Arthropods (Arthropoda) – LIFE IN THE LOWER SUSQUEHANNA RIVER WATERSHED

October 4, 2025

A Sixth Species of Annual Cicada in the Lower Susquehanna Basin: Neotibicen robinsonianus

Just as we were very pleased last month to have the opportunity to hear the sounds of the rarest of the Periodical Cicadas—the Little Seventeen-year Cicada—in the Conewago Hills of York County to thus provide our only record of the species during the Brood XIV emergence in the Lower Susquehanna River Watershed, we were this week delighted to find and record a population of what may be the valley’s rarest cicada experiencing annual flights—the Robinson’s Cicada—just a few miles away at Gifford Pinchot State Park.

Like the Little Seventeen-year Cicada, Robinson’s Cicada (Neotibicen robinsonianus) is a species found more commonly in the southern United States, occurring with scattered distribution in a range that extends west into Missouri, Kansas, and Texas. They are of rare occurrence in the lands of the Chesapeake drainage basin in Virginia and Maryland.

Populations of Swamp Cicadas, also known as Morning Cicadas, are presently in the midst of their yearly courtship rituals. Their songs are easily heard in woods, edge habitat, and suburbia throughout the region during the AM hours. It was while recording the sounds of these common insects in the Conewago Day-use Area at Gifford Pinchot State Park that we noticed a very unique song from high in the oaks, hickories, and other hardwoods along the lake. Listen to this sound clip featuring a group of serenading male Swamp Cicadas. In the background, a Robinson’s Cicada’s song consisting of a pulsing series of raspy buzzes, each about one second in duration, can be heard, particularly starting at 00:40.

The Lower Susquehanna River Watershed, and Gifford Pinchot State Park in particular, may currently represent the northern limit of the geographic range of the Robinson’s Cicada. If you’re in the area during the coming weeks, drop by the park and have a listen. And don’t forget to check out our “Cicadas” page for sound clips of all the species found in our area.

July 21, 2025July 21, 2025

Sunny Swarms of Insects

It begins on a sunny morning in spring each year, just as the ground temperature reaches sixty degrees or more…

Eastern Subterranean Termites emerge in unison from a nest located in the soil beneath a log. Each of these swarming “alates” is a potential king or queen seeking to find a location with an ample supply of fallen timber to provide food for establishment of a new colony. They are escorted to the surface by a soldier (lower center) equipped with powerful jaws for protecting the existing nest. Similar-looking worker termites tend the nest, the queen, her eggs, and their siblings, but usually remain hidden from view. The workers feed upon wood, hosting cellulose-digesting protozoa and bacteria in their guts to break down the fibers. This symbiotic relationship is an important mode of decomposition in the forest, the process that turns wood into the organic matter that enriches soil and helps it to retain more moisture.

Fungi on Fallen Log — Termites are among the numerous arthropods that join fungi and a variety of microbes to decompose dead wood and other plant matter into the nutrients and organic materials used by living plants to thrive and grow.

Green Frog Stalking a Eastern Subterranean Termite — Soon after lift off, the majority of swarming termites are consumed by swallows, swifts, and other birds, but some are discovered at ground level.

Green Frog Eating Termites — A lightning-fast strike with its tongue and this Green frog has snatched up yet another termite. Those that slip by the dragnet of terrestrial and aerial predators can sometimes start a new colony in the ground beneath a dead tree or in a vulnerable house or other wooden structure. To keep a small clan from invading your home, be certain the wood elements of your building(s) are kept dry and are not in contact with dirt, soil, bark mulch, etc. Regular inspections for evidence of their presence can head off the long-term damage termites can inflict on the stuff we construct with tree skeletons.

Of course, termites aren’t the only groups of insects to swarm. As heated runoff from slow-moving thundershowers has increased stream temperatures during the past couple of weeks, there have occurred a number of seasonal mayfly “hatches” on the Susquehanna and its tributaries. These “hatches” are actually the nuptial flights of newly emerged imago and adult mayflies. The most conspicuous of these is the Great Brown Drake.

Seen here with a much smaller and more typical regional mayfly to its left, the Great Brown Drake was for several years infamous for swarming the lights and creating traffic hazards on bridges spanning the Susquehanna. During the past two weeks, nighttime flights of these giants have ventured out to gather at well lit locations in housing and business districts more than a dozen miles from the river. Earlier this century, this proclivity to wander probably led the Great Brown Drake to first invade silty segments of the Susquehanna as a colonizer from its native range in the Mississippi watershed.

Swarms of another storm-related visitor are being seen throughout the lower Susquehanna valley right now. Have you noticed the Wandering Gliders?

Throughout the month, swarms of Wandering Gliders, the most widespread dragonflies in the world, descended on areas hit by localized slow-moving thundershowers. Large numbers of these global travelers are known to get swept up within the thermal air masses that lead to these storms. In suitable terrain within the path of the downpours, they linger to search for flooded places where they can mate and deposit eggs. Wandering Gliders frequently mistake large parking lots at shopping malls, grocery stores, etc. for wetlands and will be seen in these areas depositing eggs upon the hoods and roofs of shiny motor vehicles, surfaces which appear puddle-like in their eyes.

Mating Common Green Darners — More dragonfly swarms are yet to come. Adult Common Green Darners are presently in the Lower Susquehanna River Watershed mating and depositing eggs within vegetated ponds, lakes, and wetlands. Beginning in August, adult Common Green Darners and other migratory species including Black Saddlebags, Carolina Saddlebags, Wandering Gliders, and Twelve-spotted Skimmers will begin swarming as they feed on flying insects (including lots of mosquitos and gnats) and start working their way toward the Atlantic Coastal Plain. Along the barrier islands by September, concentrations of southbound dragonflies can reach the thousands, particularly at choke points like Cape May, New Jersey, and Cape Charles, Virginia. So be sure to keep an eye on the sky for swarms of dragonflies during coming weeks. And don’t forget to check out our “Damselflies and Dragonflies” page by clicking the tab at the top of this page.

July 11, 2025July 25, 2025

With Periodical Cicadas Gone, Here Come the Annual Cicadas…Right on Schedule!

Eastern Scissor Grinder — As the choruses of Brood XIV Periodical Cicadas fall silent for another seventeen years, the sounds of the more widespread annual Neotibicen cicadas are starting to be heard throughout the lower Susquehanna valley. These latter insects have a more abbreviated life cycle, spending just two to five years in the subterranean nymphal stage before appearing during mid-summer to breed. The emergence of these annual insects is not synchronized into broods, so some adults are found taking flight, singing, and mating every year. Over the centuries, male annual cicadas that emerged and commenced courtship songs earlier than July have certainly failed to successfully reproduce during Periodical Cicada years. So to avoid competition with the overwhelming drone of the seventeen-year cicadas that may emerge along with them, natural selection has delayed the maturation of the annual species until just after the periodicals have gone quiet. To see pictures and hear the sounds of our five (now 6 as of July 24th) species of annual cicadas, click the “Cicadas” tab at the top of this page.

July 6, 2025

Sorry, But We Have No Steamed Crabs Today

Earlier today, we collected some mint leaves from the garden to make a batch of iced tea. Just prior to plunging them into boiling water, we noticed this Goldenrod Crab Spider (Misumena vatia) hiding in the foliage.

June 28, 2025June 28, 2025

Brood XIV Periodical Cicada Wrap Up

While the heat and humidity of early summer blankets the region, Brood XIV Periodical Cicadas are wrapping up their courtship and breeding cycle for 2025. We’ve spent the past week visiting additional sites in and near the Lower Susquehanna River Watershed where their emergence is evident.

We begin in York County just to the west of the river and Conewago Falls in mostly forested terrain located just southeast of Gifford Pinchot State Park. Within this area, often called the Conewago Hills, a very localized population of cicadas could be heard in the woodlands surrounding the scattered homes along Bull Road. Despite the dominant drone of an abundance of singing Pharaoh Periodical Cicadas, we were able to hear and record the courtship song of a small number of the rare Little Seventeen-year Cicadas. Their lawn sprinkler-like pulsating songs help mate-seeking males penetrate the otherwise overwhelming chorus of the Pharaoh cicadas in the area.

