Some comparative attributes of unspoiled lotic vs. lentic freshwater ecosystems. Low-gradient (slow-moving) lotic waters often create, and remain connected to, accompanying riverine wetlands (a lentic freshwater ecosystem). These swamps, marshes, and ditches absorb, purify, and infiltrate flood waters while supporting a diverse number of plant and animal species.
We frequently perceive all waterfowl migration to be synchronized with the conspicuous movements of familiar species like Snow Geese, Tundra Swans, and Canada Geese—big flights coming south in October and November, then a return to the north in late February and March. And we’re all quite aware of the occurrence of large gatherings of some of these migrants while they make stopovers on some of our largest lentic (still) waters—the man-made lakes and reservoirs created by damming local streams. But did you know that there are populations of colorful waterfowl with dynamic migrations that extend throughout the winter and early spring with movements that are often continuous. Under favorable conditions, these birds favor the lotic (flowing) waters of the river and its larger tributaries as they transit the lower Susquehanna valley. That’s because unpolluted lotic freshwater ecosystems support a greater diversity of plants and animals than lentic waters, and therefore offer more opportunities for hungry migrating waterfowl to find food. Let’s have a look at some of the species that visit the river during their seasonal journeys…
While the urge to head south in the autumn is largely stimulated by the shortening of the photoperiod, it is the presence of ice, particularly on glacial lakes throughout the lands to the north of the lower Susquehanna River basin, that pushes many diving ducks to finally make their way south toward the guarantee of open-water feeding areas along the coast. This movement may occur at anytime between November and late February or, as we have seen during some of the mild winters of recent decades, it may scarcely be noticed at all.Common Mergansers often lead they way when it comes to migratory diving ducks. They regularly move south in conspicuous numbers by late November and December and are regularly pushing north as soon as the ice begins to melt. During a typical year, it is not unusual for some populations of these large diving birds to remain north of us during the winter. Then, in the days after a sudden rush of frigid polar air, an appreciable increase in ice cover will force a mid-winter movement of birds down the Susquehanna.Temperatures in lotic fresh waters vary over the length of the stream or river. They are largely determined by the collective impact of the numerous sources of heat flux depicted in this graphic. (Environmental Protection Agency image)Buffleheads begin passing through the lower Susquehanna region in Novemeber on their way to coastal saltwater bays for the winter. Lingering populations feed by diving in the river’s pools and riffles for benthic invertebrates including snails and insect larvae. Lesser quantities of aquatic plant matter supplement their diet.Many Common Goldeneyes will remain on shallow, ice-free waters of the northern lakes and rivers sculpted by the most recent glacial event, but only until they are forced south into and through the lower Susquehanna valley by the encroachment of freezing conditions. On the river, they are among the dozen or so species of diving ducks we see visiting or passing through during the typical late fall and early river.
During their visits to the lotic (also known as riverine) fresh waters of the Susquehanna and its largest tributaries, benthic-feeding waterfowl make short dives to take advantage of the plants, small fish, invertebrates, and other food sources inhabiting the stream bottom in the riffles and pools of the free-flowing waterway. Substrates, listed here by size (in descending order), along with other parameters influenced by this zonation determine the variety and abundance of the forage available to migrating waterfowl and other consumers. Ice or high water and poor visibility due to flooding can render the riffles and pools of the channel unusable for feeding. The birds must then choose to either linger and rest without feeding or leave the lotic freshwater habitat to seek sustenance. During a flood, this may require relocation to a nearby lentic (still) body of fresh water such as a lake or reservoir. The presence of ice will almost invariably force the birds to fly on to the Atlantic Coastal Plain and the tidal waters of its bays and estuaries.Hooded Mergansers are one of the few species of diving ducks likely to utilize flooded shoreline timber and riverine (fluvial) wetlands as refuge from high water on Susquehanna.A Pied-billed Grebe and a pair of Canvasbacks on an ice-free stretch of the lower Susquehanna in mid-winter feed and loaf in a riffle-flanked pool where a large mat of American Eelgrass, a submerged aquatic plant also known as Tapegrass or Wild Celery, grows during the summer. The vast mosaic of riffles and pools in a river this size offers tremendous opportunities for a diverse array of aquatic species to find their niche wherein they can survive and flourish.The presence of ice forces Buffleheads and other diving ducks to gather in turbulent open water, often below a riffle or dam. Another alternative is to continue on toward the salty bays and estuaries of the coast. High water may push these birds into the shallows among the flooded woods to feed, but they seldom utilize heavily forested riparian wetlands as a refuge due to their need for a running start to get airborne.
