the male Cassin’s cicada penetrate the songs of the more numerous Pharaoh Periodical Cicadas in the area (second sound clip).  When synchronized into a chorus that surges in volume, the songs of Cassin’s Periodical Cicadas can overcome the overwhelming drone of the nearby Pharaoh cicadas (third sound clip).
Pharaoh Periodical Cicada Depositing Eggs
After mating and before the lives of these seventeen-year cicadas draw to a close, the females need to deposit their fertilized eggs into the small end twigs of suitable trees.  On a small hawthorn tree (Cretaegus species) along the edge of the forest at Greenwood Furnace State Park, this Pharaoh Periodical Cicada is using her ovipositor to make a slit in a twig and place her eggs.
Cassin's Periodical Cicada Ovipositing
Simultaneously on the same little hawthorn tree, this female Cassin’s Periodical Cicada is depositing her fertilized eggs.

Within the last 48 hours, we visited one last location in the Lower Susquehanna River Watershed where Brood XIV Periodical Cicadas have emerged during 2025.  In the anthracite coal country of Northumberland County, a flight of Pharaoh Periodical Cicadas is nearing its end.  We found them to be quite abundant in forested areas of Zerbe Run between Big and Little Mountains around Trevorton and on the wooded slopes of Mahanoy Mountain south of nearby Shamokin.  Line Mountain south of Gowen City had a substantial emergence as well.

Pharaoh Periodical Cicada
A Brood XIV Pharaoh Periodical Cicada near Zerbe Run west of Trevorton, Northumberland County, Pennsylvania.  The following sound clip features the fading chorus of these cicadas and some of the nesting birds that may actually be preying upon them: Yellow-billed Cuckoo, Red-eyed Vireo, and Northern Cardinal.   
Foliage with Evidence of Ovipositing
Brown leaves reveal the end twigs where female Pharaoh Periodical Cicadas have deposited eggs during the last several weeks.  During July, the larvae will hatch and drop to the ground to start a new generation of Brood XIV cicadas.  As subterranean nymphs, they’ll spend the coming seventeen years feeding on small amounts of xylem sap from tree roots.  In 2042, during the next Brood XIV emergence, these nymphs will come to the surface and take flight as adults.
Foliage with Evidence of Ovipositing
Evidence of egg deposition among foliage on Line Mountain at State Game Lands 229.
Dead Pharaoh Periodical Cicadas
Accumulations of deceased Pharaoh Periodical Cicadas on Line Mountain.
Eastern Gartersnake
Fallen cicadas that show any sign of life are being snatched up by predators such as this Eastern Gartersnake (Thamnophis sirtalis sirtalis).  Meanwhile, the remainder of the biomass is picked apart by scavengers or is left to reducers for breakdown into fertilizer and organic matter for the forest.  Nothing goes to waste.

To chart our travels, we’ve put together this map plotting the occurrence of significant flights of Periodical Cicadas during the 2025 emergence.  Unlike the more densely distributed Brood X cicadas of 2021, the range of Brood XIV insects is noticeably fragmented, even in areas that are forested.  We found it interesting how frequently we found Brood XIV cicadas on lands used as sources of lumber to make charcoal for fueling nineteenth-century iron furnace operations.

Greenwood Furnace
The furnaces at Greenwood Furnace State Park required the daily cutting of one acre of timber to make enough charcoal to fuel the iron-making process.  Did keeping thousands of acres in various stages of forest succession to supply the charcoal needs of these operations aid the survival of earlier generations of Periodical Cicadas on these lands?  Or, after the furnaces converted to coal for fuel, did the preservation of many of these parcels as state, federal, and private forests allow the cicadas to find refuge from the widespread impacts of agriculture and expanding urbanization in adjacent lands?  Maybe it’s a little of both.  We always bear in mind that annual insects and other animals are more than one hundred generations removed from the negative or positive impacts of the early years of the industrial age, but only about ten generations have passed since populations of seventeen-year Periodical Cicadas were directly influenced by these factors.  What do you think?

Well, that’s a wrap.  Please don’t forget to check out our new Cicadas page by clicking the “Cicadas” tab at the top of this page.  Soon after the Periodical Cicadas are gone, the annual cicadas will be emerging and our page can help you identify the five species found regularly in the lower Susquehanna valley.  ‘Til next time, keep buzzing!

Brood XIV Periodical Cicadas in the Lower Susquehanna River Watershed

Here are some sights and sounds from the ongoing emergence of Brood XIV Periodical Cicadas in the Lower Susquehanna River Watershed.

We begin in the easternmost spur of the lower basin where a sizeable emergence of cicadas can be seen and heard in the woodlands surrounding the headwaters of the Conestoga River in Berks County north of Morgantown.  This flight extends east into Chester County and the French Creek drainage of the Schuylkill River watershed on State Game Lands 43 north of Elverson and consists of Pharaoh Periodical Cicadas (Magicicada septendecim), the most common species among 17-year broods.

Pharaoh Periodical Cicada
A Brood XIV Pharaoh Periodical Cicada at State Game Lands 43 identified by the red bar extending from the eye to the wing root.
Pharaoh Periodical Cicada
The underside of the abdomen on a male Pharaoh Periodical Cicada showing the wide orange bars on each segment.
Pharaoh Periodical Cicadas exuvia
Exuvia of a recently emerged Pharaoh Periodical Cicada.
Pharaoh Periodical Cicada singing
Soon after landing on a perch, a male Pharaoh Periodical Cicada will usually announce his presence by singing.  It’s an attempt to quickly attract potential mates that may be in the vicinity.
Pharaoh Periodical Cicadas
Pharaoh Periodical Cicadas ascending the branches of an oak.
Pharaoh Periodical Cicadas singing chorus
Gatherings of thousands of singing male Pharaoh Periodical Cicadas create a distinctive droning chorus.
Pharaoh Periodical Cicadas
A receptive female will make a click sound with her wings to summon a suitable singing male for mating.
Pharaoh Periodical Cicadas
While usually occurring in the safety of the trees, the breeding frenzy can spill over onto the ground where we happened to find this copulating pair.
Chestnut Oak Hosting Cicadas
After mating, female cicadas make slits in the end twigs of selected trees into which they lay their eggs.  The process of egg-laying and larval emergence will usually wilt and kill end growth on the affected branches, causing little harm to healthy trees.  It’s similar to the trim you might give to a bonsai plant to keep it stout and sturdy.
Pharaoh Periodical Cicadas
How long do Periodical Cicadas live?  Well, by last week, we were already finding dead specimens by the thousands.  Most of them had already completed their breeding cycle and planted the seeds for a new generation of Brood XIV cicadas.
A deceased Pharaoh Periodical Cicada at the Fire Tower Parking Area at French Creek State Park.  This specimen and a chorus on the hill’s forested south slope were the northeastern-most evidence of Brood XIV Periodical Cicadas we could find for the population cluster in portions of Berks, Chester, and Lancaster Counties around Morgantown.
Second-year Mississippi Kite Feeding on Cicadas
The abundance of Pharaoh Periodical Cicadas on State Game Lands 43 in Chester County has attracted numerous raptors, particularly wandering one-year-old birds that aren’t quite mature enough to nest.  Among the sightings have been Red-tailed Hawks, Red-shouldered Hawks, Broad-winged Hawks, and at least three Mississippi Kites, a rarity on the Piedmont this far inland from the coastal plain.  (See the post from June 5, 2021, for details on the occurrence of Mississippi Kites in northernmost Delaware during the Brood X emergence.)

