A special message from your local Chinese Mantis (Tenodera sinensis).
Hey you! Yes, you. I pray you’re paying attention to what’s flying around out there, otherwise it’ll all pass you by.
This summer’s hot humid breezes from the south have not only carried swarms of dragonflies into the lower Susquehanna valley, but butterflies too.
So check out these extravagant visitors from south of the Mason-Dixon Line—before my appetite gets the better of me.
In some years, the Sachem, a vagrant from the south, can become the most frequently observed orange skipper in the Susquehanna valley, far outnumbering faltering populations of resident species around lawns, gardens, meadows, and roadsides. 2020 appears to be one of those years.The Fiery Skipper (Hylephila phyleus) sometimes strays north along the lower Susquehanna River in late summer. This one was photographed recently at Conewago Falls.The Cloudless Sulphur is a large fast-flying brilliant yellow butterfly. Here in the Susquehanna valley, it is usually seen singly, though colonizers will breed in patches of Partridge Pea, Wild Senna (Senna hebecarpa), and Maryland Senna (Senna marilandica). Cloudless Sulphurs are really widespread this year. In areas where intensive farming practices are in use, your chances of seeing one of these wandering around right now might be just as good as your chances of seeing one of the diminished flights of the resident Clouded Sulphurs (Colias philodice) and Orange Sulphurs (Colias eurytheme).The Red-banded Hairstreak (Calycopis cecrops) is the least conspicuous of the stray butterflies shown here. This beauty was photographed several days ago along the edge of an oak forest in northern Lancaster County, PA.
There you have it. Get out there and have a look around. These species won’t be active much longer. In just a matter of weeks, our migratory butterflies, including Monarchs, will be heading south and our visiting strays will either follow their lead or risk succumbing to frosty weather.
For more photographs of butterflies, be sure to click the “Butterflies” tab at the top of the page. We’re adding more as we get them.
After nearly a full week of record-breaking cold, including two nights with a widespread freeze, warm weather has returned. Today, for the first time this year, the temperature was above eighty degrees Fahrenheit throughout the lower Susquehanna region. Not only can the growing season now resume, but the northward movement of Neotropical birds can again take flight—much to our delight.
A rainy day on Friday, May 8, preceded the arrival of a cold arctic air mass in the eastern United States. It initiated a sustained layover for many migrating birds.
Bobolinks (Dolichonyx oryzivorus) in flocks of as many as fifty birds gathered in weedy meadows and alfalfa fields for the week.A Bobolink sheltering in a field of Sweet Vernal Grass (Anthoxanthum odoratum) during the rain on Friday, May 8th.Two of seven Solitary Sandpipers (Tringa solitaria) in a wet field on Friday, May 8. Not-so-solitary after all.Grounded by inclement weather, Rose-breasted Grosbeaks (Pheucticus ludovicianus) made visits to suburban bird feeders in the lower Susquehanna valley. (Charles A. Fox image)
Freeze warnings were issued for five of the next six mornings. The nocturnal flights of migrating birds, most of them consisting of Neotropical species by now, appeared to be impacted. Even on clear moonlit nights, these birds wisely remained grounded. Unlike the more hardy species that moved north during the preceding weeks, Neotropical birds rely heavily on insects as a food source. For them, burning excessive energy by flying through cold air into areas that may be void of food upon arrival could be a death sentence. So they wait.
A freeze warning was issued for Saturday morning, May 9, in the counties colored dark blue on the map. (NOAA/National Weather Service image)This radar image from 3:28 A.M. Saturday morning, May 9, indicates a minor movement of birds in the Great Plains, but there are no notable returns shown around weather radar sites in the freeze area, including the lower Susquehanna valley. (NOAA/National Weather Service image)To avoid the cold wind on Saturday, May 9, this Veery was staying low to the ground within a thicket of shrubs in the forest.This Black-throated Blue Warbler avoided the treetops and spent time in the woodland understory. He sang not a note. With birds conserving energy for the cold night(s) ahead, it was uncharacteristically quiet for the second Saturday in May.A secretive Northern Waterthrush (Parkesia noveboracensis) remained in a wetland thicket.A Scarlet Tanager (Piranga olivacea) tucks his bill beneath a wing and fluffs-up to fight off the cold during a brief May 9th snow flurry.In open country, gusty winds kept Eastern Kingbirds, a species of flycatcher, near the ground in search of the insects they need to sustain them.Horned Larks are one of the few birds that attempt to scratch out an existence in cultivated fields. The application of herbicides and the use of systemic insecticides (including neonicotinoids) eliminates nearly all weed seeds and insects in land subjected to high-intensity farming. For most birds, including Neotropical migrants, cropland in the lower Susquehanna valley has become a dead zone. Birds and other animals might visit, but they really don’t “live” there anymore.Unable to find flying insects over upland fields during the cold snap, swallows concentrated over bodies of water to feed. Some Tree Swallows may have abandoned their nests to survive this week’s cold. Fragmentation of habitats in the lower Susquehanna valley reduces the abundance and diversity of natural food sources for wildlife. For birds like swallows, events like late-season freezes, heat waves, or droughts can easily disrupt their limited food supply and cause brood failure.For this Barn Swallow, attempting to hunt insects above the warm pavement of a roadway had fatal consequences.Another freeze warning was issued for Sunday morning, May 10, in the counties colored dark blue on this map. (NOAA/National Weather Service image)This radar image from 4:58 A.M. Sunday morning, May 10, again indicates the absence of a flight of migrating birds in the area subjected to freezing temperatures. Unlike migrants earlier in the season, the Neotropical species that move north during the May exodus appear unwilling to resume their trek during freezing weather. (NOAA/National Weather Service image)On Sunday evening, May 10, a liftoff of nocturnal migrants is indicated around radar sites along the Atlantic Coastal Plain and, to a lesser degree, in central Pennsylvania. The approaching rain and yet another cold front quickly grounded this flight.After a one day respite, yet another freeze warning was issued for Tuesday morning, May 12. (NOAA/National Weather Service image)And again, no flight in the freeze area. (NOAA/National Weather Service image)The freeze warning for Wednesday morning, May 13. (NOAA/National Weather Service image)And the nocturnal flight: heavy in the Mississippi valley and minimal in the freeze area. (NOAA/National Weather Service image)The freeze on Thursday morning, May 14. (NOAA/National Weather Service image)At 3:08 A.M. on May 14th, a flight is indicated streaming north through central Texas and dispersing into the eastern half of the United States, but not progressing into New England. (NOAA/National Weather Service image)The flight at eight minutes after midnight this morning. Note the stormy cold front diving southeast across the upper Mississippi valley. As is often the case, the concentration of migrating birds is densest in the warm air ahead of the front. (NOAA/National Weather Service image)
Today throughout the lower Susquehanna region, bird songs again fill the air and it seems to be mid-May as we remember it. The flights have resumed.
Indigo Bunting numbers are increasing as breeding populations arrive and migrants continue through. Look for them in thickets along utility and railroad right-of-ways.Common Yellowthroats and other colorful warblers are among the May migrants currently resuming their northward flights.The echoes of the songs of tropical birds are beginning to fill the forests of the lower Susquehanna watershed. The flute-like harmonies of the Wood Thrush are among the most impressive.Ovenbirds are ground-nesting warblers with a surprisingly explosive song for their size. Many arrived within the last two days to stake out a territory for breeding. Listen for “teacher-teacher-teacher” emanating from a woodland near you.