The Little Seventeen-year Cicada’s (Magicicada septendecula) thorax is black between the eye and the origin of the wings. It is the rarest of the three species of seventeen-year cicadas.

From the Conewago Hills we moved northwest into the section of southern Cumberland County known as South Mountain. Here, Pharaoh Periodical Cicadas were widespread in ridgetop forests along the Appalachian Trail, particularly in the area extending from Long Mountain in the east through Mount Holly to forests south of King’s Gap Environmental Education Center in the west.

Pharaoh Periodical Cicadas along the Appalachian Trail on South Mountain, Cumberland County, Pennsylvania. Like the cicadas we visited last week on the east side of the Susquehanna, this population is surviving on lands with a history of timber harvest and charcoal production to fuel nearby iron furnaces during the nineteenth century.

While on South Mountain, we opted for a side trip into the neighboring Potomac watershed of Frederick County, Maryland, where these hills ascend to greater altitude and are known as the Blue Ridge Mountains, a name that sticks with them all the way through Shenandoah National Park, the Great Smoky Mountains, and to their southern terminus in northwestern Georgia. We found a fragmented emergence of Pharaoh Periodical Cicadas atop the Catoctin Mountain section of the Blue Ridge just above the remains of Catoctin Furnace, again on lands that had been timbered to make charcoal to fuel iron production prior to their protection as vast expanses of forest.

A Pharaoh Periodical Cicada on Catoctin Mountain near Catoctin Furnace south of Thurmont, Maryland. These cicadas are not part of a Brood XIV emergence, but are instead a population of Brood X (2021) stragglers.

Back in Pennsylvania, we’re on our way to the watersheds of the northernmost tributaries of the lower Susquehanna’s largest tributary, the Juniata River. There, we found Brood XIV cicadas more widespread and in larger numbers than occurred at previous sites. Both Pharaoh and Cassin’s Periodical Cicadas were seen and heard along Jack’s Mountain and the Kishacoquillas Creek north of Lewistown/Burnham in Mifflin County. To the north of the Kishacoquillas Valley and Stone Mountain in northernmost Huntingdon County, the choruses of the two species were again widespread, particularly along the forest edges in Greenwood Furnace State Park, Rothrock State Forest, and adjacent areas of the Standing Stone Creek watershed.

Tymbal on the male Pharaoh Periodical Cicada — A view of the sound-generating tymbal on a male Pharaoh Periodical Cicada at Greenwood Furnace State Park. Rapid vibration of the tymbals by a set of specialized muscles generates the distinctive calls and courtship songs of the various cicada species. When handled, these tymbals can produce a harsh “panic call”. This distress sound could startle a would-be predator and provide the cicada with an opportunity to escape.

Tymbal on a Cassin's Periodical Cicada — The sound organs comprised of ribbed tymbals and specialized muscles on the male Cassin’s Periodical Cicada generate a “panic call” as well as the distinctive calls and songs used to penetrate the droning choruses of the more numerous Pharaoh Periodical Cicadas with which it shares a seventeen-year flight.

Cassin's Periodical Cicadas — Using their specialized sound organs, Cassin’s Periodical Cicadas generate a courtship song that usually includes buzzy phrases and ticking notes (first sound clip). The buzzing and ticking helps the male Cassin’s cicada penetrate the songs of the more numerous Pharaoh Periodical Cicadas in the area (second sound clip). When synchronized into a chorus that surges in volume, the songs of Cassin’s Periodical Cicadas can overcome the overwhelming drone of the nearby Pharaoh cicadas (third sound clip).

Pharaoh Periodical Cicada Depositing Eggs — After mating and before the lives of these seventeen-year cicadas draw to a close, the females need to deposit their fertilized eggs into the small end twigs of suitable trees. On a small hawthorn tree (Cretaegus species) along the edge of the forest at Greenwood Furnace State Park, this Pharaoh Periodical Cicada is using her ovipositor to make a slit in a twig and place her eggs.

Cassin's Periodical Cicada Ovipositing — Simultaneously on the same little hawthorn tree, this female Cassin’s Periodical Cicada is depositing her fertilized eggs.

Within the last 48 hours, we visited one last location in the Lower Susquehanna River Watershed where Brood XIV Periodical Cicadas have emerged during 2025. In the anthracite coal country of Northumberland County, a flight of Pharaoh Periodical Cicadas is nearing its end. We found them to be quite abundant in forested areas of Zerbe Run between Big and Little Mountains around Trevorton and on the wooded slopes of Mahanoy Mountain south of nearby Shamokin. Line Mountain south of Gowen City had a substantial emergence as well.

Foliage with Evidence of Ovipositing — Brown leaves reveal the end twigs where female Pharaoh Periodical Cicadas have deposited eggs during the last several weeks. During July, the larvae will hatch and drop to the ground to start a new generation of Brood XIV cicadas. As subterranean nymphs, they’ll spend the coming seventeen years feeding on small amounts of xylem sap from tree roots. In 2042, during the next Brood XIV emergence, these nymphs will come to the surface and take flight as adults.

Dead Pharaoh Periodical Cicadas — Accumulations of deceased Pharaoh Periodical Cicadas on Line Mountain.

Fallen cicadas that show any sign of life are being snatched up by predators such as this Eastern Gartersnake (Thamnophis sirtalis sirtalis). Meanwhile, the remainder of the biomass is picked apart by scavengers or is left to reducers for breakdown into fertilizer and organic matter for the forest. Nothing goes to waste.

To chart our travels, we’ve put together this map plotting the occurrence of significant flights of Periodical Cicadas during the 2025 emergence. Unlike the more densely distributed Brood X cicadas of 2021, the range of Brood XIV insects is noticeably fragmented, even in areas that are forested. We found it interesting how frequently we found Brood XIV cicadas on lands used as sources of lumber to make charcoal for fueling nineteenth-century iron furnace operations.

The furnaces at Greenwood Furnace State Park required the daily cutting of one acre of timber to make enough charcoal to fuel the iron-making process. Did keeping thousands of acres in various stages of forest succession to supply the charcoal needs of these operations aid the survival of earlier generations of Periodical Cicadas on these lands? Or, after the furnaces converted to coal for fuel, did the preservation of many of these parcels as state, federal, and private forests allow the cicadas to find refuge from the widespread impacts of agriculture and expanding urbanization in adjacent lands? Maybe it’s a little of both. We always bear in mind that annual insects and other animals are more than one hundred generations removed from the negative or positive impacts of the early years of the industrial age, but only about ten generations have passed since populations of seventeen-year Periodical Cicadas were directly influenced by these factors. What do you think?

Well, that’s a wrap. Please don’t forget to check out our new Cicadas page by clicking the “Cicadas” tab at the top of this page. Soon after the Periodical Cicadas are gone, the annual cicadas will be emerging and our page can help you identify the five species found regularly in the lower Susquehanna valley. ‘Til next time, keep buzzing!

June 16, 2025June 20, 2025

Brood XIV Periodical Cicadas in the Lower Susquehanna River Watershed

Here are some sights and sounds from the ongoing emergence of Brood XIV Periodical Cicadas in the Lower Susquehanna River Watershed.

We begin in the easternmost spur of the lower basin where a sizeable emergence of cicadas can be seen and heard in the woodlands surrounding the headwaters of the Conestoga River in Berks County north of Morgantown. This flight extends east into Chester County and the French Creek drainage of the Schuylkill River watershed on State Game Lands 43 north of Elverson and consists of Pharaoh Periodical Cicadas (Magicicada septendecim), the most common species among 17-year broods.

Pharaoh Periodical Cicadas exuvia — Exuvia of a recently emerged Pharaoh Periodical Cicada.

Pharaoh Periodical Cicada singing — Soon after landing on a perch, a male Pharaoh Periodical Cicada will usually announce his presence by singing. It’s an attempt to quickly attract potential mates that may be in the vicinity.

Pharaoh Periodical Cicadas singing chorus — Gatherings of thousands of singing male Pharaoh Periodical Cicadas create a distinctive droning chorus.

