Migrating Waterfowl on Lotic Fresh Waters

Lotic vs. Lentic Freshwater Ecosystems — Some comparative attributes of unspoiled lotic vs. lentic freshwater ecosystems. Low-gradient (slow-moving) lotic waters often create, and remain connected to, accompanying riverine wetlands (a lentic freshwater ecosystem). These swamps, marshes, and ditches absorb, purify, and infiltrate flood waters while supporting a diverse number of plant and animal species.

We frequently perceive all waterfowl migration to be synchronized with the conspicuous movements of familiar species like Snow Geese, Tundra Swans, and Canada Geese—big flights coming south in October and November, then a return to the north in late February and March. And we’re all quite aware of the occurrence of large gatherings of some of these migrants while they make stopovers on some of our largest lentic (still) waters—the man-made lakes and reservoirs created by damming local streams. But did you know that there are populations of colorful waterfowl with dynamic migrations that extend throughout the winter and early spring with movements that are often continuous. Under favorable conditions, these birds favor the lotic (flowing) waters of the river and its larger tributaries as they transit the lower Susquehanna valley. That’s because unpolluted lotic freshwater ecosystems support a greater diversity of plants and animals than lentic waters, and therefore offer more opportunities for hungry migrating waterfowl to find food. Let’s have a look at some of the species that visit the river during their seasonal journeys…

Ice on the Susquehanna — While the urge to head south in the autumn is largely stimulated by the shortening of the photoperiod, it is the presence of ice, particularly on glacial lakes throughout the lands to the north of the lower Susquehanna River basin, that pushes many diving ducks to finally make their way south toward the guarantee of open-water feeding areas along the coast. This movement may occur at anytime between November and late February or, as we have seen during some of the mild winters of recent decades, it may scarcely be noticed at all.

Common Mergansers often lead they way when it comes to migratory diving ducks. They regularly move south in conspicuous numbers by late November and December and are regularly pushing north as soon as the ice begins to melt. During a typical year, it is not unusual for some populations of these large diving birds to remain north of us during the winter. Then, in the days after a sudden rush of frigid polar air, an appreciable increase in ice cover will force a mid-winter movement of birds down the Susquehanna.

Heat Flux Sources Impacting Temperature in Lotic Fresh Waters — Temperatures in lotic fresh waters vary over the length of the stream or river. They are largely determined by the collective impact of the numerous sources of heat flux depicted in this graphic. (Environmental Protection Agency image)

Buffleheads begin passing through the lower Susquehanna region in Novemeber on their way to coastal saltwater bays for the winter. Lingering populations feed by diving in the river’s pools and riffles for benthic invertebrates including snails and insect larvae. Lesser quantities of aquatic plant matter supplement their diet.

Many Common Goldeneyes will remain on shallow, ice-free waters of the northern lakes and rivers sculpted by the most recent glacial event, but only until they are forced south into and through the lower Susquehanna valley by the encroachment of freezing conditions. On the river, they are among the dozen or so species of diving ducks we see visiting or passing through during the typical late fall and early river.

Geomorphological Zonation of a Stream or River — During their visits to the lotic (also known as riverine) fresh waters of the Susquehanna and its largest tributaries, benthic-feeding waterfowl make short dives to take advantage of the plants, small fish, invertebrates, and other food sources inhabiting the stream bottom in the riffles and pools of the free-flowing waterway. Substrates, listed here by size (in descending order), along with other parameters influenced by this zonation determine the variety and abundance of the forage available to migrating waterfowl and other consumers. Ice or high water and poor visibility due to flooding can render the riffles and pools of the channel unusable for feeding. The birds must then choose to either linger and rest without feeding or leave the lotic freshwater habitat to seek sustenance. During a flood, this may require relocation to a nearby lentic (still) body of fresh water such as a lake or reservoir. The presence of ice will almost invariably force the birds to fly on to the Atlantic Coastal Plain and the tidal waters of its bays and estuaries.

Hooded Mergansers are one of the few species of diving ducks likely to utilize flooded shoreline timber and riverine (fluvial) wetlands as refuge from high water on Susquehanna.

Pied-billed Grebe and Canvasbacks — A Pied-billed Grebe and a pair of Canvasbacks on an ice-free stretch of the lower Susquehanna in mid-winter feed and loaf in a riffle-flanked pool where a large mat of American Eelgrass, a submerged aquatic plant also known as Tapegrass or Wild Celery, grows during the summer. The vast mosaic of riffles and pools in a river this size offers tremendous opportunities for a diverse array of aquatic species to find their niche wherein they can survive and flourish.

