Here are some sights and sounds from the ongoing emergence of Brood XIV Periodical Cicadas in the Lower Susquehanna River Watershed.
We begin in the easternmost spur of the lower basin where a sizeable emergence of cicadas can be seen and heard in the woodlands surrounding the headwaters of the Conestoga River in Berks County north of Morgantown. This flight extends east into Chester County and the French Creek drainage of the Schuylkill River watershed on State Game Lands 43 north of Elverson and consists of Pharaoh Periodical Cicadas (Magicicada septendecim), the most common species among 17-year broods.
A Brood XIV Pharaoh Periodical Cicada at State Game Lands 43 identified by the red bar extending from the eye to the wing root.The underside of the abdomen on a male Pharaoh Periodical Cicada showing the wide orange bars on each segment.Exuvia of a recently emerged Pharaoh Periodical Cicada.Soon after landing on a perch, a male Pharaoh Periodical Cicada will usually announce his presence by singing. It’s an attempt to quickly attract potential mates that may be in the vicinity.Pharaoh Periodical Cicadas ascending the branches of an oak.Gatherings of thousands of singing male Pharaoh Periodical Cicadas create a distinctive droning chorus.A receptive female will make a click sound with her wings to summon a suitable singing male for mating.While usually occurring in the safety of the trees, the breeding frenzy can spill over onto the ground where we happened to find this copulating pair.After mating, female cicadas make slits in the end twigs of selected trees into which they lay their eggs. The process of egg-laying and larval emergence will usually wilt and kill end growth on the affected branches, causing little harm to healthy trees. It’s similar to the trim you might give to a bonsai plant to keep it stout and sturdy.How long do Periodical Cicadas live? Well, by last week, we were already finding dead specimens by the thousands. Most of them had already completed their breeding cycle and planted the seeds for a new generation of Brood XIV cicadas.A deceased Pharaoh Periodical Cicada at the Fire Tower Parking Area at French Creek State Park. This specimen and a chorus on the hill’s forested south slope were the northeastern-most evidence of Brood XIV Periodical Cicadas we could find for the population cluster in portions of Berks, Chester, and Lancaster Counties around Morgantown.The abundance of Pharaoh Periodical Cicadas on State Game Lands 43 in Chester County has attracted numerous raptors, particularly wandering one-year-old birds that aren’t quite mature enough to nest. Among the sightings have been Red-tailed Hawks, Red-shouldered Hawks, Broad-winged Hawks, and at least three Mississippi Kites, a rarity on the Piedmont this far inland from the coastal plain. (See the post from June 5, 2021, for details on the occurrence of Mississippi Kites in northernmost Delaware during the Brood X emergence.)
From Route 82 north of Elverson to the west through the forested areas along Route 10 north of Morgantown and the Pennsylvania Turnpike, we found an abundance of Cassin’s Periodical Cicadas (Magicicada cassini) calling among the Pharaohs. This mix of Pharaoh and Cassin’s Periodical Cicadas extends west along the north side of the turnpike into Lancaster County and State Game Lands 52 on Black Creek north of Churchtown.
A Brood XIV Cassin’s Periodical Cicada at State Game Lands 52 is identified by the all-black margin between the eye and the wing root and……the black underside of the abdomen with no orange stripes.To penetrate the sounds of the more common Pharaoh Periodical Cicadas, male Cassin’s Periodical Cicadas gather in large concentrations to generate a loud, oscillating chorus. Its surging volume will usually exceed that of the Pharaohs singing in the vicinity.Mated Cassin’s Periodical Cicadas copulating at State Game Lands 52.The underside of copulating Cassin’s Periodical Cicadas.A pair of Cassin’s Periodical Cicadas at state Game Lands 52 in Lancaster County.
Further west in Cornwall, Lebanon County, a Brood XIV emergence can be found on similar forested terrain: the Triassic hills of the Newark Basin—rich in iron ore and renowned for furnace operations during the eighteenth and nineteenth centuries. Pharaoh Periodical Cicadas were the only species heard among this population that extends from Route 72 east through the woodlands along Route 322 into the northern edge of State Game Lands 156 in Lancaster County.
On the west side of the Susquehanna, yet another isolated population of Brood XIV Periodical Cicadas can be found in Perry County, just south of Duncannon on State Game Lands 170 on the slopes of “Cove Mountain”, the canoe-shaped convergence of the western termini of Peters and Second Mountains.
Pharaoh Periodical Cicadas dominated this Perry County chorus,…
…but we did detect at least one Cassin’s Cicada trying to find a mate.