From Route 82 north of Elverson to the west through the forested areas along Route 10 north of Morgantown and the Pennsylvania Turnpike, we found an abundance of Cassin’s Periodical Cicadas (Magicicada cassini) calling among the Pharaohs.  This mix of Pharaoh and Cassin’s Periodical Cicadas extends west along the north side of the turnpike into Lancaster County and State Game Lands 52 on Black Creek north of Churchtown.

Cassin's Periodical Cicada
A Brood XIV Cassin’s Periodical Cicada at State Game Lands 52 is identified by the all-black margin between the eye and the wing root and…
Cassin's Periodical Cicada
…the black underside of the abdomen with no orange stripes.
Cassin's Periodical Cicadas singing chorus
To penetrate the sounds of the more common Pharaoh Periodical Cicadas, male Cassin’s Periodical Cicadas gather in large concentrations to generate a loud, oscillating chorus.  Its surging volume will usually exceed that of the Pharaohs singing in the vicinity.
Cassin's Periodical Cicadas
Mated Cassin’s Periodical Cicadas copulating at State Game Lands 52.
Cassin's Periodical Cicadas mating
The underside of copulating Cassin’s Periodical Cicadas.
Cassin's Periodical Cicadas mating
A pair of Cassin’s Periodical Cicadas at state Game Lands 52 in Lancaster County.

Further west in Cornwall, Lebanon County, a Brood XIV emergence can be found on similar forested terrain: the Triassic hills of the Newark Basin—rich in iron ore and renowned for furnace operations during the eighteenth and nineteenth centuries.  Pharaoh Periodical Cicadas were the only species heard among this population that extends from Route 72 east through the woodlands along Route 322 into the northern edge of State Game Lands 156 in Lancaster County.

On the west side of the Susquehanna, yet another isolated population of Brood XIV Periodical Cicadas can be found in Perry County, just south of Duncannon on State Game Lands 170 on the slopes of “Cove Mountain”, the canoe-shaped convergence of the western termini of Peters and Second Mountains.

Pharaoh Periodical Cicadas dominated this Perry County chorus,…

…but we did detect at least one Cassin’s Cicada trying to find a mate.

Cassin's Periodical Cicada
A solitary Cassin’s Periodical Cicada issues a lonely song of short buzzes and ticking notes on State Game Lands 170.  Fragmented populations, especially those that are only able to fly and increase their distribution every 17 years, often have a challenging time expanding and reuniting their disjointed ranges.

Not to say they aren’t present, but we have yet to detect the rarest species, Magicicada septendecula, the “Little Seventeen-year Cicada”, among the various populations of Brood XIV Periodical Cicadas emerging in the lower Susquehanna valley.  For the coming two weeks or so until this brood is gone for another 17 years, the search continues.

For more on both annual and periodical cicada species in the Lower Susquehanna River Watershed, be sure to click the “Cicadas” tab at the top of this page!

The All-American Dad

Just in time for the Flag Day/Father’s Day weekend, the Jordanella floridae we raise here at susquehannawildlife.net headquarters to help control hair algae in our planted aquaria are beginning to spawn.

Jordanella floridae Courtship
A male Jordanella floridae displaying for a prospective female mate.  A receptive female will nip the fins of a courting male prompting him to chase her.

In the wild, Jordanella floridae inhabits a variety of vegetated tropical wetlands and backwaters on the Florida peninsula.  The species was first described there on Lake Munroe in 1879 by George Brown Goode and Tarleton Hoffman Bean, both of whom spent time working for the Smithsonian National Museum and the United States Fish Commission, the latter a forerunner of the U. S. Fish and Wildlife Service.  Goode and Bean collaborated frequently, chronicling the occurrence of freshwater species like J. floridae as well as marine ichthyofauna.  In 1896, the duo published Oceanic Ichthyology, an extensive study of pelagic and deep-sea fishes.  Each experienced a prestigious career and has had numerous fish species named after him.

Tarleton Hoffman Bean
Nineteenth-century ichthyologist Tarleton Hoffman Bean (1846-1916) was born along the shores of the Susquehanna below Conewago Falls in Bainbridge, Lancaster County, Pennsylvania.  He became a renowned expert on methods of fish culture.  (Public Domain image)
American Flagfish
To the female, the breeding colors of the dominant male Jordanella floridae are irresistible.  To us, the star-spangled banner pattern of this native killifish (more specifically a pupfish) justifies the species’ oft-used common name: American Flagfish.
American Flagfish Courtship
After being chased by the male, the female American Flagfish turns pale in color, indicating to the male that she’s ready to mate.  The pair then face each other and begin dancing around the nest site which has been prepared by the male.  The dance often consists of the female maneuvering so as not to again face in the direction of her partner.
American Flagfish Courtship
.To get into spawning position alongside his mate, the male flagfish must commence a series of rolling turns.
American Flagfish Spawning
Once the male is successful and the female is receptive, spawning quickly ensues and the pair swims sideways in a tight circle around the center of the nest site…
American Flagfish Spawning
…as the eggs are deposited and fertilized.
Male American Flagfish Guarding Nest
Female American Flagfish lay fewer than two dozens eggs, so the male must diligently guard the nest from potential outside threats to his progeny including snails, dragonfly nymphs, and other fishes.  Periodically, he’ll use his fins to fan a current across the eggs to clear them of debris and assure an exchange of fresh water through the nest.
Male American Flagfish Tending Nest
For the male American Flagfish, obsessive patrol of the nest site and care of the eggs continues day and night.  The American Flagfish, a dandy that dresses for his role as an All-American Dad.