For those of you who dare to shed that filthy contaminated rag you’ve been told to breathe through so that you might instead get out and enjoy some clean air in a cherished place of solitude, here’s what’s around—go have a look.
Northern Flickers have arrived. Look for them anywhere there are mature trees. Despite the fact that flickers are woodpeckers, they often feed on the ground. You’ll notice the white rump and yellow wing linings when they fly away.The tiny Chipping Sparrow frequently nests in small trees in suburban gardens. Lay off the lawn treatments to assure their success.Field Sparrows (Spizella fusilla) are a breeding species in abandoned fields where successional growth is underway.White-throated Sparrows spend the winter in the lower Susquehanna valley. Their numbers are increasing now as waves of migrants pass through on their way north.Northbound flocks of Rusty Blackbirds (Euphagus carolinus) are currently found feeding in forest swamps along the Susquehanna. Their noisy calls sound like a chorus of squeaking hinges.Migratory Red-shouldered Hawks are also making feeding stops at area wetlands.The Palm Warbler (Setophaga palmarum) is easily identified by its tail pumping behavior. Look for it in shrubs along the river shoreline or near lakes and streams. Palm Warblers are among the earliest of the warblers to move through in the spring.
The springtime show on the water continues…
Common Loons will continue migrating through the area during the upcoming month.Buffleheads are still transiting the watershed.Horned Grebes are occurring on the river and on local lakes.Seeing these one-year-old male Hooded Mergansers, the bachelors, wandering around without any adult males or females is a good sign. The adults should have moved on to the breeding grounds and local pairs should be well into a nesting cycle by now. Hatching could occur any day.Like Hooded Mergansers, Wood Ducks are cavity nesters, but their egg laying, incubation, and hatching often occurs a month or more later than that of the hoodies. Judging by the attentiveness of the drake, this pair of woodies is probably in the egg-laying stage of its breeding cycle right now.Redheads (Aythya americana) are stopping for a rest on their way north.In spring, Double-crested Cormorants proceed up the river in goose-like flocks with adult birds like these leading the way.
Hey, what are those showy flowers?
That’s Lesser Celandine (Ficaria verna). It’s often called Fig Buttercup. In early April it blankets stream banks throughout the lower Susquehanna region. If you don’t remember seeing it growing like that when you were younger, there’s a reason. Lesser Celandine is an escape from cultivation that has become invasive. While the appearance is tolerable; it’s the palatability that ruins everything. It’s poisonous if eaten by people or livestock.The Eastern Spring Beauty (Claytonia virginica) is a dainty native wildflower of riparian forests and other woodlands throughout the lower Susquehanna valley.The Trout Lily (Erythronium americanum) is beginning to bloom now. It’s a native of the region’s damp forests.Virginia Bluebells (Mertensia virginica) is not native to the Susquehanna watershed, but neither is it considered invasive. It creates colorful patches in riparian forests.Dutchman’s Breeches (Dicentra cucullaria) is a strikingly beautiful native wildflower that grows on undisturbed forested slopes throughout the Susquehanna valley.
Inside the doorway that leads to your editor’s 3,500 square foot garden hangs a small chalkboard upon which he records the common names of the species of birds that are seen there—or from there—during the year. If he remembers to, he records the date when the species was first seen during that particular year. On New Year’s Day, the results from the freshly ended year are transcribed onto a sheet of notebook paper. On the reverse, the names of butterflies, mammals, and other animals that visited the garden are copied from a second chalkboard that hangs nearby. The piece of paper is then inserted into a folder to join those from previous New Year’s Days. The folder then gets placed back into the editor’s desk drawer beneath a circular saw blade and an old scratched up set of sunglasses—so that he knows exactly where to find it if he wishes to.
A quick glance at this year’s list calls to mind a few recollections.
The 2019 bird list included 48 species, the 47 on the board plus Ruby-throated Hummingbird, which was logged on a slip of paper found tucked into the edge of the frame.
This Green Frog, photographed on New Year’s Day 2019, was “out and about” along the edge of the editor’s garden pond. Due to the recent mild weather, Green Frogs were active during the current New Year’s holiday as well.On a day with strong south winds in late February or during the first two weeks of March, there is often a conspicuous northbound spring flight of migrating waterfowl, gulls, and songbirds that crosses the lower Susquehanna valley as it departs Chesapeake Bay. These Tundra Swans were among the three thousand seen from the garden patio on March 13, 2019. A thousand migrating Canada Geese, 500 Red-winged Blackbirds, numerous Ring-billed Gulls, and some Herring Gulls were seen during the same afternoon.This juvenile Cooper’s Hawk was photographed through the editor’s kitchen window. From its favorite perch on this arbor it would occasionally find success snagging a House Sparrow from the large local flock. It first visited the garden in November, the species being absent there since early spring. Unlike previous years, there was no evidence of a breeding pair in the vicinity during 2019.Plantings that provide food and cover for wildlife are essential to their survival. Native flowers including Trumpet Vine (Campsis radicans) and Partridge Pea provide nourishment for the Ruby-throated Hummingbirds that visit the editor’s garden, but they really love a basket or pot filled with Mexican Cigar (Cuphea ignea) too. The latter (seen here) can be grown as a houseplant and moved outdoors to a semi-shaded location in summer and early fall. But remember, it’s tropical, so you’ll need to bring it back inside when frost threatens.A Swamp Sparrow is an unusual visitor to a small property surrounded by paved parking lots and treeless lawns. Nevertheless, aquatic gardens and native plants helped to attract this nocturnal migrant, seen here eating seeds from Indiangrass. It arrived on September 30 and was gone on October 2.
Before putting the folder back into the drawer for another year, the editor decided to count up the species totals on each of the sheets and load them into the chart maker in the computer.
Despite the habitat improvements in the garden, the trend is apparent. Bird diversity has not cracked the 50 species mark in 6 years. Despite native host plants and nectar species in abundance, butterfly diversity has not exceeded 10 species in 6 years.
It appears that, at the very least, the garden habitat has been disconnected from the home ranges of many species by fragmentation. His little oasis is now isolated in a landscape that becomes increasingly hostile to native wildlife with each passing year. The paving of more parking areas, the elimination of trees, shrubs, and herbaceous growth from the large number of rental properties in the area, the alteration of the biology of the nearby stream by hand-fed domestic ducks, light pollution, and the outdoor use of pesticides have all contributed to the separation of the editor’s tiny sanctuary from the travel lanes and core habitats of many of the species that formerly visited, fed, or bred there. In 2019, migrants, particularly “fly-overs”, were nearly the only sightings aside from several woodpeckers, invasive House Sparrows (Passer domesticus), and hardy Mourning Doves. Even rascally European Starlings became sporadic in occurrence—imagine that! It was the most lackluster year in memory.
The Tufted Titmouse was a daily visitor to the garden through 2018. This one was photographed investigating holes in an old magnolia there during the spring of that year. There were no Tufted Titmouse sightings in the garden in 2019. This and other resident species, especially cavity-nesters, appear to be experiencing at least a temporary decline.Breeding birds including Northern Cardinals may have had a difficult year. In the editor’s garden, a pair were still feeding and escorting one of their young in early October. The infestation of the editor’s town by domestic house and feral cats may have contributed to the failure of earlier broods, but a lack of food is also a likely factor.