The presence of ice forces Buffleheads and other diving ducks to gather in turbulent open water, often below a riffle or dam. Another alternative is to continue on toward the salty bays and estuaries of the coast. High water may push these birds into the shallows among the flooded woods to feed, but they seldom utilize heavily forested riparian wetlands as a refuge due to their need for a running start to get airborne.

**Riffle and Pool Characteristics of High Gradient (A) and Low Gradient (B) Streams**. This graphic illustrates the change in deposition characteristics of a stream or river as its gradient decreases. A high-gradient stream (A) has a rapid velocity, often forms falls, and tends to carry away a high volume of all but the largest of particles of potential substrates as they erode from the surrounding landscape. On a low-gradient stream, the loss in water velocity reduces the water column’s ability to transport even the smallest of substrate particles. Deposition of this gravel, sand, silt, and clay forms lateral bars that over time create the familiar meandering path of a naturally flowing lowland stream. (National Park Service image)

Benthic Life in Lotic Freshwaters — Benthic substrates in lotic freshwater pools and riffles support an abundance of life forms ranging from colorful diatoms on rocks and cobble, to invertebrates including snails and insect larvae, to fishes like this young Channel Catfish. Free of accumulations of sediment, this river bottom not only provides habitat for a healthy fishery, it facilitates the bidirectional exchange of water between the Susquehanna and its underlying aquifer.

On the lower Susquehanna, populations of young Quillback suckers are found almost exclusively in clear, high-gradient pools.

The Harlequin Duck winters along the rocky shores and man-made jetties of the Atlantic coast. In summer, they nest on fast-moving, headwater streams well to our north. Very rare on the Susquehanna, this is the first of two individuals found during March and April of 2025. It was observed feeding in the swift waters of the high-gradient riffles and pools where the river cuts through Blue Mountain north of Harrisburg. During previous weeks, Harlequin Ducks were being seen along the coast as far south as the mouth of the Chesapeake at Cape Charles, Virginia. It’s very possible that some of these birds traveled north through the bay area and up the Susquehanna on their way north.

**Fluvial Geomorphology of a Stream**. Many of the Susquehanna’s tributaries pass through each of these three erosional zones. Along the way, they carry out the process of breaking down the mountains formed by the Allegheny orogeny, the collision of North America and Africa that created the supercontinent Pangea about 325 to 260 million years ago (during portions of both the Carboniferous and Permian periods). Today’s main stem of the lower Susquehanna passes through a transfer zone (Zone 2), carrying eroded materials to a depositional zone (Zone 3) located within the ancient Susquehanna canyon stretching from Havre de Grace, Maryland, to Norfolk Canyon on the edge of the continental shelf. Within this zone, more than a 10,000-year accumulation of post-glacial sediments lies submerged by the rising waters of the Atlantic Ocean and Chesapeake Bay. Although the present-day lower Susquehanna is largely a transfer zone, some deposition occurs along low-gradient segments of the river, particularly where its course parallels the watershed’s ridges both above and below the high-gradient rapids where its path has eroded passage through the highlands. (National Park Service image)

Although the present-day lower Susquehanna is largely a transfer zone, some deposition occurs along low-gradient segments of the river, particularly where its course parallels the watershed’s ridges both above and below the high-gradient rapids where its path continues to erode passage through the bedrock. On this 1908 map of the Susquehanna at Conewago Falls, alluvial terraces of gravel, sand, silt, and clay can readily been seen as pale areas nearly lacking brown contour lines along the shorelines and islands of the river. Deposits within most of these terraces date back to the melt period following the most recent glacial event and beyond. The delta shown at the mouth of Conewago Creek (west) includes massive volumes of material deposited by both the creek and the river. This delta is currently known as Brunner Island, much of it developed as the site of a coal/gas-fired electric generating station. Terrace deposits along the Susquehanna’s shorelines created extensive perched marshes and swamps (wooded marshes) fed by rains, high river water, small streams, and springs, the latter often seeping from the base of the rocky escarpments carved by the ancient Susquehanna and defining the present-day inland border of the floodplain. We call these sites “Alluvial Terrace Wetlands”. Few of these critical components of river morphology survive. Those not drained for farmland were obliterated by urbanization and canal, railroad, and highway construction. (United States Geological Survey base image: Middletown, PA, quadrangle, 1908)

Water Willow is a familiar emergent plant that colonizes lateral bars and other alluvial deposits in low-gradient segments of the Susquehanna.