Photo of the Day

Great Spangled Fritillary
Along a quiet forest road, a Great Spangled Fritillary visits Red Clover (Trifolium pratense) to feed on nectar.

Photo of the Day

Common Carp
We came across this photo from a dive we did back in 1999 and thought it timely.  Here a large non-native Common Carp churns up a cloud of nutrient-charged sediment as it roots its way through a bed of American Eelgrass and Water Stargrass in the Susquehanna below Conewago Falls.  (Vintage 35 mm image)

Common Carp: A Menace Meant for the Dinner Plate

One of the earliest non-native fish species to be widely released into North American waterways was the Common Carp.  Stocks brought to the United States were likely sourced from populations already naturalized throughout much of western Europe after introductions originating from the fish’s native range in Eurasia, probably including the Danube and other watersheds east through the Volga.  In western Europe, the species promised to be an abundant and easily cultivated food source.  Under the same premise, carp were transported to the United States during the early 1800s and widely introduced into streams, lakes, and rivers throughout the country.

Common Carp thrive in nutrient-rich waters, particularly those subjected to sewage discharge and agricultural runoff, conditions which were already prevalent during the Common Carp’s initial introduction and have remained widespread ever since.  Within these polluted streams, lakes, and ponds, introduced carp feed aggressively on benthic organisms and plants, stirring up decaying organic matter (mulm) from the substrate.  This process raises turbidity in the water column and releases excessive amounts of the nutrient phosphorus resulting in unusually large algal blooms.  Algal blooms can block sunlight from the longer-lived oxygen-producing vascular plants that grow in submerged environs.  Growing beneath a dense cloud or blanket of algae can compromise the vigor of oxygen-producing vascular plants and disable their biochemical functions within the aquatic ecosystem.  As the short-lived algae die, the bacteria that decay them begin to place increased oxygen demands on the water.  With less oxygen being produced by both the vascular plants and the algae, and with oxygen consumption increased by the activity of decomposers, conditions can become fatal for fish and other organisms.  This process is known as eutrophication.  Because Common Carp are among the species most tolerant of eutrophic conditions, they tend to thrive in the conditions they create while the native fishes perish.

Common Carp spawn in the spring, usually from late April through June, when the water temperature is as low as 58 degrees and as high as 83 degrees Fahrenheit.  This activity is often triggered by a rapid increase in water temperature.  In a small lake, this may be brought on by a string of sunny days in late April or May.  On larger streams and rivers, the temperature spike that initiates the spawn may not occur until warm rains and runoff enter the stream during June.

Common Carp
Seeing the exposed backs of Common Carp as they stir up mulm and other sediments while feeding along the edges of a body of water is not at all unusual.
Common Carp
But carp pursuing other carp into the shallows is a sign that spawning has commenced.
Common Carp Spawning
In water that is often less than a foot in depth, male carp follow the breeding females into egg-laying areas among debris and emergent vegetation.
Common Carp Spawning
A fountain of splashes can ensue as males try to outdo one another for a chance to fertilize the female’s eggs.
Common Carp Spawning
The males’ aggressive pursuit can even forced a large female to temporarily ground herself on the beach.

Common Carp are one of the most widely farmed and eaten fish in all the world.  Here in the United States, they were introduced beginning two hundred years ago because they were favorable to the palate, grew to large size quickly, and were a source of much needed food.  Today, the Common Carp is seldom found on the American dinner plate.  Yet, pound for pound, it is one of the most abundant fish in many of our waters, particularly in man-made lakes.  Like some of our other most invasive species—including Blue Catfish, Flathead Catfish, and Northern Snakehead—Common Carp are perhaps the most edible of our freshwater fishes.  For many cultures, they are an important staple.  For others, they are a delicacy or holiday treat.  In America, they do horrendous damage to aquatic ecosystems following establishment as a food crop that almost never gets harvested.  Did you realize that on the internet, there are literally hundreds of recipes and culinary videos available to show you how to prepare delicious dishes made with Common Carp?  It’s true.  And for the cost of a fishing license, you can catch all you want, usually several pounds at a time.  So why not give the marine fisheries a break?  Take the big leap and learn to eat invasive freshwater species instead.

Common Carp Breaching
Jumpin’ gefilte fish, it’s what’s for dinner!

Soakin’ Up the Smoky Sun

With temperatures finally climbing to seasonable levels and with stormy sun filtering through the yellow-brown smoke coming our way courtesy of wildfires in Alberta and other parts of central Canada, we ventured out to see what might be basking in our local star’s refracted rays…

Black Saddlebags
Dragonflies including this Black Saddlebags are now actively patrolling the edges of waterways and wetlands for prey and mates.
Common Green Darners
Here we see a pair of Common Green Darners flying in tandem…
Common Green Darners
…and, having already mated, stopping at a suitable location for the female to oviposit the fertilized eggs onto submerged plant stems.
Painted Turtles
A sunny day almost always brings out the reptiles, including these Painted Turtles…
Red-eared Slider
and the invasive Red-eared Slider, a native transplant from the American midwest.
Snapping Turtle
A really big Snapping Turtle will prey on almost anything, including other Snapping Turtles…
Golden Shiners
…but this one seems to be fascinated by something a lot smaller.  Something like these juvenile Golden Shiners seen here schooling in the sun-drenched shallows.
Northern Water Snakes
Turtles aren’t the only reptiles thriving in the heat.  Northern Water Snakes (Nerodia sipedon sipedon) take full advantage of a sun-drenched rock to warm up after spending time in the chilly water of a stream.
Northern Water Snakes
You know, no one loves a snake like another snake…
Northern Water Snakes
…and when it comes to these two snakes, it looks like love is in the air!
Spicebush Swallowtail
Butterflies like this Spicebush Swallowtail enjoy time in the sun, even while seeking out minerals in a patch of moist soil.
Woodchucks
After its siblings darted into the familial burrow upon our approach, this juvenile Woodchuck instead sought the attention of its nurturing mother.  Unlike its brothers and sisters, perhaps this little groundhog isn’t afraid of its own shadow.  Or does the smoky haze have the youngster all confused about what does and doesn’t constitute as a shadow?  Well, we can’t help you there, but you have a whole eight months to figure it out!