Riffle and Pool Characteristics of High Gradient (A) and Low Gradient (B) Streams. This graphic illustrates the change in deposition characteristics of a stream or river as its gradient decreases. A high-gradient stream (A) has a rapid velocity, often forms falls, and tends to carry away a high volume of all but the largest of particles of potential substrates as they erode from the surrounding landscape. On a low-gradient stream, the loss in water velocity reduces the water column’s ability to transport even the smallest of substrate particles. Deposition of this gravel, sand, silt, and clay forms lateral bars that over time create the familiar meandering path of a naturally flowing lowland stream. (National Park Service image)Benthic substrates in lotic freshwater pools and riffles support an abundance of life forms ranging from colorful diatoms on rocks and cobble, to invertebrates including snails and insect larvae, to fishes like this young Channel Catfish. Free of accumulations of sediment, this river bottom not only provides habitat for a healthy fishery, it facilitates the bidirectional exchange of water between the Susquehanna and its underlying aquifer.On the lower Susquehanna, populations of young Quillback suckers are found almost exclusively in clear, high-gradient pools.The Harlequin Duck winters along the rocky shores and man-made jetties of the Atlantic coast. In summer, they nest on fast-moving, headwater streams well to our north. Very rare on the Susquehanna, this is the first of two individuals found during March and April of 2025. It was observed feeding in the swift waters of the high-gradient riffles and pools where the river cuts through Blue Mountain north of Harrisburg. During previous weeks, Harlequin Ducks were being seen along the coast as far south as the mouth of the Chesapeake at Cape Charles, Virginia. It’s very possible that some of these birds traveled north through the bay area and up the Susquehanna on their way north.
Fluvial Geomorphology of a Stream. Many of the Susquehanna’s tributaries pass through each of these three erosional zones. Along the way, they carry out the process of breaking down the mountains formed by the Allegheny orogeny, the collision of North America and Africa that created the supercontinent Pangea about 325 to 260 million years ago (during portions of both the Carboniferous and Permian periods). Today’s main stem of the lower Susquehanna passes through a transfer zone (Zone 2), carrying eroded materials to a depositional zone (Zone 3) located within the ancient Susquehanna canyon stretching from Havre de Grace, Maryland, to Norfolk Canyon on the edge of the continental shelf. Within this zone, more than a 10,000-year accumulation of post-glacial sediments lies submerged by the rising waters of the Atlantic Ocean and Chesapeake Bay. Although the present-day lower Susquehanna is largely a transfer zone, some deposition occurs along low-gradient segments of the river, particularly where its course parallels the watershed’s ridges both above and below the high-gradient rapids where its path has eroded passage through the highlands. (National Park Service image)Although the present-day lower Susquehanna is largely a transfer zone, some deposition occurs along low-gradient segments of the river, particularly where its course parallels the watershed’s ridges both above and below the high-gradient rapids where its path continues to erode passage through the bedrock. On this 1908 map of the Susquehanna at Conewago Falls, alluvial terraces of gravel, sand, silt, and clay can readily been seen as pale areas nearly lacking brown contour lines along the shorelines and islands of the river. Deposits within most of these terraces date back to the melt period following the most recent glacial event and beyond. The delta shown at the mouth of Conewago Creek (west) includes massive volumes of material deposited by both the creek and the river. This delta is currently known as Brunner Island, much of it developed as the site of a coal/gas-fired electric generating station. Terrace deposits along the Susquehanna’s shorelines created extensive perched marshes and swamps (wooded marshes) fed by rains, high river water, small streams, and springs, the latter often seeping from the base of the rocky escarpments carved by the ancient Susquehanna and defining the present-day inland border of the floodplain. We call these sites “Alluvial Terrace Wetlands”. Few of these critical components of river morphology survive. Those not drained for farmland were obliterated by urbanization and canal, railroad, and highway construction. (United States Geological Survey base image: Middletown, PA, quadrangle, 1908)Water Willow is a familiar emergent plant that colonizes lateral bars and other alluvial deposits in low-gradient segments of the Susquehanna.The fine roots of Water Willow collect sediment and absorb nutrients while creating dense cover for young fish and numerous species of invertebrates including the Virginian River Horn Snails seen here.
