It begins on a sunny morning in spring each year, just as the ground temperature reaches sixty degrees or more…
Eastern Subterranean Termites emerge in unison from a nest located in the soil beneath a log. Each of these swarming “alates” is a potential king or queen seeking to find a location with an ample supply of fallen timber to provide food for establishment of a new colony. They are escorted to the surface by a soldier (lower center) equipped with powerful jaws for protecting the existing nest. Similar-looking worker termites tend the nest, the queen, her eggs, and their siblings, but usually remain hidden from view. The workers feed upon wood, hosting cellulose-digesting protozoa and bacteria in their guts to break down the fibers. This symbiotic relationship is an important mode of decomposition in the forest, the process that turns wood into the organic matter that enriches soil and helps it to retain more moisture.Termites are among the numerous arthropods that join fungi and a variety of microbes to decompose dead wood and other plant matter into the nutrients and organic materials used by living plants to thrive and grow.Within moments of emerging, swarming “alates” ascend a tree trunk or other vertical surface from which they can take flight.Eastern Subterranean Termite “alates” gather atop a stump before launching skyward.Termites swarm in massive numbers in an attempt to overwhelm the predators that are inevitably attracted to their sudden appearance. The few “alates” that survive to find a source of rotting wood in which to begin a new colony are the only hope for continuing their king and queen’s legacy.Soon after lift off, the majority of swarming termites are consumed by swallows, swifts, and other birds, but some are discovered at ground level.A lightning-fast strike with its tongue and this Green frog has snatched up yet another termite. Those that slip by the dragnet of terrestrial and aerial predators can sometimes start a new colony in the ground beneath a dead tree or in a vulnerable house or other wooden structure. To keep a small clan from invading your home, be certain the wood elements of your building(s) are kept dry and are not in contact with dirt, soil, bark mulch, etc. Regular inspections for evidence of their presence can head off the long-term damage termites can inflict on the stuff we construct with tree skeletons.A Green Frog wearing its breakfast. Its next chance for a termite feast may come during the autumn when the Drywood Termites (Kalotermitidae) swarm.
Of course, termites aren’t the only groups of insects to swarm. As heated runoff from slow-moving thundershowers has increased stream temperatures during the past couple of weeks, there have occurred a number of seasonal mayfly “hatches” on the Susquehanna and its tributaries. These “hatches” are actually the nuptial flights of newly emerged imago and adult mayflies. The most conspicuous of these is the Great Brown Drake.
Seen here with a much smaller and more typical regional mayfly to its left, the Great Brown Drake was for several years infamous for swarming the lights and creating traffic hazards on bridges spanning the Susquehanna. During the past two weeks, nighttime flights of these giants have ventured out to gather at well lit locations in housing and business districts more than a dozen miles from the river. Earlier this century, this proclivity to wander probably led the Great Brown Drake to first invade silty segments of the Susquehanna as a colonizer from its native range in the Mississippi watershed.
Swarms of another storm-related visitor are being seen throughout the lower Susquehanna valley right now. Have you noticed the Wandering Gliders?
Throughout the month, swarms of Wandering Gliders, the most widespread dragonflies in the world, descended on areas hit by localized slow-moving thundershowers. Large numbers of these global travelers are known to get swept up within the thermal air masses that lead to these storms. In suitable terrain within the path of the downpours, they linger to search for flooded places where they can mate and deposit eggs. Wandering Gliders frequently mistake large parking lots at shopping malls, grocery stores, etc. for wetlands and will be seen in these areas depositing eggs upon the hoods and roofs of shiny motor vehicles, surfaces which appear puddle-like in their eyes.More dragonfly swarms are yet to come. Adult Common Green Darners are presently in the Lower Susquehanna River Watershed mating and depositing eggs within vegetated ponds, lakes, and wetlands. Beginning in August, adult Common Green Darners and other migratory species including Black Saddlebags, Carolina Saddlebags, Wandering Gliders, and Twelve-spotted Skimmers will begin swarming as they feed on flying insects (including lots of mosquitos and gnats) and start working their way toward the Atlantic Coastal Plain. Along the barrier islands by September, concentrations of southbound dragonflies can reach the thousands, particularly at choke points like Cape May, New Jersey, and Cape Charles, Virginia. So be sure to keep an eye on the sky for swarms of dragonflies during coming weeks. And don’t forget to check out our “Damselflies and Dragonflies” page by clicking the tab at the top of this page.
Even in flight, the Twelve-spotted Skimmer is easily identified by the conspicuous color pattern in its wings. Look for it now around vegetated ponds and lakes. Later, during late summer and early fall, this widespread species can often be seen among southbound movements of other migratory dragonflies.
To pass the afternoon, we sat quietly along the edge of a pond created recently by North American Beavers (Castor canadensis). They first constructed their dam on this small stream about five years ago. Since then, a flourishing wetland has become established. Have a look.
