Living in the Shadows

They get a touch of it here, and a sparkle or two there.  Maybe, for a couple of hours each day, the glorious life-giving glow of the sun finds an opening in the canopy to warm and nourish their leaves, then the rays of light creep away across the forest floor, and it’s shade for the remainder of the day.

The flowering plants which thrive in the understory of the Riparian Woodlands often escape much notice.  They gather only a fraction of the daylight collected by species growing in full exposure to the sun.  Yet, by season’s end, many produce showy flowers or nourishing fruits of great import to wildlife.  While light may be sparingly rationed through the leaves of the tall trees overhead, moisture is nearly always assured in the damp soils of the riverside forest.  For these plants, growth is slow, but continuous.  And now, it’s show time.

So let’s take a late-summer stroll through the Riparian Woodlands of Conewago Falls, minus the face full of cobwebs, and have a look at some of the strikingly beautiful plants found living in the shadows.

Oxeye (Heliopsis helianthoides) is common on the interior and along the edges of Riparian Woodland.  Specimens in deep shade flower less profusely and average less than half the height of the five feet tall inhabitants of edge environs.
Pale Touch-Me-Not (Impatiens pallida) is one of two species of native Impatiens found in the river floodplain.  Both are known as Jewelweed.  The stems and leaves of the indigenous Impatiens retain a great quantity of water, so life in filtered sunlight is essential to prevent desiccation.  Contrary to popular folklore, extracts of Jewelweed plants are not effective treatments of Poison Ivy (Toxicodendron radicans) contact dermatitis.
Spotted Touch-Me-Not (Impatiens capensis) is typically found in wetter soil than I. pallida.  Both Jewelweeds develop popping capsules which help to distribute the seeds of these annual wildflowers.  “Touch Me Not”, or you’ll be wearing tiny seeds.
Green-headed Coneflower (Rudbeckia laciniata) grows to heights of eight feet in full sun, hence its alternate common name, Tall Coneflower.  In deep shade, it may not exceed two feet in height.  Floodplains are the prime domain of this perennial.
Wingstem (Verbesina alternifolia) normally flowers no earlier than late August.  The bases of the leaves are continued onto the stem of the plant to form wings which extend downward along its length.  This wildflower tolerates shade, but flowers more profusely along the woodland edge.
Great Lobelia (Lobelia siphilitica), or Great Blue Lobelia, is a magnificent wetland and moist woodland wildflower, usually attaining three feet in height and adorned with a plant-topping spike of blossoms.  Invasive Japanese Stiltgrass (Microstegium vimineum) can be seen here competing with this plant, resulting in a shorter, less productive Lobelia.  Stiltgrass was not found in the Susquehanna River floodplain at Conewago Falls until sometime after 1997.  It has spread to all areas of woodland shade, its tiny seeds being blown and translocated along roads, mowed lots, trails, and streams to quickly colonize and overtake new ground.
American Bladdernut (Staphylea trifolia), a shrub of shaded woods, develops inflated capsules which easily float away during high water to distribute the seeds contained inside.
Buttonbush (Cephalanthus occidentalis) is a shrub of wet soils which produces a strange spherical flower, followed by this globular seed cluster.
Common Pawpaw (Asimina triloba) is a colony-forming small tree which produces a fleshy fruit.  It is the host plant for the caterpillars of the Zebra Swallowtail.  The plant and the butterfly approach the northern limit of their geographic range at Conewago Falls.
Common Spicebush (Lindera benzoin) is a widespread understory shrub in wet floodplain soils.  It is the host plant for the caterpillars of the Spicebush Swallowtail (Papilio troilus).
Sweet Autumn Virgin’s Bower (Clematis terniflora) is an escape from cultivation which has recently naturalized in the edge areas of the Conewago Falls Riparian Woodlands.  This vine is very showy when flowering and producing seed, but can be detrimental to some of the understory shrubs upon which it tends to climb.

SOURCES

Long, David; Ballentine, Noel H.; and Marks, James G., Jr.  1997.  Treatment of Poison Ivy/Oak Allergic Contact Dermatitis With an Extract of Jewelweed.  American Journal of Contact Dermatitis.  8(3): pp. 150-153.

Newcomb, Lawrence.  1977.  Newcomb’s Wildflower Guide.  Little, Brown and Company.  Boston, Massachusetts.