Prior to the nineteenth century, the low-gradient flow regime of the river both above and below the riffles at Chiques Rock (lower right on map) created prime wildlife habitat. The natural accumulations of nutrients and substrates carried into and through the lotic waterway’s pools and riffles were cycled into an ideal growing medium for extensive mats of American Eelgrass and other aquatic plants. This underwater forest hosted a seemingly endless abundance of invertebrates and fishes (both resident and migratory)—supporting a variety of consumer species including various populations of humans. But soon after the mass clearing of much of the watershed’s land for farming and lumber, the mill ponds created by dams constructed on streams to power saw and grain mills became brimful with sediments eroded from the unprotected ground. During storm events, torrents of these sediments then flowed full bore toward the Susquehanna, and began accumulating in the low-gradient segments of the river.
Sediments left behind after the removal of mill dams are known as legacy sediments. They disconnect the stream from its historic floodplain and riverine (fluvial) wetlands, thus intensifying the impact of high water in the surrounding landscape. As these nutrient-charged deposits wash away, they become a source of pollution in the waters of the Susquehanna and Chesapeake.A mat of American Eelgrass growing in the flowing waters of the Susquehanna below Conewago Falls. Eelgrass and other submerged aquatic plants provide essential habitat for a wide variety of small fish and invertebrates while also consuming nutrients deposited in the cobble, gravel, and sand substrate of the river’s pools. Excess quantities of smaller particles of silt and clay can clog the substrate and thus inhibit the hyporheic exchange of water between the stream channel and the underlying aquifer, often diminishing the biomass and diversity of organisms inhabiting this benthic habitat. Buried in these life-choking sediments, the river bottom becomes inhospitable to growth of submerged aquatics including eelgrass.This low-gradient stretch of the Susquehanna at Marietta flows parallel to the Chickies Quartzite “Hellam Hills” before making a sharp right turn to punch through the ridge as a series of rapids at Chiques Rock. Formerly a fully functional lotic ecosystem and a paradise for migrating waterfowl, this river segment is now impaired by accumulations of nutrient-laden sediments from agricultural and urban runoff.Nutrient and sediment-loaded flood waters roar across the diabase boulders at the York Haven Dam and Conewago Falls, a high-gradient segment of the Susquehanna.They continue past Brunner Island (left power plant stacks) and through Haldeman Riffles and the Shocks Mills Railroad Bridge……into the stretch of river known as the “Marietta broads” along the base of the Chickies Quartzite “Hellam Hills”. The low stream gradient here produces a slower current and increased deposition of sediments.As the flood surges through the riffle and pool complex at Chiques Rock, the high stream gradient maintains a velocity in the water column sufficient to keep additional sediments in suspension until they reach the low-gradient river segment just downstream at Washington Boro, site of a naturally occurring lateral bar area known as the Conejohela Flats. These bars now lie within the man-made depositional zone known as “Lake Clarke”. Created nearly a century ago by construction of the Safe Harbor Dam, this impoundment is accumulating astounding volumes of nutrient-loaded sediments that continue to encapsulate the flats within a stream-impairing delta.Anytime from November to April on the lower Susquehanna, a group of Redheads, Canvasbacks (3 birds to right of the middle of the picture), and a Horned Grebe (lower left) is a welcome sight in a riverine pool known to have a summertime growth of American Eelgrass. Noted Dr. Herbert Beck in 1924 when describing the Canvasback, “Like all ducks, …, it stops to feed within the county (Lancaster) less frequently than formerly, principally because the vast beds of wild celery which