Vegetation surrounding the inundated floodplain helps sequester nutrients and sediments to purify the water while also providing excellent wildlife habitat.The beaver lodge was built among shrubs growing in shallow water in the middle of the pond.Woolgrass (Scirpus cyperinus) is a bulrush that thrives as an emergent and as a terrestrial plant in moist soils bordering the pond.A male Common Whitetail dragonfly keeping watch over his territory.A Twelve-spotted Skimmer perched on Soft Rush.A Blue Dasher dragonfly seizing a Fall Field Cricket (Gryllus pennsylvanicus).A Spicebush Swallowtail visiting a Cardinal Flower.A Green Heron looking for small fish, crayfish, frogs, and tadpoles.The Green Heron stalking potential prey.A Wood Duck feeding on the tiny floating plant known as Lesser Duckweed (Lemna minor).A Least Sandpiper poking at small invertebrates along the muddy edge of the beaver pond.A Solitary Sandpiper.A Solitary Sandpiper testing the waters for proper feeding depth.A Pectoral Sandpiper searches for its next morsel of sustenance.The Sora (Porzana carolina) is a seldom seen rail of marshlands including those created by North American Beavers. Common Cattails, sedges, and rushes provide these chicken-shaped wetland birds with nesting and loafing cover.
Isn’t that amazing? North American Beavers build and maintain what human engineers struggle to master—dams and ponds that reduce pollution, allow fish passage, and support self-sustaining ecosystems. Want to clean up the streams and floodplains of your local watershed? Let the beavers do the job!
Your best bet for finding migrating shorebirds in the lower Susquehanna region is certainly a visit to a sandbar or mudflat in the river. The Conejohela Flats off Washington Boro just south of Columbia is a renowned location. Some man-made lakes including the one at Middle Creek Wildlife Management Area are purposely drawn down during the weeks of fall migration to provide exposed mud and silt for feeding and resting sandpipers and plovers. But with the Susquehanna running high due to recent rains and the cost of fuel trending high as well, maybe you want to stay closer to home to do your observing.
Fortunately for us, migratory shorebirds will drop in on almost any biologically active pool of shallow water and mud that they happen to find. This includes flooded portions of fields, construction sites, and especially stormwater retention basins. We stopped by a new basin just west of Hershey, Pennsylvania, and found more than two dozen shorebirds feeding and loafing there. We took each of these photographs from the sidewalk paralleling the south shore of the pool, thus never flushing or disturbing a single bird.
Designed to prevent stream flooding and pollution, this recently installed stormwater retention basin along US 322 west of Hershey, Pennsylvania, has already attracted a variety of migrating plovers and sandpipers.Killdeer stick close to exposed mud as they feed.Two of more than a dozen Least Sandpipers found busily feeding in the inch-deep water.A Lesser Yellowlegs searching for small invertebrates.Two Lesser Yellowlegs work out a disagreement.Male Twelve-spotted Skimmers patrol the airspace above a pair of Least Sandpipers. Dragonflies and other aquatic insects are quick to colonize the waters held in well-engineered retention basins. Proper construction and establishment of a functioning food chain/web in these man-made wetlands prevents them from becoming merely temporary cesspools for breeding mosquitos.
So don’t just drive by those big puddles, stop and have a look. You never know what you might find.
A Semipalmated Sandpiper (middle right) joins a flock of Least Sandpipers.Pectoral Sandpipers (two birds in the center) are regular fall migrants on the Susquehanna at this time of year. They are most frequently seen on gravel and sand bars adjacent to the river’s grassy islands, but unusually high water for this time of year prevents them from using this favored habitat. As a result, you might be lucky enough to discover Pectoral Sandpipers on almost any mudflat in the area.Two Pectoral Sandpipers and five smaller, but otherwise very similar, Least Sandpipers.A Killdeer (right), a Semipalmated Plover (upper right), and Least and Pectoral Sandpipers (left).
During the coming two weeks, peak numbers of migrating Neotropical birds will be passing through the northeastern United States including the lower Susquehanna valley. Hawk watches are staffed and observers are awaiting big flights of Broad-winged Hawks—hoping to see a thousand birds or more in a single day.
During its passage through the Lower Susquehanna River Watershed, an adult Broad-winged Hawk sails over Second Mountain Hawk Watch in Lebanon County, Pennsylvania.A hatch-year/juvenile Broad-winged Hawk gazes toward hawk watchers on the ground.
Broad-winged hawks feed on rodents, amphibians, and a variety of large insects while on their breeding grounds in the forests of the northern United States and Canada. They depart early, journeying to wintering areas in Central and South America before frost robs them of a reliable food supply.