Yellowlegs for Breakfast

A few nocturnal migrants flew through the moonlit night to arrive at Conewago Falls for a sunrise showing this morning.  A dozen warblers were in the treetops and a Wood Thrush (Hylocichla mustelina) chattered away in the understory of the Riparian Woodlands.  Three species of shorebirds were in the falls and on the Pothole Rocks: Least Sandpiper (Calidris minutilla), Lesser Yellowlegs (Tringa flavipes), and Greater Yellowlegs (Tringa melanoleuca).

A Greater Yellowlegs (right) and two Lesser Yellowlegs sandpipers dropped by for breakfast.

The diurnal migration was highlighted by a Merlin (Falco columbarius), an Osprey, and a Bald Eagle, each flying down the river.  Most of the other birds in the falls seemed content to linger and feed.  There’s no need to hurry folks, only trouble lurks down there in paradise at the moment.

A light to moderate flight of nocturnal migrants in the eastern United States is displayed on NOAA National Weather Service NEXRAD radar at 4:58 AM EDT.  The eye of Hurricane Irma can be seen approaching the Florida Keys.  (NOAA/National Weather Service image)
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Piles of Green Tape

A couple of inches of rain this week caused a small increase in the flow of the river, just a burp, nothing major.  This higher water coincided with some breezy days that kicked up some chop on the open waters of the Susquehanna upstream of Conewago Falls.  Apparently it was just enough turbulence to uproot some aquatic plants and send them floating into the falls.

Piled against and upon the upstream side of many of the Pothole Rocks were thousands of two to three feet-long flat ribbon-like opaque green leaves of Tapegrass, also called Wild Celery, but better known as American Eelgrass (Vallisneria americana).  Some leaves were still attached to a short set of clustered roots.  It appears that most of the plants broke free from creeping rootstock along the edge of one of this species’ spreading masses which happened to thrive during the second half of the summer.  You’ll recall that persistent high water through much of the growing season kept aquatic plants beneath a blanket of muddy current.  The American Eelgrass colonies from which these specimens originated must have grown vigorously during the favorable conditions in the month of August.  A few plants bore the long thread-like pistillate flower stems with a fruit cluster still intact.  During the recent few weeks, there have been mats of American Eelgrass visible, the tops of their leaves floating on the shallow river surface, near the east and west shorelines of the Susquehanna where it begins its pass through the Gettysburg Basin near the Pennsylvania Turnpike bridge at Highspire.  This location is a probable source of the plants found in the falls today.

Uprooted American Eelgrass floating into the Pothole Rocks under the power of a north wind.  Note the white thread-like pistillate flower stem to the left and the small rooted specimen to the upper right.  The latter is likely a plant from the creeping rootstock on the edge of a colony.  As a native aquatic species, American Eelgrass is a critical link in the Susquehanna River and Chesapeake Bay food chain.  Its decimation by pollution during the twentieth century led to migration pattern alterations and severe population losses for the Canvasback (Aythya valisineria) duck.
American Eelgrass, a very small specimen, found growing in a low-lying Pothole Rock alongside the accumulations of freshly arriving material from upstream.  Note that the creeping rootstock has leaves growing from at least three nodes on this plant.  Eelgrass dislocations are regular occurrences which sometimes begin new colonies, like the small one seen here in this Diabase Pothole Rock Microhabitat.

The cool breeze from the north was a perfect fit for today’s migration count.  Nocturnal migrants settling down for the day in the Riparian Woodlands at sunrise included more than a dozen warblers and some Gray Catbirds (Dumetella carolinensis).  Diurnal migration was underway shortly thereafter.

A moderate flight of nocturnal migrants is indicated around NEXRAD sites in the northeastern states at 3:18 AM EDT.  The outer rain bands of Hurricane Irma can be seen approaching the Florida Keys as the storm closes in on the peninsula.  (NOAA/National Weather Service image)

Four Bald Eagles were counted as migrants this morning.  Based on plumage, two were first-year eagles (Juvenile) seen up high and flying the river downstream, one was a second-year bird (Basic I) with a jagged-looking wing molt, and a third was probably a fourth year (Basic III) eagle looking much like an adult with the exception of a black terminal band on the tail.  These birds were the only ones which could safely be differentiated from the seven or more Bald Eagles of varying ages found within the past few weeks to be lingering at Conewago Falls.  There were as many as a dozen eagles which appeared to be moving through the falls area that may have been migrating, but the four counted were the only ones readily separable from the locals.

