It begins on a sunny morning in spring each year, just as the ground temperature reaches sixty degrees or more…
Eastern Subterranean Termites emerge in unison from a nest located in the soil beneath a log. Each of these swarming “alates” is a potential king or queen seeking to find a location with an ample supply of fallen timber to provide food for establishment of a new colony. They are escorted to the surface by a soldier (lower center) equipped with powerful jaws for protecting the existing nest. Similar-looking worker termites tend the nest, the queen, her eggs, and their siblings, but usually remain hidden from view. The workers feed upon wood, hosting cellulose-digesting protozoa and bacteria in their guts to break down the fibers. This symbiotic relationship is an important mode of decomposition in the forest, the process that turns wood into the organic matter that enriches soil and helps it to retain more moisture.Termites are among the numerous arthropods that join fungi and a variety of microbes to decompose dead wood and other plant matter into the nutrients and organic materials used by living plants to thrive and grow.Within moments of emerging, swarming “alates” ascend a tree trunk or other vertical surface from which they can take flight.Eastern Subterranean Termite “alates” gather atop a stump before launching skyward.Termites swarm in massive numbers in an attempt to overwhelm the predators that are inevitably attracted to their sudden appearance. The few “alates” that survive to find a source of rotting wood in which to begin a new colony are the only hope for continuing their king and queen’s legacy.Soon after lift off, the majority of swarming termites are consumed by swallows, swifts, and other birds, but some are discovered at ground level.A lightning-fast strike with its tongue and this Green frog has snatched up yet another termite. Those that slip by the dragnet of terrestrial and aerial predators can sometimes start a new colony in the ground beneath a dead tree or in a vulnerable house or other wooden structure. To keep a small clan from invading your home, be certain the wood elements of your building(s) are kept dry and are not in contact with dirt, soil, bark mulch, etc. Regular inspections for evidence of their presence can head off the long-term damage termites can inflict on the stuff we construct with tree skeletons.A Green Frog wearing its breakfast. Its next chance for a termite feast may come during the autumn when the Drywood Termites (Kalotermitidae) swarm.
Of course, termites aren’t the only groups of insects to swarm. As heated runoff from slow-moving thundershowers has increased stream temperatures during the past couple of weeks, there have occurred a number of seasonal mayfly “hatches” on the Susquehanna and its tributaries. These “hatches” are actually the nuptial flights of newly emerged imago and adult mayflies. The most conspicuous of these is the Great Brown Drake.
Seen here with a much smaller and more typical regional mayfly to its left, the Great Brown Drake was for several years infamous for swarming the lights and creating traffic hazards on bridges spanning the Susquehanna. During the past two weeks, nighttime flights of these giants have ventured out to gather at well lit locations in housing and business districts more than a dozen miles from the river. Earlier this century, this proclivity to wander probably led the Great Brown Drake to first invade silty segments of the Susquehanna as a colonizer from its native range in the Mississippi watershed.
Swarms of another storm-related visitor are being seen throughout the lower Susquehanna valley right now. Have you noticed the Wandering Gliders?
Throughout the month, swarms of Wandering Gliders, the most widespread dragonflies in the world, descended on areas hit by localized slow-moving thundershowers. Large numbers of these global travelers are known to get swept up within the thermal air masses that lead to these storms. In suitable terrain within the path of the downpours, they linger to search for flooded places where they can mate and deposit eggs. Wandering Gliders frequently mistake large parking lots at shopping malls, grocery stores, etc. for wetlands and will be seen in these areas depositing eggs upon the hoods and roofs of shiny motor vehicles, surfaces which appear puddle-like in their eyes.More dragonfly swarms are yet to come. Adult Common Green Darners are presently in the Lower Susquehanna River Watershed mating and depositing eggs within vegetated ponds, lakes, and wetlands. Beginning in August, adult Common Green Darners and other migratory species including Black Saddlebags, Carolina Saddlebags, Wandering Gliders, and Twelve-spotted Skimmers will begin swarming as they feed on flying insects (including lots of mosquitos and gnats) and start working their way toward the Atlantic Coastal Plain. Along the barrier islands by September, concentrations of southbound dragonflies can reach the thousands, particularly at choke points like Cape May, New Jersey, and Cape Charles, Virginia. So be sure to keep an eye on the sky for swarms of dragonflies during coming weeks. And don’t forget to check out our “Damselflies and Dragonflies” page by clicking the tab at the top of this page.