If habitat fragmentation were the sole cause for the downward trend in numbers and species, it would be disappointing, but comprehendible. There would be no cause for greater alarm. It would be a matter of cause and effect. But the problem is more widespread.
Although the editor spent a great deal of time in the garden this year, he was also out and about, traveling hundreds of miles per week through lands on both the east and the west shores of the lower Susquehanna. And on each journey, the number of birds seen could be counted on fingers and toes. A decade earlier, there were thousands of birds in these same locations, particularly during the late summer.
At about the time of summer solstice in June each year, Common Grackles begin congregating into roving summer flocks that will grow in size to assure their survival during the autumn migration, winter season, and return north in the spring. From his garden, the editor saw just one flock of less than a dozen birds during the summer of 2019. He saw none during his journeys through other areas of the Susquehanna valley. Flocks of one hundred birds or more did not materialize until the southbound movements of grackles passed through the region in October and November.
In the lower Susquehanna valley, something has drastically reduced the population of birds during breeding season, post-breeding dispersal, and the staging period preceding autumn migration. In much of the region, their late-spring through summer absence was, in 2019, conspicuous. What happened to the tens of thousands of swallows that used to gather on wires along rural roads in August and September before moving south? The groups of dozens of Eastern Kingbirds (Tyrannus tyrannus) that did their fly-catching from perches in willows alongside meadows and shorelines—where are they?
Several studies published during the autumn of 2019 have documented and/or predicted losses in bird populations in the eastern half of the United States and elsewhere. These studies looked at data samples collected during recent decades to either arrive at conclusions or project future trends. They cite climate change, the feline infestation, and habitat loss/degradation among the factors contributing to alterations in range, migration, and overall numbers.
There’s not much need for analysis to determine if bird numbers have plummeted in certain Lower Susquehanna Watershed habitats during the aforementioned seasons—the birds are gone. None of these studies documented or forecast such an abrupt decline. Is there a mysterious cause for the loss of the valley’s birds? Did they die off? Is there a disease or chemical killing them or inhibiting their reproduction? Is it global warming? Is it Three Mile Island? Is it plastic straws, wind turbines, or vehicle traffic?
The answer might not be so cryptic. It might be right before our eyes. And we’ll explore it during 2020.
A clean slate for 2020.
In the meantime, Uncle Ty and I going to the Pennsylvania Farm Show in Harrisburg. You should go too. They have lots of food there.
With autumn coming to a close, let’s have a look at some of the fascinating insects (and a spider) that put on a show during some mild afternoons in the late months of 2019.
Bush Katydids (Scudderia species) are found in brushy habitats and along rural roadsides. Their green summer color fades to brown, maroon, and gold to match the autumn foliage where they hide. Bush katydids often remain active until a hard freeze finally does them in.The Eastern Buck Moth (Hemileuca maia) is fuzzy, appearing to wear a warm coat for its autumn expeditions. Adults emerge in October and may fly as late as December. Females deposit their eggs on the twigs of Bear Oak (Quercus ilicifolia), Blackjack Oak (Q. marilandica), or Chestnut Oak (Q. montana), trees that, in our region, seem most favorable for the moth’s use when growing on burned barrens and mountain slopes. The spiny caterpillars are known to feed only on the foliage of these few trees. In the lower Susquehanna valley, the Eastern Buck Moth is rare because its specialized habitat is in short supply, and it’s all Smokey The Bear’s fault.The Sachem (Atalopedes campestris) wanders north from the Atlantic Coastal Plain into the Susquehanna valley each summer. In some years they become the most numerous small orange butterfly of all, particularly around home gardens. The larvae will feed on Crabgrass (Digitaria species), but have not found success overwintering this far north. By November, adults begin to look pretty drab.From 1978 through 1982, the Asian Multicolored Lady Beetle (Harmonia axyridis) was introduced into the eastern states by the United States Department of Agriculture. It has become a nuisance in many areas where it swarms, sometimes bites, and often overwinters in large smelly masses within homes and other warm buildings. As you may have guessed, it’s possibly displacing some of the less aggressive native lady beetle species.On a chilly afternoon, a sun-warmed Bold Jumping Spider (Phidippus audax) pounced and dispatched this sluggish worker Honey Bee (Apis mellifera) that was trying to gather pollen from a late-season Purple Coneflower bloom. This spider is bold indeed.Under bridges, inside bird nest boxes, and sometimes beneath porches, the female Pipe Organ Mud Dauber (Trypoxylon politum), a predatory wasp, builds these elaborate nests composed of long rows (pipes) of nursery cells. Into each cell one or more paralyzed spiders is deposited along with one of the female’s eggs. When hatched, each larva will feed upon the paralyzed spider(s) inside its cell, then pupate. The pupae overwinter, then emerge from their cells as adults during the following spring. In the autumn, males often stand guard at an entrance to the nest to prevent parasitic species, including some flies (look at the fifth pipe from the right), from laying eggs on the pupae. These wasps are not aggressive toward humans.A Black-and-yellow Mud Dauber (Sceliphron caementarium) observes a neighboring nest of Common Paper Wasps (Polistes exclamans). The Common Paper Wasp, a species also known as the Guinea Paper Wasp, is a native of the southern United States. It is currently expanding its range into the lower Susquehanna valley from the Atlantic Coastal Plain. These two wasp species and the Pipe Organ Mud Dauber are known to regularly coexist. All three will take advantage of man-made structures for their nest sites. People using the picnic tables beneath this pavilion roof never noticed the hundreds of docile wasps above.Those moody Eastern Yellowjackets (Vespula maculifrons) can get very temperamental during warm autumn days. These wasps may appear to have no enemies, but away from areas impacted by man’s everyday activities, they do. The Robber Fly (Promachus species) hunts like a flycatcher or other woodland bird, waiting on a perch along the forest’s edge for prey to pass by, then ambushing it, yellowjackets included.The invasive Spotted Lanternfly, a native of eastern Asia, continues to spread destruction. It established itself throughout much of the east side of the lower Susquehanna River during the summer and fall of 2019. Their route of travel across the farmlands of the region intersects with plenty of vineyards to obliterate and few, if any, natural enemies. Expect them to begin colonizing the west shore en masse during 2020.In 2020, plan to roll a few Spotted Lanternflies over, enjoy the view, and wait for the crimson tide to pass. With any luck, they’ll peak in a year or two.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York, NY.
It’s sprayed with herbicides. It’s mowed and mangled. It’s ground to shreds with noisy weed-trimmers. It’s scorned and maligned. It’s been targeted for elimination by some governments because it’s undesirable and “noxious”. And it has that four letter word in its name which dooms the fate of any plant that possesses it. It’s the Common Milkweed, and it’s the center of activity in our garden at this time of year. Yep, we said milk-WEED.
Now, you need to understand that our garden is small—less than 2,500 square feet. There is no lawn, and there will be no lawn. We’ll have nothing to do with the lawn nonsense. Those of you who know us, know that the lawn, or anything that looks like lawn, are through.