Red-tailed Hawks (Buteo jamaicensis) were observed riding the wind to journey not on a course following the river, but flying across it and riding the updraft on the York Haven Diabase ridge from northeast to southwest.

Bank Swallows (Riparia riparia) seem to have moved on.  None were discovered among the swarms of other species today.

Ruby-throated Hummingbirds, Caspian Terns, Cedar Waxwings (Bombycilla cedrorum), and Chimney Swifts (Chaetura pelagica) were migrating today, as were Monarch butterflies.

Not migrating, but always fun to have around, all four wise guys were here today.  I’m referring to the four members of the Corvid family regularly found in the Mid-Atlantic states: Blue Jay (Cyanocitta cristata), American Crow (Corvus brachyrhynchos), Fish Crow (Corvus ossifragus), and Common Raven (Corvus corax).

It looks like a big Blue Jay, but it’s not.  This Belted Kingfisher (Megaceryle alcyon) takes a break after flying around the falls trying to shake a marauding Ruby-throated Hummingbird off its tail.
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SOURCES

Klots, Elsie B.  1966.  The New Field Book of Freshwater Life.  G. P. Putnam’s Sons.  New York, NY.

Suggestive Selling

A Baltimore Oriole (Icterus galbula) glowed in the first sunlight of the day as it began illuminating the treetops.  I’m not certain of the cause, but I often have the urge to dig into a bowl of orange sherbet after seeing one these magnificent blackbirds.  That’s right, in the Americas, orioles and blackbirds are members of the same family, Icteridae.  Look at blackbirds more carefully, you might see the resemblance.

Sunshine at dawn and migrating warblers were again active in the foliage.  Eight species were identified today.  Off to the tropics they go.  To the land of palm and citrus, yes citrus…limes, lemons, grapefruits, and oranges.

The Ruby-throated Hummingbirds (Archilochus colubris) are on the way toward the gulf states, then on to Central and South America.  Five dashed by the rocky lookout in the falls this morning.  Remember, keep your feeders clean, wash and rinse all the parts, and refill them with a fresh batch of “nectar”, four or five parts water to one part sugar.  Repeating this process daily during hot weather should keep contamination from overtaking your feeder.  It’s not a bad idea to rotate two feeders.  Have one cleaned, rinsed, and air drying while the second is filled and in use at your feeding station, then just swap them around.  Your equipment will be just as clean as it is at the sanitary dairy…you know, where they make sherbet.

The first of the season Caspian Terns (Hydroprogne caspia), giant freshwater versions of the terns you see at the seashore, passed through the falls late this morning.  Their bills are blood-red, not orange like the more familiar terns on the coast.  They’re stunning.

Ring-billed Gulls (Larus delawarensis) have been at the falls for several weeks.  Total numbers and the composition of the age groups in the flock change over the days, so birds appear to be trickling through and are then replaced by others coming south.  The big push of southbound migrants for this and many other species that winter locally in the Mid-Atlantic region and in the southern United States is still more than a month away.  There are still plenty more birds to come after the hours of daylight are reduced and the temperatures take a dip.

A Ring-billed Gull on the lookout for a morning snack.  They’ll eat almost anything and do a good job of keeping the river picked clean of the remains of animals that have met misfortune.  They’ll linger around landfills, hydroelectric dams, and fast-food restaurant parking lots through the winter.
Turkey Vultures (Cathartes aura) are common around the falls due to the abundance of carrion in the vicinity and because of the strong thermal updrafts of air over the sun-heated Pothole Rocks.  These rising currents provide lift for circling vultures.  We would expect migrating birds of a number of species will also take advantage of these thermals to gain altitude and extend the distance of their glides.

Some migrating butterflies were counted today.  Cloudless Sulphurs, more of a vagrant than a migrant, and, of course, Monarchs.  I’ll bet you know the Monarch, it’s black and orange.  How can you miss them, colored orange.

That’s it, that’s all for now, I bid you adieu…I’m going to have a dip of orange sherbet, or two.