existed earlier on broads of the Susquehanna, as at Marietta and Washington Borough, have now been almost entirely wiped out by sedimentation of culm. Prior to 1875 the four or five square miles of quiet water off Marietta were often as abundantly spread with wild fowl as the Susquehanna Flats are now. Sometimes there were as many as 500,000 ducks of various kinds on the Marietta broad at one time.”Today, seeing just dozens of Aythya genus ducks (Redheads, Canvasbacks, scaup) on the lower Susquehanna is a notable event. If they happen to be forced down by inclement weather while migrating through, you might get lucky enough to see several hundred.While the recovery of eelgrass/wild celery beds on the Susquehanna is trivial in scale and offers little support for numbers of waterfowl to return to historic levels, restorations on the upper Chesapeake in the vicinity of the Susquehanna Flats between Havre de Grace and Aberdeen Proving Grounds may have helped refuel a gathering of mostly Aythya genus ducks during the final days of February. This mass of ducks, many of which were forced south from the frozen Great Lakes during the previous weeks (some by way of the ice-choked Susquehanna) were apparently making an abrupt turn to make their way back north. Their stay was brief, but estimates by local birders put their numbers as high as one half million. The vast majority of the concentration consisted of Aythya species: Redheads, Canvasbacks, Ring-necked Ducks, and both Lesser and Greater Scaup. It probably included a mix of birds including both northbound migrants from further down the coast and the aforementioned refugees that had just arrived to pay a quick visit while escaping the late-season ice before turning around.During the past two centuries, as food supplies in the Susquehanna grew increasingly compromised for benthic feeders like these Lesser Scaup and other diving ducks, a change in distribution was necessary for survival.As individual species, Lesser Scaup and other waterfowl that fail to adapt to natural or man-made changes in their habitats and food supplies may see their overall global numbers falter.
Despite being located in the transfer zone, the lower Susquehanna has become a significant depositional zone along much of its length, mostly courtesy of the placement of sediment-trapping man-made dams.
Following construction of the mill dams and ponds on nearly every mile of the lower Susquehanna’s low-gradient tributary streams, enterprising parties moved on to the river. The first significant spans were constructed using wide timber cribs filled with large rock. They were placed to create water deep enough to allow canal boats to cross the Susquehanna at both Clark’s Ferry at the mouth of the Juniata River in Dauphin County and at Columbia/Wrightsville. These dams also diverted water into the newly excavated canals—the Pennsylvania Eastern Division Canal (completed in 1833) which followed the river’s east shore from Clark’s Ferry to Columbia, and the Susquehanna and Tidewater Canal (completed in 1840) along the west shore from Wrightsville to Havre de Grace, Maryland. Placement of these sediment-trapping man-made dams began a process of converting vast mileage of the lower Susquehanna from a transfer zone into a deposition zone. In addition, layout of the canals and locks followed the contours along the base of the riverside ridges, seriously altering most of the alluvial terrace wetlands that the river had created as a feature of its floodplain during the post-glacial period.
Construction of the canal dams was just the beginning. During the twentieth century, more massive dams would be added to the main stem of the river for hydroelectric energy production at York Haven, Safe Harbor, Holtwood, and Conowingo. Upon their completion, the days of unassisted anadromous fish migrations were over. On both the river and its tributaries, smaller dams including dangerous low-head dams maintain water levels for boating and recreation. They too create current-diminishing, pseudo-lentic waters that blanket the lotic riffle and pool substrates with polluted sediments.