Anyway, most of the plants in the garden are native species. There are trees, numerous shrubs, some water features with aquatic plants, and filling the sunny margins is a mix of native grassland plants including Common Milkweed. The unusually wet growing season in 2018 has been very kind to these plants. They are still very green and lush. And the animals that rely on them are having a banner year. Have a look…
The flowers of the Common Milkweed were exceptionally fragrant this year. At their peak in early June, their hyacinth-or lilac-like aroma was so prevalent, it drifted into our building and overwhelmed the stink of the neighbor’s filthy dumpster that he had placed 12 feet away from our walls (100 feet from his).Common Milkweed attracts a pollinating Greenbottle Fly (Lucilia species). The dumpster attracts the invasive House Fly (Musca domestica), carrier of dysentery, typhoid, and other wonderful diseases. Are you following this? Remember as we proceed, milkweed is “noxious”.Busy Eastern Carpenter Bees (Xylocopa virginica) load up with pollen from the flowers of the Common Milkweed.A Red Milkweed Beetle (Tetraopes tetrophthalmus) munches on a tender fresh Common Milkweed leaf in mid-June.Following the pollination of the flowers, seed pods will begin to grow. We trim these off the plants. The removal of the extra weight allows most of the stems to remain erect through stormy weather. You’ll still get new plants from underground runners. As you may have guessed, we’re trying to keep these plants upright and strong to host Monarch butterfly larvae.
We’ve planted a variety of native grassland species to help support the milkweed structurally and to provide a more complete habitat for Monarch butterflies and other native insects. This year, these plants are exceptionally colorful for late-August due to the abundance of rain. The warm season grasses shown below are the four primary species found in the American tall-grass prairies and elsewhere.
Big Bluestem, a native warm-season grass in flower.Little Bluestem (Schizachyrium scoparium “Fort Indiantown Gap”) in flower. This variety grows on the tank range at the military base where the armored vehicles and prescribed burns substitute for the herd animals and fires of the prairie to prevent succession and allow it to thrive.Partridge Pea can tolerate sandy soil and is a host plant for vagrant Cloudless Sulphur butterflies.Black-eyed Susan (Rudbeckia hirta) is a popular native grassland wildflower.Indiangrass (Sorghastrum nutans) in flower. This and the other native plants shown here are available as seed from Ernst Seed Company in Meadville (PA). They have an unbelievably large selection of indigenous species. You can plant a small plot or acres and acres using really good mixes blended for purposes ranging from reclaiming pipeline right-of-ways and strip mines to naturalizing backyard gardens.A Red Admiral (Vanessa atalanta) butterfly, a migratory species like the Monarch, on Purple Coneflower (Echinacea purpurea). Yes, it is that Echinacea, the same one used as a supplement and home remedy.
There was Monarch activity in the garden today like we’ve never seen before—and it revolved around milkweed and the companion plants.
A female Monarch laying eggs on a Common Milkweed leaf.A third instar Monarch caterpillar with Oleander Aphids (Aphis nerii) on a Common Milkweed leaf. Both of these insect species absorb toxins from the milkweed which makes them distasteful to predators.Fifth instar (left and center) and fourth instar (right) monarch caterpillars devour a Common Milkweed leaf. There were over thirty of these caterpillars in just a ten by ten feet area this morning. We hope that if you’re keeping a habitat for Monarchs, you’re enjoying the same fortune right now!A slow-moving Monarch stopped for a break after making the circuit to deposit eggs on milkweed throughout the garden.Third instar (top), fourth instar (right), and fifth instar (left) Monarch caterpillars quickly consume the leaf of a Common Milkweed plant. Caterpillars emerging from eggs deposited today may not have sufficient late-season food to complete the larval segment of their life cycle. Need more milkweed!After benefiting from the nourishment of the Common Milkweed plant, a fifth instar Monarch caterpillar begins pupation on Big Bluestem grass.Two hours later, the chrysalis is complete.Another chrysalis, this one on flowering Switchgrass just two feet away from the previous one. An adult Monarch will emerge from this pupa to become part of what we hope will be the most populated southbound exodus for the species in over five years.There it is, soon ready to fly away. And all courtesy of the noxious milkweed.A chrysalis can often be found on man-made objects too. This one is on the rim of a flower pot.Ornamental flowers can attract adult Monarch butterflies seeking nectar. We’re now more careful to select seeds and plants that have not been treated with neonicotinoid insecticides. There’s growing concern over the impact these compounds may be having on pollinating species of animals. Oh…and we don’t mow, whack, cut, mutilate, or spray herbicides on our milkweed, but you probably figured that out already.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York.
At the moment there is a heavy snow falling, not an unusual occurrence for mid-February, nevertheless, it is a change in weather. Forty-eight hours ago we were in the midst of a steady rain and temperatures were in the sixties. The snow and ice had melted away and a touch of spring was in the air.
Big Bluestem in the Riverine Grasslands is inundated by the rising waters of the Susquehanna at Conewago Falls. The river ice has been dispersed by the recent mild temperatures and rains.
Anyone casually looking about while outdoors during these last several days may have noticed that birds are indeed beginning to migrate north in the lower Susquehanna valley. Killdeer, American Robins, Eastern Bluebirds, Red-winged Blackbirds, and Common Grackles are easily seen or heard in most of the area now.
Just hours ago, between nine o’clock this morning and one o’clock this afternoon, there was a spectacular flight of birds following the river north, their spring migration well underway. In the blue skies above Conewago Falls, a steady parade of Ring-billed Gulls was utilizing thermals and riding a tailwind from the south-southeast to cruise high overhead on a course toward their breeding range.
Ring-billed Gulls swarm in a thermal updraft above Conewago Falls to gain altitude prior to streaming off to the north and continuing their journey.Ring-billed Gulls climbing to heights sometimes exceeding 1,000 feet before breaking off and gliding away to the north.
The swirling hoards of Ring-billed Gulls attracted other migrants to take advantage of the thermals and glide paths on the breeze. Right among them were 44 Herring Gulls, 3 Great Black-backed Gulls, 12 Tundra Swans (Cygnuscolumbianus), 10 Canada Geese, 3 Northern Pintails (Anas acuta), 6 Common Mergansers, 3 Red-tailed Hawks, a Red-shouldered Hawk, 6 Bald Eagles (non-adults), 8 Black Vultures, and 5 Turkey Vultures.
A first-year Herring Gull (top center) is a standout in a “kettle” of Ring-billed Gulls.How many Ring-billed Gulls passed by today? More than 18,000…with emphasis on MORE THAN. You see, early this afternoon, the handy-dandy clicker-counter used to tick off and tally the big flights of birds as they pass by quit clicking and counting. Therefore, 18,000 is the absolute minimum number of Ring-billed Gulls seen migrating north today. Hopefully the trusty old oil can will get the clicker working again soon.
In the afternoon, the clouds closed in quickly, the flight ended, and by dusk more than an inch of snow was on the ground. Looks like spring to me.
A very light fog lifted quickly at sunrise. Afterward, there was a minor movement of migrants: forty-nine Ring-billed Gulls, a few Herring Gulls, a Red-shouldered Hawk following the river to the southeast, and small flocks totaling nine Cedar Waxwings and twenty-eight Red-winged Blackbirds.
A Belted Kingfisher in the morning fog.A Ring-billed Gull calls as active migrants pass overhead on their way downriver.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
In the Riparian Woodland, small mixed flocks of winter resident and year-round resident birds were actively feeding. They must build and maintain a layer of body fat to survive blustery cold nights and the possible lack of access to food during snowstorms. There’s no time to waste; nasty weather could bring fatal hardship to these birds soon.