Water Willow Roots — The fine roots of Water Willow collect sediment and absorb nutrients while creating dense cover for young fish and numerous species of invertebrates including the Virginian River Horn Snails seen here.

Prior to the nineteenth century, the low-gradient flow regime of the river both above and below the riffles at Chiques Rock (lower right on map) created prime wildlife habitat. The natural accumulations of nutrients and substrates carried into and through the lotic waterway’s pools and riffles were cycled into an ideal growing medium for extensive mats of American Eelgrass and other aquatic plants. This underwater forest hosted a seemingly endless abundance of invertebrates and fishes (both resident and migratory)—supporting a variety of consumer species including various populations of humans. But soon after the mass clearing of much of the watershed’s land for farming and lumber, the mill ponds created by dams constructed on streams to power saw and grain mills became brimful with sediments eroded from the unprotected ground. During storm events, torrents of these sediments then flowed full bore toward the Susquehanna, and began accumulating in the low-gradient segments of the river.

Legacy Sediments — Sediments left behind after the removal of mill dams are known as legacy sediments. They disconnect the stream from its historic floodplain and riverine (fluvial) wetlands, thus intensifying the impact of high water in the surrounding landscape. As these nutrient-charged deposits wash away, they become a source of pollution in the waters of the Susquehanna and Chesapeake.

A mat of American Eelgrass growing in the flowing waters of the Susquehanna below Conewago Falls. Eelgrass and other submerged aquatic plants provide essential habitat for a wide variety of small fish and invertebrates while also consuming nutrients deposited in the cobble, gravel, and sand substrate of the river’s pools. Excess quantities of smaller particles of silt and clay can clog the substrate and thus inhibit the hyporheic exchange of water between the stream channel and the underlying aquifer, often diminishing the biomass and diversity of organisms inhabiting this benthic habitat. Buried in these life-choking sediments, the river bottom becomes inhospitable to growth of submerged aquatics including eelgrass.

This low-gradient stretch of the Susquehanna at Marietta flows parallel to the Chickies Quartzite “Hellam Hills” before making a sharp right turn to punch through the ridge as a series of rapids at Chiques Rock. Formerly a fully functional lotic ecosystem and a paradise for migrating waterfowl, this river segment is now impaired by accumulations of nutrient-laden sediments from agricultural and urban runoff.

Conewago Falls Flood Waters — Nutrient and sediment-loaded flood waters roar across the diabase boulders at the York Haven Dam and Conewago Falls, a high-gradient segment of the Susquehanna.

Flood Waters at Haldeman Riffles — They continue past Brunner Island (left power plant stacks) and through Haldeman Riffles and the Shocks Mills Railroad Bridge…

Flooding on the Marietta Broads — …into the stretch of river known as the “Marietta broads” along the base of the Chickies Quartzite “Hellam Hills”. The low stream gradient here produces a slower current and increased deposition of sediments.

Susquehanna Flooding at Chiques Rock — As the flood surges through the riffle and pool complex at Chiques Rock, the high stream gradient maintains a velocity in the water column sufficient to keep additional sediments in suspension until they reach the low-gradient river segment just downstream at Washington Boro, site of a naturally occurring lateral bar area known as the Conejohela Flats. These bars now lie within the man-made depositional zone known as “Lake Clarke”. Created nearly a century ago by construction of the Safe Harbor Dam, this impoundment is accumulating astounding volumes of nutrient-loaded sediments that continue to encapsulate the flats within a stream-impairing delta.

Redheads, Canvasbacks, and a Horned Grebe — Anytime from November to April on the lower Susquehanna, a group of Redheads, Canvasbacks (3 birds to right of the middle of the picture), and a Horned Grebe (lower left) is a welcome sight in a riverine pool known to have a summertime growth of American Eelgrass. Noted Dr. Herbert Beck in 1924 when describing the Canvasback, “Like all ducks, …, it stops to feed within the county (Lancaster) less frequently than formerly, principally because the vast beds of wild celery which existed earlier on broads of the Susquehanna, as at Marietta and Washington Borough, have now been almost entirely wiped out by sedimentation of culm. Prior to 1875 the four or five square miles of quiet water off Marietta were often as abundantly spread with wild fowl as the Susquehanna Flats are now. Sometimes there were as many as 500,000 ducks of various kinds on the Marietta broad at one time.”