A solitary Cassin’s Periodical Cicada issues a lonely song of short buzzes and ticking notes on State Game Lands 170. Fragmented populations, especially those that are only able to fly and increase their distribution every 17 years, often have a challenging time expanding and reuniting their disjointed ranges.
Not to say they aren’t present, but we have yet to detect the rarest species, Magicicada septendecula, the “Little Seventeen-year Cicada”, among the various populations of Brood XIV Periodical Cicadas emerging in the lower Susquehanna valley. For the coming two weeks or so until this brood is gone for another 17 years, the search continues.
For more on both annual and periodical cicada species in the Lower Susquehanna River Watershed, be sure to click the “Cicadas” tab at the top of this page!
After repeatedly hearing the songs of these Neotropical migrants from among the foliage, we were finally able to get a look at them—but it required persistent effort.
Often found closer to the ground among shrubs and small trees, this Magnolia Warbler was a challenge to find in the upper reaches of a Chestnut Oak.Not far away, we spied this Yellow-rumped Warbler.Despite its status as probably the most common Neotropical songbird to nest in the deciduous woodlands of the lower Susquehanna valley, the Red-eyed Vireo is nevertheless notoriously difficult to locate among the leaves.The Ovenbird spends much of its time on the forest floor where it builds a domed, oven-like nest.A fortunate observer may catch a glimpse of one perched in an understory shrub or small tree.But hearing the Ovenbird’s song, “teacher-teacher-teacher”, is frequently the only way to detect it.The Worm-eating Warbler nests in understory thickets on steep forested slopes. Its rich chipping song is often the only indication of its presence.Frequently easier to locate is the American Redstart, a Neotropical warbler that calls out its territory in damp woodlands from perches atop shrubs or among the lower limbs of trees.Warbling Vireos nest near streams or other bodies of water in large deciduous trees like this Northern Hackberry.Gray Catbirds are abundant in shrubby gardens and thickets. Most will come out of hiding to investigate disturbances like an observer making a pishing sound.Another inhabitant of brushy successional growth is the Common Yellowthroat. It too is curious and responds quickly to squeaky sounds made by human visitors to their home ground.The Yellow-breasted Chat is a bird of early successional growth. To establish and defend a breeding territory, this one adopted a perch along the edge of a woodlands overlooking a field in which prescribed fire was administered less than two months ago.Like the mockingbird and thrasher, the Yellow-breasted Chat is a mimic. Its song is a repertoire of the calls and songs of the bird species with which it may compete for food and nesting space. Unique to the chat is an occasional pause, whistle, or “chuck” note that creates a bridge between many of its song’s elements.The intensely territorial chat dropping down to look us over.
Sometimes we have to count ourselves lucky if we see just one in five, ten, or even twenty of the birds we hear in the cover of the forest canopy or thicket. But that’s what makes this time of year so rewarding for the dedicated observer. The more time you spend out there, the more you’ll eventually discover. See you afield!
A singing Scarlet Tanager lurking in the shade of an oak.Seeing is believing.
Despite what seemed to be a chilly early spring, the bright green leaves that unfold to close the canopy of our deciduous forests were dense and casting shade by the last days of April. For northbound migrants, this fresh foliage provides the cover they need for foraging, resting, and, for those that will stick around to breed in the lower Susquehanna valley, nesting.
We recall many occasions when sparse foliage during the first days of May seemed to delay the big push of Neotropical species, but the seasonal arrival of these birds in 2025 is thus far mostly ahead of schedule. This absence of delay is due in part to the lushness of the oaks. Some stands have not only leafed out, but are finished flowering and have added up to 12 inches of new branch growth. We spent these early hours of May among the oaks. Here’s a look at the Neotropical migrants and other species we found…
Dozens of species of Neotropical warblers have arrived in the Lower Susquehanna River Watershed. Some will nest locally while others will continue along flight paths that will ultimately take them far to our north. We spotted this Cape May Warbler in the upper reaches of a Northern Red Oak feeding among its spent flowers. Its destination: the spruce forests of northern New England and Canada.Slightly larger than the warblers are the vireos, including this Yellow-throated Vireo that has arrived to nest in an oak-maple bottomland.Always a fan favorite, we found this marvelously tropical Baltimore Oriole among the foliage of a flowering White Oak.This young Red-tailed Hawk has survived its first winter. It appears to have learned from experience how to hunt from advantageous locations such as this Chestnut Oak along a utility right-of-way where prey may include numerous squirrels as well as mice and other small mammals.We were taking a break beneath this Pin Oak when suddenly an American Kestrel arrived to begin scolding a trespasser that was straying a bit too close to its nest;……the intruder, a Northern Harrier, soon took the hint and continued on its way.While among the oaks, it pays to check the understory where Neotropical thrushes including the Veery are arriving to fill the forests with their melodious songs.Following their nocturnal flights, the last of the season’s northbound White-throated Sparrows may presently be found spending the day in the cover of the oak woodland understory.