Chestnut Oak Hosting Cicadas — After mating, female cicadas make slits in the end twigs of selected trees into which they lay their eggs. The process of egg-laying and larval emergence will usually wilt and kill end growth on the affected branches, causing little harm to healthy trees. It’s similar to the trim you might give to a bonsai plant to keep it stout and sturdy.

A deceased Pharaoh Periodical Cicada at the Fire Tower Parking Area at French Creek State Park. This specimen and a chorus on the hill’s forested south slope were the northeastern-most evidence of Brood XIV Periodical Cicadas we could find for the population cluster in portions of Berks, Chester, and Lancaster Counties around Morgantown.

Second-year Mississippi Kite Feeding on Cicadas — The abundance of Pharaoh Periodical Cicadas on State Game Lands 43 in Chester County has attracted numerous raptors, particularly wandering one-year-old birds that aren’t quite mature enough to nest. Among the sightings have been Red-tailed Hawks, Red-shouldered Hawks, Broad-winged Hawks, and at least three Mississippi Kites, a rarity on the Piedmont this far inland from the coastal plain. (See the post from June 5, 2021, for details on the occurrence of Mississippi Kites in northernmost Delaware during the Brood X emergence.)

From Route 82 north of Elverson to the west through the forested areas along Route 10 north of Morgantown and the Pennsylvania Turnpike, we found an abundance of Cassin’s Periodical Cicadas (Magicicada cassini) calling among the Pharaohs. This mix of Pharaoh and Cassin’s Periodical Cicadas extends west along the north side of the turnpike into Lancaster County and State Game Lands 52 on Black Creek north of Churchtown.

Cassin's Periodical Cicada — A Brood XIV Cassin’s Periodical Cicada at State Game Lands 52 is identified by the all-black margin between the eye and the wing root and…

Cassin's Periodical Cicadas singing chorus — To penetrate the sounds of the more common Pharaoh Periodical Cicadas, male Cassin’s Periodical Cicadas gather in large concentrations to generate a loud, oscillating chorus. Its surging volume will usually exceed that of the Pharaohs singing in the vicinity.

Cassin's Periodical Cicadas mating — The underside of copulating Cassin’s Periodical Cicadas.

Further west in Cornwall, Lebanon County, a Brood XIV emergence can be found on similar forested terrain: the Triassic hills of the Newark Basin—rich in iron ore and renowned for furnace operations during the eighteenth and nineteenth centuries. Pharaoh Periodical Cicadas were the only species heard among this population that extends from Route 72 east through the woodlands along Route 322 into the northern edge of State Game Lands 156 in Lancaster County.

On the west side of the Susquehanna, yet another isolated population of Brood XIV Periodical Cicadas can be found in Perry County, just south of Duncannon on State Game Lands 170 on the slopes of “Cove Mountain”, the canoe-shaped convergence of the western termini of Peters and Second Mountains.

Pharaoh Periodical Cicadas dominated this Perry County chorus,…

…but we did detect at least one Cassin’s Cicada trying to find a mate.

Not to say they aren’t present, but we have yet to detect the rarest species, Magicicada septendecula, the “Little Seventeen-year Cicada”, among the various populations of Brood XIV Periodical Cicadas emerging in the lower Susquehanna valley. For the coming two weeks or so until this brood is gone for another 17 years, the search continues.

For more on both annual and periodical cicada species in the Lower Susquehanna River Watershed, be sure to click the “Cicadas” tab at the top of this page!

June 12, 2025

Photo of the Day

Along a quiet forest road, a Great Spangled Fritillary visits Red Clover (Trifolium pratense) to feed on nectar.

June 4, 2025June 4, 2025

Soakin’ Up the Smoky Sun

With temperatures finally climbing to seasonable levels and with stormy sun filtering through the yellow-brown smoke coming our way courtesy of wildfires in Alberta and other parts of central Canada, we ventured out to see what might be basking in our local star’s refracted rays…

Dragonflies including this Black Saddlebags are now actively patrolling the edges of waterways and wetlands for prey and mates.

Here we see a pair of Common Green Darners flying in tandem…

A sunny day almost always brings out the reptiles, including these Painted Turtles…

and the invasive Red-eared Slider, a native transplant from the American midwest.

A really big Snapping Turtle will prey on almost anything, including other Snapping Turtles…

…but this one seems to be fascinated by something a lot smaller. Something like these juvenile Golden Shiners seen here schooling in the sun-drenched shallows.

Turtles aren’t the only reptiles thriving in the heat. Northern Water Snakes (Nerodia sipedon sipedon) take full advantage of a sun-drenched rock to warm up after spending time in the chilly water of a stream.

Butterflies like this Spicebush Swallowtail enjoy time in the sun, even while seeking out minerals in a patch of moist soil.

Woodchucks — After its siblings darted into the familial burrow upon our approach, this juvenile Woodchuck instead sought the attention of its nurturing mother. Unlike its brothers and sisters, perhaps this little groundhog isn’t afraid of its own shadow. Or does the smoky haze have the youngster all confused about what does and doesn’t constitute as a shadow? Well, we can’t help you there, but you have a whole eight months to figure it out!

June 3, 2025June 12, 2025

Brood XIV Periodical Cicada Emergence

Its been four years to the day since we posted our account of the big Brood X Periodical Cicada flight of 2021 in the Lower Susquehanna River Watershed. With the Brood XIV clan of 17-year Periodical Cicadas getting ready fill the June air with their choruses in scattered parts of Pennsylvania’s Berks, Centre, Clinton, Franklin, Huntingdon, Juniata, Lancaster, Mifflin, Perry, Schuylkill, Snyder, Union, and York Counties right now, we thought it an appropriate time to open a new “Cicada” page here on the website. Included is a list of both the annually and periodically emerging species found in the lower Susquehanna valley, as well as a copy of the article and ID photos from the 2021 occurrence of Brood X. In coming months, we’ll be adding photos and maybe some sound clips of the different annual cicadas as well, so remember to check back from time to time for more content.

Periodical Cicada Nymph — Note the wings and red eyes of this periodical cicada nymph. Within weeks it will join billions of others in a brief adult emergence to fly, mate, and then die.

In the short term, we’re going to pay a visit to the Brood XIV territory and will bring you updates as we get them. Until then, be sure to click the new “Cicadas” tab at the top of this page to brush up on your ID of the three species of 17-year Periodical Cicadas.

Adult Periodical Cicada — Not to worry. Cicadas are native, harmless, docile creatures when handled and they pose no threat to the long-term health of your healthy, well-established trees and other vegetation, so enjoy them while they’re here!

June 2, 2025June 3, 2025

Photographs in Living Color: Black and White is Beautiful

Here at susquehannawildlife.net headquarters, we really enjoy looking back in time at old black-and-white pictures. We even have an old black-and-white television that still operates quite well. But on a nice late-spring day, there’s no sense sitting around looking at that stuff when we could be outside tracking down some sightings of a few wonderful animals.

American Toad Tadpoles — American Toad tadpoles have hatched from clusters of eggs deposited in this wet roadside ditch furnished with a clean supply of runoff filtered through a wide shoulder of early successional growth. Recent rains have kept their vernal nursery flooded, giving them the time they need to quickly mature into tiny toads and hop away before scorching summer heat dries up their natal home.

Water Striders mating. — Weekend rains and creek flooding haven’t stopped these Water Striders from pairing up to begin their breeding cycle.

Around streams, lakes, ponds, and wetlands, the Common Whitetail is one of our most conspicuous dragonflies.

Now that’s what we call a big beautiful bill, on a Great Blue Heron stalking fish.

These mating Golden-backed Snipe Flies (Chrysopilus thoracicus) are predatory insects, as are their larvae. They are most frequently found in bottomland woods.

About three feet in length, this Eastern Ratsnake is unusual because it still shows conspicuous remnants of the diamond-patterned markings it sported as a juvenile.

The plumage of the Black-and-white Warbler lacks any of the vibrant colors found in the rainbow, but is nevertheless strikingly beautiful.

May 25, 2025June 27, 2025

Deer Tick or Dog Tick?