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Beauties

It’s tough being good-looking and liked by so many.  You’ve got to watch out, because popularity makes you a target.  Others get jealous and begin a crusade to have you neutralized and removed from the spotlight.  They’ll start digging to find your little weaknesses and flaws, then they’ll exploit them to destroy your reputation.  Next thing you know, people look at you as some kind of hideous scoundrel.

Today, bright afternoon sunshine and a profusion of blooming wildflowers coaxed butterflies into action.  It was one of those days when you don’t know where to look first.

A Pearl Crescent (Phyciodes tharos) sipping nectar from Rough Boneset (Eupatorium pilosum) flowers.  Asters (Aster) are the host plants for the larvae of this butterfly.
A Common Buckeye (Junonia coenia) on Rough Boneset.  Its caterpillars are known to feed on members of the Acanthus family, possibly including the Water Willow (Justicia americana) which is so abundant in Conewago Falls.
Visitors from south of the Mason-Dixon Line arrived on the recent warm winds.  Two Cloudless Sulphurs (Phoebis sennae) patrolled the Riverine Grasslands, especially near the stands of Partridge Pea, a possible host plant.  One is seen here visiting a Halbred-leaved Rose Mallow blossom.  These large yellow butterflies are always a standout.

Purple Loosestrife (Lythrum salicaria) has a bad reputation.  Not native to the Americas, this prolific seed producer began spreading aggressively into many wetlands following its introduction.  It crowds out native plant species and can have a detrimental impact on other aquatic life.  Stands of loosestrife in slow-moving waters can alter flows, trap sediment, and adversely modify the morphology of waterways.  Expensive removal and biological control are often needed to protect critical habitat.

The dastardly Purple Loosestrife may have only two positive attributes.  First, it’s a beautiful plant.  And second, it’s popular; butterflies and other pollinators find it to be irresistible and go wild over the nectar.

A Silver-spotted Skipper (Epargyreus clarus) feeding on Purple Loosestrife nectar.  The host plants for this common butterfly’s caterpillars are a wide variety of Legumes.
A Cabbage White (Pieris rapae), a butterfly introduced from Europe in the 1800s, feeds on introduced Purple Loosestrife.
A Wild Indigo Duskywing (Erynnis baptisiae) feeding on Purple Loosestrife. This butterfly has expanded its population and range by using the introduced Crown Vetch as a host plant.

Don’t you just adore the wonderful butterflies.  Everybody does.  Just don’t tell anyone that they’re pollinating those dirty filthy no-good Purple Loosestrife plants.

SOURCES

Brock, Jim P., and Kenn Kaufman.  2003.  Butterflies of North America.  Houghton Mifflin Company.  New York.

Newcomb, Lawrence.  1977.  Newcomb’s Wildflower Guide.  Little, Brown and Company.  Boston, Massachusetts.

The Dungeon

There’s something frightening going on down there.  In the sand, beneath the plants on the shoreline, there’s a pile of soil next to a hole it’s been digging.  Now, it’s dragging something toward the tunnel it made.  What does it have?  Is that alive?

We know how the system works, the food chain that is.  The small stuff is eaten by the progressively bigger things, and there are fewer of the latter than there are of the former, thus the whole network keeps operating long-term.  Some things chew plants, others devour animals whole or in part, and then there are those, like us, that do both.  In the natural ecosystem, predators keep the numerous little critters from getting out of control and decimating certain other plant or animal populations and wrecking the whole business.  When man brings an invasive and potentially destructive species to a new area, occasionally we’re fortunate enough to have a native species adapt and begin to keep the invader under control by eating it.  It maintains the balance.  It’s easy enough to understand.

Japanese Beetles (Popillia japonica) seen here on Halberd-leaved Rose Mallow.  Without predation, exploding numbers of this invasive non-indigenous insect can defoliate and kill numerous species of plants in a given area.
The Song Sparrow (Melospiza melodia) is a generalist feeder, eating seeds and invertebrates including Japanese Beetles.  This species is the omnipresent year-round occupant of shoreline vegetation along the lower Susquehanna River.