A Carolina Chickadee (Poecile carolinensis) feeds on the seeds of an Eastern Sycamore (Platanus occidentalis), also known as American Sycamore. Chickadees are generalist feeders, eating invertebrates and suet at feeding stations in addition to the seeds of many plants. Carolina Chickadees are year-round residents at Conewago Falls.A fast-moving Golden-crowned Kinglet zips from limb to limb to grab tiny insects and other invertebrates. During the winter, these petite birds will carefully probe the bark and crevices of trees to glean enough food to survive. Golden-crowned Kinglets are winter residents at Conewago Falls. In spring, they will depart to nest in coniferous forests.A Downy Woodpecker (Picoides pubescens) searches an infected tree for insects. They are year-round residents.Carolina Wrens (Thryothorus ludovicianus) are considered year-round residents at Conewago Falls, though they may withdraw to the south during severe winters. Carolina Wrens sing year-round. Today, their loud melody echoed through the Riparian Woodland all morning.The tiny bob-tailed Winter Wren is an elusive ground-dwelling winter resident at the falls. You may hear their scolding chatter from rocky areas and tree logs where they climb around mouse-like in search of small invertebrates. Their song is a fast jumble of dainty musical trills that can sometimes be heard echoing through the Riparian Woodland in winter. In spring, they’ll depart to nest in damp coniferous forests.
They get a touch of it here, and a sparkle or two there. Maybe, for a couple of hours each day, the glorious life-giving glow of the sun finds an opening in the canopy to warm and nourish their leaves, then the rays of light creep away across the forest floor, and it’s shade for the remainder of the day.
The flowering plants which thrive in the understory of the Riparian Woodlands often escape much notice. They gather only a fraction of the daylight collected by species growing in full exposure to the sun. Yet, by season’s end, many produce showy flowers or nourishing fruits of great import to wildlife. While light may be sparingly rationed through the leaves of the tall trees overhead, moisture is nearly always assured in the damp soils of the riverside forest. For these plants, growth is slow, but continuous. And now, it’s show time.
So let’s take a late-summer stroll through the Riparian Woodlands of Conewago Falls, minus the face full of cobwebs, and have a look at some of the strikingly beautiful plants found living in the shadows.
Oxeye (Heliopsis helianthoides) is common on the interior and along the edges of Riparian Woodland. Specimens in deep shade flower less profusely and average less than half the height of the five feet tall inhabitants of edge environs.Pale Touch-Me-Not (Impatiens pallida) is one of two species of native Impatiens foundin the river floodplain. Both are known as Jewelweed. The stems and leaves of the indigenous Impatiens retain a great quantity of water, so life in filtered sunlight is essential to prevent desiccation. Contrary to popular folklore, extracts of Jewelweed plants are not effective treatments of Poison Ivy (Toxicodendron radicans) contact dermatitis.Spotted Touch-Me-Not (Impatiens capensis) is typically found in wetter soil than I. pallida. Both Jewelweeds develop popping capsules which help to distribute the seeds of these annual wildflowers. “Touch Me Not”, or you’ll be wearing tiny seeds.Green-headed Coneflower (Rudbeckia laciniata) grows to heights of eight feet in full sun, hence its alternate common name, Tall Coneflower. In deep shade, it may not exceed two feet in height. Floodplains are the prime domain of this perennial.Wingstem (Verbesina alternifolia) normally flowers no earlier than late August. The bases of the leaves are continued onto the stem of the plant to form wings which extend downward along its length. This wildflower tolerates shade, but flowers more profusely along the woodland edge.Great Lobelia (Lobelia siphilitica), or Great Blue Lobelia, is a magnificent wetland and moist woodland wildflower, usually attaining three feet in height and adorned with a plant-topping spike of blossoms. Invasive Japanese Stiltgrass (Microstegium vimineum) can be seen here competing with this plant, resulting in a shorter, less productive Lobelia. Stiltgrass was not found in the Susquehanna River floodplain at Conewago Falls until sometime after 1997. It has spread to all areas of woodland shade, its tiny seeds being blown and translocated along roads, mowed lots, trails, and streams to quickly colonize and overtake new ground.American Bladdernut (Staphylea trifolia), a shrub of shaded woods, develops inflated capsules which easily float away during high water to distribute the seeds contained inside.Buttonbush (Cephalanthus occidentalis) is a shrub of wet soils which produces a strange spherical flower, followed by this globular seed cluster.Common Pawpaw (Asimina triloba) is a colony-forming small tree which produces a fleshy fruit. It is the host plant for the caterpillars of the Zebra Swallowtail. The plant and the butterfly approach the northern limit of their geographic range at Conewago Falls.Common Spicebush (Lindera benzoin) is a widespread understory shrub in wet floodplain soils. It is the host plant for the caterpillars of the Spicebush Swallowtail (Papilio troilus).Sweet Autumn Virgin’s Bower (Clematis terniflora) is an escape from cultivation which has recently naturalized in the edge areas of the Conewago Falls Riparian Woodlands. This vine is very showy when flowering and producing seed, but can be detrimental to some of the understory shrubs upon which it tends to climb.
SOURCES
Long, David; Ballentine, Noel H.; and Marks, James G., Jr. 1997. Treatment of Poison Ivy/Oak Allergic Contact Dermatitis With an Extract of Jewelweed. American Journal of Contact Dermatitis. 8(3): pp. 150-153.
Newcomb, Lawrence. 1977. Newcomb’s Wildflower Guide. Little, Brown and Company. Boston, Massachusetts.
A couple of inches of rain this week caused a small increase in the flow of the river, just a burp, nothing major. This higher water coincided with some breezy days that kicked up some chop on the open waters of the Susquehanna upstream of Conewago Falls. Apparently it was just enough turbulence to uproot some aquatic plants and send them floating into the falls.
Piled against and upon the upstream side of many of the Pothole Rocks were thousands of two to three feet-long flat ribbon-like opaque green leaves of Tapegrass, also called Wild Celery, but better known as American Eelgrass (Vallisneria americana). Some leaves were still attached to a short set of clustered roots. It appears that most of the plants broke free from creeping rootstock along the edge of one of this species’ spreading masses which happened to thrive during the second half of the summer. You’ll recall that persistent high water through much of the growing season kept aquatic plants beneath a blanket of muddy current. The American Eelgrass colonies from which these specimens originated must have grown vigorously during the favorable conditions in the month of August. A few plants bore the long thread-like pistillate flower stems with a fruit cluster still intact. During the recent few weeks, there have been mats of American Eelgrass visible, the tops of their leaves floating on the shallow river surface, near the east and west shorelines of the Susquehanna where it begins its pass through the Gettysburg Basin near the Pennsylvania Turnpike bridge at Highspire. This location is a probable source of the plants found in the falls today.
Uprooted American Eelgrass floating into the Pothole Rocks under the power of a north wind. Note the white thread-like pistillate flower stem to the left and the small rooted specimen to the upper right. The latter is likely a plant from the creeping rootstock on the edge of a colony. As a native aquatic species, American Eelgrass is a critical link in the Susquehanna River and Chesapeake Bay food chain. Its decimation by pollution during the twentieth century led to migration pattern alterations and severe population losses for the Canvasback (Aythya valisineria) duck.American Eelgrass, a very small specimen, found growing in a low-lying Pothole Rock alongside the accumulations of freshly arriving material from upstream. Note that the creeping rootstock has leaves growing from at least three nodes on this plant. Eelgrass dislocations are regular occurrences which sometimes begin new colonies, like the small one seen here in this Diabase Pothole Rock Microhabitat.