Deer Tick vs. Dog Tick — Several weeks back, we removed the tick on the left from the editor’s neck. Upon returning from an outing earlier today, he discovered the one on the right crawling up his sock. It’s almost like they’re out to get him, so after tossing the day’s apparel into the washer, it was time to hit the showers posthaste. Because where there’s one, there are more. And you absolutely never want to spend the night with a tick.

May 25, 2025May 25, 2025

Late May Action in the Forest

Here’s a short preview of some of the finds you can expect during an outing in the Lower Susquehanna River Watershed’s forests this week…

The Mountain Laurel, designated as Pennsylvania’s state flower, is now in bloom.

Mountain Laurel Flower Buds — The buds of the Mountain Laurel remind us of a sugary frosting freshly squeezed from a baker’s pastry bag.

Mountain Laurel Flowers — The flowers of the Mountain Laurel, an evergreen understory shrub, invite pollinators to stop by for a sweet treat.

Little Wood-Satyrs (Megisto cymela) are patrolling forest edges looking for mates and, to host their eggs and larvae, the stands of grasses they find most suitable.

Many of the species of small butterflies we call skippers are now active. The Zabulon Skipper can be found patrolling grassy forest edges, particularly near streams, ponds, and wetlands.

Among the showiest of our butterflies, the Red-spotted Purple (Limenitis arthemis astyanax) is seen here on the leaves of a Black Cherry, its favored host plant.

Butterfly observers will do well to pay a visit to the new Susquehanna Riverlands State Park on Furnace Road north of Hellam in York County, Pennsylvania.

Trail to Schull's Rock Overlook — Upon arrival at Susquehanna Riverlands, drive back the gravel road for about a mile to the parking area at the edge of the agricultural field. Then, hike the trail through the woods and farm hedgerow to the Schull’s Rock overlook on the river. In the forests along this route, the understory is dominated by colonial stands of Common Pawpaw trees.

Along the lookout trail to Schull’s Rock, the Common Pawpaws’ large leaves help them to shade out potentially fast-growing competition. In proper growing situations, pawpaws develop clonal suckers that mature over time to create colonial stands of a single genetic plant.

Common Pawpaw understory. — A Common Pawpaw understory along the approach to Schull’s Rock.

During our recent visit to Schull’s Rock, dozens of Zebra Swallowtails were seen along the trail, many in the vicinity of their sole host plant, the Common Pawpaw. But even more were observed along the edges of the fields and woods where nectar sources like this Multiflora Rose were being visited by numbers of butterflies we normally see only among abundant species like Cabbage Whites. Absolutely amazing!

Scull's Rock Overlook — And the view of the Susquehanna and the Shock’s Mills railroad bridge at the mouth of Codorus Creek is pretty good too!

Common Pawpaw and Hooded Warbler — You can look for colonial stands of Common Pawpaw at other parks and preserves along the lower Susquehanna as well. Birds like this Hooded Warbler can sometimes be found among them in mature riparian forests along the steep slopes of the river gorge.

Common Pawpaw and Baltimore Oriole — A Baltimore Oriole in a Common Pawpaw along a forest edge.

Meanwhile in the treetops, the spring thrust of Neotropical migrants is drawing to a close. The Blackpoll Warbler is typically one of the last to transit the lower Susquehanna valley on its way to northern coniferous forests for summer. They’ve had an unusually protracted movement through the region this spring, the earliest individuals reported during late April. Though very difficult to see in the canopy of the mature trees where it feeds and sings, hearing one is often a benchmark for senior birders each spring. Older observers have often said of the Blackpoll Warbler’s high-pitched song, a rapid series of insect-like staccato “tseet” notes, that it was the first they could no longer detect as their ears started losing sensitivity.

In many tracts along the lower Susquehanna this spring, the American Redstart is turning out to be the most common nesting warbler. Conditions favoring their reproductive success in recent cycles, as well as good survival rates during their migrations and stays on wintering grounds, have filled many lowland forests with redstart songs in 2025. Is this the start of a trend or just an exceptionally good year? Time will tell.

Yet one more reason for a stroll in local forests this week is the chance to see and hear the Rose-breasted Grosbeak. Look for these Neotropical relatives of the cardinal nesting on territories in mature stands of deciduous trees like this Yellow Poplar, a species also known as the Tuliptree.

They spend nearly all their time among the canopy foliage of the largest timber…

May 19, 2025May 26, 2025

Back in the Day: Down on the Farm

Let us travel through time for just a little while to recall those sunny, late-spring days down on the farm—back when the rural landscape was a quiet, semi-secluded realm with little in the way of traffic, housing projects, or industrialized agriculture. Those among us who grew up on one of these family homesteads, or had friends who did, remember the joy of exploring the meadows, thickets, soggy springs, and woodlots they protected.

Low-intensity Farming — During much of the twentieth century, low-intensity agriculture provided a haven for wildlife. Periodic disturbances helped maintain cool-season grassland and early successional habitat for a number of species we currently find in decline.

For many of us, farmland was the first place we encountered and began to understand wildlife. Vast acreage provided an abundance of space to explore. And the discovery of each new creature provided an exciting experience.

Ned Smith's Birds of Field and Garden — Distributed by the Pennsylvania Game Commission, artist Ned Smith’s wildlife posters introduced many residents of the lower Susquehanna region to its birds and mammals. This poster of “Birds of Field and Garden” helped us learn what to expect and search for during our forays to the farm.

Today, high-intensity agriculture, relentless mowing, urban sprawl, and the increasing costs and demand for land have all conspired to seriously deplete habitat quality and quantity for many of the species we used to see on the local farm. Unfortunately for them, farm wildlife has largely been the victim of modern economics.

For old time’s sake, we recently passed a nostalgic afternoon at Middle Creek Wildlife Management Area examining what maintenance of traditional farm habitat has done and can do for breeding birds. Join us for a quick tour to remember how it used to be at the farm next door…

Always found nesting under the forebay of the barn, the Barn Swallow relentlessly pursued flying insects over the pond and meadow.

Eastern Meadowlarks arrived during March and April to begin nesting in their namesake. Their song, “spring-of-the-year”, heralded the new season.

Arriving in meadows and pastures during early May, the Eastern Kingbird provided for its nestlings by ambushing a variety of flying insects. By August, congregations of these birds could be found gathering along ponds and streams ahead of their fall migration.

In the cherry grove down by the creek, the Orchard Oriole would be singing incessantly to defend its territory.

Normally seed eaters through the colder months, American Goldfinches would regularly find a source of protein in the occupants of Eastern Tent Caterpillar nests.

Along the wet margins of the creek, Yellow Warblers would nest in the shrubs and small trees.

The “Traill’s Flycatcher” was a familiar find in low-lying areas of successional shrubs and small trees. Today, “Traill’s Flycatcher” is recognized as two distinct species, the Alder Flycatcher (Empidonax alnorum) and the Willow Flycatcher. In the lower Susquehanna valley, the latter (seen here) is by far the most common of the two.

During the nineteenth century, Eastern Bluebirds became a rarity on lower Susquehanna farms due to a combination of factors: pesticide (DDT) use, habitat loss, and competition with other birds for nest sites. The species saw a resurgence beginning in the 1970s with discontinuation of DDT applications and widespread provision of nesting boxes. Around human habitations, competition with invasive House Sparrows continues to be detrimental to their success.

Purple Martins and Tree Swallows — Purple Martins suffered a similar fate to the bluebirds. The potential for their recovery remains dubious and they continue to be very local breeders, fussy about selection of suitable man-made provisions for nesting. After considerable effort, Purple Martins have at last been attracted to nest in the condos placed for their use at Middle Creek Wildlife Management Area. In the artificial gourds, there are nesting Tree Swallows, a species which also benefits from the placement of boxes intended for bluebirds.

Abandoned fields and other successional habitats were and continue to be favored homes for Field Sparrows.

At almost any time of year, roving bands of Cedar Waxwings would suddenly visit old field habitat looking for berries among the shrubs and other pioneering woody growth. In early summer, after most species have already hatched their young, nesting would commence and these fruit eaters would transform into accomplished fly catchers.

