Photo of the Day

Male Blue Dasher at Memorial Lake State Park
An ever-vigilant male Blue Dasher (Pachydiplax longipennis) perches on a small twig overlooking the vegetated shallows along the shoreline of Memorial Lake in Lebanon County, Pennsylvania.

Photo of the Day

An "Annual Cicada" (Neotibicen) dries its wings after emerging from its exuvia.
The Eastern Scissor Grinder (Neotibicen winnemanna), a species of “Annual Cicada” or “Dog-day Cicada”, dries its wings after emerging from its exoskeleton (exuvia).  After two to five years of subterranean life as a nymph, this adult will soon be ready to fly into the treetops in search of a mate.  There is a sense of urgency.  Summer’s end will bring its life to an end as well.

Butterflies and More at Boyd Big Tree Preserve Conservation Area

If you’re feeling the need to see summertime butterflies and their numbers just don’t seem to be what they used to be in your garden, then plan an afternoon visit to the Boyd Big Tree Preserve along Fishing Creek Valley Road (PA 443) just east of U.S. 22/322 and the Susquehanna River north of Harrisburg.  The Pennsylvania Department of Conservation and Natural Resources manages the park’s 1,025 acres mostly as forested land with more than ten miles of trails.  While located predominately on the north slope of Blue Mountain, a portion of the preserve straddles the crest of the ridge to include the upper reaches of the southern exposure.

American Chestnut at Boyd Big Tree Preserve
A grove of American Chestnuts (Castanea dentata) planted at Boyd Big Tree Preserve is part of a propagation program working to restore blight-resistant trees to Pennsylvania and other areas of their former range which included the Appalachians and the upper Ohio River watershed.

Fortunately, one need not take a strenuous hike up Blue Mountain to observe butterflies.  Open space along the park’s quarter-mile-long entrance road is maintained as a rolling meadow of wildflowers and cool-season grasses that provide nectar for adult butterflies and host plants for their larvae.

Butterfly Meadow at Boyd Big Tree Preserve
A view looking north at the butterfly meadow and entrance road at Boyd Big Tree Preserve Conservation Area.  Second Mountain is in the background.
Walking a Meadow Path
Mowed paths follow the entrance road and a portion of the perimeter of the meadow allowing visitors a chance to wander among the waist-high growth to see butterflies, birds, and blooming plants at close range without trampling the vegetation or risking exposure to ticks.
A Silver-spotted Skipper feeding on nectar from Dogbane (Apocynum cannabinum) flowers.
A Silver-spotted Skipper feeding on nectar from the flowers of Indian Hemp (Apocynum cannabinum).  Like the milkweeds, Indian Hemp is a member of the dogbane family (Apocynaceae).
An Eastern Tiger Swallowtail feeding on Common Milkweed.
An Eastern Tiger Swallowtail feeding on Common Milkweed.
Great Spangled Fritillary on Common Milkweed.
A Great Spangled Fritillary (Speyeria cybele) on Common Milkweed.
A Black Swallowtail feeding on Common Milkweed nectar.
A Black Swallowtail (Papilio polyxenes) feeding on Common Milkweed nectar.
A Pipevine Swallowtail on Common Milkweed.
A Pipevine Swallowtail (Battus philenor) on Common Milkweed.
A Pipevine Swallowtail on Common Milkweed.
Another Pipevine Swallowtail on Common Milkweed.  Note the hook-shaped row of red-orange spots on the underside of the hindwing.
A Pipevine Swallowtail on visiting Butterfly Weed.
A Pipevine Swallowtail visiting the brilliant blooms of Butterfly Weed, a favorite of a wide variety of pollinators.
A Black Swallowtail on Butterfly Weed
A Black Swallowtail with damaged wings alights atop a Butterfly Weed flower cluster.  Note the pair of parallel rows of red-orange spot on the underside of the hindwing.
A Monarch on Butterfly Weed
A Monarch feeding on nectar from the flowers of Butterfly Weed.
A mating pair of Eastern Tailed Blues.
A mating pair of Eastern Tailed Blues on a Timothy (Phleum pratense) spike.
A female (left) and male Great Spangled Fritillary.
A male Great Spangled Fritillary (right) pursuing a female.
Common Green Darner
Butterflies aren’t the only colorful insects patrolling the meadows at Boyd Big Tree Preserve.  Dragonflies including Common Green Darners are busily pursuing prey, particularly small flying insects like mosquitos, gnats, and flies.
Juvenile Broad-winged Hawk
Dragonflies themselves can become prey and are much sought after by Broad-winged Hawks. This very vocal juvenile gave us several good looks as it ventured from the forest into the skies above the upper meadow during midday.  It wasn’t yet a good enough flier to snag a dragonfly, but it will have plenty of opportunities for practice during its upcoming fall migration which, for these Neotropical raptors, will get underway later this month.

Do yourself a favor and take a trip to the Boyd Big Tree Preserve Conservation Area.  Who knows?  It might actually inspire you to convert that lawn or other mowed space into much-needed butterfly/pollinator habitat.

While you’re out, you can identify your sightings using our photographic guide—Butterflies of the Lower Susquehanna River Watershed—by clicking the “Butterflies” tab at the top this page.  And while you’re at it, you can brush up on your hawk identification skills ahead of the upcoming migration by clicking the “Hawkwatcher’s Helper: Identifying Bald Eagles and other Diurnal Raptors” tab.  Therein you’ll find a listing and descriptions of hawk watch locations in and around the lower Susquehanna region.  Plan to visit one or more this autumn!

Shorebirds and More at Bombay Hook National Wildlife Refuge

Have you purchased your 2023-2024 Federal Duck Stamp?  Nearly every penny of the 25 dollars you spend for a duck stamp goes toward habitat acquisition and improvements for waterfowl and the hundreds of other animal species that use wetlands for breeding, feeding, and as migration stopover points.  Duck stamps aren’t just for hunters, purchasers get free admission to National Wildlife Refuges all over the United States.  So do something good for conservation—stop by your local post office and get your Federal Duck Stamp.

2023-2024 Federal Duck Stamp. Your Federal Duck Stamp is your free pass to visit the nation's National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.
Your Federal Duck Stamp is your admission ticket for entry into many of the country’s National Wildlife Refuges including Bombay Hook National Wildlife Refuge on Delaware Bay near Smyrna, Delaware.

Still not convinced that a Federal Duck Stamp is worth the money?  Well then, follow along as we take a photo tour of Bombay Hook National Wildlife Refuge.  Numbers of southbound shorebirds are on the rise in the refuge’s saltwater marshes and freshwater pools, so we timed a visit earlier this week to coincide with a late-morning high tide.