Aythya genus ducks. — Today, seeing just dozens of Aythya genus ducks (Redheads, Canvasbacks, scaup) on the lower Susquehanna is a notable event. If they happen to be forced down by inclement weather while migrating through, you might get lucky enough to see several hundred.

During the past two centuries, as food supplies in the Susquehanna grew increasingly compromised for benthic feeders like these Lesser Scaup and other diving ducks, a change in distribution was necessary for survival.

Lesser Scaup in flight. — As individual species, Lesser Scaup and other waterfowl that fail to adapt to natural or man-made changes in their habitats and food supplies may see their overall global numbers falter.

Despite being located in the transfer zone, the lower Susquehanna has become a significant depositional zone along much of its length, mostly courtesy of the placement of sediment-trapping man-made dams.

Following construction of the mill dams and ponds on nearly every mile of the lower Susquehanna’s low-gradient tributary streams, enterprising parties moved on to the river. The first significant spans were constructed using wide timber cribs filled with large rock. They were placed to create water deep enough to allow canal boats to cross the Susquehanna at both Clark’s Ferry at the mouth of the Juniata River in Dauphin County and at Columbia/Wrightsville. These dams also diverted water into the newly excavated canals—the Pennsylvania Eastern Division Canal (completed in 1833) which followed the river’s east shore from Clark’s Ferry to Columbia, and the Susquehanna and Tidewater Canal (completed in 1840) along the west shore from Wrightsville to Havre de Grace, Maryland. Placement of these sediment-trapping man-made dams began a process of converting vast mileage of the lower Susquehanna from a transfer zone into a deposition zone. In addition, layout of the canals and locks followed the contours along the base of the riverside ridges, seriously altering most of the alluvial terrace wetlands that the river had created as a feature of its floodplain during the post-glacial period.

Construction of the canal dams was just the beginning. During the twentieth century, more massive dams would be added to the main stem of the river for hydroelectric energy production at York Haven, Safe Harbor, Holtwood, and Conowingo. Upon their completion, the days of unassisted anadromous fish migrations were over. On both the river and its tributaries, smaller dams including dangerous low-head dams maintain water levels for boating and recreation. They too create current-diminishing, pseudo-lentic waters that blanket the lotic riffle and pool substrates with polluted sediments.

MAN-MADE DAMS TURN LOTIC WATERS INTO UNFLUSHED TOILETS

Sediment Deposition Behind Lower Susquehanna Dams — The construction of dams on the lower Susquehanna has converted vast mileage of the river from a lotic freshwater system into a series of man-made lentic freshwater lakes. These areas have lost their function as a lotic transfer zone and are now a sort of dysfunctional series of depositional zones collecting vast volumes of sediment containing nutrients and other pollutants. Within each impoundment, the reduced velocity of the river causes it to drop suspended sand first, then the finer particles of silt and clay closer to the dam. The flow regime of riffles and pools is lost and the hyporheic zone that exchanges water between the river and the underlying aquifer is clogged. These impaired segments of river become ripe for eutrophication: algal blooms followed by die offs that can lead to a fatal reduction of dissolved oxygen in the water column.

Deposits of lateral bars of sediment in low-gradient segments of the Susquehanna can create shallow water feeding habitat for puddle/dabbling ducks like these Gadwall. Where sediment pollution is severe, benthic foods in these areas often consist mostly of invertebrates and plant matter deposited by the current, the buried substrates devoid of a functioning ecosystem and the waters subject to eutrophication.

Common Goldeneyes on a patch of open water on an otherwise ice-covered “Lake Clarke”, the impoundment created by Safe Harbor Dam. While they may find this spot advantageous for loafing, the food supply over the sediment-buried substrate will be limited.