During the twentieth century prior to the 1980s, Ring-necked Pheasant populations in the Lower Susquehanna River Watershed were comprised of breeding descendants of introduced birds supplemented by additional releases to maintain numbers sufficient for hunting. Year-round populations did and can reside in mosaic landscapes of early successional and grassland habitats, the latter including hay fields left unmowed through the nesting season.

Red-winged Blackbirds have always been a fixture of hay fields and meadows on farms. While the increase in mowing frequency has reduced their nesting success, they have persevered as a species by nesting earlier than other birds and by utilizing other landscape features such as densely vegetated stormwater basins for breeding sites.

Do you recall the last time you saw a Bobolink nesting in a hay field near you? Arriving in early May as a Neotropical migrant, the Bobolink requires a cool-season grassland such as hay field through at least July to complete its nesting cycle. Even earlier this century, we remember nesting Bobolinks being more widespread on farms throughout the region. Now, you almost have to go to Middle Creek if you want to see them.

Formerly more widespread in hay fields throughout the lower Susquehanna valley, the native Grasshopper Sparrow is yet another species falling victim to early mowing and intensive farming.

Cool-season Grasses — **Beautiful cool-season grasses waving in the spring breeze.** Meadows and hay fields can be managed to function as cool-season grasslands to provide nesting opportunities for many of the species we used to find down on the farm.

May 15, 2025May 15, 2025

Photo of the Day

White-tailed Deer Button Buck Hosting Engorged Ticks — When it comes to sitting down and watching an episode of Hollywood Squares or indulging in a game of Tic-Tac-Toe, those of us here at susquehannawildlife.net are quick to play right along. But when it comes to mixing it up with a Tick-Backed Doe or a Tick-Backed Buck like this one, you can count us out. Upon finding this button buck infested with engorged Deer Ticks and heading our way on a grassy trail, we decided to turn around and limit our walk to the gravel roadway where there was less of a chance of picking up any hitchhikers from the vegetation where this guy has been spending his time.

May 6, 2025May 6, 2025

Singing in the Rain

Neotropical birds are fairly well acquainted with repetitive periods of thundershowers. With that in mind, we decided not to waste this stormy Tuesday by remaining indoors.

Utility Right-of-way with Successional Habitat — We hiked this utility right-of-way to the north of a heavy thunderstorm and found plenty of activity in the shrubby successional habitat there.

Rain or shine, male Indigo Buntings were busy singing. All this exuberance is intended not only to establish and defend a nesting territory…

Prior to the implementation of the intensive manicuring practices we see currently applied to most utility right-of-ways, shrubby thickets filled miles of these linear corridors to create a webbed network of early successional growth throughout the lower Susquehanna valley. Loss of this specialized habitat has led to the almost total elimination of the formerly common Blue-winged Warbler as a breeding species here.

Another Neotropical species that nests in successional thickets is the Prairie Warbler (Setophaga discolor), seen here during one of this morning’s downpours.

The Hooded Warbler is fond of wooded thickets along the edge of forested land such as those in the Third/Stony Mountain utility right-of-way.

We hope you enjoyed our walk in the rain as much as we did. If you venture out on a similar excursion, please remember this. The majority of the wild animals around us have busy lives, particularly at this time of year. Most don’t take a day off just because it rains—that includes ticks.

Female Deer Tick — Be certain to check yourself for ticks, especially these very small Deer Ticks (Ixodes scapularis), also known as Black-legged Ticks. Deer Ticks are vigilantly looking for something to latch onto, even in the rain, and they can be vectors of Lyme disease. We found this adult female as a stowaway on the editor’s neck just before heading home from today’s stroll.

May 4, 2025May 5, 2025

Six Flowering Woody Plants

Here’s a look at six native shrubs and trees you can find blooming along forest edges in the lower Susquehanna valley right now.

The Black Locust (Robinia pseudoacacia), a member of the pea or legume family (Fabaceae), can be a dominant pioneer plant of sunny successional habitats, particularly on poor soils. Nodules along its roots fix nitrogen to help facilitate the growth of the intermediate and climax species of trees and shrubs that replace the pioneers. Initially armed with protective spines to reduce browsing, the Black Locust’s branches become mostly thornless as the tree matures.

The Pinxter Flower, also known as the Pink Azalea, is certainly a contender for our most spectacular native blossoming shrub;…

Spicebush Swallowtail on Pinxter Flower — …this Spicebush Swallowtail seems to like it too.

When in bloom, the Black Cherry is a common sight in regional woodlands. Often seen already covered with blossoms when young and shrub-like, many of these trees will continue flowering and producing fruit throughout the decades required to reach a mature height of 60 feet or more.

The Blackhaw is an understory shrub preferential to sun-dappled areas beneath a break in the forest canopy. Pollinated flowers later produce clusters of blue-black berries for birds and other wildlife.

Local old timers might remember hearing folklore that equates the northward advance of the blooming of the Flowering Dogwoods with the progress of the American Shad’s spring spawning run up the river. While this is hardly a scientific proclamation, it is likely predicated on what had been some rather consistent observation prior to the construction of the lower Susquehanna’s hydroelectric dams. In fact, we’ve found it to be a useful way to remind us that it’s time for a trip to the river shoreline below Conowingo Dam to witness signs of the spring fish migration each year. We’re headed that way now and will summarize our sightings for you in days to come.

April 29, 2025May 5, 2025

Busy Bees

During a warm spell back in late March, adult Mason Bees began emerging from nest sites to begin mating. During recent days, we photographed the fertile females as they continued the process that will produce the next generation of these fruit-friendly pollinators.

Mason Bee Nest Box — Mason Bees visiting a box stocked with blocks of wood wherein holes have been drilled to provide nesting cavities for the egg-laying females.

Osmia Mason Bee — Mason Bee populations using man-made nesting structures in our region include two non-native species. Both show horn-like structures on the face, denoted here by the white hairline. To pollinate orchard crops, Osmia cornifrons, the Horn-faced Bee, was intentionally introduced to the eastern United States from northern Asia in 1978. Another early season pollinator, Osmia taurus, was accidentally introduced to North America from eastern Asia. O. taurus was first reported in Maryland and West Virginia in 2002 and had reached Pennsylvania by 2006-2008.

Female Mason Bees at Nest — Inside a nest cavity, a female Mason Bee will deposit pollen balls and an egg within each cell she creates using mud partitions. Cell by cell she fills the cavity with eggs and pollen. When the cavity is full, a thicker mud plug is constructed to seal off the entrance. After hatching, each larval bee will remain inside a cocoon within its individual cell consuming the enclosed pollen ball and maturing. After passing the winter in the nest, the new generation of adults will emerge in the spring…outermost cell occupants first.

Mason Bee Excavating Nest — When reusing a nest cavity from a previous year, a female Mason Bee will excavate old partitions and other debris before starting her own nest. This female is removing saw dust and is possibly enlarging the hole on this new block of wood that we drilled just a few weeks ago.

Mason Bees at Small Holes — The majority of our Mason Bees prefer 1/4-inch holes, but some use the smaller 3/16-inch ones. Holes up to 3/8 of an inch in diameter are readily accepted by both Osmia cornifrons and taurus.

Packing Pollen and Eggs — A look at several female Mason Bees backed into their nest cavities to deposit eggs and pollen. Note the load of pollen adhering to the abdomen of the head-first bee at the bottom of the image and the yellow residue around the entrance to the cavity above it.

Drosophila Fly at Mason Bee Nest Entrance — Before a female Mason Bee has a chance to seal up a cell or to plug up the entrance to the cavity, a Drosophila fly (left) or other nest raider may show up to plunder a portion of the stash of pollen.

Mason Bee Sealing a Nest Cavity — Once filled with cells, each containing an egg and a pollen cache, the mother bee seals the nest cavity with a wall of mud.

Mason Bee Sealing Cavity — A bee seals up a 3/16-inch cavity with a mud barricade. Adult female Mason Bees may select smaller than usual holes to deposit eggs that will produce their smaller male young.

Mason Bee Closing Nest — Upon finishing the masonry job here, this female will move on to yet another cavity to deposit the balance of her fertilized eggs.

High-occupancy Mason Bee Nest — One week later, the Mason Bees’ work is done.