Northern Bobwhite
This pair of Northern Bobwhite, a species now extirpated from the Lower Susquehanna River Watershed and the rest of Pennsylvania, escorted us into the refuge.  At Bombay Hook, they don’t waste your money mowing grass.  Instead, a mosaic of warm-season grasses and early successional growth creates ideal habitat for Northern Bobwhite and other wildlife.
Shearness Pool at Bombay Hook N.W.R.
Twice each day, high tide inundates mudflats in the saltwater tidal marshes at Bombay Hook prompting shorebirds to move into the four man-made freshwater pools.  Birds there can often be observed at close range.  The auto tour route through the refuge primarily follows a path atop the dikes that create these freshwater pools.  Morning light is best when viewing birds on the freshwater side of the road, late-afternoon light is best for observing birds on the tidal saltwater side.
Great Blue Heron
A Great Blue Heron at high tide on the edge of a tidal creek that borders Bombay Hook’s tour route at Raymond Pool.
Semipalmated Sandpipers
Semipalmated Sandpipers stream into Raymond Pool to escape the rising tide in the salt marsh.
Semipalmated Sandpipers and Short-billed Dowitcher
More Semipalmated Sandpipers and a single Short-billed Dowitcher (Limnodromus griseus) arrive at Raymond Pool.
Short-billed Dowitchers
Two more Short-billed Dowitchers on the way in.
Sandpipers, Avocets, Egrets, and Mallards
Recent rains have flooded some of the mudflats in Bombay Hook’s freshwater pools. During our visit, birds were often clustered in areas where bare ground was exposed or where water was shallow enough to feed.  Here, Short-billed Dowitchers in the foreground wade in deeper water to probe the bottom while Semipalmated Sandpipers arrive to feed along the pool’s edge.  Mallards, American Avocets, and egrets are gathered on the shore.
Short-billed Dowitchers
More Short-billed Dowitchers arriving to feed in Raymond Pool.
Semipalmated Sandpipers
Hundreds of Semipalmated Sandpipers gathered in shallow water where mudflats are usually exposed during mid-summer in Raymond Pool.
Hundreds of Semipalmated Sandpipers, several Short-billed Dowitchers, and some Forster’s Terns (Sterna forsteri) crowd onto a mud bar at Bear Swamp Pool.
Semipalmated Sandpipers, Forster's Terns, and a Short-billed Dowitcher
A zoomed-in view of the previous image showing a tightly packed crowd of Semipalmated Sandpipers, Forster’s Terns, and a Short-billed Dowitcher (upper left).
Short-billed Dowitchers
Short-billed Dowitchers wading to feed in the unusually high waters of Raymond Pool.
Short-billed Dowitchers, American Avocets, and a Snowy Egret
Short-billed Dowitchers, American Avocets, and a Snowy Egret in Raymond Pool.  A single Stilt Sandpiper (Calidris himantopus) can been seen flying near the top of the flock of dowitchers just below the egret.
Stilt Sandpiper among Short-billed Dowitchers
Zoomed-in view of a Stilt Sandpiper (Calidris himantopus), the bird with white wing linings.
American Avocets
American Avocets probe the muddy bottom of Raymond Pool.
Dunlin and Short-billed Dowitchers
Among these Short-billed Dowitchers, the second bird from the bottom is a Dunlin. This sandpiper, still in breeding plumage, is a little bit early.  Many migrating Dunlin linger at Bombay Hook into October and even November.
Least Sandpiper
This Least Sandpiper found a nice little feeding area all to itself at Bear Swamp Pool.
Lesser Yellowlegs
Lesser Yellowlegs at Bear Swamp Pool.
Lesser Yellowlegs
Lesser Yellowlegs at Bear Swamp Pool
Greater Yellowlegs
A Greater Yellowlegs at Bear Swamp Pool.
Caspian Tern
A Caspian Tern patrolling Raymond Pool.
Marsh Wren singing
The chattering notes of the Marsh Wren’s (Cistothorus palustris) song can be heard along the tour road wherever it borders tidal waters.
Marsh Wren Nest
This dome-shaped Marsh Wren nest is supported by the stems of Saltwater Cordgrass (Sporobolus alterniflorus), a plant also known as Smooth Cordgrass.  High tide licks at the roots of the cordgrass supporting the temporary domicile.
Seaside Dragonlet
By far the most common dragonfly at Bombay Hook is the Seaside Dragonlet (Erythrodiplax berenice).  It is our only dragonfly able to breed in saltwater.  Seaside Dragonlets are in constant view along the impoundment dikes in the refuge.
Red-winged Blackbird
Red-winged Blackbirds are still nesting at Bombay Hook, probably tending a second brood.
Bobolink
Look up!   A migrating Bobolink passes over the dike at Shearness Pool.
Mute Swans and Canada Geese
Non-native Mute Swans and resident-type Canada Geese in the rain-swollen Shearness Pool.
Trumpeter Swans
A pair of Trumpeter Swans (Cygnus buccinator) as seen from the observation tower at Shearness Pool.  Unlike gregarious Tundra and Mute Swans, pairs of Trumpeter Swans prefer to nest alone, one pair to a pond, lake, or sluggish stretch of river.  The range of these enormous birds was restricted to western North America and their numbers were believed to be as low as 70 birds during the early twentieth century.  An isolated population consisting of several thousand birds was discovered in a remote area of Alaska during the 1930s allowing conservation practices to protect and restore their numbers.  Trumpeter Swans are slowly repopulating scattered east coast locations following recent re-introduction into suitable habitats in the Great Lakes region.
Great Egret
A Great Egret prowling Shearness Pool.
Snowy Egret
A Snowy Egret in Bear Swamp Pool.
A hen Wood Duck (second from right) escorts her young.
Wood Ducks in Bear Swamp Pool.
Black-necked Stilt and young.
A Bombay Hook N.W.R. specialty, a Black-necked Stilt and young at Bear Swamp Pool.

As the tide recedes, shorebirds leave the freshwater pools to begin feeding on the vast mudflats exposed within the saltwater marshes.  Most birds are far from view, but that won’t stop a dedicated observer from finding other spectacular creatures on the bay side of the tour route road.

Bombay Hook National Wildlife Refuge protects a vast parcel of tidal salt marsh and an extensive network of tidal creeks. These areas are not only essential wildlife habitat, but are critical components for maintaining water quality in Delaware Bay and the Atlantic.
Atlantic Horseshoe Crab
The shells of expired Atlantic Horseshoe Crabs were formerly widespread and common among the naturally occurring flotsam along the high tide line on Delaware Bay.  We found just this one during our visit to Bombay Hook.  Man has certainly decimated populations of this ancient crustacean during recent decades.
As the tide goes out, it’s a good time for a quick walk into the salt marsh on the boardwalk trail opposite Raymond Pool.
Atlantic Marsh Fiddler Crabs
Among the Saltmarsh Cordgrass along the trail and on the banks of the tidal creek there, a visitor will find thousands and thousands of Atlantic Marsh Fiddler Crabs (Minuca pugnax).
Atlantic Marsh Fiddler Crabs
Atlantic Marsh Fiddler Crabs and their extensive system of burrows help prevent the compaction of tidal soils and thus help maintain ideal conditions for the pure stands of Saltwater Cordgrass that trap sediments and sequester nutrients in coastal wetlands.
Atlantic Marsh Fiddler Crab
A male Atlantic Marsh Fiddler Crab peers from its den.
Great Egret
Herons and egrets including this Great Egret are quite fond of fiddler crabs.  As the tide goes out, many will venture away from the freshwater pools into the salt marshes to find them.
Green Heron
A Green Heron seen just before descending into the cordgrass to find fiddler crabs for dinner.
Clapper Rail
A juvenile Clapper Rail (Rallus crepitans crepitans) emerges from the cover of the cordgrass along a tidal creek to search for a meal.
Glossy Ibis
Glossy Ibis leave their high-tide hiding place in Shearness Pool to head out into the tidal marshes for the afternoon.
Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide.
Great Black-backed Gulls, Herring Gulls, and possibly other species feed on the mudflats exposed by low tide in the marshes opposite Shearness Pool.
Ospey
An Osprey patrols the vast tidal areas opposite Shearness Pool.

No visit to Bombay Hook is complete without at least a quick loop through the upland habitats at the far end of the tour route.

Indigo Bunting
Indigo Buntings nest in areas of successional growth and yes, that is a Spotted Lanternfly on the grape vine at the far right side of the image.
Blue Grosbeak
Blue Grosbeaks (Passerina caerulea) are common nesting birds at Bombay Hook.  This one was in shrubby growth along the dike at the north end of Shearness Pool.
Trumpet Creeper and Poison Ivy
These two native vines are widespread at Bombay Hook and are an excellent source of food for birds. The orange flowers of the Trumpet Vine are a hummingbird favorite and the Poison Ivy provides berries for numerous species of wintering birds.
Pileated Woodpecker in Sweet Gum
The Pileated Woodpecker is one of the numerous birds that supplements its diet with Poison Ivy berries.  The tree this individual is visiting is an American Sweetgum (Liquidambar styraciflua), a species native to the Atlantic Coastal Plain in Delaware.  The seed balls are a favorite winter food of goldfinches and siskins.
Red-bellied Slider and Painted Turtle
Finis Pool has no frontage on the tidal marsh but is still worth a visit.  It lies along a spur road on the tour route and is located within a deciduous coastal plain forest.  Check the waters there for basking turtles like this giant Northern Red-bellied Cooter (Pseudemys rubiventris) and much smaller Painted Turtle.
White-tailed Deity
The White-tailed Deity is common along the road to Finis Pool.
Fowler's Toad
Fowler’s Toads (Anaxyrus fowleri) breed in the vernal ponds found in the vicinity of Finis Pool and elsewhere throughout the refuge.
Turk's Cap Lily
The National Wildlife Refuge System not only protects animal species, it sustains rare and unusual plants as well.  This beauty is a Turk’s Cap Lily (Lilium superbum), a native wildflower of wet woods and swamps.
Wild Turkey
Just as quail led us into the refuge this morning, this Wild Turkey did us the courtesy of leading us to the way out in the afternoon.

We hope you’ve been convinced to visit Bombay Hook National Wildlife Refuge sometime soon.  And we hope too that you’ll help fund additional conservation acquisitions and improvements by visiting your local post office and buying a Federal Duck Stamp.

A Few Plants with Wildlife Impact in June

Here’s a look at some native plants you can grow in your garden to really help wildlife in late spring and early summer.