MAN-MADE DAMS TURN LOTIC WATERS INTO UNFLUSHED TOILETS
The construction of dams on the lower Susquehanna has converted vast mileage of the river from a lotic freshwater system into a series of man-made lentic freshwater lakes. These areas have lost their function as a lotic transfer zone and are now a sort of dysfunctional series of depositional zones collecting vast volumes of sediment containing nutrients and other pollutants. Within each impoundment, the reduced velocity of the river causes it to drop suspended sand first, then the finer particles of silt and clay closer to the dam. The flow regime of riffles and pools is lost and the hyporheic zone that exchanges water between the river and the underlying aquifer is clogged. These impaired segments of river become ripe for eutrophication: algal blooms followed by die offs that can lead to a fatal reduction of dissolved oxygen in the water column.Deposits of lateral bars of sediment in low-gradient segments of the Susquehanna can create shallow water feeding habitat for puddle/dabbling ducks like these Gadwall. Where sediment pollution is severe, benthic foods in these areas often consist mostly of invertebrates and plant matter deposited by the current, the buried substrates devoid of a functioning ecosystem and the waters subject to eutrophication.Common Goldeneyes on a patch of open water on an otherwise ice-covered “Lake Clarke”, the impoundment created by Safe Harbor Dam. While they may find this spot advantageous for loafing, the food supply over the sediment-buried substrate will be limited.By the end of the twentieth century, accumulations of polluted sediments behind lower Susquehanna dams were nearing capacity. There is no working plan to attenuate the massive release of these pollutants that may be triggered by a catastrophic flood. The effort to reduce nutrient and sediment runoff remains the focus so that new loading is kept to a minimum and won’t add to the capacity problems at the dams or continue downriver to the Chesapeake at full strength when the dams are full. Alleviating the sediment aggregation problem within the river’s impoundments is a tall order and a dilemma not easily solved. (United States Geological Survey image)Common Mergansers will feed where benthic substrate supports the small fish and invertebrates they prefer. They will, however, gather in extraordinary numbers on the “lakes” created by riverine dams. Though they can only feed on what floats in with the current, hundreds or sometimes thousands of Common Mergansers will concentrate on “Conowingo Pond” during the late fall or early winter. There, safety in numbers gives them some guarantee of protection against the multitudes of eagles that simultaneously frequent the vicinity. Another advantage of staging on “Conowingo Pond” is its close proximity to favorable feeding areas on upper Chesapeake Bay and stretches of the Susquehanna where lotic riffles and pools offer abundant opportunities below the river’s dams.Fortunately for everything else living in the benthos, Common Mergansers are big enough to devour invasive, non-native Rusty Crayfish when they find them in our lotic waterways.
TIME TO CLEAN UP OUR ACT
WHERE DOES YOUR STORMWATER GO?
Channelized Urban/Suburban Streams Function as Sewers. They have no attached lowlands or floodplains to absorb, purify, and infiltrate runoff from rain events. Pollutants including litter, pet waste, lawn chemicals, tire-wear particles, hazardous fluids, and sometimes untreated human excrement flush unchecked from the municipal storm drainage system into the waterway. Thermal shock from summer downpours washing across sun-heated pavements can kill temperature sensitive fishes and other aquatic life. Nutrient and sediment loads from these impaired tributaries later accumulate downstream in low-gradient segments of the Susquehanna, turning the river into an open-air cesspool. Aggressively working to implement projects that eliminate these sources of pollution are the only effective way to keep the problem from getting worse. Making things better requires a lot more dedication and effort. (United States Geological Survey image by Frank Ippolito)
RIPARIAN BUFFERS MAKE A DIFFERENCE…WIDER IS BETTER
To sequester sediment and cycle nutrients (primarily nitrogen and phosphorus) contained in farm runoff, the U.S.D.A. recommends installing riparian buffers between streams and lands used for grazing and raising crops. To protect pollinating species including bees and butterflies from pesticide drift and eroding soils rich in fertilizers, they further recommend installing a stand of wind-pollinating plants such as conifers, oaks, and birches between the field and streamside plantings. These same conservation practices improve water quality and wildlife habitat on waterways located in residential and commercial areas as well. (United States Department of Agriculture image, click to enlarge)
FLOODPLAIN RESTORATION
Regardless OF HOW LONG YOU’VE BEEN CONDITIONED TO THINK OTHERWISE, THIS IS A DYSFUNCTIONAL, POLLUTED CREEK—AND IT NEEDS HELP
A channelized low-gradient stream eroding a path through deposits of legacy sediments displaces flood waters into previously unaffected areas and provides a continuing source of nutrient and sediment pollution during storm events. These impaired waters have a diminished capacity for supporting aquatic life including fishes.
RESTORED TO ITS HISTORIC FUNCTIONS
On an adjacent segment of the same creek, this legacy sediment removal project restored a braided meandering channel and connected it to its newly liberated, historic floodplain. In just their second year, the fluvial wetlands are effectively absorbing stormwater and sequestering nutrients as an attached component of the stream’s riffle and pool complex. During our visit earlier this week, we found American Toads, Northern Leopard Frogs (Lithobates pipiens), and Northern Spring Peepers breeding here. It’s just as Castor canadensis would have it!