The movements of our migratory birds typically continue through much of the month of May. And peak numbers of Neotropical species often occur sometime during the second week of the month. But with habitat at the ready, favorable flight conditions could facilitate quick arrival and/or passage of the bulk of the remaining migrants during the coming week. You may want to venture out sooner rather than later—but watch your step!
The Mayapple, a native wildflower also known as the Mandrake, is now in bloom among the oak leaf litter.And the Canada Mayflower (Maianthemum canadense), a native species also known as Wild Lily-of-the-valley, is blooming in moist mixed-oak forests.At the edge of an oak-maple woodland, we found this lingering White-crowned Sparrow foraging among the leaf litter and fresh, shiny foliage of Poison Ivy. Songbirds can spend their days scratching the ground among these leaves of three. If you do it, you’ll be scratching something else, and you may need an ocean of calamine lotion to boot!
Our wildlife has been having a tough winter. The local species not only contend with cold and stormy weather, but they also need to find food and shelter in a landscape that we’ve rendered sterile of these essentials throughout much of the lower Susquehanna valley’s farmlands, suburbs, and cities.
Planting trees, shrubs, flowers, and grasses that benefit our animals can go a long way, often turning a ho-hum parcel of property into a privately owned oasis. Providing places for wildlife to feed, rest, and raise their young can help assure the survival of many of our indigenous species. With a little dedication, you can be liberated from the chore of manicuring a lawn and instead spend your time enjoying the birds, mammals, insects, and other creatures that will visit your custom-made habitat.
What makes some neighborhoods so appealing? It’s the foresight property owners had a half a century or more ago when they planted their lawns and gardens with a variety of sturdy, long-lived trees and shrubs. They’ve not only minimized the need for mowing grass, they’ve provided the present-day residents of their home with added thermal stability during both the blazing heat of summer and the chilling cold of winter.
Fortunately for us, our local county conservation districts are again conducting springtime tree sales offering a variety of native and beneficial cultivated plants at discount prices. Listed here are links to information on how to pre-order your plants for pickup in April. Click away to check out the species each county is offering in 2025!
Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025
During its 2025 tree seedling sale, the Cumberland County Conservation District is offering Northeast Native Wildflower seed mix for four dollars per ounce. One ounce plants approximately 200 square feet of bare soil. This is a Zebra Swallowtail visiting nectar-rich flowers during July of the first year after sowing this mix at a site along the Susquehanna.
Pickup on: Thursday, April 24, 2025 or Friday, April 25, 2025
Able to thrive in wet soils, Red-osier and Silky Dogwood shrubs are ideal plants for intercepting and polishing stormwater in swales, detention/retention basins, and rain gardens. With their crimson twigs in winter, they look great along borders among clusters of cedars, pines, spruces, and other evergreens. They make an excellent choice for soil stabilization along the shorelines of streams, ponds, and other bodies of water too. Buy a dozen or more to create a showy mass planting in your soggy spot.
The Franklin County Conservation District is offering American Elm seedlings in bundles of 25 for 36 dollars. Start them in pots for a couple of years to really get ’em going, then find places with damp soil and plenty of room to give ’em a try. During autumn, they look great in the company of spruces, white pines, and other large evergreens.
We purchased these Eastern White Pine, Norway Spruce, and Common Winterberry plants from the Lancaster County Conservation District Tree Sale about four years ago. They’re filling in as understory growth in the margins beneath some thirty-year-old Eastern Hemlocks to create dense cover for resident and visiting fauna at susquehannawildlife.net headquarters.