The Larger Blue Flag (Iris versicolor) and Soft Rush (Juncus effusus) in flower in mid-June.
The showy bloom of a Larger Blue Flag (Iris versicolor) and the drooping inflorescence of Soft Rush (Juncus effusus).  These plants favor moist soils in wetlands and damp meadows where they form essential cover and feeding areas for insects, amphibians, and marsh birds.  Each is an excellent choice for helping to absorb nutrients in a rain garden or stream-side planting.  They do well in wet soil or shallow water along the edges of garden ponds too.
Smooth Shadbush
The fruits of Smooth Shadbush (Amelanchier laevis), also known as Allegheny Serviceberry, Smooth Serviceberry, or Smooth Juneberry, ripen in mid-June and are an irresistible treat for catbirds, robins, bluebirds, mockingbirds, and roving flocks of Cedar Waxwings.
Common Milkweed and Eastern Carpenter Bee
Also in mid-June, the fragrant blooms of Common Milkweed attract pollinators like Eastern Carpenter Bees,…
Common Milkweed and Honey Bee
…Honey Bees,…
Common Milkweed and Banded Hairstreak
…and butterflies including the Banded Hairstreak (Satyrium calanus).  In coming weeks, Monarch butterflies will find these Common Milkweed plants and begin laying their eggs on the leaves.  You can lend them a hand by planting milkweed species (Asclepias) in your garden.  Then watch the show as the eggs hatch and the caterpillars begin devouring the foliage.  Soon, they’ll pupate and, if you’re lucky, you’ll be able to watch an adult Monarch emerge from a chrysalis!

The Value of Water

Are you worried about your well running dry this summer?  Are you wondering if your public water supply is going to implement use restrictions in coming months?  If we do suddenly enter a wet spell again, are you concerned about losing valuable rainfall to flooding?  A sensible person should be curious about these issues, but here in the Lower Susquehanna River Watershed, we tend to take for granted the water we use on a daily basis.

This Wednesday, June 7,  you can learn more about the numerous measures we can take, both individually and as a community, to recharge our aquifers while at the same time improving water quality and wildlife habitat in and around our streams and rivers.  From 5:30 to 8:00 P.M., the Chiques Creek Watershed Alliance will be hosting its annual Watershed Expo at the Manheim Farm Show grounds adjacent to the Manheim Central High School in Lancaster County.  According to the organization’s web page, more than twenty organizations will be there with displays featuring conservation, aquatic wildlife, stream restoration, Honey Bees, and much more.  There will be games and custom-made fish-print t-shirts for the youngsters, plus music to relax by for those a little older.  Look for rain barrel painting and a rain barrel giveaway.  And you’ll like this—admission and ice cream are free.  Vendors including food trucks will be onsite preparing fare for sale.

And there’s much more.

To help recharge groundwater supplies, you can learn how to infiltrate stormwater from your downspouts, parking area, or driveway…

Urban Runoff
Does your local stream flood every time there’s a downpour, then sometimes dry up during the heat of summer?  Has this problem gotten worse over the years?  If so, you may be in big trouble during a drought.  Loss of base flow in a stream or river is a sure sign of depleted groundwater levels in at least a portion of its drainage basin.  Landowners, both public and private, in such a watershed need to start infiltrating stormwater into the ground instead of allowing it to become surface runoff.
Rain Garden Model
You can direct the stormwater from your downspout, parking area, or driveway into a rain garden to help recharge the aquifer that supplies your private or public well and nearby natural springs.  Displays including this model provided by Rapho Township show you how.

…there will be a tour of a comprehensive stream and floodplain rehabilitation project in Manheim Memorial Park adjacent to the fair grounds…

Legacy Sediments
Have you seen banks like these on your local stream?  On waterways throughout the Lower Susquehanna River Watershed, mill dams have trapped accumulations of sediments that eroded from farm fields prior to the implementation of soil conservation practices.  These legacy sediments channelize creeks and disconnect them from their now buried floodplains.  During storms, water that would have been absorbed by the floodplain is now displaced into areas of higher ground not historically inundated by a similar event.
Adjacent to the Manheim Farm Show grounds, the Chiques Creek Stream Restoration Project in Manheim Memorial Park has reconnected the waterway to its historic floodplain by removing a dam and the legacy sediments that accumulated behind it.
Legacy Sediments Removed
Chiques Creek in Manheim following removal of hundreds of truck loads of legacy sediments.  High water can again be absorbed by the wetlands and riparian forest of the floodplain surrounding this segment of stream.  There are no incised banks creating an unnatural channel or crumbling away to pollute downstream waters with nutrients and sediment.  Projects similar to this are critical to improving water quality in both the Susquehanna River and Chesapeake Bay.  Closer to home, they can help municipalities meet their stormwater management (MS4) requirements.
Bank-full Bench
Mark Metzler of Rettew Associates guides a tour of the Chiques Creek rehabilitation.  Here, cross vanes, stone structures that provide grade control along the stream’s course, were installed to gently steer the center of the channel away from existing structures.   Cross vanes manipulate the velocity of the creek’s flow across its breadth to dissipate potentially erosive energy and more precisely direct the deposition of gravel and sediment.

…and a highlight of the evening will be using an electrofishing apparatus to collect a sample of the fish now populating the rehabilitated segment of stream…

Electrofishing
Matt Kofroth, Lancaster County Conservation District Watershed Specialist, operates a backpack electrofishing apparatus while the netting crew prepares to capture the temporarily stunned specimens.  The catch is then brought to shore for identification and counting.

…so don’t miss it.  We can hardly wait to see you there!

The 2023 Watershed Expo is part of Lancaster Conservancy Water Week.

Termite Terminators

With temperatures on the rise in this morning’s bright sunshine, a flight of Eastern Subterranean Termites (Reticulitermes flavipes) streamed from the shelter of this tree snag to ascend skyward in a spreading cloud of kings and queens looking to start new colonies of wood chompers.

Eastern Subterranean Termites swarming.
A nuptial flight of Eastern Subterranean Termites taking to the wing after ascending a dead tree alongside a small pond.

As the swarm cleared the treetops, it immediately attracted the attention of some recently-arrived migratory birds…

Barn Swallows and Chimney Swifts Attack an Ant Swarm
Barn Swallows and Chimney Swifts seized the opportunity to devour an easy meal…hundreds of flying termites.  Their timing was perfect.  The first Barn swallows of the season arrived here just two days ago, the first appearance of the swifts was concurrent with this morning’s swarm.
Barn Swallow and Chimney Swifts
Our squadron of predators quickly devoured the airborne insects.  The entire event was concluded in just five minutes.

Photo of the Day

A Reforestation Project
Visible in the background of this image, an infestation of invasive Emerald Ash Borer larvae has killed the trees in a woodlot composed exclusively of Green Ash.  Left standing, the dead snags provide excellent habitat for a number of animal species including cavity-nesting birds like woodpeckers, known consumers of these destructive larvae.  To reforest the mowed field in the foreground, a variety of native deciduous trees have been planted.  In areas where a diversity of trees are not present to furnish a source of seeds for natural succession, manually planting an array of seedlings provides some insurance against the risk of allowing establishment of a single pioneer species such as the vulnerable Green Ash.  The white plastic tubes on the young trees offer protection from the ever-browsing White-tailed Deity.

Hidden Surprises

If you’re like us, you’re forgoing this year’s egg hunt due to the prices, and, well, because you’re a little bit too old for such a thing.

Instead, we took a closer look at some of our wildlife photographs from earlier in the week.  We’ve learned from experience that we don’t always see the finer details through the viewfinder, so it often pays to give each shot a second glance on a full-size screen.  Here are a few of our images that contained some hidden surprises.


Blue-winged Teal and American Black Ducks
We photographed these Blue-winged Teal and American Black Ducks as they were feeding in a meadow wetland…

..but upon closer inspection we located…

Common Green Darner
…a Common Green Darner patrolling for mosquitoes and other prey.  The Common Green Darner is a migratory species of dragonfly.  After mating, they deposit their eggs in wetland pools, ponds, and slow-moving streams.

Water Strider
We photographed this Water Strider as it was “walking” across a pool in a small stream that meanders through a marshy meadow…

..but after zooming in a little closer we found…

Water Strider with a Mosquito
…a mosquito coming to deposit its eggs had been seized as a mid-day meal.   Look at how the legs of the Water Strider use the surface tension of the pool to allow it to “walk on water”, even while clutching and subduing its prey.

Canada Geese
We photographed these resident Canada Geese in a small plowed cornfield in an area managed mostly as a mix of cool-season and warm-season grassland…

..but then, following further examination, we discovered…

Ring-necked Pheasant
…a hen Ring-necked Pheasant on a nest.