During the coming two weeks, peak numbers of migrating Neotropical birds will be passing through the northeastern United States including the lower Susquehanna valley. Hawk watches are staffed and observers are awaiting big flights of Broad-winged Hawks—hoping to see a thousand birds or more in a single day.
During its passage through the Lower Susquehanna River Watershed, an adult Broad-winged Hawk sails over Second Mountain Hawk Watch in Lebanon County, Pennsylvania.A hatch-year/juvenile Broad-winged Hawk gazes toward hawk watchers on the ground.
Broad-winged hawks feed on rodents, amphibians, and a variety of large insects while on their breeding grounds in the forests of the northern United States and Canada. They depart early, journeying to wintering areas in Central and South America before frost robs them of a reliable food supply.
The Carolina Saddlebags (Tramea carolina), this one photographed at Second Mountain Hawk Watch on September 8th, is the rarest of the lower Susquehanna region’s migratory dragonflies. Autumn Broad-winged Hawk movements coincide with southbound flights of the Carolina Saddlebags and the more numerous migratory dragonfly species: Common Green Darner, Wandering Glider, Twelve-spotted Skimmer, and Black Saddlebags. “Broad-wings” will often eat these and other dragonflies during migration and can sometimes be seen catching and feeding upon them while still soaring high overhead.
While migrating, Broad-winged Hawks climb to great altitudes on thermal updrafts and are notoriously difficult to see from ground level. Bright sunny skies with no clouds to serve as a backdrop further complicate a hawk counter’s ability to spot passing birds. Throughout the Lower Susquehanna River Watershed, the coming week promises to be especially challenging for those trying to observe and census the passage of high-flying Broad-winged Hawks. The forecast of hot and humid weather is not so unusual, but the addition of smoke from fires in the western states promises to intensify the haze and create an especially irritating glare for those searching the skies for raptors.
Smoke from fires along the California coast and in central Utah can be seen streaming east this morning. (NOAA/GOES image)Smoke from western fires and humid air creates a band of haze in Ohio, Pennsylvania, New Jersey, and states to the south this morning. (NOAA/GOES image)
A migrating Broad-winged Hawk in the glare of a hazy sky. In addition to visibility problems, swarms of Spotted Lanternflies above the treetops make distant hawks difficult to discern for hawk watchers scanning the horizon with binoculars.
It may seem gloomy for the mid-September flights in 2021, but hawk watchers are hardy types. They know that the birds won’t wait. So if you want to see migrating “Broad-wings” and other species, you’ve got to get out there and look up while they’re passing through.
Migrating Ospreys typically fly low enough and are large enough to be spotted even during the haziest of conditions.Bald Eagles like this fourth-year bird can ascend to great altitude, but their size usually prevents them from sneaking past a lookout unnoticed.Peregrines escape notice not due to hazy sky conditions, but because they pass by so quickly. They’re being seen at local hawk watches now through October.
These hawk watches in the Lower Susquehanna River Watershed are currently staffed by official counters and all welcome visitors:
Rocky Ridge County Park Hawk Watch—3699 Deininger Road off Mount Zion Road (Route 24) northeast of York, Pennsylvania.
Second Mountain Hawk Watch—off Cold Spring Road on the grounds of Fort Indiantown Gap in Lebanon County, Pennsylvania.
Waggoner’s Gap Hawk Watch—where Route 74 crosses Blue Mountain north of Carlisle, Pennsylvania.
—or you can just keep an eye on the sky from wherever you happen to be. And don’t forget to check the trees and shrubs because warbler numbers are peaking too! During recent days…
Northern Parula at Chiques Rock County Park in Lancaster County.Black-and-white Warbler at Rocky Ridge County Park in York County.Cape May Warbler at Chiques Rock County Park in Lancaster County.Bay-breasted Warbler at Rocky Ridge County Park in York County.