“STOP HEMMING AND HAWING AROUND ALREADY”
“HEY COWBOY, HOW ‘BOUT GETTIN’ THEM FILTHY LITTLE DOGIES OUTTA DAT CRICK?”
Here’s a polluted stream in a pasture with grazing livestock. The site is a former mill pond within which the creek eroded a channel following removal of the dam. The animals defecate and urinate where access to water is gained at a broken down embankment of the nutrient-loaded legacy sediments deposited in the pond more than a century ago. It’s a haphazard form of animal husbandry and a reminder that all it takes is just one stubborn jackass to foul up the whole waterway.
DIRECT SOURCES OF NITROGEN (AMMONIA) POLLUTION IN STREAMS
DID YOU KNOW that a dairy cow produces about 80 pounds of waste (excrement and urine) every day?
DID YOU KNOW that a horse produces about 50 pounds of waste (excrement and urine) every day?
DID YOU KNOW that a human produces about 3 to 4 pounds of waste (excrement and urine) every day? The exceptions, of course, are those who continue to insist that raising farm animals in and alongside a body of water is okey-doke—a harmless practice. These individuals tend to retain the former constituent of human waste and are thus full of it.
“ATTABOY TEX, THAT’S MORE LIKE IT!”
Now that’s better. Legacy sediments have been removed to reconnect the stream to its floodplain. A livestock crossing and exclusion fencing has been installed, and a nutrient-consuming riparian buffer has been planted. This creek segment’s pollution woes have been mitigated. Do you have a neighbor needing this type of remedial work on their farm? Have them call your local conservation district office for advice. Some programs include financial assistance covering the costs of installation as well as monetary incentives for helping to clean up the water.
AND FINALLY
WHEN IT COMES TO BUILDING DAMS ON LOTIC FRESH WATERS…
…LEAVE IT TO THE BEAVERS
North American Beavers (Castor canadensis) create habitats that connect the riffle and pool regime of a low-gradient stream to a surrounding fluvial wetland that retains sediments, cycles nutrients, and provides essential habitat for hundreds of plant and animal species. Floodplains are for flooding. And if a beaver floods an area, you can be guaranteed that it was already part of a floodplain. You see, beavers don’t encroach upon humans, it’s humans doing the encroaching upon beavers. (National Park Service image)
MAD HATTERS
DID YOU KNOW that even before the landscape was cleared for farms and a supply of timber, and before mill dams on local creeks began accumulating soil runoff from the consequently barren hillsides, all the North American Beavers, the keystone species of lower Susquehanna stream ecology, were killed and sold to make hats? It’s no wonder things are fubar!
COMING SOON…
Horned Grebes are regular migrants and sometimes winter residents on ice-free stretches of the lower Susquehanna. They spend their time plying the benthic substrate of the river’s clear riffles and pools for a variety of invertebrates and small fish. Look for them moving north in coming days sporting this beautiful breeding plumage.April and early May are prime time for observing Common Loons on the Susquehanna as they undertake a journey from the Atlantic surf where they spent the winter to nesting sites on northern lakes. For this migrant in breeding plumage, clear water for sighting plenty of benthic life in the river’s riffles and pools assures a successful dive in search of energy-replenishing forage.
You may find this hard to believe, but during the colder months in the lower Susquehanna valley, gulls aren’t as numerous as they used to be. In the years since their heyday in the late twentieth century, many of these birds have chosen to congregate in other areas of the Mid-Atlantic region where the foods they crave are more readily available.
As you may have guessed, the population boom of the 1980s and 1990s was largely predicated on human activities. These four factors were particularly beneficial for wintering gulls…
Disposal of food-bearing waste in open landfills
High-intensity agriculture with disc plowing
Gizzard Shad population boom in nutrient-impaired river/Hydroelectric power generation
Fumbled Fast Food
Earthworms lifted to the soil’s surface by plowing attracted Ring-billed Gulls by the thousands to Susquehanna Valley farmlands during the late twentieth century. (Vintage 35 mm image)Filter-feeding Gizzard Shad populations thrive in nutrient-rich waters like the Susquehanna. Their rambunctious feeding style stirs up benthic sediment deposits to release more nutrients into the water column and promote the algal blooms that often lead to detrimental eutrophic conditions. Decades ago, hundreds and sometimes thousands of gulls gathered below the river’s hydroelectric dams to feed on the seemingly endless supply of small Gizzard Shad disoriented by their passage through the turbines during electric generation. (Vintage 35 mm image)
So what happened? Why are wintering gulls going elsewhere and no longer concentrating on the Susquehanna? Well, let’s look at what has changed with our four man-made factors…
A larger percentage of the lower Susquehanna basin’s household and food industry waste is now incinerated/Landfills practice “cover as you go” waste burial.