THE BAD EGG

Red-winged Blackbirds
We photographed this small group of migrating Red-winged Blackbirds while it was feeding among corn stubble in a plowed field…

…but a careful search of the flock revealed…

Brown-headed Cowbirds and Red-winged Blackbirds
…three female Brown-headed Cowbirds among them (the unstreaked brown birds, two to the far left and one among the “red-wings” to the right).  Cowbirds practice nest parasitism as a means of putting their young up for adoption.  Red-winged Blackbirds and numerous other species are the unknowing victims.  The female cowbird discreetly deposits her egg(s) in the adopting party’s nest and abandons it.  The cowbird egg and the hatchling that follows is cared for by the victim species, often at the expense of their own young.

Purple Haze Across the Fields

Have you noticed a purple haze across the fields right now?  If so, you may have wondered, “What kind of flowers are they?”

A purple haze of color stretches across fields not already green with cold-season crops like winter wheat.

Say hello to Purple Dead Nettle (Lamium purpureum), a non-native invasive species that has increased its prevalence in recent years by finding an improved niche in no-till cropland.  Purple Dead Nettle, also known as Red Dead Nettle, is native to Asia and Europe.  It has been a familiar early spring “weed” in gardens, along roadsides, and in other disturbed ground for decades.

Purple Dead Nettle owes its new-found success to the timing of its compressed growing season.  Its tiny seeds germinate during the fall and winter, after crops have been harvested and herbicide application has ended for the season.  The plants flower early in the spring and are thus particularly attractive to Honey Bees and other pollinators looking for a source of energy-rich nectar as they ramp up activity after winter lock down.  In many cases, Purple Dead Nettle has already completed its flowering cycle and produced seeds before there is any activity in the field to prepare for planting the summer crop.  The seeds spend the warmer months in dormancy, avoiding the hazards of modern cultivation that expel most other species of native and non-native plants from the agricultural landscape.

Purple Dead Nettle in Bloom
Flowering Purple Dead Nettle as a volunteer cover crop among last year’s corn stubble in a no-till field.
Purple Dead Nettle
Like the flowers of orchids, Purple Hedge Nettle blossoms are described as yoke shaped or bilateral (zygomorphic).  Psychedelic experiences are produced only through observation, not by ingestion.  A member of the mint family, its edible young leaves and tops have nutritional value, making a unique addition to salads and soups.
Purple Dead Nettle, Common Dandelion, and Shepherd's Purse
We call them “weeds”, but what do we know?  Purple Dead Nettle, Common Dandelion, and Shepherd’s Purse (Capsella bursa-pastoris), three edible non-native invasive species with similar life cycles seen here flowering along a rural road among fields where intensive farming is practiced.  Shepard’s Purse, like many members of the mustard family, is already producing seeds at the bottom of the flower cluster by the time the uppermost buds open for business.
Spraying Herbicide
In preparation for seeding of a warm-season crop,  herbicide is applied on a no-till field to kill Shepherd’s Purse and other cool-season plants.  To help prevent sediment and nutrient discharge from lands where high-intensity agriculture is practiced, no-till methods are used to reduce runoff from the areas of bare soil that would otherwise be created by traditional plowing.

While modern farming has eliminated a majority of native plant and animal species from agricultural lands of the lower Susquehanna valley, its crop management practices have simultaneously invited vigorous invasion by a select few non-native species.   High-intensity farming devotes its acreage to providing food for a growing population of people—not to providing wildlife habitat.  That’s why it’s so important to minimize our impact on non-farm lands throughout the remainder of the watershed.  If we continue subdividing, paving, and mowing more and more space, we’ll eventually be living in a polluted semi-arid landscape populated by little else but non-native invasive plants and animals.  We can certainly do better than that.

Photo of the Day

A male Deer Tick
It’s so tiny, you can’t even feel it crawling on your skin.  It’s the Deer Tick (Ixodes scapularis), also known as the Black-legged Tick, and it’s already looking for someone to latch on to.   Don’t let it be you.  After your foray outdoors, be certain to shower and conduct a full body search for these little hitchhikers.  Then, remember to launder the clothing you were wearing.  Deer Ticks in the Lower Susquehanna River Watershed frequently transmit Borrelia burgdorferi, the bacteria that causes Lyme disease, and can be vectors of other microbiological pathogens as well.  The specimen in this photograph is an adult male.  Adult females are similar, but the edge of the abdomen has a crescent-shaped reddish border.  The nymphal stage tick, which is half the size of an adult, is even more likely to infect humans with the Lyme disease bacteria, so be extra vigilant. 

Underwater View of Life in a Vernal Pool

It may look like just a puddle in the woods, but this is a very specialized wetland habitat, a habitat that is quickly disappearing from the Lower Susquehanna River Watershed.  It’s a vernal pool—also known as a vernal pond or an ephemeral (lasting a short time) pool or pond.

Viable vernal pools have several traits in common…

      • They contain water in the spring (hence the name vernal).
      • They have no permanent inflow or outflow of water.
      • They typically dry up during part of the year—usually in late summer.
      • They are fish-free.
      • They provide breeding habitat for certain indicator species of forest-dwelling amphibians and other animals.
      • They are surrounded by forest habitat that supports the amphibians and other vernal pool species during the terrestrial portion of their life cycle.

To have a closer look at what is presently living in this “black leaf” vernal pool, we’re calling on the crew of the S. S. Haldeman to go down under and investigate.

Along the surface of the pool we’re seeing clusters of amphibian eggs, a sign that this pond has been visited by breeding adult frogs and/or salamanders during recent weeks.
Amphibian eggs and the white tail filaments of an invertebrate of interest, Springtime Fairy Shrimp (Eubranchipus vernalis), an endemic of vernal ponds.

Let’s take it down for a better look.  Dive, all dive!

Algae provides food for the shrimp and other inhabitants of the pool.  Leaf litter furnishes hiding places for the pool’s many inhabitants.
These loose clusters of eggs appears to be those of Wood Frogs, a vernal pool indicator species.
Clusters of Wood Frog eggs, the embryos within those in the center of the image less developed than those to the left.
More Wood Frog eggs.  Hatching can take anywhere from two weeks to two months, depending on temperature.
Wood Frog eggs with developing larvae (tadpoles) plainly visible.  The green color of the eggs is created by a symbiotic algae, Oophila amblystomatis, a species unique to vernal pools.  The algae utilizes the waste produced by the developing embryos to fuel its growth and in return releases oxygen into the water during photosynthesis.  Upon hatching, the tadpoles rely upon the algae as one of their principle food sources.
A zoomed-in view showing development of the larvae and what appears to be a tiny invertebrate clinging on the white egg in the upper right.  White eggs don’t hatch and may be infected by a fungus.
Wood Frog eggs and Springtime Fairy Shrimp.
Wood Frog eggs and Springtime Fairy Shrimp.
Springtime Fairy Shrimp swim upside-down.  Note the small, bluish clusters of eggs attached to the abdomens of these females.  Springtime Fairy Shrimp live their entire lives in the vernal pool.  After being deposited in the debris at the bottom of the pool, the eggs will dry out during the summer, then freeze and re-hydrate before hatching during the late winter.
A damselfly larva consuming fairy shrimp.  (Visible in the margin between the uppermost lobes of the dark-colored oak leaf to the right.)
Getting in close we see A) the damselfly larva eating a Springtime Fairy Shrimp and B) one of several discarded exoskeletons of consumed shrimp near this predator.
A fishfly larva (Chauliodinae).  Mosquito numbers are kept in check by the abundance of predators in these pools.
Springtime Fairy Shrimp and a Marbled Salamander (Ambystoma opacum) larva.  The presence of these species confirms this small body of water is a fully-functioning vernal pool.
Springtime Fairy Shrimp and two more larval Marbled Salamanders.  The salamanders’ enlarged gills are necessary to extract sufficient oxygen from the still waters of the pool.
The Marbled Salamander is one of three species of mole salamanders found in the Lower Susquehanna River Watershed.  All breed in vernal pools and live their air-breathing adult lives under the leaves of the forest in subterranean tunnels where they feed on worms and other invertebrates.  Photos of an adult Marbled Salamander and the other two species, Spotted Salamander (Ambystoma maculatum) and Jefferson Salamander (Ambystoma jeffersonianum), can be found by clicking the “Amphibians” tab at the top of this page.
Marbled Salamanders lay their eggs during the fall.  If the bed of the pool is dry at breeding time, the adult female will remain to guard the eggs until rain floods the pool.  The eggs hatch upon inundation, sometimes during the winter.
Marbled Salamanders, like all amphibians that develop in vernal pools, must complete transformation into their air-breathing terrestrial life stage before the pool dries up in the summer heat.
A larval Marbled Salamander explores the bottom of the pool.
A larval Marbled Salamander, Wood Frog eggs, and Springtime Fairy Shrimp, it’s an abundance of life in what at first glance may appear to be just a mud puddle.