It’s been more than a week since Tropical Storm Isaias moved swiftly up the Atlantic seaboard leaving wind and flood damage in its wake. Here in the Lower Susquehanna River Watershed, the brevity of its presence minimized the effects.
Tropical Storm Isaias moves north-northeast across the Delmarva Peninsula.
You may have noticed some summertime visitors flying about during these hot humid days that followed Isaias’ passing. They’re the dragonflies.
Our familiar friend the Wandering Glider is widespread throughout the valley right now—dropping eggs on shiny automobile hoods that look to them like a nice quiet puddle of water.
The Wandering Glider is a global traveler. Here in the lower Susquehanna valley, it is currently abundant around still water and in large parking lots.
Each of the other common migratory species is here too. Look for them patrolling the skies over large bodies of water and over adjacent fallow land and meadows where tiny flying insects abound. Did these dragonflies arrive on the winds associated with the tropical storm, or did they move in with the waves of warm air that followed it? Probably a little of both.
The Common Green Darner, a large dragonfly, can be the most abundant of the migratory species. Watch for high-flying swarms in the coming weeks.The Black Saddlebags is recognized in flight by the black base of each hindwing. These patches give the appearance of a pair of saddlebags draped across the dragonfly’s thorax.The Twelve-spotted Skimmer(Libellula pulchella) is a regular migrant.The Prince Baskettail (Epitheca princeps) can occur among mixed groups of dragonflies. Despite it rarely being mentioned as a migrant, its proclivity for non-stop day-long flight makes it a likely sighting among sizeable swarms.
Big swarms of dragonflies don’t go unnoticed by predators—particularly birds. The southbound migration of kites, Broad-winged Hawks, American Kestrels, and Merlins often coincides with the swarming of migratory dragonflies in late summer. Each of these raptors will grab and feed upon these insects while on the wing—so keep an eye on the sky.
This Peregrine Falcon found the congregations of hundreds of dragonflies worthy of a closer look……and an acrobatic fly-by to disrupt the swarms. Fun, fun, fun, fun, fun.
Second Mountain Hawk Watch is located on a ridge top along the northern edge of the Fort Indiantown Gap Military Reservation and the southern edge of State Game Lands 211 in Lebanon County, Pennsylvania. The valley on the north side of the ridge, also known as St. Anthony’s Wilderness, is drained to the Susquehanna by Stony Creek. The valley to the south is drained toward the river by Indiantown Run, a tributary of Swatara Creek.
The hawk watch is able to operate at this prime location for observing the autumn migration of birds, butterflies, dragonflies, and bats through the courtesy of the Pennsylvania Game Commission and the Garrison Commander at Fort Indiantown Gap. The Second Mountain Hawk Watch Association is a non-profit organization that staffs the count site daily throughout the season and reports data to the North American Hawk Watch Association (posted daily at hawkcount.org).
Today, Second Mountain Hawk Watch was populated by observers who enjoyed today’s break in the rainy weather with a visit to the lookout to see what birds might be on the move. All were anxiously awaiting a big flight of Broad-winged Hawks, a forest-dwelling Neotropical species that often travels back to its wintering grounds in groups exceeding one hundred birds. Each autumn, many inland hawk watches in the northeast experience at least one day in mid-September with a Broad-winged Hawk count exceeding 1,000 birds. They are an early-season migrant and today’s southeast winds ahead of the remnants of Hurricane Florence (currently in the Carolinas) could push southwest-heading “Broad-wings” out of the Piedmont Province and into the Ridge and Valley Province for a pass by the Second Mountain lookout.
The flight turned out to be steady through the day with over three hundred Broad-winged Hawks sighted. The largest group consisted of several dozen birds. We would hope there are probably many more yet to come after the Florence rains pass through the northeast and out to sea by mid-week. Also seen today were Bald Eagles, Ospreys, American Kestrels, and a migrating Red-headed Woodpecker.