Brood XIV Periodical Cicada Emergence

Its been four years to the day since we posted our account of the big Brood X Periodical Cicada flight of 2021 in the Lower Susquehanna River Watershed.  With the Brood XIV clan of 17-year Periodical Cicadas getting ready fill the June air with their choruses in scattered parts of Pennsylvania’s Berks, Centre, Clinton, Franklin, Huntingdon, Juniata, Lancaster, Mifflin, Perry, Schuylkill, Snyder, Union, and York Counties right now, we thought it an appropriate time to open a new “Cicada” page here on the website.  Included is a list of both the annually and periodically emerging species found in the lower Susquehanna valley, as well as a copy of the article and ID photos from the 2021 occurrence of Brood X.  In coming months, we’ll be adding photos and maybe some sound clips of the different annual cicadas as well, so remember to check back from time to time for more content.

Periodical Cicada Nymph
Note the wings and red eyes of this periodical cicada nymph.  Within weeks it will join billions of others in a brief adult emergence to fly, mate, and then die.

In the short term, we’re going to pay a visit to the Brood XIV territory and will bring you updates as we get them.  Until then, be sure to click the new “Cicadas” tab at the top of this page to brush up on your ID of the three species of 17-year Periodical Cicadas.

Adult Periodical Cicada
Not to worry.  Cicadas are native, harmless, docile creatures when handled and they pose no threat to the long-term health of your healthy, well-established trees and other vegetation, so enjoy them while they’re here!

Photographs in Living Color: Black and White is Beautiful

Here at susquehannawildlife.net headquarters, we really enjoy looking back in time at old black-and-white pictures.  We even have an old black-and-white television that still operates quite well.  But on a nice late-spring day, there’s no sense sitting around looking at that stuff when we could be outside tracking down some sightings of a few wonderful animals.

American Toad Tadpoles
American Toad tadpoles have hatched from clusters of eggs deposited in this wet roadside ditch furnished with a clean supply of runoff filtered through a wide shoulder of early successional growth.  Recent rains have kept their vernal nursery flooded, giving them the time they need to quickly mature into tiny toads and hop away before scorching summer heat dries up their natal home.
Water Striders mating.
Weekend rains and creek flooding haven’t stopped these Water Striders from pairing up to begin their breeding cycle.
Common Whitetail
Around streams, lakes, ponds, and wetlands, the Common Whitetail is one of our most conspicuous dragonflies.
Great Blue Heron
Now that’s what we call a big beautiful bill, on a Great Blue Heron stalking fish.
Golden-backed Snipe Flies
These mating Golden-backed Snipe Flies (Chrysopilus thoracicus) are predatory insects, as are their larvae.  They are most frequently found in bottomland woods.
Eastern Ratsnake
About three feet in length, this Eastern Ratsnake is unusual because it still shows conspicuous remnants of the diamond-patterned markings it sported as a juvenile.
Black-and-white Warbler
The plumage of the Black-and-white Warbler lacks any of the vibrant colors found in the rainbow, but is nevertheless strikingly beautiful.
Black-and-white Warbler
This male Black-and-white Warbler appears a little bit ruffled as he dries out his feathers following a brief afternoon downpour. 
Black-and-white Warbler
But as the sunshine returns, he bursts into song from a forest perch within the nesting territory he has chosen to defend.  In addition to the vocalizations, this eye-catching plumage pattern helps advertise his presence to both prospective mates and would-be trespassers alike.  But against the peeling bark of massive trees where this bird can often be found quietly feeding in a manner reminiscent of a nuthatch, the feathers can also provide a surprisingly effective means of camouflage.  

Photo of the Day

Cedar Waxwing Feeding on Juneberries
One of dozens of Cedar Waxwings seen descending upon ripe juneberries in a mini grove consisting of either Smooth Shadbush (Amelanchier laevis) or the allied and very similar-looking hybrid juneberry Amelanchier x lamarkii.  Smooth Shadbush can be grown as a shrub or small tree and is also known as Smooth Serviceberry, Allegheny Serviceberry, or Smooth Juneberry.  The hybrid Amelanchier x lamarkii is believed to be a naturally occurring cross between Smooth Shadbush (A. laevis) and either Canadian Serviceberry (A. canadensis) or Downy Serviceberry (A. arborea).  Juneberries/serviceberries/shadbushes, including a number of man-made cultivars, produce white flowers in early spring and can be obtained through numerous suppliers for inclusion in conservation projects, home gardens, or for use as street trees.  Believe it or not, the very productive planting seen here was located in a parking lot island at a busy Walmart store.

Beechnuts in a Jam

Renowned for its smooth, light-gray bark and its large size, the American Beech is one the most easily recognized trees found in climax forests throughout the lower Susquehanna valley.  Preferring rich soils, this shade tolerant native produces an abundance of nutritious nuts for wildlife including deer, turkey, grouse, squirrels, woodpeckers, and a variety of songbirds.

If you’ve visited a stand of beech trees lately, you may have noticed that the canopy seems a little sparse in comparison to the foliage of the oaks, poplars, and other hardwood species in the vicinity.

American Beech Canopy
Sunlight reaching the forest floor through gaps in the canopy of American Beech trees.

A closer look reveals the cause.  And yes, it’s big, big trouble.

American Beech with Beech Leaf Disease
Many newly emerged leaves on this American Beech are thickened and curling in comparison to apparently healthy leaves on the same branch.  These sickly leaves are being afflicted by Beech Leaf Disease (BLD), the result of infection by a parasitic roundworm, Litylenchus crenatae mccannii, a microscopic nematode that reaches its greatest abundance within the victim’s new buds.  After increasing their population density there during the summer and fall, the nematodes overwinter within the dormant buds, then cause progressive cell damage in the foliage that emerges during the spring to make the leaves appear “striped” before withering and falling away.
American Beech with Beech Leaf Disease
Early season leaves on an American Beech perishing from infection by Litylenchus crenatae mccannii, the nematode responsible for Beech Leaf Disease.  BLD also affects Oriental Beech (Fagus orientalis) and European Beech (F. sylvatica), both imported to North America for ornamental cultivation.  BLD was first detected Lake County, Ohio, during 2012 and has already spread to every county in Pennsylvania and into much of the northeastern United States and neighboring portions of Canada.  The nematode responsible for BLD possibly originated among F. orientalis stocks in Japan, but its exact origins remain unconfirmed.
Beech Infected by Litylenchus crenatae mccannii
Cell damage from Litylenchus crenatae mccannii makes the new foliage on this American Beech appear “striped” before total deterioration.
Stand of American Beech with BLD
A sparse canopy in a stand of native American Beech trees being destroyed by the nematode, Litylenchus crenatae mccannii.  Defoliation over a period of 5 to 8 years will probably prove fatal to trees of this size.  Smaller trees may succumb in just 2 to 4 years.  Other sources of stress such as the extended period of drought we recently experienced may hasten the demise of afflicted trees.