Late summer days are marked by a change in the sounds coming from the forests surrounding the falls.  For birds, breeding season is ending, so the males cease their chorus of songs and insects take over the musical duties.  The buzzing calls of male “annual cicadas” (Neotibicen species) are the most familiar.  The female “annual cicada” lays her eggs in the twigs of trees.  After hatching, the nymphs drop to the ground and burrow into the soil to live and feed along tree roots for the next two to five years.  A dry exoskeleton clinging to a tree trunk is evidence that a nymph has emerged from its subterranean haunts and flown away as an adult to breed and soon thereafter die.  Flights of adult “annual cicadas” occur every year, but never come anywhere close to reaching the enormous numbers of “periodical cicadas” (Magicicada species).  The three species of “periodical cicadas” synchronize their life cycles throughout their combined regional populations to create broods that emerge as spectacular flights once every 13 or 17 years.

An “annual cicada” also known as a “dog-day cicada”, clings to the stem of a Halberd-leaved Rose Mallow at Conewago Falls.

For the adult cicada, there is danger, and that danger resembles an enormous bee.  It’s an Eastern Cicada Killer (Specius speciosus) wasp, and it will latch onto a cicada and begin stinging while both are in flight.  The stings soon paralyze the screeching, panicked cicada.  The Cicada Killer then begins the task of airlifting and/or dragging its victim to the lair it has prepared.  The cicada is placed in one of more than a dozen cells in the tunnel complex where it will serve as food for the wasp’s larvae.  The wasp lays an egg on the cicada, then leaves and pushes the hole closed.  The egg hatches in a several days and the larval grub is on its own to feast upon the hapless cicada.

An Eastern Cicada Killer (Sphecius speciosus) along the river shoreline. Despite their intimidating appearance, they do not sting humans and can be quite docile when approached.

Other species in the Solitary Wasp family (Sphecidae) have similar life cycles using specific prey which they incapacitate to serve as sustenance for their larvae.

A Solitary Wasp, one of the Thread-waisted Wasps (Ammophila species), drags a paralyzed moth caterpillar to its breeding dungeon in the sandy soil at Conewago Falls.  For the victim, there is no escape from the crypt.

The Solitary Wasps are an important control on the populations of their respective prey.  Additionally, the wasp’s bizarre life cycle ensures a greater survival rate for its own offspring by providing sufficient food for each of its progeny before the egg beginning its life is ever put in place.  It’s complete family planning.

The cicadas reproduce quickly and, as a species, seem to endure the assault by Cicada Killers, birds, and other predators.  The periodical cicadas (Magicicada), with adult flights occurring as a massive swarm of an entire population every thirteen or seventeen years, survive as species by providing predators with so ample a supply of food that most of the adults go unmolested to complete reproduction.  Stay tuned, 2021 is due to be the next periodical cicada year in the vicinity of Conewago Falls.

SOURCES

Eaton, Eric R., and Kenn Kaufman.  2007.  Kaufman Field Guide to Insects of North America.  Houghton Mifflin Company.  New York.

Fussy Eaters

She ate only toaster pastries…that’s it…nothing else.  Every now and then, on special occasions, when a big dinner was served, she’d have a small helping of mashed potatoes, no gravy, just plain, thank you.  She received all her nutrition from several meals a week of macaroni and cheese assembled from processed ingredients found in a cardboard box.  It contains eight essential vitamins and minerals, don’t you know?  You remember her, don’t you?

Adult female butterflies must lay their eggs where the hatched larvae will promptly find the precise food needed to fuel their growth.  These caterpillars are fussy eaters, with some able to feed upon only one particular species or genus of plant to grow through the five stages, the instars, of larval life.  The energy for their fifth molt into a pupa, known as a chrysalis, and metamorphosis into an adult butterfly requires mass consumption of the required plant matter.  Their life cycle causes most butterflies to be very habitat specific.  These splendid insects may visit the urban or suburban garden as adults to feed on nectar plants, however, successful reproduction relies upon environs which include suitable, thriving, pesticide-free host plants for the caterpillars.  Their survival depends upon more than the vegetation surrounding the typical lawn will provide.

The Monarch (Danaus plexippus), a butterfly familiar in North America for its conspicuous autumn migrations to forests in Mexico, uses the milkweeds (Asclepias) almost exclusively as a host plant.  Here at  Conewago Falls, wetlands with Swamp Milkweed (Asclepias incarnata) and unsprayed clearings with Common Milkweed (A. syriaca) are essential to the successful reproduction of the species.  Human disturbance, including liberal use of herbicides, and invasive plant species can diminish the biomass of the Monarch’s favored nourishment, thus reducing significantly the abundance of the migratory late-season generation.