Migrating Broad-winged Hawks circle on a thermal updraft above Second Mountain Hawk Watch to gain altitude before gliding away to the southwest.
Migrating insects included Monarch butterflies, and the three commonest species of migratory dragonflies: Wandering Glider, Black Saddlebags, and Common Green Darner. The Common Green Darners swarmed the lookout by the dozens late in the afternoon and attracted a couple of American Kestrels, which had apparently set down from a day of migration. American Kestrels and Broad-winged Hawks feed upon dragonflies and often migrate in tandem with them for at least a portion of their journey.
Still later, as the last of the Broad-winged Hawks descended from great heights and began passing by just above the trees looking for a place to settle down, a most unwelcome visitor arrived at the lookout. It glided in from the St. Anthony’s Wilderness side of the ridge on showy crimson-red wings, then became nearly indiscernible from gray tree bark when it landed on a limb. It was the dreaded and potentially invasive Spotted Lanternfly (Lycorma delicatula). This large leafhopper is native to Asia and was first discovered in North America in the Oley Valley of eastern Berks County, Pennsylvania in 2014. The larval stage is exceptionally damaging to cultivated grape and orchard crops. It poses a threat to forest trees as well. Despite efforts to contain the species through quarantine and other methods, it’s obviously spreading quickly. Here on the Second Mountain lookout, we know that wind has a huge influence on the movement of birds and insects. The east and southeast winds we’ve experienced for nearly a week may be carrying Spotted Lanternflies well out of their most recent range and into the forests of the Ridge and Valley Province. We do know for certain that the Spotted Lanternfly has found its way into the Lower Susquehanna River Watershed.
This adult Spotted Lanternfly landed in a birch tree behind the observers at the Second Mountain Hawk Watch late this afternoon. It was first recognized by its bright red wings as it glided from treetops on the north side of the lookout.
It’s been an atypical summer. The lower Susquehanna River valley has been in a cycle of heavy rains for over a month and stream flooding has been a recurring event. At Conewago Falls, the Pothole Rocks have been inundated for weeks. The location used as a lookout for the Autumn Migration Count last fall is at the moment submerged in ten feet of roaring water. Any attempt to tally the migrants which are passing thru in 2018 will thus be delayed indefinitely. Of greater import, the flooding at Conewago Falls is impacting many of the animals and plants there at a critical time in their annual life cycle. Having been displaced from its usual breeding sites on the river, one insect species in particular seems to be omnipresent in upland areas right now, and few people have ever heard of it.
So, you take a cruise in the motorcar to your favorite store and arrive at the sprawling parking lot. Not wishing to have your doors dented or paint chipped because you settled for a space tightly packed among other shopper’s conveyances, you park out there in the “boondocks”. You know the place, the lightly-used portion of the lot where sometimes brush grows from cracks in the asphalt and you must be on alert for impatient consumers who throttle-up to high speeds and dash diagonally across the carefully painted grids on the pavement to reach their favorite parking destination in the front row. Coming to a stop, you take the car out of gear, set the brake, disengage the safety belt, and gather your shopping list. You grasp the door handle and, not wanting to be flattened by one of the aforementioned motorists, you have a look around before exiting.
It was then that you saw the thing, hovering above your shiny bright hood. For a brief moment, it seemed to be peering right through the windshield at you with big reddish-brown eyes. In just a second or two, it turned its whole bronze body ninety degrees to the left and darted away on its cellophane wings. Maybe you didn’t really get a good look at it. It was so fast. But it certainly was odd. Oh well, time to walk inside a grab a few provisions. Away you go.
Upon completion of your shopping, you’re taking the long stroll back to your car and you notice more of these peculiar creatures. Two are coupled together and are hovering above someone’s automobile hood, then they drop down, and the lower of the two taps its abdomen on the paint. You ask yourself, “What are these bizarre things?”
Meet the Wandering Glider (Pantala flavescens), also known as the Globe Wanderer or Globe Skimmer, a wide-ranging dragonfly known to occur on every continent with the exception of Antarctica.