By the end of the twentieth century, accumulations of polluted sediments behind lower Susquehanna dams were nearing capacity. There is no working plan to attenuate the massive release of these pollutants that may be triggered by a catastrophic flood. The effort to reduce nutrient and sediment runoff remains the focus so that new loading is kept to a minimum and won’t add to the capacity problems at the dams or continue downriver to the Chesapeake at full strength when the dams are full. Alleviating the sediment aggregation problem within the river’s impoundments is a tall order and a dilemma not easily solved. (United States Geological Survey image)

Fortunately for everything else living in the benthos, Common Mergansers are big enough to devour invasive, non-native Rusty Crayfish when they find them in our lotic waterways.

TIME TO CLEAN UP OUR ACT

WHERE DOES YOUR STORMWATER GO?

Channelized Urban Stream — **Channelized Urban/Suburban Streams Function as Sewers.** They have no attached lowlands or floodplains to absorb, purify, and infiltrate runoff from rain events. Pollutants including litter, pet waste, lawn chemicals, tire-wear particles, hazardous fluids, and sometimes untreated human excrement flush unchecked from the municipal storm drainage system into the waterway. Thermal shock from summer downpours washing across sun-heated pavements can kill temperature sensitive fishes and other aquatic life. Nutrient and sediment loads from these impaired tributaries later accumulate downstream in low-gradient segments of the Susquehanna, turning the river into an open-air cesspool. Aggressively working to implement projects that eliminate these sources of pollution are the only effective way to keep the problem from getting worse. Making things better requires a lot more dedication and effort. (United States Geological Survey image by Frank Ippolito)

RIPARIAN BUFFERS MAKE A DIFFERENCE…WIDER IS BETTER

FLOODPLAIN RESTORATION

Regardless OF HOW LONG YOU’VE BEEN CONDITIONED TO THINK OTHERWISE, THIS IS A DYSFUNCTIONAL, POLLUTED CREEK—AND IT NEEDS HELP

RESTORED TO ITS HISTORIC FUNCTIONS

Floodplain and Stream Restoration — On an adjacent segment of the same creek, this legacy sediment removal project restored a braided meandering channel and connected it to its newly liberated, historic floodplain. In just their second year, the fluvial wetlands are effectively absorbing stormwater and sequestering nutrients as an attached component of the stream’s riffle and pool complex. During our visit earlier this week, we found American Toads, Northern Leopard Frogs (Lithobates pipiens), and Northern Spring Peepers breeding here. It’s just as Castor canadensis would have it!

“STOP HEMMING AND HAWING AROUND ALREADY”

“HEY COWBOY, HOW ‘BOUT GETTIN’ THEM FILTHY LITTLE DOGIES OUTTA DAT CRICK?”

Impaired Stream — Here’s a polluted stream in a pasture with grazing livestock. The site is a former mill pond within which the creek eroded a channel following removal of the dam. The animals defecate and urinate where access to water is gained at a broken down embankment of the nutrient-loaded legacy sediments deposited in the pond more than a century ago. It’s a haphazard form of animal husbandry and a reminder that all it takes is just one stubborn jackass to foul up the whole waterway.

DIRECT SOURCES OF NITROGEN (AMMONIA) POLLUTION IN STREAMS

DID YOU KNOW that a dairy cow produces about 80 pounds of waste (excrement and urine) every day?

DID YOU KNOW that a horse produces about 50 pounds of waste (excrement and urine) every day?

DID YOU KNOW that a human produces about 3 to 4 pounds of waste (excrement and urine) every day? The exceptions, of course, are those who continue to insist that raising farm animals in and alongside a body of water is okey-doke—a harmless practice. These individuals tend to retain the former constituent of human waste and are thus full of it.

“ATTABOY TEX, THAT’S MORE LIKE IT!”

Stream Fencing and Livestock Crossing — Now that’s better. Legacy sediments have been removed to reconnect the stream to its floodplain. A livestock crossing and exclusion fencing has been installed, and a nutrient-consuming riparian buffer has been planted. This creek segment’s pollution woes have been mitigated. Do you have a neighbor needing this type of remedial work on their farm? Have them call your local conservation district office for advice. Some programs include financial assistance covering the costs of installation as well as monetary incentives for helping to clean up the water.