Implementation of “no-till” farming has practically eliminated availability of earthworms and other sub-surface foods for gulls.
The population explosion of invasive Asiatic Clams has reduced the Gizzard Shad’s relative abundance and biomass among filter feeders.
Hold on tight! Fast food has become too expensive to waste.
The non-native population of Asiatic Clams in the Susquehanna and most of its larger tributaries exploded during the last two decades of the twentieth century. Its present-day biomass in the river is exceeded by no other macroinvertebrate species. The share of plankton and other tiny foods that the Asiatic Clam harvests from the water column is no longer available to native filter-feeders including Gizzard Shad. Hence, Gizzard Shad biomass has been reduced and far fewer are available to attract amazingly enormous flocks of hungry gulls to hydroelectric dams. (Vintage 35 mm image)
GULLS THIS WINTER
Despite larid abundance on the lower Susquehanna not being the spectacle it was during the man-made boom days, an observer can still find a variety of medium and large-sized gulls wintering in the region. We ventured out to catch a glimpse of some of the species being seen both within the watershed and very nearby.
By far, most of the gulls you’ll encounter in the Lower Susquehanna River Watershed right now are Ring-billed Gulls. Some of them still drop by at the business end of the drive-thru lane looking for a lost order of fries and maybe a cheeseburger in paradise.First-winter (left), second-winter (center), and adult Ring-billed Gulls on the Susquehanna.Nearly every flock of gulls found in our area right now is composed exclusively of Ring-billed Gulls. The trick to finding other species, particularly rarities, is to look for slightly larger birds mixed among them, particularly American Herring Gulls like the first-winter (back row left) and third-winter (back row right) birds seen here. Herring and other species of similar-sized gulls seem to prefer each other’s company while on the wintering grounds.Five American Herring Gulls with smaller Ring-billed Gulls. The bird to the upper left and the bird hunkered down to its right are adult American Herring Gulls, the three brownish birds are in their first winter.Non-adult American Herring Gulls in flight. Always look for birds with all-pale wing tips when encountering herring gulls in flight.Midway in size between the Ring-billed Gulls in the foreground and the first-winter American Herring Gulls in the upper left and middle right of this image is a Lesser Black-backed Gull (dark-mantled bird resting at center). The similar-sized bird in the water behind it is a first-winter Iceland Gull (Larus glaucoides), a rare visitor from the arctic.Another look at the first-winter Iceland Gull from the previous image. Did you notice the all-white primary feathers and compare them to the dark wing tips on the Ring-billed Gulls seen here in its company?The conspicuously pale wings of the first-winter Iceland Gull, seen here bathing in the presence of two first-winter American Herring Gulls.Both this first-winter Iceland Gull and the bird from the previous three images are currently being seen just east of the Susquehanna watershed at Blue Marsh Lake in Berks County, Pennsylvania. Additional Iceland Gulls are currently being reported in Maryland on the Susquehanna at Conowingo Dam; on upper Chesapeake Bay in Baltimore County, Maryland; and in southeastern-most Bucks County, near Tullytown, Pennsylvania, at a busy landfill site that attracts tens of thousands of gulls each winter. Double-digit numbers of Iceland Gulls have been reported at this latter site during recent weeks.Lesser Black-backed Gulls like this one observed at Blue Marsh Lake are uncommon in the Lower Susquehanna River Watershed. They are progressively more likely as you travel east toward the aforementioned Bucks County landfills where hundreds of these birds make up a core Atlantic seaboard population.Great Black-backed Gulls are the most frequently encountered large gull on the Susquehanna. They’re easily identified by their enormous size and, as adults, their dark mantle.Rivaling the Great Black-backed Gull in size is the Glaucous Gull (Larus hyperboreus), another rare arctic visitor with pale wing tips. There are numerous reports of these unusual winter visitors from sites on Delaware Bay north to the Tullytown landfills on Delaware River where half a dozen or more have been occurring. Seen near the mouth of the Susquehanna on Chesapeake Bay at North East, Maryland, has been a first-winter bird similar to this one that we photographed at Blue Marsh Lake in Berks County, Pennsylvania.