The cool breeze from the north was a perfect fit for today’s migration count. Nocturnal migrants settling down for the day in the Riparian Woodlands at sunrise included more than a dozen warblers and some Gray Catbirds (Dumetella carolinensis). Diurnal migration was underway shortly thereafter.
A moderate flight of nocturnal migrants is indicated around NEXRAD sites in the northeastern states at 3:18 AM EDT. The outer rain bands of Hurricane Irma can be seen approaching the Florida Keys as the storm closes in on the peninsula. (NOAA/National Weather Service image)
Four Bald Eagles were counted as migrants this morning. Based on plumage, two were first-year eagles (Juvenile) seen up high and flying the river downstream, one was a second-year bird (Basic I) with a jagged-looking wing molt, and a third was probably a fourth year (Basic III) eagle looking much like an adult with the exception of a black terminal band on the tail. These birds were the only ones which could safely be differentiated from the seven or more Bald Eagles of varying ages found within the past few weeks to be lingering at Conewago Falls. There were as many as a dozen eagles which appeared to be moving through the falls area that may have been migrating, but the four counted were the only ones readily separable from the locals.
Red-tailed Hawks (Buteo jamaicensis) were observed riding the wind to journey not on a course following the river, but flying across it and riding the updraft on the York Haven Diabase ridge from northeast to southwest.
Bank Swallows (Riparia riparia) seem to have moved on. None were discovered among the swarms of other species today.
Ruby-throated Hummingbirds, Caspian Terns, Cedar Waxwings (Bombycilla cedrorum), and Chimney Swifts (Chaetura pelagica) were migrating today, as were Monarch butterflies.
Not migrating, but always fun to have around, all four wise guys were here today. I’m referring to the four members of the Corvid family regularly found in the Mid-Atlantic states: Blue Jay (Cyanocitta cristata), American Crow (Corvus brachyrhynchos), Fish Crow (Corvus ossifragus), and Common Raven (Corvus corax).
It looks like a big Blue Jay, but it’s not. This Belted Kingfisher (Megaceryle alcyon) takes a break after flying around the falls trying to shake a marauding Ruby-throated Hummingbird off its tail.CLICK ON THE LOGO FOR TODAY’S MIGRATION COUNT TOTALS
SOURCES
Klots, Elsie B. 1966. The New Field Book of Freshwater Life. G. P. Putnam’s Sons. New York, NY.
It’s tough being good-looking and liked by so many. You’ve got to watch out, because popularity makes you a target. Others get jealous and begin a crusade to have you neutralized and removed from the spotlight. They’ll start digging to find your little weaknesses and flaws, then they’ll exploit them to destroy your reputation. Next thing you know, people look at you as some kind of hideous scoundrel.
Today, bright afternoon sunshine and a profusion of blooming wildflowers coaxed butterflies into action. It was one of those days when you don’t know where to look first.
A Pearl Crescent (Phyciodes tharos) sipping nectar from Rough Boneset (Eupatorium pilosum) flowers. Asters (Aster) are the host plants for the larvae of this butterfly.A Buckeye (Junonia coenia) on Rough Boneset. Its caterpillars are known to feed on members of the Acanthus family, possibly including the Water Willow (Justicia americana) which is so abundant in Conewago Falls.Visitors from south of the Mason-Dixon Line arrived on the recent warm winds. Two Cloudless Sulphurs (Phoebis sennae) patrolled the Riverine Grasslands, especially near the stands of Partridge Pea, a possible host plant. One is seen here visiting a Halbred-leaved Rose Mallow blossom. These large yellow butterflies are always a standout.
Purple Loosestrife (Lythrum salicaria) has a bad reputation. Not native to the Americas, this prolific seed producer began spreading aggressively into many wetlands following its introduction. It crowds out native plant species and can have a detrimental impact on other aquatic life. Stands of loosestrife in slow-moving waters can alter flows, trap sediment, and adversely modify the morphology of waterways. Expensive removal and biological control are often needed to protect critical habitat.
The dastardly Purple Loosestrife may have only two positive attributes. First, it’s a beautiful plant. And second, it’s popular; butterflies and other pollinators find it to be irresistible and go wild over the nectar.
A Silver-spotted Skipper (Epargyreus clarus) feeding on Purple Loosestrife nectar. The host plants for this common butterfly’s caterpillars are a wide variety of Legumes.A Cabbage White (Pieris rapae), a butterfly introduced from Europe in the 1800s, feeds on introduced Purple Loosestrife.A Wild Indigo Duskywing (Erynnis baptisiae) feeding on Purple Loosestrife. This butterfly has expanded its population and range by using the introduced Crown Vetch as a host plant.
Don’t you just adore the wonderful butterflies. Everybody does. Just don’t tell anyone that they’re pollinating those dirty filthy no-good Purple Loosestrife plants.
SOURCES
Brock, Jim P., and Kenn Kaufman. 2003. Butterflies of North America. Houghton Mifflin Company. New York.
Newcomb, Lawrence. 1977. Newcomb’s Wildflower Guide. Little, Brown and Company. Boston, Massachusetts.
There’s something frightening going on down there. In the sand, beneath the plants on the shoreline, there’s a pile of soil next to a hole it’s been digging. Now, it’s dragging something toward the tunnel it made. What does it have? Is that alive?
We know how the system works, the food chain that is. The small stuff is eaten by the progressively bigger things, and there are fewer of the latter than there are of the former, thus the whole network keeps operating long-term. Some things chew plants, others devour animals whole or in part, and then there are those, like us, that do both. In the natural ecosystem, predators keep the numerous little critters from getting out of control and decimating certain other plant or animal populations and wrecking the whole business. When man brings an invasive and potentially destructive species to a new area, occasionally we’re fortunate enough to have a native species adapt and begin to keep the invader under control by eating it. It maintains the balance. It’s easy enough to understand.
Japanese Beetles (Popillia japonica) seen here on Halberd-leaved Rose Mallow. Without predation, exploding numbers of this invasive non-indigenous insect can defoliate and kill numerous species of plants in a given area.The Song Sparrow (Melospiza melodia) is a generalist feeder, eating seeds and invertebrates including Japanese Beetles. This species is the omnipresent year-round occupant of shoreline vegetation along the lower Susquehanna River.
Late summer days are marked by a change in the sounds coming from the forests surrounding the falls. For birds, breeding season is ending, so the males cease their chorus of songs and insects take over the musical duties. The buzzing calls of male “Annual Cicadas” (Neotibicen species) are the most familiar. The female “Annual Cicada” lays her eggs in the twigs of trees. After hatching, the nymphs drop to the ground and burrow into the soil to live and feed along tree roots for the next two to five years. A dry exoskeleton clinging to a tree trunk is evidence that a nymph has emerged from its subterranean haunts and flown away as an adult to breed and soon thereafter die. Flights of adult “Annual Cicadas” occur every year, but never come anywhere close to reaching the enormous numbers of “Periodical Cicadas” (Magicicada species). The three species of “Periodical Cicadas” synchronize their life cycles throughout their combined regional populations to create broods that emerge as spectacular flights once every 13 or 17 years.