We hope you enjoyed this quick look at life in a vernal pool.  While the crew of the S. S. Haldeman decontaminates the vessel (we always scrub and disinfect the ship before moving between bodies of water) and prepares for its next voyage, you can learn more about vernal pools and the forest ecosystems of which they are such a vital component.  Be sure to check out…

If you are a landowner or a land manager, you can find materials specifically providing guidance for protecting, restoring, and re-establishing vernal pool habitats at…

Wood Frogs mating
Wasted Effort-A pair of Wood Frogs mating in a dried-up vernal pool.

Plantings for Wet Lowlands

This linear grove of mature trees, many of them nearly one hundred years old, is a planting of native White Oaks (Quercus alba) and Swamp White Oaks (Quercus bicolor).

Imagine the benefit of trees like this along that section of stream you’re mowing or grazing right now.  The Swamp White Oak in particular thrives in wet soils and is available now for just a couple of bucks per tree from several of the lower Susquehanna’s County Conservation District Tree Sales.  These and other trees and shrubs planted along creeks and rivers to create a riparian buffer help reduce sediment and nutrient pollution.  In addition, these vegetated borders protect against soil erosion, they provide shade to otherwise sun-scorched waters, and they provide essential wildlife habitat.  What’s not to love?

Swamp White Oak
Autumn leaf of a Swamp White Oak

The following native species make great companions for Swamp White Oaks in a lowland setting and are available at bargain prices from one or more of the County Conservation District Tree Sales now underway…

Red Maple
The Red Maple is an ideal tree for a stream buffer project. They do so well that you should limit them to 10% or less of the plants in your project so that they don’t overwhelm slower-growing species.
River Birch
The River Birch (Betula nigra) is a multi-trunked tree of lowlands.  Large specimens with arching trunks help shade waterways and provide a source of falling insects for surface-feeding fish.  Its peeling bark is a distinctive feature.
Common Winterberry
The Common Winterberry with its showy red winter-time fruit is a slow-growing shrub of wet soils.  Only female specimens of this deciduous holly produce berries, so you need to plant a bunch to make sure you have both genders for successful pollination.
American Robins feeding on Common Winterberry.
An American Robin feeding on Common Winterberry.
Common Spicebush
Common Spicebush is a shrub of moist lowland soils.  It is the host plant for the Spicebush Swallowtail butterfly and produces small red berries for birds and other wildlife.  Plant it widely among taller trees to provide native vegetation in the understory of your forest.
Common Spicebush foliage and berries.
Common Spicebush foliage and berries in the shade beneath a canopy of tall trees.
Common Pawpaw
The Common Pawpaw a small shade-loving tree of the forest understory.
Common Pawpaw
Common Pawpaw is a colony-forming small tree which produces a fleshy fruit.  It is the host plant for the caterpillars of the Zebra Swallowtail.
Buttonbush
The Buttonbush is a shrub of wet soils.  It produces a round flower cluster, followed by this globular seed cluster.
Eastern Sycamore
And don’t forget the Eastern Sycamore, the giant of the lowlands.  At maturity, the white-and-tan-colored bark on massive specimens makes them a spectacular sight along stream courses and river shores.  Birds ranging from owls, eagles, and herons to smaller species including the Yellow-throated Warbler rely upon them for nesting sites.
Yellow-crowned Night Herons Nesting in an Eastern Sycamore
Yellow-crowned Night Herons, an endangered species in Pennsylvania, nesting in an Eastern Sycamore.

So don’t mow, do something positive and plant a buffer!

Act now to order your plants because deadlines are approaching fast.  For links to the County Conservation District Tree Sales in the Lower Susquehanna River Watershed, see our February 18th post.

These Geese are Outta Here!

Flies?  Cabbage White butterflies?  Can it really be a late-February day?  It certainly is.  Here are a few more signs of an early spring.

Green Frog
Green Frogs were out and about on this balmy February day trying to latch on to one of those flying insects.  Their long winter’s nap lasted just over six weeks.
Common Grackle
Approximately two hundred Common Grackles passed by susquehannawildlife.net headquarters today.  This one stopped to have a look around before continuing its northbound expedition.
Migrating Canada Geese
Difficult to spot, hundreds of high-flying Canada Geese were seen in the hazy sky above the headquarters garden during the late morning.  These migrants are working their way north from Chesapeake Bay and won’t be seen again in our region until fall.

Time to Order Your Trees for Spring Planting

County Conservation District Tree Sales are underway throughout the Lower Susquehanna River Watershed.  Now is the time to order for pickup in April.  The prices are a bargain and the selection is fabulous.  For species descriptions and more details, visit each tree sale web page (click the sale name highlighted in blue).  And don’t forget to order bundles of evergreens for planting in mixed clumps and groves to provide winter shelter and summertime nesting sites for our local birds.  They’re only $12.00 for a bundle of 10—can’t beat that deal!

Cumberland County Conservation District Annual Tree Seedling Sale—

Orders due by: Friday, March 24, 2023

Pickup on: Thursday, April 20, 2023 or Friday, April 21, 2023

Showy Northeast Meadow Mix
Don’t mow it.  Plant a meadow or pollinator garden instead.
Showy Northeast Meadow Mix
Both Cumberland and Perry Counties are offering a native warm-season grass and wildflower seed mix for planting your own meadow or pollinator garden.  Perry County is also taking orders for a seed mix specifically formulated to grow plants for attracting hummingbirds and butterflies.

Dauphin County Conservation District Seedling Sale—

Orders due by: Monday, March 20, 2023

Pickup on: Thursday, April 20, 2023 or Friday, April 21, 2023

American Goldfinch atop an Eastern Hemlock
The Eastern Hemlock, Pennsylvania’s official state tree, is an excellent choice for addition to your landscape or reforestation project.  It tolerates rocky soils and its cones are an prime source of food for birds ranging from chickadees to finches.

Lancaster County Annual Tree Seedling Sale—

Orders due by: Friday, March 10, 2023

Pickup on: Thursday, April 13, 2023

Northern Red Oak
The handsome yet underused Northern Red Oak is a sturdy long-lived native tree that is ideal for street-side, lawn, and reforestation plantings.  In spring, it can be a magnet for migrating Neotropical birds when its flowers attract a wide variety of tiny insects to its upper reaches.  Unlike many other oaks, this species is a relatively fast grower.

Lebanon County Conservation District Tree and Plant Sale—

Orders due by: Wednesday, March 8, 2023

Pickup on: Friday, April 7, 2023

Pileated Woodpecker feeding on Black Gum berries.
In autumn, even after the bright red foliage is gone, the berries of mature Black Gum (Nyssa sylvatica) trees attract a wide variety of birds like this Pileated Woodpecker.  The Lebanon County Conservation District is offering Black Gum, also known as Black Tupelo, during their 2023 tree sale.  Why not order and plant a half dozen or more?

Perry County Conservation District Tree Sale—(click on 2023 Tree Sale Brochure tab when it scrolls across the page)

Orders due by: March 22, 2023

Pickup on: Thursday, April 13, 2023

Female Eastern Bluebird with Food for Young
The Perry County Conservation District is not only offering plants during this year’s sale, you can also purchase bluebird nest boxes for just $12.00 each!
Riparian Buffer at 15 Years
For less than the cost of one year of mowing, this stream corridor in Conewago Township, Dauphin County was reforested by the owner with hundreds of native trees, the majority purchased through County Conservation District Tree Sale events spanning a period of several years.  By replacing bare soil and mowed areas, the riparian buffer created by these plantings has significantly reduced the nutrient and sediment loads that were polluting the small stream therein known as Brill’s Run.  With determination and not a lot of money, you can do it too.
Maples, Pin Oaks, Eastern White Pines, and other trees in the Brill's Run riparian buffer.
But don’t forget the Eastern White Pines!

Photo of the Day

Eastern Comma
The frosted wing edges let you know that this is the winter form of the Eastern Comma butterfly.  Adult commas, also known as anglewings, will soon enter crevices in dead and living trees to begin hibernation.  On some occasions, they’ll seek shelter from the cold among voids in man-made structures.  Look for commas on mild winter days when they awaken to zip around sunny woodlands and gardens.  You may even see these speedy little fliers pausing to feed on tree sap or carrion.

Photo of the Day

Shadow Darner
If you happen to see a large dark dragonfly cruising around on a sunny November afternoon, it’s a pretty safe bet that it’s a Shadow Darner.  These hardy odonates often persist even after the flights of migratory species have ended for the year.  Shadow Darners are particularly fond of patrolling the shady edges of woodlands where they snag insects and devour them in midair.  Though they are most often found breeding in beaver ponds and other quiet waters, we’ve had them successfully reproduce in the fish-free frog and tadpole pond at susquehannawildlife.net headquarters.