Monarch caterpillar after a fourth molt.  The fifth instar feeding on Swamp Milkweed.
A fifth molt begets the Monarch pupa, the chrysalis, from which the showy adult butterfly will emerge.
Adult Monarch feeding on Goldenrod (Solidago) nectar.

Butterflies are good indicators of the ecological health of a given environment.  A diversity of butterfly species in a given area requires a wide array of mostly indigenous plants to provide food for reproduction.  Let’s have a look at some of the species seen around Conewago Falls this week…

An adult Silvery Checkerspot (Chlosyne nycteis) visiting a nectar plant, Partridge Pea (Chamaecrista fasciculata).  Wingstem (Verbesina alternifolia), a plant of the Riparian Woodlands, is among the probable hosts for the caterpillars.
A Gray Hairstreak (Strymon melinus) visits Crown Vetch, a possible host plant.  Other potential larval food in the area includes Partridge Pea, Halberd-leaved Rose Mallow (Hibiscus laevis) of the river shoreline, and Swamp Rose Mallow (Hibiscus moscheutos), a plant of wetlands.
The Eastern Tailed Blue (Cupido comyntas) may use Partridge Pea , a native wildflower species, and the introduced Crown Vetch (Securigera/Coronilla varia) as host and nectar plants at Conewago Falls.
The Least Skipper (Ancyloxypha numitor) is at home among tall grasses in woodland openings, at riverside, and in the scoured grassland habitat of the Pothole Rocks in the falls.  Host plants available include Switchgrass (Panicum vigatum), Freshwater Cordgrass (Spartina pectinata), and Foxtails (Setaria).
The Zabulon Skipper (Poanes zabulon) is an inhabitant of moist clearings where the caterpillars may feed upon Lovegrasses (Eragrostis) and Purpletop (Tridens flavus).
The Eastern Tiger Swallowtail (Papilio glaucus), a female seen here gathering nectar from Joe-Pye Weed (Eutrochium), relies upon several forest trees as hosts. Black Cherry (Prunus serotina), Willow (Salix), Yellow Poplar (Liriodendron tulipifera), also known as Tuliptree, and Green Ash (Fraxinus pennsylvanica) are among the local species known to be used.  The future of the latter food species at Conewago Falls is doubtful.  Fortunately for the Eastern Tiger Swallowtail, the “generalist” feeding requirements of this butterfly’s larvae enable the species to survive the loss of a host plant.
A female Eastern Tiger Swallowtail, black morph, gathering nectar from Joe-Pye Weed.
The Zebra Swallowtail (Protographium marcellus), an adult seen here on Joe-Pye Weed, feeds exclusively upon Pawpaw (Asimina) trees as a caterpillar.  This butterfly species may wander, but its breeding range is limited to the moist Riparian Woodlands where colonial groves of Pawpaw may be found.  The Common Pawpaw (Asimina triloba), our native species in Pennsylvania, and the Zebra Swallowtail occur at the northern edge of their geographic ranges in the Lower Susquehanna River Watershed.  Planting Pawpaw trees as an element of streamside reforestation projects certainly benefits this marvelous butterfly.

The spectacularly colorful butterflies are a real treat on a hot summer day.  Their affinity for showy plants doubles the pleasure.

By the way, I’m certain by now you’ve recalled that fussy eater…and how beautiful she grew up to be.

SOURCES

Brock, Jim P., and Kaufman, Kenn.  2003.  Butterflies of North America.  Houghton Mifflin Company.  New York, NY.

Digging In

If you visit the shores of the Susquehanna River during the warmer months of the year, there’s a pretty good probability that you’ll be taking a visitor along home with you.  Not to worry, it won’t raid the icebox or change the television channels when you leave the room to get a snack.  It won’t put you in the doghouse with the landlord for having a forbidden pet.  As a matter of fact, you may not even notice your new companion.  Sure enough though, it’s there, crawling through the luxurious warm fabric of your clothing and seeking out a good place to dig in and chow down.  O.K., so now you’re worried.