An “Annual Cicada” also known as a “Dog-day Cicada”, clings to the stem of a Halberd-leaved Rose Mallow at Conewago Falls.
For the adult cicada, there is danger, and that danger resembles an enormous bee. It’s an Eastern Cicada Killer (Specius speciosus) wasp, and it will latch onto a cicada and begin stinging while both are in flight. The stings soon paralyze the screeching, panicked cicada. The Cicada Killer then begins the task of airlifting and/or dragging its victim to the lair it has prepared. The cicada is placed in one of more than a dozen cells in the tunnel complex where it will serve as food for the wasp’s larvae. The wasp lays an egg on the cicada, then leaves and pushes the hole closed. The egg hatches in a several days and the larval grub is on its own to feast upon the hapless cicada.
An Eastern Cicada Killer (Sphecius speciosus) along the river shoreline. Despite their intimidating appearance, they do not sting humans and can be quite docile when approached.
Other species in the Solitary Wasp family (Sphecidae) have similar life cycles using specific prey which they incapacitate to serve as sustenance for their larvae.
A Solitary Wasp, one of the Thread-waisted Wasps (Ammophila species), drags a paralyzed moth caterpillar to its breeding dungeon in the sandy soil at Conewago Falls. For the victim, there is no escape from the crypt.
The Solitary Wasps are an important control on the populations of their respective prey. Additionally, the wasp’s bizarre life cycle ensures a greater survival rate for its own offspring by providing sufficient food for each of its progeny before the egg beginning its life is ever put in place. It’s complete family planning.
The cicadas reproduce quickly and, as a species, seem to endure the assault by Cicada Killers, birds, and other predators. The Periodical Cicadas (Magicicada), with adult flights occurring as a massive swarm of an entire population every thirteen or seventeen years, survive as species by providing predators with so ample a supply of food that most of the adults go unmolested to complete reproduction. Stay tuned, 2021 is due to be the next Periodical Cicada year in the vicinity of Conewago Falls.
SOURCES
Eaton, Eric R., and Kenn Kaufman. 2007. Kaufman Field Guide to Insects of North America. Houghton Mifflin Company. New York.
The tall seed-topped stems swaying in a summer breeze are a pleasant scene. And the colorful autumn shades of blue, orange, purple, red, and, of course, green leaves on these clumping plants are nice. But of the multitude of flowering plants, Big Bluestem, Freshwater Cordgrass, and Switchgrass aren’t much of a draw. No self-respecting bloom addict is going out of their way to have a gander at any grass that hasn’t been subjugated and tamed by a hideous set of spinning steel blades. Grass flowers…are you kidding?
Big Bluestem in flower in the Riverine Grasslands at Conewago Falls.
O.K., so you need something more. Here’s more.
Meet the Partridge Pea (Chamaecrista fasciculata). It’s an annual plant growing in the Riverine Grasslands at Conewago Falls as a companion to Big Bluestem. It has a special niche growing in the sandy and, in summertime, dry soils left behind by earlier flooding and ice scour. The divided leaves close upon contact and also at nightfall. Bees and other pollinators are drawn to the abundance of butter-yellow blossoms. Like the familiar pea of the vegetable garden, the flowers are followed by flat seed pods.
The Partridge Pea can tolerate dry sandy soils.
But wait, here’s more.
In addition to its abundance in Conewago Falls, the Halberd-leaved Rose Mallow (Hibiscus laevis) is the ubiquitous water’s edge plant along the free-flowing Susquehanna River for miles downstream. It grows in large clumps, often defining the border between the emergent zone and shore-rooted plants. It is particularly successful in accumulations of alluvium interspersed with heavier pebbles and stone into which the roots will anchor to endure flooding and scour. Such substrate buildup around the falls, along mid-river islands, and along the shores of the low-lying Riparian Woodlands immediately below the falls are often quite hospitable to the species.
Halberd-leaved Rose Mallow is a durable inhabitant of the falls. Regular flooding keeps competing species at bay. A taproot helps to safeguard against dislocation, allowing plants to grow in places subjected to turbulent currents.Halberd-leaved Rose Mallow in bloom. The similarity to cultivated members of the Hibiscus genus can readily be seen. It is one of the showiest of perennial wildflowers in the floodplain. Note the lobed, halberd-shaped leaves, source of its former species name militaris.The seeds of Halberd-leaved Rose Mallow are contained in bladders which can float to assist in their distribution. Some of these bladders cling to the dead leafless stems in winter, making it an easy plant to identify in nearly any season.
A second native wildflower species in the genus Hibiscus is found in the Conewago Falls floodplain, this one in wetlands. The Swamp Rose Mallow (H. moscheutos) is similar to Halberd-leaved Rose Mallow, but sports more variable and colorful blooms. The leaves are toothed without the deep halberd-style lobes and, like the stems, are downy. As the common name implies, it requires swampy habitat with ample water and sunlight.
Swamp Rose Mallow in a sunny wetland. This variety with solid-colored flowers (without dark centers) and pale green leaves and stems was formerly known as a separate species of Swamp Rose Mallow, H. palustris. Note that the flowers are terminal on the stems.A few scattered specimens of a more typical variety of Swamp Rose Mallow are found on the shoreline and in the Riverine Grasslands of Conewago Falls. The blooms are bright pink with darker centers and the leaf stems are robust and reddish. This one is seen growing among Halberd-leaved Rose Mallow, with which it shares the characteristic of having flower stems growing from some of the upper leaf axils. A variety with red-centered white flowers is often found throughout the plant’s range.
In summary, we find Partridge Pea in the Riverine Grasslands growing in sandy deposits left by flood and ice scour. We find Halberd-leaved Rose Mallow rooted at the border between shore and the emergent zone. We find Swamp Rose Mallow as an emergent in the wetlands of the floodplain. And finally, we find marshmallows in only one location in the area of Conewago Falls. Bon ap’.
Here’s S’more
SOURCES
Newcomb, Lawrence. 1977. Newcomb’s Wildflower Guide. Little, Brown and Company. Boston, Massachusetts.
It has not been a good summer if you happen to be a submerged plant species in the lower Susquehanna River. Regularly occurring showers and thunderstorms have produced torrents of rain and higher than usual river stages. The high water alone wouldn’t prevent you from growing, colonizing a wider area, and floating several small flowers on the surface, however, the turbidity, the suspended sediment, would. The muddy current casts a dirty shadow on the benthic zone preventing bottom-rooted plants from getting much headway. There will be smaller floating mats of the uppermost leaves of these species. Fish and invertebrates which rely upon this habitat for food and shelter will find sparse accommodation…better luck next year.
Due to the dirty water, fish-eating birds are having a challenging season as they try to catch sufficient quantities of prey to feed themselves and their offspring. A family of Ospreys (Pandion haliaetus) at Conewago Falls, including recently fledged young, were observed throughout this morning and had no successful catches. Of the hundred or more individual piscivores of various species present, none were seen retrieving fish from the river. The visibility in the water column needs to improve before fishing is a viable enterprise again.
Ospreys competing for a suitable fishing perch. Improving water conditions in the coming week should increase their success as predators.Versatile at finding food, adult Bald Eagles are experienced and know to be on the lookout for Ospreys with fish, a meal they can steal through intimidation.