Happy Halloween

Here’s wishing you and yours a Happy Halloween.  It’s a much-anticipated day of excitement capped by surprise visits from strange-looking hideous creatures you’ve never seen before.  They don’t stop by for a chat.  Nope, not a word.  Just a little bit of nearly imperceptible buzzing when the move around.  You see, the little sneaks have hatched a plan.  They want to eat your stuff and maybe trash the place before they go.  And when you finally get rid of them, more start showing up—dozens and dozens, then hundreds.  The more you have, the more you attract.  You’ll be shocked that there are that many living in your neighborhood.  It’s like a scene from “Nightmare on Maple Street”.  The invasion drives some people mad, but you’re just going to love it.  So, get ready, because here they come.  Trick or Treat!

Brown Marmorated Stink Bug
Q:  What’s scariest thing about having Brown Marmorated Stink Bugs over for Halloween?
Brown Marmorated Stink Bug and Spotted Lanternfly
A:  They bring along their friends.

Photo of the Day

Northern Walkingstick
The harmless, herbivorous Northern Walkingstick (Diapheromera femorata), also known as the Common Walkingstick, is well camouflaged when among the twigs and stems of deciduous trees.  As foliage drops in autumn, these wingless insects often descend to ground level and sometimes venture into view.  This individual was found in the vicinity of ideal habitat: a portion of forest including Black Cherry and Black Oak (Quercus velutina) trees.  Juvenile Northern Walkingsticks feed on the leaves of the former, adults on those of the latter.

Photo of the Day

Wild Turkey
A warm weekend will give foraging Wild Turkeys a chance to continue finding protein-rich grasshoppers and crickets before freezing weather forces them to transition to seeds, berries, acorns, hickory nuts, and other fare for sustenance.  Watch for gobblers and hens in forest clearings while you’re out and about having a gander at the fall foliage.

Photo of the Day

Monarch
After somehow finding refuge from freezing temperatures during recent nights, Monarch butterflies were on the move in the Lower Susquehanna River Watershed this afternoon. This one was photographed taking a break to recharge on nectar from flowering Frost Aster, also known as Heath Aster.

Photo of the Day

Eastern Phoebe
This migrating Eastern Phoebe is on the lookout for flying insects, a scarce find on cold autumn mornings.  Eastern Phoebes are the hardiest of our flycatchers and are among the last of our insectivorous birds to head south in the fall.  They’ll return in mid-March feeding on stoneflies and other early-emerging insects along streams and rivers.

Photo of the Day

Lesser Angle-winged Katydid
The Lesser Angle-winged Katydid (Microcentrum retinerve) is a common late-summer and early-autumn inhabitant of treetops.  Males like this one are recognized by the brown spot seen dorsally just behind the head.  Most active at night, they are sometimes attracted to artificial light.  But owing to their affinity for arboreal life and their superb leaf-like camouflage, these and other katydids are more often heard than seen.  The song of the Lesser Angle-winged Katydid is a set of two or three rattles given in quick succession and repeated at one second intervals.  In response to decreasing temperatures during the fall, the song becomes progressively slower and the interval between sets of rattles increases.  A hard frost ends the chorus for the year.

Photo of the Day

Hickory Tussock Moth Caterpillar
During autumn, fuzzy caterpillars crossing roads and trails are a familiar sight throughout the Lower Susquehanna River Watershed.  In oak-hickory forests you may encounter the stunningly beautiful Hickory Tussock Moth (Lophocampa caryae) caterpillar, but don’t touch it; the spine-like bristles are mildly venomous.  Some people react with skin irritation and rashes after contact.  This native species rarely defoliates its host trees.  When it does, it’s usually just weeks before the deciduous leaves would fall, so the overall health of the plant is little affected.  To balance the population, there are numerous avian and terrestrial predators that feed upon both the larval and adult stages of the Hickory Tussock Moth.

Photo of the Day

Butterflies of the Lower Susquehanna River Watershed: Horace's Duskywing
A small butterfly with flights extending into October, the Horace’s Duskywing (Erynnis horatius) is an easily overlooked inhabitant of forest clearings in the vicinity of oaks, particularly Chinkapin Oak (Quercus muehlenbergii), a favorite larval foodplant.

Photo of the Day

Yellowjacket Hover Fly
With today’s abundance of sunshine, the storm-related cold snap has come to an end and autumn insects were again out and about in the lower Susquehanna region.  Yellowjacket Hover Flies (Milesia virginiensis) are important pollinators, feeding on flower nectar along the edges of forests and in meadows.  These harmless mimics of yellowjacket wasps not only resemble their ill-tempered lookalikes in appearance, but they produce similar buzzy wing sounds too.  Yellowjacket Hover Flies are known colloquially as “news bees” due to their habit of lingering around people and constantly “giving them the news”.  The larvae are as beneficial as the adults, feeding on decaying plant matter in rotting wood.  Larvae of some other members of their family, the Syrphidae, feed on aphids.

Photo of the Day

Purple Finch feeding on Green Ash seeds.
It seems a bit early, but Purple Finches are indeed beginning to transit the Lower Susquehanna River Watershed on their way south. This one is feeding on the abundance of seeds produced by a Green Ash that has, at least thus far, survived the Emerald Ash Borer invasion.

Photo of the Day

Zabulon Skipper on Pickerelweed
A female Zabulon Skipper visits a cluster of Pickerelweed blossoms.  The Zabulon Skipper is a small butterfly of our streamsides, riversides, damp meadows, and other moist grassy spaces.  The Pickerelweed is an emergent plant of lakes, ponds, and wetlands.  Add it to your next project to improve water quality and help pollinators like the Zabulon Skipper.  You may even attract a hummingbird or two!   

Shakedown Cruise of the S. S. Haldeman

First there was the Nautilus.  Then there was the Seaview.  And who can forget the Yellow Submarine?  Well, now there’s the S. S. Haldeman, and today we celebrated her shakedown cruise and maiden voyage.  The Haldeman is powered by spent fuel that first saw light of day near Conewago Falls at a dismantled site that presently amounts to nothing more than an electrical substation.  Though antique in appearance, the vessel discharges few emissions, provided there aren’t any burps or hiccups while underway.  So, climb aboard as we take a cruise up the Susquehanna at periscope depth to have a quick look around!

Brunner Island as seen from the east channel.
Close-in approach to emergent Water Willow growing on an alluvial Island.
The approach to York Haven Dam and Conewago Falls from the west channel.
A pair of Powdered Dancers on a midriver log.

Watertight and working fine.  Let’s flood the tanks and have a peek at the benthos.  Dive, all dive!

American Eelgrass, also known as Tapegrass, looks to be growing well in the channels.  Historically, vast mats of this plant were the primary food source for the thousands of Canvasback ducks that once visited the lower Susquehanna each autumn.
As is Water Stargrass (Heteranthera dubia).  When mature, both of these native plants provide excellent cover for young fish.  Note the abundance of shells from deceased Asiatic Clams (Corbicula fluminea) covering the substrate.
Mayfly nymph
A three-tailed mayfly (Ephemeroptera) nymph and a several exoskeletons cling to the downstream side of a rock.
Comb-lipped Casemaker Caddisfly larva and case.
This hollowed-out stick may be a portable protective shelter belonging to a Comb-lipped Casemaker Caddisfly larva (Calamoceratidae).  The larva itself appears to be extending from the end of the “case” in the upper right of the image.  Heteroplectron americanum, a species known for such behavior, is a possibility. 
Rusty Crayfish
In the Susquehanna and its tributaries, the Rusty Crayfish (Faxonius rusticus) is an introduced invasive species.  It has little difficulty displacing native species due to its size and aggressiveness.
Rusty Crayfish
A Rusty Crayfish.
Freshwater Snails Susquehanna: Virginian River Horn Snail
Summers with conditions that promote eelgrass and stargrass growth tend to be big years for Virginian River Horn Snails (Elimia virginica).  2022 appears to be one of those years.  They’re abundant and they’re everywhere on the rocks and gravel substrate in midriver.  Feeding almost incessantly on algae and detritus, these snails are an essential component of the riverine ecosystem, breaking down organic matter for final decomposition by bacteria and fungi.
Freshwater Snails Susquehanna: Virginian River Horn Snail
Bits of debris suspended in the flowing water streak by this Virginian River Horn Snail.  The spire-shaped shell is a streamlining adaptation for maneuvering and holding fast in the strong current.
Freshwater Snails Susquehanna: Virginian River Horn Snail
A young Virginian River Horn Snail following a mature adult.  Note the green algae growing among the decaying plant and animal remains that blanket the river bottom.
Freshwater Snails Susquehanna: Virginian River Horn Snail
Two of a population that may presently include millions of Virginian River Horn Snails living downstream of Conewago Falls.
Susquehanna Snails: Virginian River Horn Snails and Lesser Mystery Snails
Virginian River Horn Snails with Lesser Mystery Snails (Campeloma decisum), another native species commonly encountered at Conewago Falls and in surrounding waters.
Freshwater Snails Susquehanna: River Snail and Virginian River Horn Snail
A River Snail (Leptoxis carinata), also known as a Crested Mudalia, hitching a ride on a Virginian River Horn Snail.  The two species are frequently found together.
Mollusks of the Susquehanna: Yellow Lampmussel and River Snail
A River Snail cleaning the shell of a native freshwater Unionidae mussel, Lampsilis cariosa, commonly called the Yellow Lampmussel or Carried Lampmussel.  Because of their general decline in abundance and range, all Unionidae mussels are protected in Pennsylvania.
Fishes of the Susquehanna: Banded Darter
The Banded Darter (Etheostoma zonale) is a member of the perch family (Percidae).
Fishes of the Susquehanna: Smallmouth Bass
A Smallmouth Bass in strong current.
Fishes of the Susquehanna: Spotfin or Satinfin Shiners
Along the edge of an alluvial island at midriver, Cyprinella (Spotfin or Satinfin) Shiners gather in the cover of an emergent stand of Water Willow.  The closely related Spotfin Shiner (Cyprinella spiloptera) and Satinfin Shiner (Cyprinella analostanus) are nearly impossible to differentiate in the field.
Fishes of the Susquehanna: Spotfin or Satinfin Shiner
A breeding condition male Cyprinella (Spotfin or Satinfin) Shiner.
Fishes of the Susquehanna; Juvenile Channel Catfish
A juvenile Channel Catfish.