Ticks, particularly the American Dog Tick (Dermacentor variabilis), are widespread in the Lower Susquehanna River Watershed.  Like spiders, they are arachnids.  They have a four-stage life cycle (egg-larva-nymph-adult) which, in the case of D. variabilis, requires a minimum of two months to complete.  Females lay up to 6,500 eggs on the ground.  Then the fun begins as the larvae with any hope of survival must attach to a small mammal to feed.  They can survive for almost a year before finding a host.  After a successful hookup and subsequent blood feast of up to two weeks duration, the larva drops to the ground, molts into a nymph, and finds another small mammal, usually a bit bigger this time, to feed upon.  A nymph can survive for up to six months before needing to feed.  Finding the second host, the nymph feeds for 3 to 10 days, then drops to the ground to molt into an adult.  Adult American Dog Ticks can endure up to two years without feeding on a host.  The adults mate and feed on larger mammals such as deer and domestic animals including, of course, dogs.  After a blood meal of five days to two weeks duration, the adult female tick drops to the ground to lay eggs and initiate a new generation.

The adult American Dog Tick attaches to a potential host by hanging from vegetation and grabbing the passing victim with its forward legs to hitch a ride.  If undetected, a female will find a nice warm spot and “dig in” to begin feeding.  This male is looking for love, and in all the wrong places.

The American Dog Tick is renowned as a carrier of the Rocky Mountain Spotted Fever bacteria (Rickettsia rickettsii).  The bacteria is vectored by the ticks from rodents to dogs and humans.  The adult tick must be attached to the victim for a minimum of six to eight hours to transmit the pathogen.  A rash spreading from the wrists and the ankles to other portions of the body begins two to fourteen days after infection.

Tularemia, caused by the bacteria Francisella tularensis, can be passed by the American Dog Tick.  Symptoms can appear in three to twenty-one days and include chills, fever, and inflammation of the lymph nodes.

American Dog Ticks which attach to dogs, particularly near the neck, and are left in place to feed and engorge themselves for longer than five days can cause Canine Tick Paralysis.  Symptoms usually begin to subside only after a recovery period following removal of the arachnid.

The American Dog Tick is exposed to Borrelia burgdorferi, the bacteria responsible for Lyme Disease, however, transmittal of this pathogen is by the smaller Deer Tick (Ixodes scapularis), also known as the Black-legged Tick.  The Deer Tick is not presently common at Conewago Falls.  In the adjacent uplands, it is widespread and is carrying Lyme Disease where the White-tailed Deity (Odocoileus virginianus), the preferred host for the ticks, is found along with mice and other small rodents, the source of B. burgdorferi bacteria.  The Deer Tick easily escapes notice and cases of Lyme Disease are frequent, so vigilance is necessary.

SOURCES

Chan, Wai-Han, and Kaufman, Phillip.  2008.  American Dog Tick.  University of Florida Featured Creatures website  entnemdept.ufl.edu/creatures/urban/medical/american_dog_tick.htm  as accessed July 30, 2017.

 

The Antagonist

They can be a pesky nuisance.  The annoying high-frequency buzzing is bad enough, but it’s the quiet ones that get you.  While you were swatting at the noisy one, the silent gender sticks you and begins to feed.  Maybe you know it, or maybe you don’t.   She could make you itch and scratch.  If she’s carrying a blood-borne pathogen, you could get sick and possibly die.

To humans, mosquitos are the most dangerous animal in the world (though not in the United States where man himself and the domestic dog are more of a threat).  Globally, the Anopheles mosquitos that spread Malaria have been responsible for millions and millions of human deaths.  Some areas of Africa are void of human habitation due to the prevalence of Malaria-spreading Anopheles mosquitos.  In the northeastern United States, the Northern House Mosquito (Culex pipiens), as the carrier of West Nile Virus, is the species of greatest concern.  Around human habitations, standing water in tires, gutters, and debris are favorite breeding areas.  Dumping stagnant water helps prevent the rapid reproduction of this mosquito.

In recent years, the global distribution of these mosquito-borne illnesses has been one of man’s inadvertent accomplishments.  An infected human is the source of pathogens which the feeding mosquito transmits to another unsuspecting victim.  Infectious humans, traveling the globe, have spread some of these diseases to new areas or reintroduced them to sectors of the world where they were thought to have been eliminated.  Additionally, where the specific mosquito carrier of a disease is absent, the mobility of man and his cargos has found a way to transport them there.  Aedes aegypti, the “Yellow Fever Mosquito”, carrier of its namesake and the Zeka Virus, has found passage to much of the world including the southern United States.  Unlike other species, Aedes aegypti dwells inside human habitationsthus transmitting disease rapidly from person to person.  Another non-native species, the Asian Tiger Mosquito (Aedes albopictus), vector of Dengue Fever in the tropics, arrived in Houston in 1985 in shipments of used tires from Japan and in Los Angeles in 2001 in wet containers of “lucky bamboo” from Taiwan…some luck.