Wandering Gliders sometimes arrive in the lower Susquehanna River valley in large numbers after catching a ride on sustained winds from southerly directions and will often fly and migrate in storm systems. Conditions for such movements have been optimal in our region since mid-July. These dragonflies will often hover above motor vehicle hoods and, after mating, females will deposit eggs upon them, apparently mistaking their glossy surface for small pools of water.
Wandering Gliders travel the globe, and as such are accomplished fliers. Adults spend most of the day on the wing, feeding upon a variety of flying insects. Days ago, I watched several intercepting a swarm of flying ants. As fast as ants left the ground they were grabbed and devoured by the gliders. Wandering Gliders are adept at taking day-flying mosquitos, often zipping stealthily past a person’s head or shoulders to grab one of the little pests—the would-be skeeter victim usually unaware of the whole affair.
Due to their nomadic life history, Wandering Gliders are opportunists when breeding and will lay eggs in most any body of freshwater. Their larvae do not overwinter prior to maturity; adults can be expected in a little more than one to two months. Repetitive flooding in the Lower Susquehanna River Watershed this summer may be reducing the availability of the best local breeding sites for this species—riverine, stream, and floodplain pools of standing water with prey. This may explain why thousands of Wandering Gliders are patrolling parking lots, farmlands, and urban areas this summer. And it’s the likely reason for their use of puddles on asphalt pavement, on rubber roofs, and in fields as places to try to deposit eggs. Unfortunately, they may be as likely to succeed there as they are on your motor vehicle hood.
At this time a year ago, the airspace above the Diabase Pothole Rocks at Conewago Falls was jammed with territorial male Wandering Gliders. Each male hovered at various locations around his breeding territory consisting of pools and water-filled potholes. Intruders would quickly be dispatched from the area, then the male would resume his patrols from a set of repetitively-used hovering positions about six feet above the rocks. Mating and egg-laying continued into late September. The larvae, also called nymphs or naiads, were readily observed in many pools and potholes in early October and the emergence of juveniles was noted in mid-October. The absence of flooding, the mild autumn weather, and the moderation of water temperatures in the pools and potholes courtesy of the sun-drenched diabase boulders helped to extend the 2017 breeding season for Wandering Gliders in Conewago Falls. They aren’t likely to experience the same favor this year, but their great ability to travel and adapt should overcome this momentary misfortune.
A male Wandering Glider aggressively patrols his territory in the Diabase Pothole Rocks Microhabitat at Conewago Falls. August 20, 2017.A mating pair of Wandering Gliders continue flying non-stop above one of thousands of suitable breeding pools among the Diabase Pothole Rocks at Conewago Falls. September 23, 2017.A female (bottom)Wandering Glider has deposited eggs in a pool while flying in tandem with a male (top). They’ll do the same thing on your automobile hood! Conewago Falls Diabase Pothole Rocks Microhabitat. September 23, 2017.Wandering Glider larvae are at the top of the food chain in flooded potholes. As they grew, these dragonfly larvae decimated the mosquito larvae which were abundant there earlier in the summer. October 7, 2017.A juvenile male Wandering Glider emerges from the pool where it fed and grew as a larva. It remained at water’s edge on the surface of a sun-warmed diabase rock for several hours to dry its wings. It soon flew away to parts unknown, possibly traveling hundreds or thousands of miles. Look carefully at the wings for the beige dash marks on the forward edge near the terminal end. Females lack this marking. Conewago Falls Diabase Pothole Rocks Microhabitat. October 14, 2017.A Wandering Glider exuviae, the shed exoskeleton of a creature gone, but not forgotten. October 14, 2017.
The humid rainy remains of Hurricane Nate have long since passed by Pennsylvania, yet mild wet weather lingers to confuse one’s sense of the seasons. This gloomy misty day was less than spectacular for watching migrating birds and insects, but some did pass by. Many resident animals of the falls are availing themselves of the opportunity to continue active behavior before the cold winds of autumn and winter force a change of lifestyle.