The Carolina Saddlebags (Tramea carolina), this one photographed at Second Mountain Hawk Watch on September 8th, is the rarest of the lower Susquehanna region’s migratory dragonflies. Autumn Broad-winged Hawk movements coincide with southbound flights of the Carolina Saddlebags and the more numerous migratory dragonfly species: Common Green Darner, Wandering Glider, Twelve-spotted Skimmer, and Black Saddlebags. “Broad-wings” will often eat these and other dragonflies during migration and can sometimes be seen catching and feeding upon them while still soaring high overhead.
While migrating, Broad-winged Hawks climb to great altitudes on thermal updrafts and are notoriously difficult to see from ground level. Bright sunny skies with no clouds to serve as a backdrop further complicate a hawk counter’s ability to spot passing birds. Throughout the Lower Susquehanna River Watershed, the coming week promises to be especially challenging for those trying to observe and census the passage of high-flying Broad-winged Hawks. The forecast of hot and humid weather is not so unusual, but the addition of smoke from fires in the western states promises to intensify the haze and create an especially irritating glare for those searching the skies for raptors.
Smoke from fires along the California coast and in central Utah can be seen streaming east this morning. (NOAA/GOES image)Smoke from western fires and humid air creates a band of haze in Ohio, Pennsylvania, New Jersey, and states to the south this morning. (NOAA/GOES image)
A migrating Broad-winged Hawk in the glare of a hazy sky. In addition to visibility problems, swarms of Spotted Lanternflies above the treetops make distant hawks difficult to discern for hawk watchers scanning the horizon with binoculars.
It may seem gloomy for the mid-September flights in 2021, but hawk watchers are hardy types. They know that the birds won’t wait. So if you want to see migrating “Broad-wings” and other species, you’ve got to get out there and look up while they’re passing through.
Migrating Ospreys typically fly low enough and are large enough to be spotted even during the haziest of conditions.Bald Eagles like this fourth-year bird can ascend to great altitude, but their size usually prevents them from sneaking past a lookout unnoticed.Peregrines escape notice not due to hazy sky conditions, but because they pass by so quickly. They’re being seen at local hawk watches now through October.
These hawk watches in the Lower Susquehanna River Watershed are currently staffed by official counters and all welcome visitors:
Rocky Ridge County Park Hawk Watch—3699 Deininger Road off Mount Zion Road (Route 24) northeast of York, Pennsylvania.
Second Mountain Hawk Watch—off Cold Spring Road on the grounds of Fort Indiantown Gap in Lebanon County, Pennsylvania.
Waggoner’s Gap Hawk Watch—where Route 74 crosses Blue Mountain north of Carlisle, Pennsylvania.
—or you can just keep an eye on the sky from wherever you happen to be. And don’t forget to check the trees and shrubs because warbler numbers are peaking too! During recent days…
Northern Parula at Chiques Rock County Park in Lancaster County.Black-and-white Warbler at Rocky Ridge County Park in York County.Cape May Warbler at Chiques Rock County Park in Lancaster County.Bay-breasted Warbler at Rocky Ridge County Park in York County.
It’s been more than a week since Tropical Storm Isaias moved swiftly up the Atlantic seaboard leaving wind and flood damage in its wake. Here in the Lower Susquehanna River Watershed, the brevity of its presence minimized the effects.
Tropical Storm Isaias moves north-northeast across the Delmarva Peninsula.
You may have noticed some summertime visitors flying about during these hot humid days that followed Isaias’ passing. They’re the dragonflies.
Our familiar friend the Wandering Glider is widespread throughout the valley right now—dropping eggs on shiny automobile hoods that look to them like a nice quiet puddle of water.
The Wandering Glider is a global traveler. Here in the lower Susquehanna valley, it is currently abundant around still water and in large parking lots.
Each of the other common migratory species is here too. Look for them patrolling the skies over large bodies of water and over adjacent fallow land and meadows where tiny flying insects abound. Did these dragonflies arrive on the winds associated with the tropical storm, or did they move in with the waves of warm air that followed it? Probably a little of both.
The Common Green Darner, a large dragonfly, can be the most abundant of the migratory species. Watch for high-flying swarms in the coming weeks.The Black Saddlebags is recognized in flight by the black base of each hindwing. These patches give the appearance of a pair of saddlebags draped across the dragonfly’s thorax.The Twelve-spotted Skimmer(Libellula pulchella) is a regular migrant.The Prince Baskettail (Epitheca princeps) can occur among mixed groups of dragonflies. Despite it rarely being mentioned as a migrant, its proclivity for non-stop day-long flight makes it a likely sighting among sizeable swarms.
Big swarms of dragonflies don’t go unnoticed by predators—particularly birds. The southbound migration of kites, Broad-winged Hawks, American Kestrels, and Merlins often coincides with the swarming of migratory dragonflies in late summer. Each of these raptors will grab and feed upon these insects while on the wing—so keep an eye on the sky.
This Peregrine Falcon found the congregations of hundreds of dragonflies worthy of a closer look……and an acrobatic fly-by to disrupt the swarms. Fun, fun, fun, fun, fun.