For 2025, the Lebanon County Conservation District is offering Dawn Redwood (Metasequoia glyptostroboides) trees in packs of ten for twelve dollars. Though native to Asia, these adaptable trees present little threat of naturalizing and have many positive attributes in a conservation planting. Given ample space, the Dawn Redwood, a relative of the sequoias, will rapidly grow to a towering giant. They create a particularly dramatic landscape feature when planted in clumps of three to five trees or more. With age, the trunks become stout and very sturdy. Don’t like raking? The finely divided deciduous foliage can be left where it falls in autumn. It usually disintegrates by spring to enrich the soil and promote more growth.The genus Metasequoia was first described in 1941 based upon fossils collected in Jurassic and earlier strata from widespread locations in the northern hemisphere. Metasequoia were believed extinct until just a few years later when a small number of living Dawn Redwoods were first discovered in southern China. Now distributed around the world for cultivation, direct descendants of this wild population of Metasequoia glyptostroboides are available for nearly anyone in a temperate climate to plant and grow to exceptional size. (National Park Service image)The Lebanon County Conservation District is also selling Bald Cypress trees. They’re offered in bundles of ten for ten dollars. These long-lived trees resemble the Dawn Redwood. Both are tolerant of damp ground, but the native Bald Cypress is the species to choose for placement along streams, in wetlands, and on other sites with standing water or saturated soil.Wildlife rich Bald Cypress swamps currently occur on the Atlantic Coastal Plain as far north as Sussex County, Delaware. Just to the south, they’re also found along Chesapeake Bay in areas that, during the last glacial maximum when sea level was 300 to 400 feet below today’s tide lines, were the Lower Susquehanna River Watershed in portions of present-day Maryland and Virginia. The northward post-glacial range extension of Bald Cypresses is now blocked by centuries of human intervention that has eliminated, isolated, or fragmented the wetland habitats where they could potentially become established. Why not lend them a hand? Plant a cypress swamp in your flood-prone bottomland. (National Park Service image by Andrew Bennett)
A privacy planting of sturdy, native Eastern White Pines and Northern Red Oaks thriving around the border of a parking area where they also provide shade from the sun and help infiltrate a share of the stormwater that would otherwise become runoff.This year, the York County Conservation District is offering a Showy Northeast Native Wildflower and Grass seed mix for $19.99 per quarter pound. Sure beats mowing!
If you live in Adams County, Pennsylvania, you may be eligible to receive free trees and shrubs for your property from the Adams County Planting Partnership (Adams County Conservation District and the Watershed Alliance of Adams County). These trees are provided by the Chesapeake Bay Foundation’s Keystone 10-Million Trees Partnership which aims to close a seven-year project in 2025 by realizing the goal of planting 10 million trees to protect streams by stabilizing soils, taking up nutrients, reducing stormwater runoff, and providing shade. If you own property located outside of Adams County, but still within the Chesapeake Bay Watershed (which includes all of the Susquehanna, Juniata, and Potomac River drainages), you still may have an opportunity to get involved. Contact your local county conservation district office or watershed organization for information.
As they mature, tree and shrub plantings along streams return pollution-controlling functions to floodplains and provide critical habitat for wildlife. These riparian buffers not only improve water quality for fisheries, they also create travel corridors that prevent terrestrial animal populations from becoming isolated.Do you own a parcel of streamside or wetland acreage that you’d like to set aside and plant for the benefit of wildlife and water quality? Contact your local county conservation district office and ask them to tell you about CREP (Conservation Reserve Enhancement Program) and other programs that may offer incentives including payment of all or a portion of the costs of plantings and other habitat improvements.
We hope you’re already shopping. Need help making your selections? Click on the “Trees, Shrubs, and Woody Vines” tab at the top of this page to check out Uncle Tyler Dyer’s leaf collection. He has most of the species labelled with their National Wetland Plant List Indicator Rating. You can consult these ratings to help find species suited to the soil moisture on your planting site(s). For example: if your site has sloped upland ground and/or the soils sometimes dry out in summer, select plants with a rating such as UPL or FACU. If your planting in soils that remain moist or wet, select plants with the OBL or FACW rating. Plants rated FAC are generally adaptable and can usually go either way, but may not thrive or survive under stressful conditions in extremely wet or dry soils.
NATIONAL WETLAND PLANT LIST INDICATOR RATING DEFINITIONS
OBL (Obligate Wetland Plants)—Almost always occur in wetlands.
FACW (Facultative Wetland Plants)—Usually occur in wetlands, but may occur in non-wetlands.
FAC (Facultative Wetland Plants)—Occur in wetlands and non-wetlands.
FACU (Facultative Upland Plants)—Usually occur in non-wetlands, but may occur in wetlands.
UPL (Upland Plants)—Almost never occur in wetlands.
Using these ratings, you might choose to plant Pin Oaks (FACW) and Swamp White Oaks (FACW) in your riparian buffer along a stream; Northern Red Oaks (FACU) and White Oaks (FACU) in the lawn or along the street, driveway, or parking area; and Chestnut Oaks (UPL) on your really dry hillside with shallow soil. Give it a try.