While treatment for BLD is possible, it must be done early.  Protecting an entire stand in a forest can be prohibitively expensive, but if you have a specimen tree or small grove you think you might like to save, click here for a Penn State Extension guide with more information.

A Doe and Newborn Fawn Get Acquainted

An encounter from yesterday afternoon—you see it just the way we did…

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

Odocoileus virginianus Doe and Fawn

You know, there’s no better time than the present to slow down and quit driving like a sociopath.  After all, you wouldn’t want to murder Bambi’s mom and leave him all alone right now, would you?

Deer Tick or Dog Tick?

Deer Tick vs. Dog Tick
Several weeks back, we removed the tick on the left from the editor’s neck.  Upon returning from an outing earlier today, he discovered the one on the right crawling up his sock.  It’s almost like they’re out to get him, so after tossing the day’s apparel into the washer, it was time to hit the showers posthaste.  Because where there’s one, there are more.  And you absolutely never want to spend the night with a tick.

Late May Action in the Forest

Here’s a short preview of some of the finds you can expect during an outing in the Lower Susquehanna River Watershed’s forests this week…

Mountain Laurel
The Mountain Laurel, designated as Pennsylvania’s state flower, is now in bloom.
Mountain Laurel Flower Buds
The buds of the Mountain Laurel remind us of a sugary frosting freshly squeezed from a baker’s pastry bag.
Mountain Laurel Flowers
The flowers of the Mountain Laurel, an evergreen understory shrub, invite pollinators to stop by for a sweet treat.
Little Wood-Satyr
Little Wood-Satyrs (Megisto cymela) are patrolling forest edges looking for mates and, to host their eggs and larvae, the stands of grasses they find most suitable.
Zabulon Skipper
Many of the species of small butterflies we call skippers are now active. The Zabulon Skipper can be found patrolling grassy forest edges, particularly near streams, ponds, and wetlands.
Red-spotted Purple
Among the showiest of our butterflies, the Red-spotted Purple (Limenitis arthemis astyanax) is seen here on the leaves of a Black Cherry, its favored host plant.
Red-spotted Purple
Another red-spotted Purple seen picking up minerals from a dried up puddle depression on a gravel road.
Susquehanna Riverlands State Park
Butterfly observers will do well to pay a visit to the new Susquehanna Riverlands State Park on Furnace Road north of Hellam in York County, Pennsylvania.
Trail to Schull's Rock Overlook
Upon arrival at Susquehanna Riverlands, drive back the gravel road for about a mile to the parking area at the edge of the agricultural field.  Then, hike the trail through the woods and farm hedgerow to the Schull’s Rock overlook on the river.  In the forests along this route, the understory is dominated by colonial stands of Common Pawpaw trees.
Common Pawpaw
Along the lookout trail to Schull’s Rock, the Common Pawpaws’ large leaves help them to shade out potentially fast-growing competition.  In proper growing situations, pawpaws develop clonal suckers that mature over time to create colonial stands of a single genetic plant.
Common Pawpaw understory.
A Common Pawpaw understory along the approach to Schull’s Rock.
Common Pawpaw
A colonial stand of Common Pawpaw along the trail leading to Schull’s Rock.
Zebra Swallowtail
During our recent visit to Schull’s Rock, dozens of Zebra Swallowtails were seen along the trail, many in the vicinity of their sole host plant, the Common Pawpaw.  But even more were observed along the edges of the fields and woods where nectar sources like this Multiflora Rose were being visited by numbers of butterflies we normally see only among abundant species like Cabbage Whites.  Absolutely amazing!
Scull's Rock Overlook
And the view of the Susquehanna and the Shock’s Mills railroad bridge at the mouth of Codorus Creek is pretty good too!
Common Pawpaw and Hooded Warbler
You can look for colonial stands of Common Pawpaw at other parks and preserves along the lower Susquehanna as well.  Birds like this Hooded Warbler can sometimes be found among them in mature riparian forests along the steep slopes of the river gorge.
Common Pawpaw and Baltimore Oriole
A Baltimore Oriole in a Common Pawpaw along a forest edge.
Blackpoll Warbler
Meanwhile in the treetops, the spring thrust of Neotropical migrants is drawing to a close.  The Blackpoll Warbler is typically one of the last to transit the lower Susquehanna valley on its way to northern coniferous forests for summer.  They’ve had an unusually protracted movement through the region this spring, the earliest individuals reported during late April.  Though very difficult to see in the canopy of the mature trees where it feeds and sings, hearing one is often a benchmark for senior birders each spring.  Older observers have often said of the Blackpoll Warbler’s high-pitched song, a rapid series of insect-like staccato “tseet” notes, that it was the first they could no longer detect as their ears started losing sensitivity.
American Redstart
In many tracts along the lower Susquehanna this spring, the American Redstart is turning out to be the most common nesting warbler.  Conditions favoring their reproductive success in recent cycles, as well as good survival rates during their migrations and stays on wintering grounds, have filled many lowland forests with redstart songs in 2025.  Is this the start of a trend or just an exceptionally good year?  Time will tell.
Rose-breasted Grosbeak
Yet one more reason for a stroll in local forests this week is the chance to see and hear the Rose-breasted Grosbeak.  Look for these Neotropical relatives of the cardinal nesting on territories in mature stands of deciduous trees like this Yellow Poplar, a species also known as the Tuliptree.
They spend nearly all their time among the canopy foliage of the largest timber…
Rose-breasted Grosbeak
…but pause frequently to repeat a song often described as something akin to that which might be performed by a robin subjected to voice lessons.  The Rose-breasted Grosbeak is certainly a bird worth seeing and hearing.

Five Flowering Plants for Cleaner Water

Currently in the Lower Susquehanna River Watershed, you can find these five species of herbaceous plants in full bloom.  As they grow, they and others like them help to purify waters within their respective ecosystems by taking up nutrients—namely, the nitrogen and phosphorus that can lead to detrimental algal blooms and eutrophication in ponds, lakes, streams, and rivers.