March 15, 2025

Photo of the Day

Late winter is hardly the time of year one would expect to find a grasshopper bouncing around a rocky woodland clearing. But earlier this week during our visit to Rothrock State Forest in Huntingdon County, we photographed this Northern Green-striped Grasshopper (Chortophaga viridifasciata viridifasciata) nymph among the talus atop Tussey Mountain. Unlike the majority of other grasshopper species in the Susquehanna basin which overwinter as eggs, the Northern Green-striped hatches in late summer and progresses through all but the final one or two of its five developmental instars before passing the colder months in a sheltered location among fallen leaves, dried grasses, and decaying plant matter. Often the earliest of our grasshoppers to emerge, the Northern Green-striped nymph soon completes its final molts and reaches adulthood just in time for the warm days of late spring and early summer. Two color forms are common, brown and green, both possessing reddish-brown abdominal segments. Fuchsia-colored erythristic individuals are rare.

November 7, 2024November 7, 2024

Photo of the Day

During the summer, the Lesser Angle-winged Katydid has little difficulty hiding among the lush green foliage of hardwood trees. Oft times, the only sign of its presence is its call, a series of short rapid rattles, “ttt—-tttt—-ttt”, issued with a second or two of silence between each rattle. The call is often part of the nocturnal chorus it shares with other species of katydids on warm evenings. Frost and freeze usually bring these annual breeding rituals to end. But during our current resurgence of warm autumn weather, we found this hardy survivor in a forest clearing where it has little hope of escaping notice among bare trees and earth-tone fallen leaves. As the afternoon temperature soared into the upper seventies, it further revealed its presence by striking up its tell-tale mating call, “ttt—-tttt—-ttt.”

October 30, 2024October 31, 2024

A Towering American Elm

During the past week, Uncle Tyler Dyer has been out searching for autumn leaves to add to his collection. One of the species he had not encountered in previous outings was the American Elm (Ulmus americana), so he made a special trip to see a rare mammoth specimen in a small neighborhood park (Park Place) along Chestnut Street between 5th and Quince Streets in Lebanon, Pennsylvania.

American Elm at Park Place in Lebanon, Pennsylvania — Possibly the largest and oldest remaining American Elm in the lower Susquehanna watershed, the Park Place tree in Lebanon exceeds 60 feet in height and may be more than 300 years of age. In the nearly one hundred years since Dutch elm disease (DED) first began killing elms in eastern North America, this tree has been spared the fatal effects of the infection. DED is caused by several species of microfungi (Ophiostoma ulmi, O. himal-ulmi, and O. novo-ulmi) spread by numerous bark beetle (Curculionidae) species. As early as 1928, infected beetles arrived in the United States from the Netherlands among shipments of logs.

There’s still time to get out and see autumn foliage. With warmer weather upon us—at least temporarily—it’s a good time to go for a stroll. Who knows, you might find some spectacular leaves like these collected by Uncle Ty earlier this week. All were found adorning native plants!

Winged Sumac (Rhus copallina), a small native tree. In case you’re wondering, it’s not poisonous.

Black Chokeberry (Photinia melanocarpa), a native shrub also known as Aronia. It produces black-colored fruits in summer.

Red-osier Dogwood (Cornus sericea), a shrub that prefers wet or damp soils. It yields white fruits.

Maple-leafed Viburnum (Viburnum acerifolium), a native shrub of upland forest understories.

Highbush Blueberry (Vaccinium corymbosum), a native shrub and the source of many cultivated forms of the popular fruit.

To identify your finds, be certain to click the “Trees, Shrubs, and Woody Vines” tab at the top of this page to check out Uncle Ty’s extensive collection.

October 12, 2024

Things to See While You’re Leaf Peeping

During your foray to view the colorful foliage of the autumn landscape, a little effort will reveal much more than meets the eye of the casual observer.

Fallen Foliage in the Forest — Don’t just stand there like a bump on a log, go for a stroll, watch your step, and have a look around. There’s a lot to see out there!

The dense understory of this forest on Blue Mountain consists almost exclusively of Witch-hazel (Hamamelis virginiana).

Witch-hazel in Flower — Witch-hazel is unique among our native flowering shrubs; it blooms in autumn.

Until we experience a heavy frost, some of our most beautiful butterflies, including this Common Buckeye, remain active along forest edges.

The arrival of large numbers of Yellow-rumped Warblers heralds the transition to movements of migrants that will spend the colder months in temperate climates like ours instead of continuing on to the tropics. In winter, small numbers of “yellow-rumps” sometimes linger in the Lower Susquehanna River Watershed, as long as a supply of wild berries is readily available.

The Palm Warbler, a persistent “tail wagger”, is another late-season migrant that occasionally spends the winter. The migration of these drab “Western Palm Warblers” will be followed in coming weeks by the more yellowish “Eastern Palm Warblers” of the eastern third of the species’ breeding range in Canada.

White-throated Sparrows are southbound and easily observed right now. Many will remain along our brushy forest edges for winter.

Another native sparrow, the familiar Dark-eyed Junco, is now arriving.

Yellow-bellied Sapsuckers — The Yellow-bellied Sapsucker is a secretive migratory woodpecker that is currently moving through the region on its way to southern pine forests for the winter. Juveniles like this one in the lower right remain close to their parents during migration and continue to beg for food.

The Pileated Woodpecker is a resident species in our mature forests and woodlands. We were showered by “sawdust” as we watched this one dismantling a rotted dead tree while searching for beetle grubs and other invertebrates.

You may see Eastern Bluebirds year-round in the lower Susquehanna valley, but many are migratory. A southbound push is currently transiting our area.

Hawk Watching on Blue Mountain at Boyd Big Tree Preserve. — Mix your leaf watching with hawk watching by visiting a ridgetop vista like this one at Boyd Big Tree Preserve Conservation Area on Blue Mountain north of Harrisburg and you can double your pleasure.

In addition to spectacular foliage, you stand a good chance of seeing an inquisitive Common Raven…

Screaming Red-tailed Hawk — …or a migrating Red-tailed Hawk that gives you a good scream as it passes by…

You may even double your fun by seeing two Common Ravens…

…or a couple of migrating Red-tailed Hawks…

You too can experience the joys of walking and chewing gum at the same time, so grab your field glasses, your camera, and your jacket, then spend lots of time outdoors this fall. You can see all of this and much more.

Don’t forget to click the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab at the top of this page to help you find a place to see both fall foliage and migrating birds of prey in coming weeks. And click the “Trees, Shrubs, and Woody Vines” tab to find a photo guide that can help you identify the autumn leaves you encounter during your outings.

September 21, 2024September 21, 2024

This Week’s Enormous Diurnal Flights

For the past several weeks, we’ve given you a look at the fallout of Neotropical songbirds on mornings following significant nocturnal flights of southbound migrants. Now let’s examine the diurnal (daytime) flights that have developed in recent days over parts of the lower Susquehanna basin and adjacent regions.

For those observing diurnal migrants, particularly at raptor-counting stations, the third week of September is prime time for large flights of tropics-bound Broad-winged Hawks. Winds from easterly directions during this year’s movements kept the greatest concentrations of these birds in the Ridge and Valley Province and upper areas of the Piedmont as they transited our region along a southwesterly heading. Counts topped 1,000 birds or more on at least one day at each of these lookouts during the past seven days. Meanwhile at hawk watches along the Piedmont/Atlantic Coastal Plain border, where flights topped 10,000 or more birds on the best days last fall, observers struggled to see 100 Broad-winged Hawks in a single day.

Among the challenges counters faced while enumerating migratory Broad-winged Hawks this week was the often clear blue skies, a glaring sun, and the high altitude at which these raptors fly during the mid-day hours.

"Kettling" Broad-winged Hawks — Large numbers of Broad-winged Hawks climbing on a thermal updraft to gain altitude can become increasingly difficult to spot in a clear blue sky.

Another factor complicating the hawk counters’ tasks this week, particularly west of the Susquehanna River, was the widespread presence of another group of diurnal fliers.

Was it the migratory Blue Jays that are beginning to move through the area? No.

Was it the increasing numbers of Sharp-shinned Hawks and other raptors? No.