For us, seeing a Glaucous Gull brought back memories of the last time we saw the species. It was forty-five years ago on New Year’s Day 1981 that we discovered two first-winter birds feeding on Gizzard Shad in open water on an otherwise ice-choked Susquehanna below the York Haven Dam powerhouse at Conewago Falls. Hey Doc Robert, do you remember that day?
One of the earliest non-native fish species to be widely released into North American waterways was the Common Carp. Stocks brought to the United States were likely sourced from populations already naturalized throughout much of western Europe after introductions originating from the fish’s native range in Eurasia, probably including the Danube and other watersheds east through the Volga. In western Europe, the species promised to be an abundant and easily cultivated food source. Under the same premise, carp were transported to the United States during the early 1800s and widely introduced into streams, lakes, and rivers throughout the country.
Common Carp thrive in nutrient-rich waters, particularly those subjected to sewage discharge and agricultural runoff, conditions which were already prevalent during the Common Carp’s initial introduction and have remained widespread ever since. Within these polluted streams, lakes, and ponds, introduced carp feed aggressively on benthic organisms and plants, stirring up decaying organic matter (mulm) from the substrate. This process raises turbidity in the water column and releases excessive amounts of the nutrient phosphorus resulting in unusually large algal blooms. Algal blooms can block sunlight from the longer-lived oxygen-producing vascular plants that grow in submerged environs. Growing beneath a dense cloud or blanket of algae can compromise the vigor of oxygen-producing vascular plants and disable their biochemical functions within the aquatic ecosystem. As the short-lived algae die, the bacteria that decay them begin to place increased oxygen demands on the water. With less oxygen being produced by both the vascular plants and the algae, and with oxygen consumption increased by the activity of decomposers, conditions can become fatal for fish and other organisms. This process is known as eutrophication. Because Common Carp are among the species most tolerant of eutrophic conditions, they tend to thrive in the conditions they create while the native fishes perish.
Common Carp spawn in the spring, usually from late April through June, when the water temperature is as low as 58 degrees and as high as 83 degrees Fahrenheit. This activity is often triggered by a rapid increase in water temperature. In a small lake, this may be brought on by a string of sunny days in late April or May. On larger streams and rivers, the temperature spike that initiates the spawn may not occur until warm rains and runoff enter the stream during June.
Seeing the exposed backs of Common Carp as they stir up mulm and other sediments while feeding along the edges of a body of water is not at all unusual.But carp pursuing other carp into the shallows is a sign that spawning has commenced.In water that is often less than a foot in depth, male carp follow the breeding females into egg-laying areas among debris and emergent vegetation.A fountain of splashes can ensue as males try to outdo one another for a chance to fertilize the female’s eggs.The males’ aggressive pursuit can even forced a large female to temporarily ground herself on the beach.
Common Carp are one of the most widely farmed and eaten fish in all the world. Here in the United States, they were introduced beginning two hundred years ago because they were favorable to the palate, grew to large size quickly, and were a source of much needed food. Today, the Common Carp is seldom found on the American dinner plate. Yet, pound for pound, it is one of the most abundant fish in many of our waters, particularly in man-made lakes. Like some of our other most invasive species—including Blue Catfish, Flathead Catfish, and Northern Snakehead—Common Carp are perhaps the most edible of our freshwater fishes. For many cultures, they are an important staple. For others, they are a delicacy or holiday treat. In America, they do horrendous damage to aquatic ecosystems following establishment as a food crop that almost never gets harvested. Did you realize that on the internet, there are literally hundreds of recipes and culinary videos available to show you how to prepare delicious dishes made with Common Carp? It’s true. And for the cost of a fishing license, you can catch all you want, usually several pounds at a time. So why not give the marine fisheries a break? Take the big leap and learn to eat invasive freshwater species instead.