(United States Geological Survey image by Virginia-West Virginia Science Center)
Marsh Blue Violet
The Marsh Blue Violet (Viola cucullata) is most frequently found growing in the wet soils of forest bottomlands, usually where springs first break the earth’s surface and begin slowly trickling away to form a small brook or join an existing stream.  The blooms are recognized by their darker purple centers and their long stems.
Marsh Blue Violet
This particular Marsh Blue Violet was found at 750′ altitude in the running water at a mountainside spring seep on a south-facing slope in the Ridge and Valley Province.
Soft Rush
The seldom-noticed flowers of the Soft Rush (Juncus effusus), also known as the Common Rush, emerge from the sides of its quill-like stems.  This wetland species is found in damp soils, sometimes in standing water, and grows in stiff, erect clumps that persist through winter.  When found in pastures, Soft Rush is seldom of interest to cattle or other livestock.  It therefore doesn’t lure these animals into muddy, puddle-prone areas.  When subjected to heavy grazing in dry weather and flooding during wet spells, these puddle sites may host nearly pure stands of Soft Rush, the only plant able to thrive there.  When it comes to nutrient uptake in these soggy sections of the meadow, the soft Rush is the lone ranger.  Soft Rush seeds are available from Ernst Conservation Seeds in Meadville, PA, and are included in many of their mixes formulated for stormwater management basins and other wet soil applications.
Larger Blue Flag
Larger Blue Flag (Iris versicolor) is a plant of wetlands and shorelines.  It can be grown as an emergent in ponds and lakes where it will help to absorb nutrients from both the water and the underlying substrate.
Larger Blue Flag
Larger Blue Flag is a native species in the lower Susquehanna valley.
Yellow Iris
The Yellow Iris (Iris pseudacorus), also known as the Water Flag, is native to Eurasia and Africa.  Seen here growing as an emergent among native Common Cattails (a superb water purifier), the Yellow Iris can easily escape cultivation and become invasive.  The showy flowers and water-cleansing benefits of this plant make it attractive for use in the garden or farm pond, but considerations must be made for its aggressive growth and proclivity to escape to neighboring habitats.  If you’re purchasing irises for transplanting, you’re probably better off sticking with the native Larger Blue Flag; it is far less vigorous and you’ll be able to grow other aquatic species along with it.
Spatterdock
In large ponds, lakes, and low-gradient streams, one of the best aquatic plants for sequestering nutrients and clarifying water is Spatterdock, also known as Cow-lily or Yellow Pond Lily.  Spatterdock does best as an emergent in shallow water along the shoreline.  It grows well in full sunshine and makes excellent habitat for wildlife.  Depending on the nutrient load from fish, waterfowl, decaying vegetation, and other sources, plant cover may need to be as high as 30% or more of the surface area to keep algae from overtaking a lake or large pond.  Spatterdock can often be used to help fulfill these needs while still offering open water beneath the leaves and between the stems for fish, amphibians, reptiles, and macroinvertebrates to thrive.
Spatterdock
Though probably not suitable for small garden ponds, Spatterdock (Nuphar advena) can be an excellent choice for helping to clear up the nutrient-loaded waters of a farm pond or lake.  You can find it, the irises, and Soft Rush available through some pond nurseries and garden centers.  If you can’t get them locally, check out retail and wholesale suppliers online, but remember to inspect any livestock you bring in from outside the area for hitchhikers like non-native snails (native snails are O.K.).  To be safe, always quarantine and monitor your aquatic plants for 30 days.  Tubers can be given a bath in 3% hydrogen peroxide solution for up to five minutes, then rinsed with water.  Repeat the treatment as needed until no snails or eggs are seen.  Another option: local pond owners who have them may be willing to divide some Iris and/or Spatterdock tubers and provide them for sale or gift to those who ask.  Just a couple will get you started.

Back in the Day: Down on the Farm

Let us travel through time for just a little while to recall those sunny, late-spring days down on the farm—back when the rural landscape was a quiet, semi-secluded realm with little in the way of traffic, housing projects, or industrialized agriculture.  Those among us who grew up on one of these family homesteads, or had friends who did, remember the joy of exploring the meadows, thickets, soggy springs, and woodlots they protected.

Low-intensity Farming
During much of the twentieth century, low-intensity agriculture provided a haven for wildlife.  Periodic disturbances helped maintain cool-season grassland and early successional habitat for a number of species we currently find in decline.

For many of us, farmland was the first place we encountered and began to understand wildlife.  Vast acreage provided an abundance of space to explore.  And the discovery of each new creature provided an exciting experience.

Distributed by the Pennsylvania Game Commission, artist Ned Smith’s wildlife posters introduced many residents of the lower Susquehanna region to its birds and mammals.  This poster of “Birds of Field and Garden” helped us learn what to expect and search for during our forays to the farm.

Today, high-intensity agriculture, relentless mowing, urban sprawl, and the increasing costs and demand for land have all conspired to seriously deplete habitat quality and quantity for many of the species we used to see on the local farm.  Unfortunately for them, farm wildlife has largely been the victim of modern economics.

For old time’s sake, we recently passed a nostalgic afternoon at Middle Creek Wildlife Management Area examining what maintenance of traditional farm habitat has done and can do for breeding birds.  Join us for a quick tour to remember how it used to be at the farm next door…