Hawkwatchers Scanning the Skies — So what was it that complicated the hawkwatchers’ efforts to find high-flying Broad-winged Hawks during this week’s flights?

Is it a bird? Is it a plane? No. And it’s nobody’s superhero either. It’s the non-native Spotted Lanternfly and it is now very numerous west of the Susquehanna and common enough to the east to make even experienced observers take a second look while scanning the skies for Broad-winged Hawks and other migrating raptors. Just how many do raptor counters have to contend with? Even where these invasive insects have become less populous in the lower Susquehanna valley, hundreds still fill the skies above forest clearings around hawk-counting stations including Waggoner’s Gap, Second Mountain, and Rocky Ridge County Park.

Spotted Lanternflies on Radar — Spotted Lanternfly populations explode with great vigor as they colonize areas with plenty of tender new growth on native, non-native, and cultivated shrubs, trees, and vines. Earlier this week, National Weather Service Doppler Radar in Pittsburgh, Pennsylvania, captured enormous mid-afternoon flights of Spotted Lanternflies in recently invaded areas of southwestern Pennsylvania and eastern Ohio. (NOAA/National Weather Service Doppler Radar image)

As the Broad-winged Hawk migration draws to a close during the coming ten days, the occurrence of the one dozen other migratory diurnal raptors will be on the upswing. They generally fly at lower altitudes, often relying upon wind updrafts on the ridges for lift instead of high-rising thermal updrafts. They therefore present better observation opportunities for visitors at hawk watch lookouts and are less frequently confused with high-flying Spotted Lanternflies.

A Merlin seen earlier this week migrating through the Lower Susquehanna River Watershed. The numbers of migrating Merlins and other falcons will peak sometime in coming weeks.

August 31, 2024August 31, 2024

A Prescribed Fire Site After the Burn

Back on March 24th, we took a detailed look at the process involved in administering prescribed fire as a tool for managing grassland and early successional habitat. Today we’re going turn back the hands of time to give you a glimpse of how the treated site fared during the five months since the controlled burn. Let’s go back to Middle Creek Wildlife Management Area for a photo tour to see how things have come along…

Prescribed Fire application in March — Pennsylvania Game Commission crews administering prescribed fire on a grassland demonstration site back on March 16.

Prescribed Fire Site in May — By late May, native herbaceous perennial wildflowers including Joe-pye Weed had re-greened the site. One of the goals of the burn was to kill fire-sensitive woody plants, thus preventing the process of succession from reforesting the site.

Yellow-breasted Chat in May — Because prescribed fire is administered in a mosaic pattern that permits some early successional growth to remain until the next burn, birds including this thicket-nesting Yellow-breasted Chat are able to take advantage of the mixed habitat during their breeding season in May and June.

Prescribed Fire Site in August — By August, the site is a haven for native plants and animals.

Native Warm-season Grasses and Wildlflowers in August — The burn has promoted the growth and late-summer bloom of fire-tolerant native wildflowers and warm-season grasses…

Indiangrass in Flower — …including Indiangrass,…

Thin-leaved Coneflower, a plant also known as Brown-eyed Susan. — …Thin-leaved Coneflower (Rudbeckia triloba), a plant also known as Brown-eyed Susan,…

…and Joe-pye Weed, a plant butterflies find irresistible.

Eastern Tiger Swallowtail collecting nectar from Joe-pye Weed.

Black-morph Eastern Tiger Swallowtail — A black-morph Eastern Tiger Swallowtail collecting nectar from Spotted Knapweed (Centaurea steobe micranthos), a non-native invasive plant found growing in an area of the burn site missed by this year’s fire. While many non-native plants are unable to survive the flames and heat produced by prescribed fire, it isn’t an absolute cure-all. It doesn’t eliminate all invasive plants, it just keeps them from dominating a landscape by out-competing native species. Left unmanaged, Spotted Knapweed is a tough perennial invasive that can easily become one the species able to overtake a vulnerable grassland. It can be a stubborn survivor of some prescribed burns. On the plus side, butterflies really like it.

By August, native grassland plants in the prescribed fire area were already providing an abundance of seeds for birds including this American Goldfinch.

For larger birds like turkeys and pheasants, an abundance of Carolina Grasshoppers are providing a protein-dense food source in managed grasslands.

Elsewhere around the refuge at Middle Creek, prescribed fire and other management techniques are providing high-quality grassland habitat for numerous species of nesting birds…

Bobolink in Hay Field — …and in hay fields where mowing was delayed until the nesting season, including the fledging process, was completed earlier this month.

We hope you enjoyed this short photo tour of grassland management practices. Now, we’d like to leave you with one last set of pictures—a set you may find as interesting as we found them. Each is of a different Eastern Cottontail, a species we found to be particularly common on prescribed fire sites when we took these images in late May. The first two are of the individuals we happened to be able to photograph in areas subjected to fire two months earlier in March. The latter two are of cottontails we happened to photograph elsewhere on the refuge in areas not in proximity to ground treated with a prescribed burn or exposed to accidental fire in recent years.

Eastern Cottontail at a site subjected to prescribed fire earlier in the spring.

These first two rabbits are living the good life in a warm-season grass wonderland.

Eastern Cottontail with ticks. — Eastern Cottontail at a site not subjected to any recent fire activity.

Oh Deer! Oh Deer! These last two rabbits have no clock to track the time; they have only ticks. Better not go for a stroll with them Alice—that’s no wonderland! I know, I know, it’s time to go. See ya later.

August 30, 2024August 31, 2024

Aphrodite Fritillary

The Aphrodite Fritillary (Speyeria aphrodite), also known simply as the Aphrodite, is a brush-footed butterfly of deciduous, coniferous, and mixed forests. We found this female in a grassland margin between woodlots where prescribed fire was administered during the autumn of 2022 to reduce accumulations of natural fuels and an overabundance of invasive vegetation. A goal of the burn was to promote the growth of native species including the violets (Viola species) favored as larval host plants by this and other fritillaries.

By the time these adult butterflies make their reproductive flights in late summer, the violets that serve as larval host plants have gone dormant. To find patches of ground where the violets will come to life in spring, the female Aphrodite Fritillary has an ability to sense the presence of dormant roots, probably by smell. Upon finding an area where suitable violets will begin greening up next year, she’ll deposit her eggs. The eggs overwinter, then hatch to feed on the tender new violet leaves of spring.

July 27, 2024July 27, 2024

Wildflower Meadow Update

Do you recall our “Photo of the Day” from seven months ago…

Riverfront Park Wildflower Meadow — Here’s something to look forward to in the new year. The good citizens of East Donegal Township in Lancaster County have partnered with Alliance for the Chesapeake Bay to establish an extensive wildflower meadow on what had been a mowed field of turf grass at Riverside Park in the Susquehanna floodplain near Marietta. As the photo shows, the lawn plants have been eliminated in preparation for seeding with a diverse assortment of native grasses and wildflowers to provide habitat for birds and pollinators including butterflies, bees, and other insects. Once established, the meadow’s extensive vegetative growth will help reduce stormwater runoff by better infiltrating rainfall to recharge the aquifer. During flood events, the plantings will provide soil stabilization and increase the ability of the acreage to uptake nutrients, thus reducing the negative impact of major storms on the quality of water in the river and in Chesapeake Bay. Check the project’s progress by stopping by from time to time in 2024!

Well, here’s what that site looks like today…

And there are pollinating insects galore, most notably butterflies…

A Cabbage White collecting nectar on Blue Vervain.

A Sulphur — A Clouded or Orange Sulphur among the grasses in the meadow.

The Least Skipper is our tiniest butterfly.

The Little Glassywing (Vernia verna), this one feeding on vervain nectar, deposits its eggs on Purpletop grass, which then functions as the host plant for this butterfly’s larvae.

A Summer Azure (Celastrina neglecta) feeding on the nectar of a Black-eyed Susan.

An Eastern Tiger Swallowtail on Blue Vervain. Nearby Yellow (Tulip) Poplars and other trees serve as host plants for this butterfly’s larvae.