Warm drizzle at daybreak prompted several Northern Spring Peepers (Pseudacris crucifer crucifer) to begin calling from the wetlands in the Riparian Woodlands of Conewago Falls. An enormous chorus of these calls normally begins with the first warm rains of early spring to usher in this tiny frog’s mating season. Today, it was just a few “peeps” among anxious friends.
The tiny Northern Spring Peeper is recognized by the dark “X” across its back. Soon, shelter must be found among loose bark and fallen logs to commence hibernation. Emergence, often prompted by warm spring rains, will quickly be followed by a growing chorus of breeding calls as sometimes hundreds of these frogs assemble in vernal pools where mating will then occur.
Any additional river flow that resulted from the rains of the previous week is scarcely noticeable among the Pothole Rocks. The water level remains low, the water column is fairly clear, and the water temperatures are in the 60s Fahrenheit.
It’s no real surprise then to see aquatic turtles climbing onto the boulders in the falls to enjoy a little warmth, if not from the sun, then from the stored heat in the rocks. As usual, they’re quick to slide into the depths soon after sensing someone approaching or moving nearby. Seldom found anywhere but on the river, these skilled divers are Common Map Turtles (Graptemys geographica), also known as Northern Map Turtles. Their paddle-like feet are well adapted to swimming in strong current. They are benthic feeders, feasting upon a wide variety of invertebrates found among the stone and substrate of the river bottom.
Adult Common Map Turtles hibernate communally on the river bottom in a location protected from ice scour and turbulent flow, often using boulders, logs, or other structures as shelter from strong current. The oxygenation of waters tumbling through Conewago Falls may be critical to the survival of the turtles overwintering downstream. Dissolved oxygen in the water is absorbed by the nearly inactive turtles as they remain submerged at their hideout through the winter. Though Common Map Turtles, particularly males, may occasionally move about in their hibernation location, they are not seen coming to the surface to breathe.
The Common Map Turtles in the Susquehanna River basin are a population disconnected from that found in the main range of the species in the Great Lakes and upper Mississippi basin. Another isolated population exists in the Delaware River.
Common Map Turtles, including this recently hatched young seen in August, are often observed climbing onto rocks in the river.Note the oversize swimming fin adaptations of the feet on this adult Common Map Turtle found among the Pothole Rocks in Conewago Falls. Young and adults are capable of navigating some strong current to feed and escape danger.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
SOURCES
Committee on the Status of Endangered Wildlife in Canada. 2002. Status Report of the Northern Map Turtle. Canadian Wildlife Service. Ottawa, Ontario.
A moderate breeze from the south placed a headwind into the face of migrants trying to wing their way to winter quarters. The urge to reach their destination overwhelmed any inclination a bird or insect may have had to stay put and try again another day.
Blue Jays were joined by increasing numbers of American Robins crossing the river in small groups to continue their migratory voyages. Killdeer (Charadrius vociferous) and a handful of sandpipers headed down the river route. Other migrants today included a Cooper’s Hawk (Astur cooperii), Eastern Bluebirds (Sialia sialis), and a few Common Mergansers (Mergus merganser), House Finches (Haemorhous mexicanus), and Common Grackles (Quiscalus quiscula).
The afternoon belonged to the insects. The warm wind blew scores of Monarchs toward the north as they persistently flapped on a southwest heading. Many may have actually lost ground today. Painted Lady (Vanessa cardui) and Cloudless Sulphur butterflies were observed battling their way south as well. All three of the common migrating dragonflies were seen: Common Green Darner (Anax junius), Wandering Glider (Pantala flavescens), and Black Saddlebags (Tramea lacerata).
The warm weather and summer breeze are expected to continue as the rain and wind from Hurricane Nate, today striking coastal Alabama and Mississippi, progresses toward the Susquehanna River watershed during the coming forty-eight hours.
This Great Blue Heron was joined by numerous other fishermen and a good number of sightseers in the falls today.A colorful young Northern Water Snake (Nerodia sipedon) takes advantage of the sun-heated surface of a Pothole Rock to remain nimble and active. Cooler weather will soon compel this and other reptiles to find shelter for winter hibernation.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