AND FINALLY

WHEN IT COMES TO BUILDING DAMS ON LOTIC FRESH WATERS…

…LEAVE IT TO THE BEAVERS

North American Beavers (Castor canadensis) create habitats that connect the riffle and pool regime of a low-gradient stream to a surrounding fluvial wetland that retains sediments, cycles nutrients, and provides essential habitat for hundreds of plant and animal species. Floodplains are for flooding. And if a beaver floods an area, you can be guaranteed that it was already part of a floodplain. You see, beavers don’t encroach upon humans, it’s humans doing the encroaching upon beavers. (National Park Service image)

MAD HATTERS

DID YOU KNOW that even before the landscape was cleared for farms and a supply of timber, and before mill dams on local creeks began accumulating soil runoff from the consequently barren hillsides, all the North American Beavers, the keystone species of lower Susquehanna stream ecology, were killed and sold to make hats? It’s no wonder things are fubar!

COMING SOON…

Horned Grebes are regular migrants and sometimes winter residents on ice-free stretches of the lower Susquehanna. They spend their time plying the benthic substrate of the river’s clear riffles and pools for a variety of invertebrates and small fish. Look for them moving north in coming days sporting this beautiful breeding plumage.

April and early May are prime time for observing Common Loons on the Susquehanna as they undertake a journey from the Atlantic surf where they spent the winter to nesting sites on northern lakes. For this migrant in breeding plumage, clear water for sighting plenty of benthic life in the river’s riffles and pools assures a successful dive in search of energy-replenishing forage.

June 2, 2025June 3, 2025

Photographs in Living Color: Black and White is Beautiful

Here at susquehannawildlife.net headquarters, we really enjoy looking back in time at old black-and-white pictures. We even have an old black-and-white television that still operates quite well. But on a nice late-spring day, there’s no sense sitting around looking at that stuff when we could be outside tracking down some sightings of a few wonderful animals.

American Toad Tadpoles — American Toad tadpoles have hatched from clusters of eggs deposited in this wet roadside ditch furnished with a clean supply of runoff filtered through a wide shoulder of early successional growth. Recent rains have kept their vernal nursery flooded, giving them the time they need to quickly mature into tiny toads and hop away before scorching summer heat dries up their natal home.

Water Striders mating. — Weekend rains and creek flooding haven’t stopped these Water Striders from pairing up to begin their breeding cycle.

Around streams, lakes, ponds, and wetlands, the Common Whitetail is one of our most conspicuous dragonflies.

Now that’s what we call a big beautiful bill, on a Great Blue Heron stalking fish.

These mating Golden-backed Snipe Flies (Chrysopilus thoracicus) are predatory insects, as are their larvae. They are most frequently found in bottomland woods.

About three feet in length, this Eastern Ratsnake is unusual because it still shows conspicuous remnants of the diamond-patterned markings it sported as a juvenile.

The plumage of the Black-and-white Warbler lacks any of the vibrant colors found in the rainbow, but is nevertheless strikingly beautiful.

May 28, 2024June 15, 2024

Pest Control at the Picnic

During one of the interludes between yesterday’s series of thunderstorms and rain showers, we had a chance to visit a wooded picnic spot to devour a little snack. Having packed lite fare, there wasn’t enough to share. Fortunately, pest control wasn’t a concern. It was handled for us…

This young American Toad (Anaxyrus americanus) patrolled the grounds of the grove gobbling up any creepy crawlies it happened to encounter.

Though spotted nearby, none of these traditional picnic crashers, possibly Big-headed Ants (Pheidole species), made it through the toad’s dragnet.

What had this Eastern Black Carpenter Ant (Camponotus pennsylvanicus) so frustrated during its attempt to spoil our affair?

Decaying Tree Stump — …so it’s back to gnawing on this rotting stump for you!

We spied this curious little Crab Spider (Tmarus species) searching the benches for tiny invertebrates.

Then, at the corner of the table, we found this Running Crab Spider (Philodromus species) clutching what appeared to be two ofttimes pesky, but harmless, flying insects called non-biting midges (Chironomidae).

After enjoying our little luncheon and watching all the sideshows, it was time to cautiously make our way home,…

…halting briefly to allow this Snapping Turtle a chance to cross the road.

July 24, 2023August 25, 2023

Shorebirds and More at Bombay Hook National Wildlife Refuge

Have you purchased your 2023-2024 Federal Duck Stamp? Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points. Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States. So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.

2023-2024 Federal Duck Stamp. Your Federal Duck Stamp is your free pass to visit the nation's National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware. — Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.

Still not convinced that a Federal Duck Stamp is worth the money? Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge. Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.

This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge. At Bombay Hook, they don’t waste your money mowing grass. Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.

Shearness Pool at Bombay Hook N.W.R. — Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools. Birds there can often be observed at close range. The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools. Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.

Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.

Semipalmated Sandpipers and Short-billed Dowitcher — More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.

Two more Short-billed Dowitchers on the way in.

Sandpipers, Avocets, Egrets, and Mallards — Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed. Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge. Mallards, American Avocets, and egrets are gathered on the shore.

Semipalmated Sandpipers, Short-billed Dowitchers, and Forster's Terns — Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.

Semipalmated Sandpipers, Forster's Terns, and a Short-billed Dowitcher — A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).

Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool. A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.

Stilt Sandpiper among Short-billed Dowitchers — Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.

American Avocets probe the muddy bottom of Raymond Pool.

Dunlin and Short-billed Dowitchers — Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early. Many migrating Dunlin linger at Bombay Hook into October and even November.

This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.

A Greater Yellowlegs at Bear Swamp Pool.

Marsh Wren singing — The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.

Marsh Wren Nest — This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass. High tide licks at the roots of the cordgrass supporting the temporary domicile.

By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice). It is our only dragonfly able to breed in saltwater. Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.

Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.

Look up! A migrating Bobolink passes over the dike at Shearness Pool.

Mute Swans and Canada Geese — Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.

A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool. Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river. The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century. An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers. Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.

A hen Wood Duck (second from right) escorts her young. — Wood Ducks in Bear Swamp Pool.

Black-necked Stilt and young. — A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.

As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes. Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.

Tidal Marshes — Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.

The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay. We found just this one during our visit to Bombay Hook. Man has certainly decimated populations of this ancient crustacean during recent decades.

Boardwalk Trail — As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.

Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).

A male Atlantic Marsh Fiddler Crab peers from its den.

A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.

A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.

Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.

Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide. — Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.

Ospey — An Osprey patrols the vast tidal areas opposite Shearness Pool.

No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.

Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.

Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook. This one was in shrubby growth along the dike at the north end of Shearness Pool.

Trumpet Creeper and Poison Ivy — These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.

Pileated Woodpecker in Sweet Gum — The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries. The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware. The seed balls are a favorite winter food of goldfinches and siskins.

Red-bellied Slider and Painted Turtle — Finis Pool has no frontage on the tidal marsh but is still worth a visit. It lies along a spur road on the tour route and is located within a deciduous coastal plain forest. Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.

The White-tailed Deity is common along the road to Finis Pool.

Fowler's Toad — Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.

Turk's Cap Lily — The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well. This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.

Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.

We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon. And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.

May 26, 2023June 1, 2023

Forty Years Ago in the Lower Rio Grande Valley: Day Six

Back in late May of 1983, four members of the Lancaster County Bird Club—Russ Markert, Harold Morrrin, Steve Santner, and your editor—embarked on an energetic trip to find, observe, and photograph birds in the Lower Rio Grande Valley of Texas. What follows is a daily account of that two-week-long expedition. Notes logged by Markert some four decades ago are quoted in italics. The images are scans of 35 mm color slide photographs taken along the way by your editor.

DAY SIX—May 26, 1983

“Bentsen State Park — Texas”

“Arose at 7:30. Birded the park and watched for the reported Roadside Hawk. No luck. A Roadrunner and Kiskadee was a welcome addition to our list.”

The appearance of Roadside Hawk (Rupornis magnirostris) north of the Rio Grande in Hidalgo County, Texas, was hot news in 1983. In the weeks leading up to our visit and while we were there, no sightings of this mega-rarity were confirmed. The reports that were received were not accompanied by photographs or sufficient details to rule out the Hook-billed Kites (Chondrohierax unicinatus) and Cooper’s Hawks that were known to be present. Roadside Hawk remains a very elusive tropical vagrant in the United States. It was seen as recently as December of 2018 along the man-made flood control levee to the east of Bentsen-Rio Grande Valley State Park near the Border Patrol Corral and the adjacent National Butterfly Center, neither of which were present in 1983.

Both the Greater Roadrunner (Geococcyx californianus), a familiar member of the cuckoo family (Cuculidae), and the Great Kiskadee (Pitangus sulphuratus), a flycatcher at the northern limit of its range in south Texas, were much anticipated finds at Bentsen-Rio Grande State Park. We would get better looks at both when we moved to the west in a couple of days.

“Traveled to Brownsville and met Harold and Steve at the airport at 12:30 P.M.”