The Sachem, this one visiting a Black-eyed Susan, is a variable species with a range that normally lies south of the 40th parallel, the line of latitude that intersects the Susquehanna in the area of the Conejohela Flats at Washington Boro.

A Sachem visiting the blooms of Oxeye sunflower. During recent weeks, scorching winds from the south and southwest have transported an abundance of these vagrant skippers into the lower Susquehanna valley and beyond.

Here and there among the meadow’s plantings we noticed one of our favorites starting to flower, the Partridge Pea.

Partridge Pea happens to be a host plant for another vagrant from the south, the big, lime-yellow Cloudless Sulphur. We saw at least half a dozen patrolling the meadow.

Why on earth would anyone waste their time, energy, and money mowing grass when they could have this? Won’t you please consider committing graminicide this fall? That’s right, kill that lawn—at least the majority of it. Then visit the Ernst Seed website, buy some “Native Northeast Wildlflower Mix” and/or other blends, and get your meadow planted in time for the 2025 growing season. Just think of all the new kinds of native plants and animals you’ll be seeing. It could change your life as well as theirs.

A Snowberry Clearwing (Hemaris diffinis), more commonly known as a hummingbird moth, visits the flowers of Blue Vervain in the Riverfront Park wildflower meadow.

July 9, 2024July 9, 2024

Photo of the Day

Similar in appearance to the invasive Brown Marmorated Stink Bug, the Western Conifer Seed Bug (Leptoglossus occidentalis) is a seldom-noticed denizen of pines. During the 1950s, their range started expanding from the American west into the eastern states. Like the stink bug, the Western Conifer Seed Bug emits a buzzing sound in flight and is capable of releasing a nasty-smelling compound from scent glands when harassed. They feed on sap from developing pine cones, often causing deformities to the seeds, but posing no real harm to the trees. The Western Conifer Seed Bug causes greatest consternation during the fall when, like the invasive stink bugs, it gathers in numbers and attempts to enter homes to spend the winter. Those that get inside are mostly just an annoyance, but there have been reports of plumbing leaks caused by individual insects piercing PEX plastic water tubing with their mouths.

July 4, 2024July 5, 2024

Colonies at War

Where does all the time go? Already in 2024, half the calendar is in the trash and the gasoline and gunpowder gang’s biggest holiday of the year is upon us. Instead of bringing you the memory-making odors of quick-burning sulfur or the noise and multi-faceted irritations that revved-up combustion engines bring, we thought it best to provide our readers with a taste of history for this Fourth of July. Join us, won’t you, for a look back at one of the many events that shaped the landscape of our present-day world.

The early morning’s sun had just begun bathing the verdant gardens of the olde towne centre with a warm glowing light. Birds were singing and the local folk were beginning to stir in preparation for their day’s chores. Then, suddenly, something was stirring afoot.

The great battle had commenced. Within minutes, thousands of colonists spilled onto the pavement to join the melee and defend their homes.

Pavement Ants Streaming from Subterranean Nest — There’s no towne crier spreading the word on horseback. Sensing aggression from a neighboring colony, a worker Pavement Ant (Tetramorium immigrans) functioning as a sentry issues the alert. The aroma of pheromones produced by the sentry warns of danger and calls other workers to drop what their doing and instead respond to defend the nest.

Pavement Ants Join the Fray — Engaged in a dispute over territory, two colonies of Pavement Ants clash. Though native to Europe, there is no evidence of Napoleonic tactics in their warfare. All maneuver seems to be by chance.

Two Colonies of Pavement Ants in Battle — The workers doing the fighting are the sterile daughters of the one queen in each colony. In addition to defending the nest, they do all the foraging and care for the queen’s eggs and young. The young in each nest are its workers’ sisters and will include one or more new fertile queens. These new queens, along with fertile males (brothers of the workers), develop wings and fly away to mate. After mating, a young queen begins a new colony, excavating her own nest wherein she raises a first brood of workers to tend her forthcoming generation of eggs and young. As the new colony grows, the queen’s workers expand the size of the underground nest by carrying particles of soil to the surface, depositing them around the entrance as telltale mounds.

Male Pavement Ant Amid Fighting Workers. — A winged male Pavement Ant gets caught in the fury of combat. His primary role in life is to make a nuptial flight and mate with a queen to start a new colony.

The fighting was at close quarters—face to face with dominant soldiers sparing no effort to prevail in the struggle.

Pavement Ants in Combat — Worker Pavement Ants, all females, assume the role of soldiers to defend their nest, their colony, and their queen.

After about an hour had passed, the tide had turned and the fighting mass drifted to the south of the battle’s starting point. The aggressors had been repelled. The dispute was resolved—at least for a little while.

Pavement Ants in Battle — Thousands of Pavement Ants at the high-water mark of their desperate struggle. It was a fierce, jaw-to-jaw contest to tear one’s opponent to pieces.

Pavement Ant Casualty — A winged Pavement Ant, probably a male and not a queen, falls victim to the fighting (upper right). This casualty will not take part in a nuptial flight and will not contribute its colony’s DNA to a new population of ants.

Aggressive Pavement Ants Repelled — The tide turns and the invaders from the south are pushed back in the direction from whence they came. Within minutes, the soldiers transitioned back to being workers. No visual signs of the fight remained; casualties were carried away.

It wasn’t a struggle for independence. And it wasn’t a fight for liberty. For the sterile Pavement Ant worker, all the exertion and all the hazard of assuming the role of a soldier had but one purpose—to raise her sisters and become an aunt. Long live the queen.

June 27, 2024

Photo of the Day

Clubbed Mydas Flies in Copula — **Love is in the Air-** The conspicuously large Clubbed Mydas Fly (Mydas clavatus) is a harmless mimic of the spider wasps (Pompilidae). They carry their masquerade to the extreme with bold behavior that includes pumping of their abdomen to simulate an ability to sting. Adults visit flowers to feed on pollen and nectar while the larvae are predatory, relying on a diet of scarab beetles (Scarabaeidae) which they find crawling within the dead and rotting wood where they reside. When mating, adults fly around in copula, reminding one of “love bugs” and other members of the fly family Bibionidae, the march flies. This mating pair was photographed along the Susquehanna at Conewago Falls.

June 18, 2024June 18, 2024

Colorful Damselflies

Check out these glistening gems—mating damselflies on a late spring afternoon.

It’s two pairs of Powdered Dancers, males clasping ovipositing females, a striped blue form female on the left and a brown form female on the right.

A male Stream Bluet (Enallagma exsulans) perched on a grass stem in a vegetated buffer along a rehabilitated creek.

A pair of Stream Bluets, male clasping female.

A male Ebony Jewelwing (Calopteryx maculata) looking for a mate.

A female Ebony Jewelwing — There she is, the shy female Ebony Jewelwing among the shelter of some streamside foliage.

A male Variable Dancer. — A male Variable Dancer (Argia fumipennis).

A mating pair of Variable Dancers, male clasping female.

Powdered Dancers and Variable Dancers — Two pairs of mating damselflies, Powdered Dancers (left) and Variable Dancers (right), with both females ovipositing.

A male Double-striped Bluet (Enallagma basidens) showing his stuff.

Powdered Dancers and Fragile Forktail — Mating Powdered Dancers, male clasping female, and an ovipositing female Fragile Forktail.

Mating Orange Bluets in wheel position. — A pair of mating Orange Bluets (Enallagma signatum) in wheel position, male above and female below.

Aren’t they precious? You bet they are.

To see these and other damselflies, as well as their larger cousins the dragonflies, be certain to visit your favorite vegetated lake, pond, stream, or wetland on a sunny afternoon. You might be surprised by the variety of colorful species you can find.

And to help identify your sightings, don’t forget to visit our “Damselflies and Dragonflies” page by clicking the tab bearing that name at the top of this page.

June 15, 2024June 15, 2024

Photo of the Day

The minuscule Long-legged Fly (Condylostylus species) is a predatory consumer of soft-bodied invertebrates. In the headquarters garden, we found this individual and others scurrying around on the leaves of Common Milkweed where they may be seeking to gobble up infesting aphids.

June 14, 2024

Photo of the Day

A nectar-seeking Honey Bee visits the fragrant flowers of a Common Milkweed.