Barn Swallow
Always found nesting under the forebay of the barn, the Barn Swallow relentlessly pursued flying insects over the pond and meadow.
Eastern Meadowlark
Eastern Meadowlarks arrived during March and April to begin nesting in their namesake.  Their song, “spring-of-the-year”, heralded the new season.
Eastern Kingbird
Arriving in meadows and pastures during early May, the Eastern Kingbird provided for its nestlings by ambushing a variety of flying insects.  By August, congregations of these birds could be found gathering along ponds and streams ahead of their fall migration.
Orchard Oriole
In the cherry grove down by the creek, the Orchard Oriole would be singing incessantly to defend its territory.
American Goldfinches
Normally seed eaters through the colder months, American Goldfinches would regularly find a source of protein in the occupants of Eastern Tent Caterpillar nests.
Yellow Warbler
Along the wet margins of the creek, Yellow Warblers would nest in the shrubs and small trees.
Willow Flycatcher
The “Traill’s Flycatcher” was a familiar find in low-lying areas of successional shrubs and small trees.  Today, “Traill’s Flycatcher” is recognized as two distinct species, the Alder Flycatcher (Empidonax alnorum) and the Willow Flycatcher.  In the lower Susquehanna valley, the latter (seen here) is by far the most common of the two.
Eastern Bluebirds
During the nineteenth century, Eastern Bluebirds became a rarity on lower Susquehanna farms due to a combination of factors: pesticide (DDT) use, habitat loss, and competition with other birds for nest sites.  The species saw a resurgence beginning in the 1970s with discontinuation of DDT applications and widespread provision of nesting boxes.  Around human habitations, competition with invasive House Sparrows continues to be detrimental to their success.
Purple Martins and Tree Swallows
Purple Martins suffered a similar fate to the bluebirds.  The potential for their recovery remains dubious and they continue to be very local breeders, fussy about selection of suitable man-made provisions for nesting.  After considerable effort, Purple Martins have at last been attracted to nest in the condos placed for their use at Middle Creek Wildlife Management Area.  In the artificial gourds, there are nesting Tree Swallows, a species which also benefits from the placement of boxes intended for bluebirds.
Field Sparrow
Abandoned fields and other successional habitats were and continue to be favored homes for Field Sparrows.
Cedar Waxwing
At almost any time of year, roving bands of Cedar Waxwings would suddenly visit old field habitat looking for berries among the shrubs and other pioneering woody growth.  In early summer, after most species have already hatched their young, nesting would commence and these fruit eaters would transform into accomplished fly catchers.
Ring-necked Pheasant
During the twentieth century prior to the 1980s, Ring-necked Pheasant populations in the Lower Susquehanna River Watershed were comprised of breeding descendants of introduced birds supplemented by additional releases to maintain numbers sufficient for hunting.  Year-round populations did and can reside in mosaic landscapes of early successional and grassland habitats, the latter including hay fields left unmowed through the nesting season.
Red-winged Blackbird
Red-winged Blackbirds have always been a fixture of hay fields and meadows on farms.  While the increase in mowing frequency has reduced their nesting success, they have persevered as a species by nesting earlier than other birds and by utilizing other landscape features such as densely vegetated stormwater basins for breeding sites.
Bobolink
Do you recall the last time you saw a Bobolink nesting in a hay field near you?  Arriving in early May as a Neotropical migrant, the Bobolink requires a cool-season grassland such as hay field through at least July to complete its nesting cycle.  Even earlier this century, we remember nesting Bobolinks being more widespread on farms throughout the region.  Now, you almost have to go to Middle Creek if you want to see them.
Grasshopper Sparrow
Formerly more widespread in hay fields throughout the lower Susquehanna valley, the native Grasshopper Sparrow is yet another species falling victim to early mowing and intensive farming.
Grasshopper Sparrow
The solution to their dilemma is as advertised.  Instead of cutting the grass, why not take heed of the example set here and cut back on the tens of thousands of acres that are excessively or needlessly mowed in the Lower Susquehanna River Watershed?  How ’bout letting a significant percentage of your property regenerate as successional habitat as well?  It can and does make a difference!
Cool-season Grasses
Beautiful cool-season grasses waving in the spring breeze.  Meadows and hay fields can be managed to function as cool-season grasslands to provide nesting opportunities for many of the species we used to find down on the farm.

Photo of the Day

White-tailed Deer Button Buck Hosting Engorged Ticks
When it comes to sitting down and watching an episode of Hollywood Squares or indulging in a game of Tic-Tac-Toe, those of us here at susquehannawildlife.net are quick to play right along.  But when it comes to mixing it up with a Tick-Backed Doe or a Tick-Backed Buck like this one, you can count us out.  Upon finding this button buck infested with engorged Deer Ticks and heading our way on a grassy trail, we decided to turn around and limit our walk to the gravel roadway where there was less of a chance of picking up any hitchhikers from the vegetation where this guy has been spending his time.

Nest Builders at Work

For many animals, an adequate shelter is paramount for their successful reproduction.  Here’s a sample of some of the lower Susquehanna valley’s nest builders in action…

Pileated Woodpecker Excavating Nest
Many of our year-round resident bird species get a head start on the breeding season as cavity nesters.  Some of these mated pairs use naturally occurring hollows, while still others take advantage of the voids left vacant by the more industrious previous occupants.  Woodpeckers in particular are responsible for excavating many of the cavities that are later used as homes by a variety of birds and mammals to both rear their young and provide winter shelter.  Pileated Woodpeckers, like other members of the family Picidae, have an almost mystic ability to locate diseased or insect-infested trees for selection as feeding and nesting sites.  In this composite image, a pair is seen already working on a potential nursery during mid-January.  After use by the woodpeckers, abandoned cavities of this size can become nesting sites for a variety of animals including bees, small owls, Great Crested Flycatchers, Wood Ducks, Hooded Mergansers, and squirrels.
Downy Woodpecker at Nest
After use as a nesting site, a void excavated by Downy Woodpeckers can be occupied in subsequent years by chickadees, titmice, nuthatches, wrens, and other cavity-dwelling species.
Muskrat with Leafy Twig
This Muskrat (Ondatra zibethicus) with a leafy twig in tow takes advantage of spring’s new growth to construct or repair its house,…
Muskrat with Leafy Twig
…a process that can be repeated or renewed as necessary throughout the year.
Muskrat House
A Muskrat house in March.  In the absence of leafy twigs, dried cattail stems will suffice.  As it ages and decays, the house’s organic matter generates heat and makes an ideal location for turtles to deposit and hatch their eggs.
Wood Thrush
Soon after Neotropical migrants begin arriving in the forests of the lower Susquehanna watershed, they begin constructing their nests.  The majority of these species build “outdoors”, not within the confines of a tree cavity.  Here we see a Wood Thrush with its bill full of dried leaves and other materials…
Wood Thrush Nest
…ready to line the cup of its nest in the fork of a small understory tree.
Blue-gray Gnatcatcher
Though it often arrives during early April after spending the winter in sub-tropical and even some temperate climes, the Blue-gray Gnatcatcher must wait to start construction of its nest until many of the Neotropical migrants arrive in early May.
Blue-gray Gnatcatcher
You see, the Blue-gray Gnatcatcher relies on plenty of web-spinning spider activity to supply the construction materials it needs.
Blue-gray Gnatcatcher
A Blue-gray Gnatcatcher pulling apart a spider’s web on a warm May morning.
Blue-gray Gnatcatchers
Back at the nest site…
Blue-gray Gnatcatcher
…the sticky spider webs bind together lichens and small bits of bark…
Blue-gray Gnatcatcher
…to form a perfect little cup for the nesting Blue-gray Gnatcatchers.
Baltimore Oriole with Nesting Material
Baltimore Orioles weave one the most unique nests of any species occurring in eastern North America.
Baltimore Oriole Nest
Unfortunately for them, man-made litter can often seem to be the ideal material for binding the nest together.  In an area only sporadically visited by anglers, this oriole had no trouble finding lots of monofilament fishing line, trash that can fatally entangle both adult and young birds of any species.  If you see any fishing line at all, please pick it up and dispose of it properly.
Baltimore Oriole
Always keep an eye open for fishing line and get it before the birds do!
Brown-headed Cowbirds
Like many other avians, male Brown-headed Cowbirds are now relentlessly pursuing females of their kind.
Brown-headed Cowbirds
All his effort is expended in an attempt to impress her and thus have a chance to mate.
Brown-headed Cowbirds
This male can indeed put all his energy into the courtship ritual because Brown-headed Cowbirds toil not to build a nest.  They instead locate and “parasitize” the nests of a variety of other songbirds.  After mating, the female will lay an egg in a host’s abode, often selecting a slightly smaller species like a Yellow Warbler or native sparrow as a suitable victim.  If undetected, the egg will be incubated by the host species.  Upon hatching, the larger cowbird nestling will dominate the brood, often ejecting the host’s young and/or eggs from the nest.  The host parents then concentrate all their efforts to feed and fledge only the young cowbird.
Indigo Bunting
Watching and waiting.  The Indigo Bunting evades cowbird parasitism by first recognizing the invader’s egg.  They then either add a new layer of nest lining over it or they abandon the nest completely and construct a new one.  Some patient buntings may delay their breeding cycle until after cowbird courting behavior ceases in coming weeks.