Recently, we found these Chestnut Oak acorns setting roots into the leaf litter to secure their place among the plants that will turn the forest understory green in the spring. Individual acorns that germinate soon after falling to the ground in autumn may avoid becoming food for squirrels, turkeys, deer, and other wildlife, thus increasing their chances of surviving to later become adult trees able to produce acorns that pass this quick-development trait to yet another generation of oaks.
With autumn coming to a close, let’s have a look at some of the fascinating insects (and a spider) that put on a show during some mild afternoons in the late months of 2019.
Bush Katydids (Scudderia species) are found in brushy habitats and along rural roadsides. Their green summer color fades to brown, maroon, and gold to match the autumn foliage where they hide. Bush katydids often remain active until a hard freeze finally does them in.The Eastern Buck Moth (Hemileuca maia) is fuzzy, appearing to wear a warm coat for its autumn expeditions. Adults emerge in October and may fly as late as December. Females deposit their eggs on the twigs of Bear Oak (Quercus ilicifolia), Blackjack Oak (Q. marilandica), or Chestnut Oak (Q. montana), trees that, in our region, seem most favorable for the moth’s use when growing on burned barrens and mountain slopes. The spiny caterpillars are known to feed only on the foliage of these few trees. In the lower Susquehanna valley, the Eastern Buck Moth is rare because its specialized habitat is in short supply, and it’s all Smokey The Bear’s fault.The Sachem (Atalopedes campestris) wanders north from the Atlantic Coastal Plain into the Susquehanna valley each summer. In some years they become the most numerous small orange butterfly of all, particularly around home gardens. The larvae will feed on Crabgrass (Digitaria species), but have not found success overwintering this far north. By November, adults begin to look pretty drab.From 1978 through 1982, the Asian Multicolored Lady Beetle (Harmonia axyridis) was introduced into the eastern states by the United States Department of Agriculture. It has become a nuisance in many areas where it swarms, sometimes bites, and often overwinters in large smelly masses within homes and other warm buildings. As you may have guessed, it’s possibly displacing some of the less aggressive native lady beetle species.On a chilly afternoon, a sun-warmed Bold Jumping Spider (Phidippus audax) pounced and dispatched this sluggish worker Honey Bee (Apis mellifera) that was trying to gather pollen from a late-season Purple Coneflower bloom. This spider is bold indeed.Under bridges, inside bird nest boxes, and sometimes beneath porches, the female Pipe Organ Mud Dauber (Trypoxylon politum), a predatory wasp, builds these elaborate nests composed of long rows (pipes) of nursery cells. Into each cell one or more paralyzed spiders is deposited along with one of the female’s eggs. When hatched, each larva will feed upon the paralyzed spider(s) inside its cell, then pupate. The pupae overwinter, then emerge from their cells as adults during the following spring. In the autumn, males often stand guard at an entrance to the nest to prevent parasitic species, including some flies (look at the fifth pipe from the right), from laying eggs on the pupae. These wasps are not aggressive toward humans.A Black-and-yellow Mud Dauber (Sceliphron caementarium) observes a neighboring nest of Common Paper Wasps (Polistes exclamans). The Common Paper Wasp, a species also known as the Guinea Paper Wasp, is a native of the southern United States. It is currently expanding its range into the lower Susquehanna valley from the Atlantic Coastal Plain. These two wasp species and the Pipe Organ Mud Dauber are known to regularly coexist. All three will take advantage of man-made structures for their nest sites. People using the picnic tables beneath this pavilion roof never noticed the hundreds of docile wasps above.Those moody Eastern Yellowjackets (Vespula maculifrons) can get very temperamental during warm autumn days. These wasps may appear to have no enemies, but away from areas impacted by man’s everyday activities, they do. The Robber Fly (Promachus species) hunts like a flycatcher or other woodland bird, waiting on a perch along the forest’s edge for prey to pass by, then ambushing it, yellowjackets included.The invasive Spotted Lanternfly, a native of eastern Asia, continues to spread destruction. It established itself throughout much of the east side of the lower Susquehanna River during the summer and fall of 2019. Their route of travel across the farmlands of the region intersects with plenty of vineyards to obliterate and few, if any, natural enemies. Expect them to begin colonizing the west shore en masse during 2020.In 2020, plan to roll a few Spotted Lanternflies over, enjoy the view, and wait for the crimson tide to pass. With any luck, they’ll peak in a year or two.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York, NY.