Back in 1983, the Brownsville Airport was the most frequently used destination for those wanting to fly into the Lower Rio Grande Valley. Today, Valley International Airport in Harlingen is the busier of the two. Both of these airports have an interesting history from the World War II years. Brownsville Airport, renamed Brownsville Army Air Field for the duration of the war, was in use as a base for antisubmarine warfare flights, pilot training, aircraft engine overhauls, and the occasional servicing of B-29 bombers. The airport in Harlingen was founded during the period as Harlingen Army Air Field and was home to the Harlingen Aerial Gunnery School. A cheesy Hollywood propaganda movie called Aerial Gunner (1943) was shot there in late 1942 and starred Richard Arlen and a local actress, Amelita (Lita) Ward, who was discovered by the picture’s producers in Harlingen during the early stages of filming. A young Robert Mitchum portrays one of the gunners. It’s one of those typical B-movies of the period with a story line that is all mixed up with what happened years ago. A climactic part of the film depicts students being taken aloft to fire machine guns at kite-like targets being towed behind other training aircraft flying over the nearby Gulf of Mexico. You can find this classic and watch it for free on the internet. Afterward, you can go join up if you want, but you can no longer get the war bonds they were selling.

“We tried for the Clay-colored Robin — No Luck.”

Now that Harold and Steve had joined us, we began putting in an earnest effort to find the five species we all needed for our life lists—those Lower Rio Grande Valley exclusives Father Tom had given us directions to find. First up was Clay-colored Robin, known today as Clay-colored Thrush (Turdus grayi), a brown-colored bird similar to our American Robin. Its native range in 1983 was from northern Colombia north to the Rio Grande River. The only pair known to be reliably nesting north of the river in the United States, thus within the A.B.A. listing area, was in Brownsville in a park-like setting on the side of a small hill with a commercial radio station transmitter and antenna on its crest. It was an easy place to find in the otherwise flat landscape, but we had no luck finding the birds. Were we too late? Was nesting season over? Had the birds already dispersed?

“Visited Santa Ana again and added Ground Dove and White-tailed (Black-shouldered) Kite.”

Father Tom provided us with a tip that Hook-billed Kites had nested in the vicinity of Santa Ana National Wildlife Refuge. We remained vigilant, but saw none. The petite Common Ground Dove (Columbina passerina) was harder to locate than I expected, possibly because its small size makes it more prone to depredation than larger doves and pigeons, so it may be a little more secretive. White-tailed Kite (Elanus leucurus) sightings were a highlight of the trip, so graceful and stunning in appearance, but very wary and distant—not a good photo opportunity. Other sightings at Santa Ana included the first Harris’s Hawk (Parabuteo unicinctus) of the trip, White-tipped Dove, Groove-billed Ani, Barn Swallow, Black-crested Titmouse, Long-billed Thrasher, a lingering Solitary Vireo, Bronzed Cowbird, Olive Sparrow, and Northern Cardinal. We heard a Dickcissel (Spiza americana) in a sunflower field by the refuge entrance, but failed to see it.

Harris's Hawk — A Harris’s Hawk at Santa Ana National Wildlife Refuge. This species is a year-round resident in much of the scrubland of the southwestern United States and Mexico. (Vintage 35mm image)

Also at Santa Ana N.W.R. a Giant Toad and a Texas Tortoise (Gopherus berlandieri), a scrubland native of south Texas and northern Mexico.

At Santa Ana National Wildlife Refuge, we stumbled upon this Giant Toad, known today as the Cane Toad, Marine Toad, or Giant Neotropical Toad (Rhinella marina). It’s range extends from Amazonia north into the Lower Rio Grande Valley. (Vintage 35 mm image)

“We arrived back at Bentsen State Park and I got supper ready while the others birded. At dusk we saw the Elf Owls again and toured the roads. Saw two Pauraques. To bed by 11:15 after a welcome shower.”

A White-eyed Vireo was found during our late afternoon walk. On a tip from Father Tom, we checked the campground in the area of the east restrooms for nesting Tropical Parulas (Setophaga pitiayumi). For each of us, it was another of the five target species for the trip. We heard nothing that resembled this warbler’s song, which sounds very much like that of the Northern Parula with which we were each familiar.

Nightjars are more often seen than heard, so having the opportunity to finish the day watching two Common Pauraques sail across a break in the forest to grab airborne insects was a real treat.