Their Songs Give Them Away

After repeatedly hearing the songs of these Neotropical migrants from among the foliage, we were finally able to get a look at them—but it required persistent effort.

Magnolia Warbler
Often found closer to the ground among shrubs and small trees, this Magnolia Warbler was a challenge to find in the upper reaches of a Chestnut Oak.
Yellow-rumped Warbler
Not far away, we spied this Yellow-rumped Warbler.
Red-eyed Vireo
Despite its status as probably the most common Neotropical songbird to nest in the deciduous woodlands of the lower Susquehanna valley, the Red-eyed Vireo is nevertheless notoriously difficult to locate among the leaves.
Ovenbird
The Ovenbird spends much of its time on the forest floor where it builds a domed, oven-like nest.
Ovenbird
A fortunate observer may catch a glimpse of one perched in an understory shrub or small tree.
Ovenbird
But hearing the Ovenbird’s song, “teacher-teacher-teacher”, is frequently the only way to detect it.
Worm-eating Warbler
The Worm-eating Warbler nests in understory thickets on steep forested slopes.  Its rich chipping song is often the only indication of its presence.
American Redstart
Frequently easier to locate is the American Redstart, a Neotropical warbler that calls out its territory in damp woodlands from perches atop shrubs or among the lower limbs of trees.
Warbling Vireo
Warbling Vireos nest near streams or other bodies of water in large deciduous trees like this Northern Hackberry.
Gray Catbird
Gray Catbirds are abundant in shrubby gardens and thickets.  Most will come out of hiding to investigate disturbances like an observer making a pishing sound.
Common Yellowthroat
Another inhabitant of brushy successional growth is the Common Yellowthroat.  It too is curious and responds quickly to squeaky sounds made by human visitors to their home ground.
Yellow-breasted Chat
The Yellow-breasted Chat is a bird of early successional growth.  To establish and defend a breeding territory, this one adopted a perch along the edge of a woodlands overlooking a field in which prescribed fire was administered less than two months ago.
Yellow-breasted Chat
Like the mockingbird and thrasher, the Yellow-breasted Chat is a mimic.  Its song is a repertoire of the calls and songs of the bird species with which it may compete for food and nesting space.  Unique to the chat is an occasional pause, whistle, or “chuck” note that creates a bridge between many of its song’s elements.
Yellow-breasted Chat
The intensely territorial chat dropping down to look us over.

Sometimes we have to count ourselves lucky if we see just one in five, ten, or even twenty of the birds we hear in the cover of the forest canopy or thicket.  But that’s what makes this time of year so rewarding for the dedicated observer.  The more time you spend out there, the more you’ll eventually discover.  See you afield!

Scarlet Tanager
A singing Scarlet Tanager lurking in the shade of an oak.
Scarlet Tanager
Seeing is believing.

Singing in the Rain

Neotropical birds are fairly well acquainted with repetitive periods of thundershowers.  With that in mind, we decided not to waste this stormy Tuesday by remaining indoors.

Thundershower
Periods of rain need not put a damper on a day outdoors observing birds that wintered in a rain forest or other tropical environment.
Utility Right-of-way with Successional Habitat
We hiked this utility right-of-way to the north of a heavy thunderstorm and found plenty of activity in the shrubby successional habitat there.
Indigo Bunting
Rain or shine, male Indigo Buntings were busy singing.  All this exuberance is intended not only to establish and defend a nesting territory…
Indigo Bunting
…but to attract the attention of a mate as well.
Blue-winged Warbler
Prior to the implementation of the intensive manicuring practices we see currently applied to most utility right-of-ways, shrubby thickets filled miles of these linear corridors to create a webbed network of early successional growth throughout the lower Susquehanna valley.  Loss of this specialized habitat has led to the almost total elimination of the formerly common Blue-winged Warbler as a breeding species here.
Blue-winged Warbler
We found three male Blue-winged Warblers singing on territories in this bushy clearing where electric transmission lines pass over Third/Stony Mountain on State Game Lands 211.
Blue-winged Warbler
Their presence at this site is testament to the importance of maintaining corridors of quality successional habitat in the landscape.
Blue-winged Warbler
The Blue-winged Warbler is a Neotropical migrant with an easy-to-learn song.  It’s a very simple, buzzy sounding “beeee-bzzz”.
Prairie Warbler
Another Neotropical species that nests in successional thickets is the Prairie Warbler (Setophaga discolor), seen here during one of this morning’s downpours.
Prairie Warbler
The Prairie Warbler can sometimes be found in stands of pioneer plants like Eastern Red Cedar on sites with barren soils or those that have been subjected to wildfire.
Hooded Warbler
The Hooded Warbler is fond of wooded thickets along the edge of forested land such as those in the Third/Stony Mountain utility right-of-way.
Hooded Warbler
The presence of a Mountain Laurel thicket also enhances a forest’s ability to host breeding Hooded Warblers.

We hope you enjoyed our walk in the rain as much as we did.  If you venture out on a similar excursion, please remember this.  The majority of the wild animals around us have busy lives, particularly at this time of year.  Most don’t take a day off just because it rains—that includes ticks.

Female Deer Tick
Be certain to check yourself for ticks, especially these very small Deer Ticks (Ixodes scapularis), also known as Black-legged Ticks.  Deer Ticks are vigilantly looking for something to latch onto, even in the rain, and they can be vectors of Lyme disease.  We found this adult female as a stowaway on the editor’s neck just before heading home from today’s stroll.