Asian Tiger Mosquito in action during the daylight hours, typical behavior of the genus. This species has been found in the area of Conewago Falls since at least 2013.

Poor mosquito, despite the death, suffering, and misery it has brought to Homo sapiens and other species around the planet, it will never be the most destructive animal on earth.  You, my bloodthirsty friends, will place second at best.  You see, mosquitos get no respect, even if they do create great wildlife sanctuaries by scaring people away.

The winner knows how to wipe out other species and environs not only to ensure its own survival, but, in many of its populations, to provide leisure, luxury, gluttony, and amusement.  This species possesses the cognitive ability to think and reason.  It can contemplate its own existence and the concepts of time.  It is aware of its history, the present, and its future, though its optimism about the latter may be its greatest delusion.  Despite possessing intellect and a capacity to empathize, it is devious, sinister, and selfish in its treatment of nearly every other living thing around it.  Its numbers expand and its consumption increases.  It travels the world carrying pest and disease to all its corners.  It pollutes the water, land, and air.  It has developed language, culture, and social hierarchies which create myths and superstitions to subdue the free will of its masses.   Ignoring the gift of insight to evaluate the future, it continues to reproduce without regard for a means of sustenance.  It is the ultimate organism, however, its numbers will overwhelm its resources.  The crowning distinction will be the extinction.

Homo sapiens will be the first animal to cause a mass extinction of life on earth.  The forces of nature and the cosmos need to wait their turn; man will take care of the species annihilation this time around.  The plants, animals, and clean environment necessary for a prosperous healthy life will cease to exist.  In the end, humans will degenerate, live in anguish, and leave no progeny.  Fate will do to man what he has done to his co-inhabitants of the planet.

The Bald Eagle (Haliaeetus leucocephalus) is again a breeding species in the Susquehanna River watershed.  It is generally believed that during the mid-twentieth century, Dichlorodiphenyltrichloroethane (DDT) pesticide residues accumulated in female top-of-the-food-chain birds including Bald Eagles.  As a result, thinner egg shells were produced.  These shells usually cracked during incubation, leading to failed reproduction in entire populations of birds, particularly those that fed upon fish or waterfowl.  In much of the developed world, DDT was used liberally during the mid-twentieth century to combat Malaria by killing mosquitos.  It was widely used throughout the United States as a general insecticide until it was banned here in 1972.  (Editors Note:  There is the possibility that polychlorinated biphenyls [P.C.B.s] and other industrial pollutants contributed to the reproductive failure of birds at the apex of aquatic food chains.  Just prior to the recovery of these troubled species, passage of the Clean Water Act in 1972 initiated reductions in toxic discharges from point sources into streams, rivers, lakes, bays, and oceans.  Production of P.C.B.s was banned in the United States in 1978.  Today, P.C.B.s from former discharge and dumping sites continue to be found in water.  Spills can still occur from sources including old electric transformers.)
To substitute any other beast would be folly.  Man, the human, Homo sapiens, the winner and champion, will repeatedly avail himself as the antagonist during our examination of the wonders of wildlife.  He is the villain.  The tragedy of his self-proclaimed dominion over the living things of the world will wash across these pages like muddy water topping a dam.  There’s nothing I can do about it, aside from fabricating a bad novel with a fictional characterization of man.  So let’s get on with it and take a look at “A Natural History of Conewago Falls”.  Let’s discover the protagonist, the heroic underdog of our story, “Life in the Lower Susquehanna River Watershed.”

Over the top today.
SOURCES

Avery, Dennis T.  1995.  Saving the Planet with Pesticides and Plastic: The Environmental Triumph of High-Yield Farming.  Hudson Institute.  Indianapolis, Indiana.

Eaton, Eric R., and Kenn Kaufman.  2007.  Kaufman Field Guide to Insects of North America.  Houghton Mifflin Co.  New York.

Newman, L.H.  1965.  Man and Insects.  The Natural History Press.  Garden City, New York.