While the submerged plant communities may be stunted by 2017’s extraordinary water levels, there is a very unique habitat in Conewago Falls which endures summer flooding and, in addition, requires the scouring effects of river ice to maintain its mosaic of unique plants. It is known as a Riverine Grassland or scour grassland.
The predominant plants of the Riverine Grasslands are perennial warm-season grasses. The deep root systems of these hardy species have evolved to survive events which prevent the grassland from reverting to woodland through succession. Fire, intense grazing by wild herd animals, poor soils, drought, and other hardships, including flooding and ice scour, will eliminate intolerant plant species and prevent an area from reforesting. In winter and early spring, scraping and grinding by flood-driven chunk ice mechanically removes large woody and poorly rooted herbaceous growth from susceptible portions of the falls. These adverse conditions clear the way for populations of species more often associated with North America’s tall grass prairies to take root. Let’s have a look at some of the common species found in the “Conewago Falls Pothole Rocks Prairie”.
Big Bluestem (Andropogon gerardi), seen heregrowing in the cracks of a pothole rock. High water nourishes the plant by filling the crevices with nutrient-loaded sediment. This species evolved with roots over three feet deep to survive fires, trampling by bison, and drought.Freshwater Cordgrass (Spartina pectinata) does well with its roots in water. It creates exceptional bird habitat and grows in the falls and on ice-scoured small islands in free-flowing segments of the Susquehanna River downstream.Switchgrass (Panicum virgatum), like Big Bluestem, is one of the tall grass prairie species and, like Freshwater Cordgrass, grows in near pure stands on ice-scoured islands. It takes flooding well and its extensive root system prevents erosion.Though not a grass, Water Willow (Justicia americana) is familiar as a flood-enduring emergent plant of river islands, gravel bars, and shorelines where its creeping rhizome root system spreads the plant into large masses. These stands are often known locally as “grass beds”. This member of the acanthus family provides habitat for fish and invertebrates among its flooded leaves and stems. Its presence is critical to aquatic life in a year such as this.
The Conewago Falls Riverine Grassland is home to numerous other very interesting plants. We’ll look at more of them next time.
SOURCES
Brown, Lauren. 1979. Grasses, An Identification Guide. Houghton Mifflin Company. New York, NY.
She ate only toaster pastries…that’s it…nothing else. Every now and then, on special occasions, when a big dinner was served, she’d have a small helping of mashed potatoes, no gravy, just plain, thank you. She received all her nutrition from several meals a week of macaroni and cheese assembled from processed ingredients found in a cardboard box. It contains eight essential vitamins and minerals, don’t you know? You remember her, don’t you?
Adult female butterflies must lay their eggs where the hatched larvae will promptly find the precise food needed to fuel their growth. These caterpillars are fussy eaters, with some able to feed upon only one particular species or genus of plant to grow through the five stages, the instars, of larval life. The energy for their fifth molt into a pupa, known as a chrysalis, and metamorphosis into an adult butterfly requires mass consumption of the required plant matter. Their life cycle causes most butterflies to be very habitat specific. These splendid insects may visit the urban or suburban garden as adults to feed on nectar plants, however, successful reproduction relies upon environs which include suitable, thriving, pesticide-free host plants for the caterpillars. Their survival depends upon more than the vegetation surrounding the typical lawn will provide.
The Monarch (Danaus plexippus), a butterfly familiar in North America for its conspicuous autumn migrations to forests in Mexico, uses the milkweeds (Asclepias) almost exclusively as a host plant. Here at Conewago Falls, wetlands with Swamp Milkweed (Asclepias incarnata) and unsprayed clearings with Common Milkweed (A. syriaca) are essential to the successful reproduction of the species. Human disturbance, including liberal use of herbicides, and invasive plant species can diminish the biomass of the Monarch’s favored nourishment, thus reducing significantly the abundance of the migratory late-season generation.
Monarch caterpillar after a fourth molt. The fifth instar feeding on Swamp Milkweed.A fifth molt begets the Monarch pupa, the chrysalis, from which the showy adult butterfly will emerge.Adult Monarch feeding on Goldenrod (Solidago) nectar.
Butterflies are good indicators of the ecological health of a given environment. A diversity of butterfly species in a given area requires a wide array of mostly indigenous plants to provide food for reproduction. Let’s have a look at some of the species seen around Conewago Falls this week…
An adult Silvery Checkerspot (Chlosyne nycteis) visiting a nectar plant, Partridge Pea (Chamaecrista fasciculata). Wingstem (Verbesina alternifolia), a plant of the Riparian Woodlands, is among the probable hosts for the caterpillars.A Gray Hairstreak (Strymon melinus) visits Crown Vetch, a possible host plant. Other potential larval food in the area includes Partridge Pea, Halberd-leaved Rose Mallow (Hibiscus laevis) of the river shoreline, and Swamp Rose Mallow (Hibiscus moscheutos), a plant of wetlands.The Eastern Tailed Blue (Cupido comyntas) may use Partridge Pea , a native wildflower species, and the introduced Crown Vetch (Securigera/Coronilla varia) as host and nectar plants at Conewago Falls.The Least Skipper (Ancyloxypha numitor) is at home among tall grasses in woodland openings, at riverside, and in the scoured grassland habitat of the Pothole Rocks in the falls. Host plants available include Switchgrass (Panicum vigatum), Freshwater Cordgrass (Spartina pectinata), and Foxtails (Setaria).The Zabulon Skipper (Poanes zabulon) is an inhabitant of moist clearings where the caterpillars may feed upon Lovegrasses (Eragrostis) and Purpletop (Tridens flavus).The Eastern Tiger Swallowtail (Papilio glaucus), a female seen here gathering nectar from Joe-Pye Weed (Eutrochium), relies upon several forest trees as hosts. Black Cherry (Prunus serotina), Willow (Salix), Yellow Poplar (Liriodendron tulipifera), also known as Tuliptree, and Green Ash (Fraxinus pennsylvanica) are among the local species known to be used. The future of the latter food species at Conewago Falls is doubtful. Fortunately for the Eastern Tiger Swallowtail, the “generalist” feeding requirements of this butterfly’s larvae enable the species to survive the loss of a host plant.A female Eastern Tiger Swallowtail, black morph, gathering nectar from Joe-Pye Weed.The Zebra Swallowtail (Protographium marcellus), an adult seen here on Joe-Pye Weed, feeds exclusively upon Pawpaw (Asimina) trees as a caterpillar. This butterfly species may wander, but its breeding range is limited to the moist Riparian Woodlands where colonial groves of Pawpaw may be found. The Common Pawpaw (Asimina triloba), our native species in Pennsylvania, and the Zebra Swallowtail occur at the northern edge of their geographic ranges in the Lower Susquehanna River Watershed. Planting Pawpaw trees as an element of streamside reforestation projects certainly benefits this marvelous butterfly.
The spectacularly colorful butterflies are a real treat on a hot summer day. Their affinity for showy plants doubles the pleasure.
By the way, I’m certain by now you’ve recalled that fussy eater…and how beautiful she grew up to be.
SOURCES
Brock, Jim P., and Kaufman, Kenn. 2003. Butterflies of North America. Houghton Mifflin Company. New York, NY.