We’re finding that a sonar “pinger” isn’t very useful while running in shallow water.  Instead, we should consider bringing along a set of Pings—for the more than a dozen golf balls seen on the river bottom.  It appears they’ve been here for a while, having rolled in from the links upstream during the floods.  Interestingly, several aquatic species were making use of them.

River Snail cleaning a golf ball.
River Snail cleaning a golf ball.
Net-spinning Caddisfly (Hydropsychidae)
A golf ball used as an anchor point for silk cases woven by Net-spinning Caddisfly (Hydropsychidae) larvae to snare food from the water column.
Freshwater Snails (Gastropods) of the Lower Susquehanna River Watershed: Creeping Ancylid (Ferrissia species)
A Creeping Ancylid (Ferrissia species), a tiny gastropod also known as a Coolie Hat Snail, River Limpet, or Brook Freshwater Limpet, inhabits the dimple on a “Top Flight”.
Freshwater Snails (Gastropods) of the Lower Susquehanna River Watershed: Creeping Ancylid (Ferrissia species)
A closeup view of the Creeping Ancylid.  The shell sits atop the snail’s body like a helmet.
We now know why your golf balls always end up in the drink, it’s where they go to have their young.

Well, it looks like the skipper’s tired and grumpy, so that’s all for now.  Until next time, bon voyage!

Monarch an Endangered Species: What You Can Do Right Now

This month, the International Union for Conservation of Nature (I.U.C.N.) added the Migratory Monarch Butterfly (Danaus plexippus plexippus) to its “Red List of Threatened Species”, classifying it as endangered.  Perhaps there is no better time than the present to have a look at the virtues of replacing areas of mowed and manicured grass with a wildflower garden or meadow that provides essential breeding and feeding habitat for Monarchs and hundreds of other species of animals.

Monarch on Common Milkweed Flower Cluster
A recently arrived Monarch visits a cluster of fragrant Common Milkweed flowers in the garden at the susquehannawildlife.net headquarters.  Milkweeds included among a wide variety of plants in a garden or meadow habitat can help local populations of Monarchs increase their numbers before the autumn flights to wintering grounds commence in the fall.  Female Monarchs lay their eggs on milkweed leaves, then, after hatching, the larvae (caterpillars) feed on them before pupating.

If you’re not quite sure about finally breaking the ties that bind you to the cult of lawn manicuring, then compare the attributes of a parcel maintained as mowed grass with those of a space planted as a wildflower garden or meadow.  In our example we’ve mixed native warm season grasses with the wildflowers and thrown in a couple of Eastern Red Cedars to create a more authentic early successional habitat.

Comparison of Mowed Grass to Wildflower Meadow
* Particularly when native warm-season grasses are included (root depth 6′-8′)

Still not ready to take the leap.  Think about this: once established, the wildflower planting can be maintained without the use of herbicides or insecticides.  There’ll be no pesticide residues leaching into the soil or running off during downpours.  Yes friends, it doesn’t matter whether you’re using a private well or a community system, a wildflower meadow is an asset to your water supply.  Not only is it free of man-made chemicals, but it also provides stormwater retention to recharge the aquifer by holding precipitation on site and guiding it into the ground.  Mowed grass on the other hand, particularly when situated on steep slopes or when the ground is frozen or dry, does little to stop or slow the sheet runoff that floods and pollutes streams during heavy rains.

What if I told you that for less than fifty bucks, you could start a wildflower garden covering 1,000 square feet of space?  That’s a nice plot 25′ x 40′ or a strip 10′ wide and 100′ long along a driveway, field margin, roadside, property line, swale, or stream.  All you need to do is cast seed evenly across bare soil in a sunny location and you’ll soon have a spectacular wildflower garden.  Here at the susquehannawildllife.net headquarters we don’t have that much space, so we just cast the seed along the margins of the driveway and around established trees and shrubs.  Look what we get for pennies a plant…

Wildflower Garden
Some of the wildflowers and warm-season grasses grown from scattered seed in the susquehannawildlife.net headquarters garden.

Here’s a closer look…

Lance-leaved Coreopsis
Lance-leaved Coreopsis (Coreopsis lanceolata), a perennial.
Black-eyed Susan
Black-eyed Susan, a biennial or short-lived perennial.
Black-eyed Susan "Gloriosa Daisy"
“Gloriosa Daisy”, a variety of Black-eyed Susan, a biennial or short-lived perennial.
Purple Coneflower
Purple Coneflower, an excellent perennial for pollinators.  The ripe seeds provide food for American Goldfinches.
Common Sunflower
A short variety of Common Sunflower, an annual and a source of free bird seed.
Common Sunflower
Another short variety of Common Sunflower, an annual.

All this and best of all, we never need to mow.

Around the garden, we’ve used a northeast wildflower mix from American Meadows.  It’s a blend of annuals and perennials that’s easy to grow.  On their website, you’ll find seeds for individual species as well as mixes and instructions for planting and maintaining your wildflower garden.  They even have a mix specifically formulated for hummingbirds and butterflies.

Annuals in bloom
When planted in spring and early summer, annuals included in a wildflower mix will provide vibrant color during the first year.  Many varieties will self-seed to supplement the display provided by biennials and perennials in subsequent years.
Wildflower Seed Mix
A northeast wildflower mix from American Meadows.  There are no fillers.  One pound of pure live seed easily plants 1,000 square feet.

Nothing does more to promote the spread and abundance of non-native plants, including invasive species, than repetitive mowing.  One of the big advantages of planting a wildflower garden or meadow is the opportunity to promote the growth of a community of diverse native plants on your property.  A single mowing is done only during the dormant season to reseed annuals and to maintain the meadow in an early successional stage—preventing reversion to forest.

For wildflower mixes containing native species, including ecotypes from locations in and near the Lower Susquehanna River Watershed, nobody beats Ernst Conservation Seeds of Meadville, Pennsylvania.  Their selection of grass and wildflower seed mixes could keep you planting new projects for a lifetime.  They craft blends for specific regions, states, physiographic provinces, habitats, soils, and uses.  Check out these examples of some of the scores of mixes offered at Ernst Conservation Seeds

      • Pipeline Mixes
      • Pasture, Grazing, and Hay Mixes
      • Cover Crops
      • Pondside Mixes
      • Warm-season Grass Mixes
      • Retention Basin Mixes
      • Wildlife Mixes
      • Pollinator Mixes
      • Wetland Mixes
      • Floodplain and Riparian Buffer Mixes
      • Rain Garden Mixes
      • Steep Slope Mixes
      • Solar Farm Mixes
      • Strip Mine Reclamation Mixes

We’ve used their “Showy Northeast Native Wildflower and Grass Mix” on streambank renewal projects with great success.  For Monarchs, we really recommend the “Butterfly and Hummingbird Garden Mix”.  It includes many of the species pictured above plus “Fort Indiantown Gap” Little Bluestem, a warm-season grass native to Lebanon County, Pennsylvania, and milkweeds (Asclepias), which are not included in their northeast native wildflower blends.  More than a dozen of the flowers and grasses currently included in this mix are derived from Pennsylvania ecotypes, so you can expect them to thrive in the Lower Susquehanna River Watershed.

Swamp Milkweed
Swamp Milkweed, a perennial species, is included in the Ernst Seed “Butterfly and Hummingbird Garden Mix”.  It is a favorite of female Monarchs seeking a location to deposit eggs.
Monarch Caterpillar feeding on Swamp Milkweed
A Monarch larva (caterpillar) feeding on Swamp Milkweed.
Butterfly Weed
Butterfly Weed (Asclepias tuberosa) is included in the Ernst Seed “Butterfly and Hummingbird Garden Mix”.  This perennial is also known as Butterfly Milkweed.
Tiger Swallowtails visiting Butterfly Weed
Eastern Tiger Swallowtails are among the dozens of species of pollinators that will visit Butterfly Weed.

In addition to the milkweeds, you’ll find these attractive plants included in Ernst Conservation Seed’s “Butterfly and Hummingbird Garden Mix”, as well as in some of their other blends.

Wild Bergamot
The perennial Wild Bergamot, also known as Bee Balm, is an excellent pollinator plant, and the tubular flowers are a favorite of hummingbirds.
Oxeye
Oxeye is adorned with showy clusters of sunflower-like blooms in mid-summer.  It is a perennial plant.
Plains Coreopsis
Plains Coreopsis (Coreopsis tinctoria), also known as Plains Tickseed, is a versatile annual that can survive occasional flooding as well as drought.
Gray-headed Coneflower
Gray-headed Coneflower (Ratibida pinnata), a tall perennial, is spectacular during its long flowering season.
Monarch on goldenrod.
Goldenrods are a favorite nectar plant for migrating Monarchs in autumn.  They seldom need to be sown into a wildflower garden; the seeds of local species usually arrive on the wind.  They are included in the “Butterfly and Hummingbird Garden Mix” from Ernst Conservation Seeds in low dose, just in case the wind doesn’t bring anything your way.
Partridge Pea
Is something missing from your seed mix?  You can purchase individual species from the selections available at American Meadows and Ernst Conservation Seeds.  Partridge Pea is a good native annual to add.  It is a host plant for the Cloudless Sulphur butterfly and hummingbirds will often visit the flowers.  It does really well in sandy soils.
Indiangrass in flower.
Indiangrass is a warm-season species that makes a great addition to any wildflower meadow mix.  Its deep roots make it resistant to drought and ideal for preventing erosion.

Why not give the Monarchs and other wildlife living around you a little help?  Plant a wildflower garden or meadow.  It’s so easy, a child can do it.

Planting a riparian buffer with wildflowers and warm-season grasses
Volunteers sow a riparian buffer on a recontoured stream bank using wildflower and warm-season grass seed blended uniformly with sand.  By casting the sand/seed mixture evenly over the planting site, participants can visually assure that seed has been distributed according to the space calculations.
Riparian Buffer of wildflowers
The same seeded site less than four months later.
Monarch Pupa
A Monarch pupa from which the adult butterfly will emerge.

Photo of the Day

Leafcutter Bee Visiting Wild Bergamot (Monarda fistulosa)
A Leafcutter Bee (Megachile species) visits the blossoms of a Wild Bergamot (Monarda fistulosa), a native wildflower also known as Bee Balm or Horsemint.  Like Mason Bees (Osmia species), leafcutters will readily nest in the tubular cavities provided inside man-made bee houses.  As their names suggest, Mason Bees use mud to construct and seal the cells for their eggs while leafcutters harvest leaves for the task.  Female Leafcutter Bees, including this individual, can be recognized by the accumulation of pollen carried on the underside of their abdomen.  Within each nest cell, a female will lay an egg atop a supply of pollen and nectar.  These provisions will nourish the larva through development and transformation into an adult.

Photo of the Day

Pickerelweed and Eastern Carpenter Bee
An Eastern Carpenter Bee visits the flowers of an emergent Pickerelweed (Pontederia cordata) in the susquehannawildlife.net headquarters pond.  Each Pickerelweed blossom has conspicuous yellow spots on its uppermost petal, an adaptation shared with the Great Rhododendrons featured in a post earlier this month (July 1).  For each of these species the purpose of these pollen look-a-likes is the same, to attract bees to the pistils and stamens of the flower.  Do these lures work?  Just take a look at the pollen accumulated on the rear leg of this bee.

Blooming in Early July: Great Rhododendron

With the gasoline and gunpowder gang’s biggest holiday of the year now upon us, wouldn’t it be nice to get away from the noise and the enduring adolescence for just a little while to see something spectacular that isn’t exploding or on fire?  Well, here’s a suggestion: head for the hills to check out the flowers of our native rhododendron, the Great Rhododendron (Rhododendron maximum), also known as Rosebay.

Great Rhododendron
The Great Rhododendron is an evergreen shrub found growing in the forest understory on slopes with consistently moist (mesic) soils.  The large, thick leaves make it easy to identify.  During really cold weather, they may droop and curl, but they still remain green and attached to the plant.

Thickets composed of our native heathers/heaths (Ericaceae) including Great Rhododendron, Mountain Laurel, and Pinxter Flower (Rhododendron periclymenoides), particularly when growing in association with Eastern Hemlock and/or Eastern White Pine, provide critical winter shelter for forest wildlife.  The flowers of native heathers/heaths attract bees and other pollinating insects and those of the deciduous Pinxter Flower, which blooms in May, are a favorite of butterflies and Ruby-throated Hummingbirds.

Pinxter Flower in bloom
A close relative of the Great Rhododendron is the Pinxter Flower, also known as the Pink Azalea.

Forests with understories that include Great Rhododendrons do not respond well to logging.  Although many Great Rhododendrons regenerate after cutting, the loss of consistent moisture levels in the soil due to the absence of a forest canopy during the sunny summertime can, over time, decimate an entire population of plants.  In addition, few rhododendrons are produced by seed, even under optimal conditions.  Great Rhododendron seeds and seedlings are very sensitive to the physical composition of forest substrate and its moisture content during both germination and growth.  A lack of humus, the damp organic matter in soil, nullifies the chances of successful recolonization of a rhododendron understory by seed.  In locations where moisture levels are adequate for their survival and regeneration after logging, impenetrable Great Rhododendron thickets will sometimes come to dominate a site.  These monocultures can, at least in the short term, cause problems for foresters by interrupting the cycle of succession and excluding the reestablishment of native trees.  In the case of forests harboring stands of Great Rhododendron, it can take a long time for a balanced ecological state to return following a disturbance as significant as logging.

Birds of Conewago Falls in the Lower Susquehanna River Watershed: Ruffed Grouse
Ruffed Grouse (Bonasa umbellus) may be particularly sensitive to the loss of winter shelter and travel lanes provided by thickets of Great Rhododendron and other members of the heather/heath family.  (Vintage 35 mm image)

In the lower Susquehanna region, the Great Rhododendron blooms from late June through the middle of July, much later than the ornamental rhododendrons and azaleas found in our gardens.   Set against a backdrop of deep green foliage, the enormous clusters of white flowers are hard to miss.

Great Rhododendron Flower Cluster
Great Rhododendrons sport an attractive blossom cluster.  The colors of the flower, especially the markings found only on the uppermost petal, guide pollinators to the stamens (male organs) and pistil (female organ).
Bumble Bee Pollinating a Great Rhododendron Flower
To this Bumble Bee (Bombus species), the yellowish spots on the uppermost petal of the Great Rhododendron may appear to be clumps of pollen and are thus an irresistible lure.  

In the Lower Susquehanna River Watershed, there are but a few remaining stands of Great Rhododendron.  One of the most extensive populations is in the Ridge and Valley Province on the north side of Second Mountain along Swatara Creek near Ravine (just off Interstate 81) in Schuylkill County, Pennsylvania.  Smaller groves are found in the Piedmont Province in the resort town of Mount Gretna in Lebanon County and in stream ravines along the lower river gorge at the Lancaster Conservancy’s Ferncliff and Wissler’s Run Preserves.  Go have a look.  You’ll be glad you did.

Great Rhododendron along Route 125 near Ravine
Great Rhododendron along Route 125 along the base of the north slope of Second Mountain north of Ravine, Schuylkill County, Pennsylvania.
Great Rhododendron along Swatara Creek
Great Rhododendrons beginning to bloom during the second week of July along Swatara Creek north of Ravine, Schuylkill County, Pennsylvania.  Note how acid mine drainage has stained the rocks in the upper reaches of this tributary of the lower Susquehanna.  Mitigation of leachate pollutants from legacy mines has facilitated the return of over two dozen species of fish to this waterway, but the telltale discoloration of the stream substrate will remain for some time.